390 lines
49 KiB
XML
390 lines
49 KiB
XML
<document id="838A39899E133A8F28FA15B4303C1628" ID-DOI="10.35249/rche.49.3.23.26" ID-ISSN="0718-8994" ID-Zenodo-Dep="13206445" ID-ZooBank="D0130490-055A-451F-A6DA-E1B664946B7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722664042887" checkinUser="felipe" docAuthor="Barria, Manuel D., Quirós, Dora Isabel & Bayfield-Farrell, William Peter" docDate="2023" docId="03EA87C7E2219B792D80F0CBFC7EFA96" docLanguage="en" docName="RevChilEntomol.49.3.649-655.pdf" docOrigin="Revista Chilena de Entomología (Rev. Chil. Entomol.) 49 (3)" docSource="http://zoobank.org/d0130490-055a-451f-a6da-e1b664946b7b" docStyle="DocumentStyle:451321A1F3358405F3D5797D4FC35865.3:RevChilEntomol.2020-.journal_article" docStyleId="451321A1F3358405F3D5797D4FC35865" docStyleName="RevChilEntomol.2020-.journal_article" docStyleVersion="3" docTitle="Phyllophaga (Phyllophaga) clavijobustosi Barria, Quiros, and Bayfield-Farrell 2023, sp. nov." docType="treatment" docUuid="83F12AA1-C068-4A10-B09F-DC40CF0A971B" docUuidSource="ZooBank" docVersion="2" lastPageNumber="653" masterDocId="FFD3FFBFE2209B7D2D0DF436FFCAFF81" masterDocTitle="Description of a new Panamanian species of Phyllophaga Harris, 1827 (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini) of the orosina species group" masterLastPageNumber="655" masterPageNumber="649" pageNumber="650" updateTime="1722703852240" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="172945ECC0FF0364E232C60B450E0304">Description of a new Panamanian species of Phyllophaga Harris, 1827 (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini) of the orosina species group</mods:title>
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<mods:name id="BADD2B805E719E61D3B8D93198BD65C4" type="personal">
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<mods:roleTerm id="1B4C74CCA712A20395D0B7A19F687E51">Author</mods:roleTerm>
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<mods:namePart id="D42BFF53526FFAC8C0F0F42D0E5E7A14">Barria, Manuel D.</mods:namePart>
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<mods:name id="09EC6B0F035D77FBA1DE3340E24FA948" type="personal">
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<mods:roleTerm id="0B8D6A9456F69C74F73E6D2FED964CD1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="51CCCBE99D439E62D068D8A712FBA3FA">Quirós, Dora Isabel</mods:namePart>
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</mods:name>
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<mods:name id="1067ED4AA7F984234B02E256A240F7DB" type="personal">
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<mods:roleTerm id="56EF3F3DB444F712F3EC757A80B44F8F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="05018578146632A9C2C644EAAFBEC5FC">Bayfield-Farrell, William Peter</mods:namePart>
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<mods:typeOfResource id="FA7D6D70A218ECF9FA5F6703E0EA839A">text</mods:typeOfResource>
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<mods:title id="4557C41C8B43BC9C6FC7EF9DF5FBE344">Revista Chilena de Entomología</mods:title>
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<mods:part id="2874F212F11E8C2C9D8971BA54BA8578">
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<mods:date id="478AE024CE95AC9FFCC4CDFF6B0C48DC">2023</mods:date>
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<mods:title id="2D440D2C6B6C63C12833682E593C047A">Rev. Chil. Entomol.</mods:title>
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<mods:number id="AD7E6BC474B6915D4BD1432F59F069DE">2023-09-29</mods:number>
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<mods:number id="0B7BBF246335591BAA4797556609545E">49</mods:number>
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<mods:number id="533B3DE09895026FF09B35075D153E66">3</mods:number>
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<mods:classification id="89AAC7FC11AB2ED649615905B2DF5444">journal article</mods:classification>
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<mods:identifier id="F6D6EB4FCEC2CB77543DE6B10ED9A952" type="ISSN">0718-8994</mods:identifier>
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<treatment id="03EA87C7E2219B792D80F0CBFC7EFA96" LSID="urn:lsid:zoobank.org:act:83F12AA1-C068-4A10-B09F-DC40CF0A971B" httpUri="http://treatment.plazi.org/id/03EA87C7E2219B792D80F0CBFC7EFA96" lastPageId="4" lastPageNumber="653" pageId="1" pageNumber="650">
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<subSubSection id="C359655AE2219B7C2D80F0CBFB81FA94" box="[141,1099,1277,1301]" pageId="1" pageNumber="650" type="nomenclature">
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<paragraph id="8BFC36D1E2219B7C2D80F0CBFB81FA94" blockId="1.[141,1099,1277,1333]" box="[141,1099,1277,1301]" pageId="1" pageNumber="650">
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<heading id="D0B481BDE2219B7C2D80F0CBFB81FA94" box="[141,1099,1277,1301]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="650" reason="4">
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<taxonomicName id="4C434D52E2219B7C2D80F0CBFC2CFA94" ID-CoL="BV625" authority="Barria, Quiros, and Bayfield-Farrell" authorityName="Barria, Quiros, and Bayfield-Farrell" authorityYear="2023" box="[141,998,1277,1301]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="650" phylum="Arthropoda" rank="species" species="clavijobustosi" status="sp. nov." subGenus="Phyllophaga">
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<emphasis id="B937EAC3E2219B7C2D80F0CBFEDBFA94" box="[141,273,1277,1301]" italics="true" pageId="1" pageNumber="650">Phyllophaga</emphasis>
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||
(
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<emphasis id="B937EAC3E2219B7C2C2DF0CBFE6EFA94" box="[288,420,1277,1301]" italics="true" pageId="1" pageNumber="650">Phyllophaga</emphasis>
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||
)
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<emphasis id="B937EAC3E2219B7C2CBEF0CBFD89FA94" box="[435,579,1277,1301]" italics="true" pageId="1" pageNumber="650">clavijobustosi</emphasis>
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||
Barria, Quirós, and Bayfield-Farrell
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</taxonomicName>
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||
,
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||
<emphasis id="B937EAC3E2219B7C2EFCF0CBFB81FA94" bold="true" box="[1009,1099,1277,1301]" pageId="1" pageNumber="650">
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||
<taxonomicNameLabel id="A20457B8E2219B7C2EFCF0CBFB81FA94" box="[1009,1099,1277,1301]" pageId="1" pageNumber="650" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C359655AE2219B7C2F3CF12BFD6DFAB4" box="[561,679,1309,1333]" pageId="1" pageNumber="650" type="description">
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<paragraph id="8BFC36D1E2219B7C2F3CF12BFD6DFAB4" blockId="1.[141,1099,1277,1333]" box="[561,679,1309,1333]" pageId="1" pageNumber="650">
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||
(
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<figureCitation id="13782A54E2219B7C2F34F12BFD4EFAB4" box="[569,644,1309,1333]" captionStart="Figure 1" captionStartId="2.[151,222,1695,1717]" captionTargetBox="[180,1186,1130,1653]" captionTargetId="figure-450@2.[151,1209,1104,1678]" captionTargetPageId="2" captionText="Figure 1a-c. Phyllophaga (s. str.) clavijobustosi sp. nov., male holotype. a-c) Dorsal, ventral, and lateral views. /Phyllophaga (s. str.) clavijobustosi sp. nov., holotipo macho. a-c) Vistas dorsal, ventral y lateral." figureDoi="http://doi.org/10.5281/zenodo.13206447" httpUri="https://zenodo.org/record/13206447/files/figure.png" pageId="1" pageNumber="650">Figs. 1</figureCitation>
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,
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<figureCitation id="13782A54E2219B7C2F9CF12BFD55FAB4" box="[657,671,1309,1333]" captionStart="Figures 2" captionStartId="3.[91,172,1111,1133]" captionTargetBox="[108,1136,123,1099]" captionTargetId="figure-273@3.[91,1149,118,1110]" captionTargetPageId="3" captionText="Figures 2a-j. Phyllophaga clavijobustosi sp. nov. a) Male antenna. b) External protarsal claw. c) Internal protarsal claw. d) External mesotarsal claw. e) Internal mesotarsal claw. f) Parameres, dorsal view. g) Parameres, lateral view. h) Internal sac of the aedeagus, lateral view. i) Parameres, ventral view. j) Gonocoxites, dorsal view. / a) Antena del macho. b) Uña protarsal externa. c) Uña protarsal interna. d) Uña mesotarsal externa. e) Uña mesotarsal interna. f) Parámeros, vista dorsal. g) Parámeros, vista lateral. h) Saco interno del edeago, vista lateral.i) Parámeros, vista ventral.j) Gonocoxitos, vista dorsal." figureDoi="http://doi.org/10.5281/zenodo.13206449" httpUri="https://zenodo.org/record/13206449/files/figure.png" pageId="1" pageNumber="650">2</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C359655AE2219B7C2D56F16AFBB6FA74" pageId="1" pageNumber="650" type="materials_examined">
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||
<paragraph id="8BFC36D1E2219B7C2D56F16AFBB6FA74" blockId="1.[90,1149,1367,1525]" pageId="1" pageNumber="650">
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||
<emphasis id="B937EAC3E2219B7C2D56F16AFE8FFAF5" bold="true" box="[91,325,1372,1396]" pageId="1" pageNumber="650">Material examined.</emphasis>
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||
<materialsCitation id="3B2B3C8CE2219B7C2C40F16BFE23FA54" collectingDate="2022-05-28" collectingDateMax="2022-05-29" collectingDateMin="2022-05-28" collectionCode="MIUP" collectorName="Male & M. Barria & W. Bayfield-Farrell & Female" country="Panama" elevation="840" latitude="8.539556" location="Division Continental" longLatPrecision="1" longitude="-81.15545" municipality="Santa Fe" pageId="1" pageNumber="650" specimenCount="2" stateProvince="Veraguas" typeStatus="holotype">
|
||
<collectorName id="26B65307E2219B7C2C40F16BFE42FAF4" box="[333,392,1373,1397]" pageId="1" pageNumber="650">Male</collectorName>
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||
<typeStatus id="54F88873E2219B7C2C82F16BFE3DFAF4" box="[399,503,1373,1397]" pageId="1" pageNumber="650" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED52AE14E2219B7C2F0BF16BFD9BFAF4" box="[518,593,1373,1397]" pageId="1" pageNumber="650">MIUP</collectionCode>
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||
), labeled:
|
||
<emphasis id="B937EAC3E2219B7C2FCBF16AFD19FAF5" bold="true" box="[710,723,1372,1396]" pageId="1" pageNumber="650">“</emphasis>
|
||
<collectingCountry id="F3547641E2219B7C2FDEF161FC9BFAF4" box="[723,849,1367,1397]" name="Panama" pageId="1" pageNumber="650">PANAMÁ</collectingCountry>
|
||
:
|
||
<collectingRegion id="4987F833E2219B7C2E56F16BFC00FAF4" box="[859,970,1373,1397]" country="Panama" name="Veraguas" pageId="1" pageNumber="650">Veraguas</collectingRegion>
|
||
: P. N.
|
||
<collectingMunicipality id="6B98ACABE2219B7C291EF16BFBB3FAF5" box="[1043,1145,1373,1397]" pageId="1" pageNumber="650">Santa Fe</collectingMunicipality>
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||
, cerca de la
|
||
<location id="8E9C600AE2219B7C2DD5F14BFE07FA14" LSID="urn:lsid:plazi:treatment:03EA87C7E2219B792D80F0CBFC7EFA96:8E9C600AE2219B7C2DD5F14BFE07FA14" box="[216,461,1405,1429]" country="Panama" latitude="8.539556" longLatPrecision="1" longitude="-81.15545" municipality="Santa Fe" name="Division Continental" pageId="1" pageNumber="650" stateProvince="Veraguas">División Continental</location>
|
||
,
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||
<quantity id="4CBB9B34E2219B7C2CDAF14BFDD7FA15" box="[471,541,1405,1429]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="1" pageNumber="650" unit="m" value="840.0">
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<elevation id="006ED1E2E2219B7C2CDAF14BFDD7FA15" box="[471,541,1405,1429]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="1" pageNumber="650" unit="m" value="840.0">840 m</elevation>
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||
</quantity>
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,
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<geoCoordinate id="EE775016E2219B7C2F2AF14BFD79FA14" box="[551,691,1405,1429]" degrees="8" direction="north" minutes="32" orientation="latitude" pageId="1" pageNumber="650" precision="1" seconds="22.4" value="8.539556">8°32’22.4”N</geoCoordinate>
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<geoCoordinate id="EE775016E2219B7C2FB5F14BFC9AFA14" box="[696,848,1405,1429]" degrees="81" direction="west" minutes="09" orientation="longitude" pageId="1" pageNumber="650" precision="1" seconds="19.6" value="-81.15545">81°09’19.6”W</geoCoordinate>
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,
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<date id="FFFD1011E2219B7C2E54F14BFBCAFA14" box="[857,1024,1405,1429]" pageId="1" pageNumber="650" value="2022-05-28" valueMax="2022-05-29" valueMin="2022-05-28">
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<collectingDate id="EFB9E9F9E2219B7C2E54F14BFBCAFA14" box="[857,1024,1405,1429]" pageId="1" pageNumber="650" value="2022-05-28" valueMax="2022-05-29" valueMin="2022-05-28">28-29/V/2022</collectingDate>
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||
</date>
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||
,
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||
<collectingMethod id="52024EC6E2219B7C2906F14BFE42FA35" pageId="1" pageNumber="650">Hg Metal Hallide and UV light trap</collectingMethod>
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||
,
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||
<collectorName id="26B65307E2219B7C2C9EF1ABFDCBFA34" box="[403,513,1437,1461]" pageId="1" pageNumber="650">M. Barria</collectorName>
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||
and
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||
<collectorName id="26B65307E2219B7C2F37F1ABFCDAFA34" box="[570,784,1437,1461]" pageId="1" pageNumber="650">W. Bayfield-Farrell</collectorName>
|
||
col.”.
|
||
<collectorName id="26B65307E2219B7C2E57F1ABFC64FA34" box="[858,942,1437,1461]" pageId="1" pageNumber="650">Female</collectorName>
|
||
<typeStatus id="54F88873E2219B7C2EB9F1ABFBD9FA34" box="[948,1043,1437,1461]" pageId="1" pageNumber="650" type="allotype">allotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED52AE14E2219B7C292CF1ABFBA6FA34" box="[1057,1132,1437,1461]" pageId="1" pageNumber="650">MIUP</collectionCode>
|
||
), labeled: same data as the holotype
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B2B3C8CE2219B7C2CF5F18BFBB3FA54" box="[504,1145,1469,1493]" collectingDate="2022-05-28" collectingDateMax="2022-05-29" collectingDateMin="2022-05-28" collectionCode="MDBC" collectorName="Male & M. Barria & W. Bayfield-Farrell & Female" country="Panama" elevation="840" latitude="8.539556" location="Division Continental" longLatPrecision="1" longitude="-81.15545" municipality="Santa Fe" pageId="1" pageNumber="650" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Veraguas" typeStatus="paratype">
|
||
<specimenCount id="9D45FD58E2219B7C2CF5F18BFD7EFA54" box="[504,692,1469,1493]" count="1" pageId="1" pageNumber="650" type="male" typeStatus="paratype">1 paratype male</specimenCount>
|
||
,
|
||
<specimenCount id="9D45FD58E2219B7C2FB3F18BFC47FA54" box="[702,909,1469,1493]" count="1" pageId="1" pageNumber="650" type="female" typeStatus="paratype">1 paratype female</specimenCount>
|
||
(
|
||
<collectionCode id="ED52AE14E2219B7C2E96F18BFC3BFA54" box="[923,1009,1469,1493]" pageId="1" pageNumber="650">MDBC</collectionCode>
|
||
): same data
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B2B3C8CE2219B7C2D57F1EBFE07FA74" box="[90,461,1501,1525]" collectingDate="2022-05-28" collectingDateMax="2022-05-29" collectingDateMin="2022-05-28" collectionCode="NHMUK" collectorName="Male & M. Barria & W. Bayfield-Farrell & Female" country="Panama" elevation="840" latitude="8.539556" location="Division Continental" longLatPrecision="1" longitude="-81.15545" municipality="Santa Fe" pageId="1" pageNumber="650" specimenCount="2" specimenCount-female="2" stateProvince="Veraguas" typeStatus="paratype">
|
||
<specimenCount id="9D45FD58E2219B7C2D57F1EBFF0CFA74" box="[90,198,1501,1525]" count="2" pageId="1" pageNumber="650" type="female">2 females</specimenCount>
|
||
(
|
||
<collectionCode id="ED52AE14E2219B7C2DDEF1EBFE8FFA74" box="[211,325,1501,1525]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="1" pageNumber="650" type="Museum">NHMUK</collectionCode>
|
||
): same data
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B2B3C8CE2219B7C2CD8F1EBFCFEFA74" box="[469,820,1501,1525]" collectingDate="2022-05-28" collectingDateMax="2022-05-29" collectingDateMin="2022-05-28" collectionCode="IAvH-E" collectorName="Male & M. Barria & W. Bayfield-Farrell & Female" country="Panama" elevation="840" latitude="8.539556" location="Division Continental" longLatPrecision="1" longitude="-81.15545" municipality="Santa Fe" pageId="1" pageNumber="650" specimenCount="3" specimenCount-female="3" stateProvince="Veraguas" typeStatus="paratype">
|
||
<specimenCount id="9D45FD58E2219B7C2CD8F1EBFD88FA74" box="[469,578,1501,1525]" count="3" pageId="1" pageNumber="650" type="female">3 females</specimenCount>
|
||
(
|
||
<collectionCode id="ED52AE14E2219B7C2F43F1EBFD66FA74" box="[590,684,1501,1525]" pageId="1" pageNumber="650">IAvH-E</collectionCode>
|
||
): same data
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B2B3C8CE2219B7C2E30F1EBFBB2FA74" box="[829,1144,1501,1525]" collectingDate="2022-05-28" collectingDateMax="2022-05-29" collectingDateMin="2022-05-28" collectionCode="CAS" collectorName="Male & M. Barria & W. Bayfield-Farrell & Female" country="Panama" elevation="840" latitude="8.539556" location="Division Continental" longLatPrecision="1" longitude="-81.15545" municipality="Santa Fe" pageId="1" pageNumber="650" specimenCount="2" specimenCount-female="2" stateProvince="Veraguas" typeStatus="paratype">
|
||
<specimenCount id="9D45FD58E2219B7C2E30F1EBFC60FA74" box="[829,938,1501,1525]" count="2" pageId="1" pageNumber="650" type="female">2 females</specimenCount>
|
||
(
|
||
<collectionCode id="ED52AE14E2219B7C2EBBF1EBFC3AFA74" box="[950,1008,1501,1525]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="1" pageNumber="650" type="Museum">CAS</collectionCode>
|
||
): same data
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359655AE2219B7E2D56F22AFCCFFA05" lastPageId="3" lastPageNumber="652" pageId="1" pageNumber="650" type="description">
|
||
<paragraph id="8BFC36D1E2219B7F2D56F22AFD74FCD4" blockId="1.[88,1150,1564,1749]" lastBlockId="2.[151,1210,125,1077]" lastPageId="2" lastPageNumber="651" pageId="1" pageNumber="650">
|
||
<emphasis id="B937EAC3E2219B7C2D56F22AFE64F9B4" bold="true" box="[91,430,1564,1589]" pageId="1" pageNumber="650">
|
||
Description. Male
|
||
<typeStatus id="54F88873E2219B7C2C4EF22BFE64F9B4" box="[323,430,1565,1589]" pageId="1" pageNumber="650" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13782A54E2219B7C2CCCF22BFDDAF9B4" box="[449,528,1565,1589]" captionStart="Figure 1" captionStartId="2.[151,222,1695,1717]" captionTargetBox="[180,1186,1130,1653]" captionTargetId="figure-450@2.[151,1209,1104,1678]" captionTargetPageId="2" captionText="Figure 1a-c. Phyllophaga (s. str.) clavijobustosi sp. nov., male holotype. a-c) Dorsal, ventral, and lateral views. /Phyllophaga (s. str.) clavijobustosi sp. nov., holotipo macho. a-c) Vistas dorsal, ventral y lateral." figureDoi="http://doi.org/10.5281/zenodo.13206447" httpUri="https://zenodo.org/record/13206447/files/figure.png" pageId="1" pageNumber="650">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="13782A54E2219B7C2F2CF22BFDE5F9B4" box="[545,559,1565,1589]" captionStart="Figures 2" captionStartId="3.[91,172,1111,1133]" captionTargetBox="[108,1136,123,1099]" captionTargetId="figure-273@3.[91,1149,118,1110]" captionTargetPageId="3" captionText="Figures 2a-j. Phyllophaga clavijobustosi sp. nov. a) Male antenna. b) External protarsal claw. c) Internal protarsal claw. d) External mesotarsal claw. e) Internal mesotarsal claw. f) Parameres, dorsal view. g) Parameres, lateral view. h) Internal sac of the aedeagus, lateral view. i) Parameres, ventral view. j) Gonocoxites, dorsal view. / a) Antena del macho. b) Uña protarsal externa. c) Uña protarsal interna. d) Uña mesotarsal externa. e) Uña mesotarsal interna. f) Parámeros, vista dorsal. g) Parámeros, vista lateral. h) Saco interno del edeago, vista lateral.i) Parámeros, vista ventral.j) Gonocoxitos, vista dorsal." figureDoi="http://doi.org/10.5281/zenodo.13206449" httpUri="https://zenodo.org/record/13206449/files/figure.png" pageId="1" pageNumber="650">2</figureCitation>
|
||
a-i). Length:
|
||
<quantity id="4CBB9B34E2219B7C2FC7F22BFCFFF9B5" box="[714,821,1565,1589]" metricMagnitude="-2" metricUnit="m" metricValue="2.31" pageId="1" pageNumber="650" unit="mm" value="23.1">23.1 mm</quantity>
|
||
. Humeral width:
|
||
<quantity id="4CBB9B34E2219B7C2902F22BFBB3F9B5" box="[1039,1145,1565,1589]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="1" pageNumber="650" unit="mm" value="10.2">10.2 mm</quantity>
|
||
. Width of pronotum:
|
||
<quantity id="4CBB9B34E2219B7C2C47F20BFE7BF9D5" box="[330,433,1597,1621]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="650" unit="mm" value="12.5">12.5 mm</quantity>
|
||
. Color: Dorsally reddish-brown, shiny. Head: Frons convex, glabrous, twice as wide as long, surface moderately punctuate, irregularly sparse; clypeus surface glabrous, three times wider than long, lateral margins rounded, apical margin sinuate slightly emarginate, almost straight; antenna with 10 antennomeres, antennal club longer than the previous seven (1:0.6) third antennomere semi-cylindrical and slightly elongate, fourth and fifth antennomeres shorter than the others, sixth more elongate posteriorly, seventh antennomere of compressed aspect, shorter than the sixth antennomere; antennal club almost twice as long as the preceding six antennomere; ocular canthus with at least 11 erect, rufous setae. Pronotum: Surface with punctures similar to the frons, glabrous, lateral margins entirely crenulated and with several erect setae; anterior angles obtuse. Scutellum: Wider than long, glabrous, surface with small, sparse and scattered punctures. Elytra: Surface generally glabrous, with a few very short setae near to the humeral and apical callus, and near to the lateral sides. Surface densely punctate, punctures separated from each other by 1-2 the puncture size and moderately deep. Apical callus poorly prominent, rounded. Pygidium: Plate convex, shiny; surface moderately and sparsely punctate, with short and rufous setae. Apical margin elevated and with some setae medially. Venter: Pro-, meso- and metasternum densely setigerous; sternites 2 to 4 with a distinct medial concavity; sternite 5 slightly convex medially, but almost inappreciable, with many erect setae in the apical border; sternite 6 (anal plate) broadly convex medially, apex distinctly sinuate and with several erect setae. Legs: Protibiae tridentate; protarsal claws with the internal denticle narrow and close to the base; external denticle longer and clearly separated from the internal denticle (
|
||
<figureCitation id="13782A54E2229B7F2C08F64BFE99FD14" box="[261,339,637,661]" captionStart="Figures 2" captionStartId="3.[91,172,1111,1133]" captionTargetBox="[108,1136,123,1099]" captionTargetId="figure-273@3.[91,1149,118,1110]" captionTargetPageId="3" captionText="Figures 2a-j. Phyllophaga clavijobustosi sp. nov. a) Male antenna. b) External protarsal claw. c) Internal protarsal claw. d) External mesotarsal claw. e) Internal mesotarsal claw. f) Parameres, dorsal view. g) Parameres, lateral view. h) Internal sac of the aedeagus, lateral view. i) Parameres, ventral view. j) Gonocoxites, dorsal view. / a) Antena del macho. b) Uña protarsal externa. c) Uña protarsal interna. d) Uña mesotarsal externa. e) Uña mesotarsal interna. f) Parámeros, vista dorsal. g) Parámeros, vista lateral. h) Saco interno del edeago, vista lateral.i) Parámeros, vista ventral.j) Gonocoxitos, vista dorsal." figureDoi="http://doi.org/10.5281/zenodo.13206449" httpUri="https://zenodo.org/record/13206449/files/figure.png" pageId="2" pageNumber="651">Figs. 2</figureCitation>
|
||
b-c); mesotarsal claws (
|
||
<figureCitation id="13782A54E2229B7F2F65F64BFD7DFD14" box="[616,695,637,661]" captionStart="Figures 2" captionStartId="3.[91,172,1111,1133]" captionTargetBox="[108,1136,123,1099]" captionTargetId="figure-273@3.[91,1149,118,1110]" captionTargetPageId="3" captionText="Figures 2a-j. Phyllophaga clavijobustosi sp. nov. a) Male antenna. b) External protarsal claw. c) Internal protarsal claw. d) External mesotarsal claw. e) Internal mesotarsal claw. f) Parameres, dorsal view. g) Parameres, lateral view. h) Internal sac of the aedeagus, lateral view. i) Parameres, ventral view. j) Gonocoxites, dorsal view. / a) Antena del macho. b) Uña protarsal externa. c) Uña protarsal interna. d) Uña mesotarsal externa. e) Uña mesotarsal interna. f) Parámeros, vista dorsal. g) Parámeros, vista lateral. h) Saco interno del edeago, vista lateral.i) Parámeros, vista ventral.j) Gonocoxitos, vista dorsal." figureDoi="http://doi.org/10.5281/zenodo.13206449" httpUri="https://zenodo.org/record/13206449/files/figure.png" pageId="2" pageNumber="651">Figs. 2</figureCitation>
|
||
d-e) with the internal denticle reduced and narrow, more reduced in the external claw (
|
||
<figureCitation id="13782A54E2229B7F2F9EF6ABFD28FD34" box="[659,738,669,693]" captionStart="Figures 2" captionStartId="3.[91,172,1111,1133]" captionTargetBox="[108,1136,123,1099]" captionTargetId="figure-273@3.[91,1149,118,1110]" captionTargetPageId="3" captionText="Figures 2a-j. Phyllophaga clavijobustosi sp. nov. a) Male antenna. b) External protarsal claw. c) Internal protarsal claw. d) External mesotarsal claw. e) Internal mesotarsal claw. f) Parameres, dorsal view. g) Parameres, lateral view. h) Internal sac of the aedeagus, lateral view. i) Parameres, ventral view. j) Gonocoxites, dorsal view. / a) Antena del macho. b) Uña protarsal externa. c) Uña protarsal interna. d) Uña mesotarsal externa. e) Uña mesotarsal interna. f) Parámeros, vista dorsal. g) Parámeros, vista lateral. h) Saco interno del edeago, vista lateral.i) Parámeros, vista ventral.j) Gonocoxitos, vista dorsal." figureDoi="http://doi.org/10.5281/zenodo.13206449" httpUri="https://zenodo.org/record/13206449/files/figure.png" pageId="2" pageNumber="651">Fig. 2e</figureCitation>
|
||
), internal denticle very close to the base; external denticle longer and clearly separated from the internal denticle, external denticles asymmetrical, upper denticle of the internal claw narrower and less recurved than the external denticle of the external claw. Genitalia (
|
||
<figureCitation id="13782A54E2229B7F2FC1F6CBFCD3FC94" box="[716,793,765,789]" captionStart="Figures 2" captionStartId="3.[91,172,1111,1133]" captionTargetBox="[108,1136,123,1099]" captionTargetId="figure-273@3.[91,1149,118,1110]" captionTargetPageId="3" captionText="Figures 2a-j. Phyllophaga clavijobustosi sp. nov. a) Male antenna. b) External protarsal claw. c) Internal protarsal claw. d) External mesotarsal claw. e) Internal mesotarsal claw. f) Parameres, dorsal view. g) Parameres, lateral view. h) Internal sac of the aedeagus, lateral view. i) Parameres, ventral view. j) Gonocoxites, dorsal view. / a) Antena del macho. b) Uña protarsal externa. c) Uña protarsal interna. d) Uña mesotarsal externa. e) Uña mesotarsal interna. f) Parámeros, vista dorsal. g) Parámeros, vista lateral. h) Saco interno del edeago, vista lateral.i) Parámeros, vista ventral.j) Gonocoxitos, vista dorsal." figureDoi="http://doi.org/10.5281/zenodo.13206449" httpUri="https://zenodo.org/record/13206449/files/figure.png" pageId="2" pageNumber="651">Figs. 2</figureCitation>
|
||
f-i): Male parameres fused dorsally; rounded apical lobes, with a short and recurved ventral process; internal sac of the aedeagus with the ring of spinules arranged horizontally.
|
||
</paragraph>
|
||
<paragraph id="8BFC36D1E2229B7E2DB0F76BFCCFFA05" blockId="2.[151,1210,125,1077]" lastBlockId="3.[91,1149,1324,1413]" lastPageId="3" lastPageNumber="652" pageId="2" pageNumber="651">
|
||
<emphasis id="B937EAC3E2229B7F2DB0F76BFE42FCF4" bold="true" box="[189,392,861,885]" pageId="2" pageNumber="651">
|
||
Female
|
||
<typeStatus id="54F88873E2229B7F2C2DF76BFE4EFCF4" box="[288,388,861,885]" pageId="2" pageNumber="651" type="allotype">allotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Similar to the male except: Length:
|
||
<quantity id="4CBB9B34E2229B7F2E42F76BFC71FCF5" box="[847,955,861,885]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="2" pageNumber="651" unit="mm" value="23.8">23.8 mm</quantity>
|
||
. Humeral width:
|
||
<quantity id="4CBB9B34E2229B7F2995F76BFF03FC15" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="2" pageNumber="651" unit="mm" value="9.6">9.6 mm</quantity>
|
||
. Width of elytra:
|
||
<quantity id="4CBB9B34E2229B7F2C9DF74BFE3CFC15" box="[400,502,893,917]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="2" pageNumber="651" unit="mm" value="11.6">11.6 mm</quantity>
|
||
. Antennal club almost as long as the preceding six antennal segments; tarsal claws unmodified as in males; abdominal sternites are not convex medially; apex of anal plate (sternite 6) without non-sinuate emargination; basal edge of pygidial plate slightly wider than in males. Genital plates well sclerotized (
|
||
<figureCitation id="13782A54E2229B7F2EBAF7EBFBC9FC74" box="[951,1027,989,1013]" captionStart="Figures 2" captionStartId="3.[91,172,1111,1133]" captionTargetBox="[108,1136,123,1099]" captionTargetId="figure-273@3.[91,1149,118,1110]" captionTargetPageId="3" captionText="Figures 2a-j. Phyllophaga clavijobustosi sp. nov. a) Male antenna. b) External protarsal claw. c) Internal protarsal claw. d) External mesotarsal claw. e) Internal mesotarsal claw. f) Parameres, dorsal view. g) Parameres, lateral view. h) Internal sac of the aedeagus, lateral view. i) Parameres, ventral view. j) Gonocoxites, dorsal view. / a) Antena del macho. b) Uña protarsal externa. c) Uña protarsal interna. d) Uña mesotarsal externa. e) Uña mesotarsal interna. f) Parámeros, vista dorsal. g) Parámeros, vista lateral. h) Saco interno del edeago, vista lateral.i) Parámeros, vista ventral.j) Gonocoxitos, vista dorsal." figureDoi="http://doi.org/10.5281/zenodo.13206449" httpUri="https://zenodo.org/record/13206449/files/figure.png" pageId="2" pageNumber="651">Fig. 2j</figureCitation>
|
||
); upper genital plates fused with the lower plates, rounded edges except for the apical edge of the upper plates which are slightly crenulate, plates surface with several short, erect and rufous setae.
|
||
<emphasis id="B937EAC3E2239B7E2D56F11AFF14FAC5" bold="true" box="[91,222,1324,1348]" pageId="3" pageNumber="652">Variations.</emphasis>
|
||
Male
|
||
<typeStatus id="54F88873E2239B7E2C2AF11BFE5BFAC4" box="[295,401,1325,1349]" pageId="3" pageNumber="652" type="paratype">paratype</typeStatus>
|
||
similar to the
|
||
<typeStatus id="54F88873E2239B7E2F37F11BFD6FFAC4" box="[570,677,1325,1349]" pageId="3" pageNumber="652" type="holotype">holotype</typeStatus>
|
||
, different only in size. Length:
|
||
<quantity id="4CBB9B34E2239B7E291FF11BFBB2FAC5" box="[1042,1144,1325,1349]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" pageId="3" pageNumber="652" unit="mm" value="22.3">22.3 mm</quantity>
|
||
; humeral width: 9.0 mm; width of elytra:
|
||
<quantity id="4CBB9B34E2239B7E2F3AF17BFD5AFAE5" box="[567,656,1357,1381]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="3" pageNumber="652" unit="mm" value="9.8">9.8 mm</quantity>
|
||
. Females
|
||
<typeStatus id="54F88873E2239B7E2E0CF17BFCB3FAE4" box="[769,889,1357,1381]" pageId="3" pageNumber="652" type="paratype">paratypes</typeStatus>
|
||
: length:
|
||
<quantity id="4CBB9B34E2239B7E2ED6F17BFBB2FAE5" box="[987,1144,1357,1381]" metricMagnitude="-2" metricUnit="m" metricValue="2.325" metricValueMax="2.38" metricValueMin="2.27" pageId="3" pageNumber="652" unit="mm" value="23.25" valueMax="23.8" valueMin="22.7">22.7-23.8 mm</quantity>
|
||
; humeral width: 9.0-
|
||
<quantity id="4CBB9B34E2239B7E2C4FF15BFE56FA05" box="[322,412,1389,1413]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="3" pageNumber="652" unit="mm" value="9.6">9.6 mm</quantity>
|
||
; width of elytra:
|
||
<quantity id="4CBB9B34E2239B7E2F69F15BFCCBFA05" box="[612,769,1389,1413]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" metricValueMax="1.16" metricValueMin="1.08" pageId="3" pageNumber="652" unit="mm" value="11.2" valueMax="11.6" valueMin="10.8">10.8-11.6 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3C6659E2229B7F2D9AF2A9FB70F953" ID-DOI="http://doi.org/10.5281/zenodo.13206447" ID-Zenodo-Dep="13206447" httpUri="https://zenodo.org/record/13206447/files/figure.png" pageId="2" pageNumber="651" startId="2.[151,222,1695,1717]" targetBox="[180,1186,1130,1653]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BFC36D1E2229B7F2D9AF2A9FB70F953" blockId="2.[151,1210,1695,1746]" pageId="2" pageNumber="651">
|
||
<emphasis id="B937EAC3E2229B7F2D9AF2A9FEDFF935" bold="true" box="[151,277,1695,1717]" pageId="2" pageNumber="651">Figure 1a-c.</emphasis>
|
||
<taxonomicName id="4C434D52E2229B7F2C16F2A9FDA3F935" authorityName="Barria & Quirós & Bayfield-Farrell" authorityYear="2023" box="[283,617,1695,1717]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="651" phylum="Arthropoda" rank="species" species="clavijobustosi" status="sp. nov.">
|
||
<emphasis id="B937EAC3E2229B7F2C16F2A9FE59F935" box="[283,403,1695,1716]" italics="true" pageId="2" pageNumber="651">Phyllophaga</emphasis>
|
||
(
|
||
<emphasis id="B937EAC3E2229B7F2CACF296FE60F934" box="[417,426,1696,1717]" italics="true" pageId="2" pageNumber="651">s</emphasis>
|
||
.
|
||
<emphasis id="B937EAC3E2229B7F2CBBF296FE1BF934" box="[438,465,1696,1717]" italics="true" pageId="2" pageNumber="651">str</emphasis>
|
||
.)
|
||
<emphasis id="B937EAC3E2229B7F2CE8F2A9FDA3F935" box="[485,617,1695,1716]" italics="true" pageId="2" pageNumber="651">clavijobustosi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B937EAC3E2229B7F2F62F2A9FD0AF935" bold="true" box="[623,704,1695,1716]" pageId="2" pageNumber="651">
|
||
<taxonomicNameLabel id="A20457B8E2229B7F2F62F2A9FD0AF935" box="[623,704,1695,1716]" pageId="2" pageNumber="651" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype.
|
||
<emphasis id="B937EAC3E2229B7F2E7CF2A9FC52F934" bold="true" box="[881,920,1695,1717]" pageId="2" pageNumber="651">a-c)</emphasis>
|
||
Dorsal, ventral, and lateral views. /
|
||
<taxonomicName id="4C434D52E2229B7F2DFEF28AFDF5F950" authorityName="Barria & Quirós & Bayfield-Farrell" authorityYear="2023" box="[243,575,1724,1746]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="651" phylum="Arthropoda" rank="species" species="clavijobustosi" status="sp. nov.">
|
||
<emphasis id="B937EAC3E2229B7F2DFEF28AFEA1F950" box="[243,363,1724,1745]" italics="true" pageId="2" pageNumber="651">Phyllophaga</emphasis>
|
||
(
|
||
<emphasis id="B937EAC3E2229B7F2C74F28BFE48F953" box="[377,386,1725,1746]" italics="true" pageId="2" pageNumber="651">s</emphasis>
|
||
.
|
||
<emphasis id="B937EAC3E2229B7F2C80F28BFE62F953" box="[397,424,1725,1746]" italics="true" pageId="2" pageNumber="651">str</emphasis>
|
||
.)
|
||
<emphasis id="B937EAC3E2229B7F2CB6F28AFDF5F950" box="[443,575,1724,1745]" italics="true" pageId="2" pageNumber="651">clavijobustosi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B937EAC3E2229B7F2F49F28BFD5EF953" bold="true" box="[580,660,1725,1746]" pageId="2" pageNumber="651">
|
||
<taxonomicNameLabel id="A20457B8E2229B7F2F49F28BFD5EF953" box="[580,660,1725,1746]" pageId="2" pageNumber="651" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotipo macho.
|
||
<emphasis id="B937EAC3E2229B7F2E5DF28AFCBDF953" bold="true" box="[848,887,1724,1746]" pageId="2" pageNumber="651">a-c)</emphasis>
|
||
Vistas dorsal, ventral y lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3C6659E2239B7E2D56F061FBB4FB7E" ID-DOI="http://doi.org/10.5281/zenodo.13206449" ID-Zenodo-Dep="13206449" httpUri="https://zenodo.org/record/13206449/files/figure.png" pageId="3" pageNumber="652" startId="3.[91,172,1111,1133]" targetBox="[108,1136,123,1099]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BFC36D1E2239B7E2D56F061FBB4FB7E" blockId="3.[91,1150,1110,1279]" pageId="3" pageNumber="652">
|
||
<emphasis id="B937EAC3E2239B7E2D56F061FF15FBEC" bold="true" box="[91,223,1111,1133]" pageId="3" pageNumber="652">Figures 2a-j.</emphasis>
|
||
<taxonomicName id="4C434D52E2239B7E2DE8F061FE2EFBED" authorityName="Barria & Quirós & Bayfield-Farrell" authorityYear="2023" box="[229,484,1111,1132]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="652" phylum="Arthropoda" rank="species" species="clavijobustosi" status="sp. nov.">
|
||
<emphasis id="B937EAC3E2239B7E2DE8F061FE2EFBED" box="[229,484,1111,1132]" italics="true" pageId="3" pageNumber="652">Phyllophaga clavijobustosi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B937EAC3E2239B7E2CE7F061FD9CFBED" bold="true" box="[490,598,1110,1132]" pageId="3" pageNumber="652">
|
||
<taxonomicNameLabel id="A20457B8E2239B7E2CE7F061FDF1FBED" box="[490,571,1111,1132]" pageId="3" pageNumber="652" rank="species">sp. nov.</taxonomicNameLabel>
|
||
a)
|
||
</emphasis>
|
||
Male antenna.
|
||
<emphasis id="B937EAC3E2239B7E2FF5F060FCC4FBED" bold="true" box="[760,782,1110,1132]" pageId="3" pageNumber="652">b)</emphasis>
|
||
External protarsal claw.
|
||
<emphasis id="B937EAC3E2239B7E291CF060FBEEFBED" bold="true" box="[1041,1060,1110,1132]" pageId="3" pageNumber="652">c)</emphasis>
|
||
Internal protarsal claw.
|
||
<emphasis id="B937EAC3E2239B7E2DF2F042FEDCFB0B" bold="true" box="[255,278,1140,1162]" pageId="3" pageNumber="652">d)</emphasis>
|
||
External mesotarsal claw.
|
||
<emphasis id="B937EAC3E2239B7E2F38F042FD80FB0B" bold="true" box="[565,586,1140,1162]" pageId="3" pageNumber="652">e)</emphasis>
|
||
Internal mesotarsal claw.
|
||
<emphasis id="B937EAC3E2239B7E2E68F042FCBCFB0B" bold="true" box="[869,886,1140,1162]" pageId="3" pageNumber="652">f)</emphasis>
|
||
Parameres, dorsal view.
|
||
<emphasis id="B937EAC3E2239B7E2D56F0A7FFBAFB26" bold="true" box="[91,112,1169,1191]" pageId="3" pageNumber="652">g)</emphasis>
|
||
Parameres, lateral view.
|
||
<emphasis id="B937EAC3E2239B7E2C7AF0A7FE47FB26" bold="true" box="[375,397,1169,1191]" pageId="3" pageNumber="652">h)</emphasis>
|
||
Internal sac of the aedeagus, lateral view.
|
||
<emphasis id="B937EAC3E2239B7E2E43F0A7FC97FB26" bold="true" box="[846,861,1169,1191]" pageId="3" pageNumber="652">i)</emphasis>
|
||
Parameres, ventral view.
|
||
<emphasis id="B937EAC3E2239B7E2960F0A7FBB7FB26" bold="true" box="[1133,1149,1169,1191]" pageId="3" pageNumber="652">j)</emphasis>
|
||
Gonocoxites, dorsal view. /
|
||
<emphasis id="B937EAC3E2239B7E2C89F098FE52FB45" bold="true" box="[388,408,1198,1220]" pageId="3" pageNumber="652">a)</emphasis>
|
||
Antena del macho.
|
||
<emphasis id="B937EAC3E2239B7E2F60F098FD4EFB45" bold="true" box="[621,644,1198,1220]" pageId="3" pageNumber="652">b)</emphasis>
|
||
Uña protarsal externa.
|
||
<emphasis id="B937EAC3E2239B7E2E76F098FC44FB45" bold="true" box="[891,910,1198,1220]" pageId="3" pageNumber="652">c)</emphasis>
|
||
Uña protarsal interna.
|
||
<emphasis id="B937EAC3E2239B7E2D56F0FAFFB8FB63" bold="true" box="[91,114,1228,1250]" pageId="3" pageNumber="652">d)</emphasis>
|
||
Uña mesotarsal externa.
|
||
<emphasis id="B937EAC3E2239B7E2C71F0FAFE5AFB63" bold="true" box="[380,400,1228,1250]" pageId="3" pageNumber="652">e)</emphasis>
|
||
Uña mesotarsal interna.
|
||
<emphasis id="B937EAC3E2239B7E2F95F0FAFD60FB63" bold="true" box="[664,682,1228,1250]" pageId="3" pageNumber="652">f)</emphasis>
|
||
Parámeros, vista dorsal.
|
||
<emphasis id="B937EAC3E2239B7E2EBEF0FAFC02FB63" bold="true" box="[947,968,1228,1250]" pageId="3" pageNumber="652">g)</emphasis>
|
||
Parámeros, vista lateral.
|
||
<emphasis id="B937EAC3E2239B7E2DAAF0DFFF77FB7E" bold="true" box="[167,189,1257,1279]" pageId="3" pageNumber="652">h)</emphasis>
|
||
Saco interno del edeago, vista lateral.
|
||
<emphasis id="B937EAC3E2239B7E2F4FF0DFFD98FB7E" bold="true" box="[578,594,1257,1279]" pageId="3" pageNumber="652">i)</emphasis>
|
||
Parámeros, vista ventral.
|
||
<emphasis id="B937EAC3E2239B7E2E50F0DFFCA7FB7E" bold="true" box="[861,877,1257,1279]" pageId="3" pageNumber="652">j)</emphasis>
|
||
Gonocoxitos, vista dorsal.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C359655AE2239B792D56F19AFDB8FF75" lastPageId="4" lastPageNumber="653" pageId="3" pageNumber="652" type="diagnosis">
|
||
<paragraph id="8BFC36D1E2239B792D56F19AFDB8FF75" blockId="3.[91,1149,1452,1733]" lastBlockId="4.[151,1209,125,245]" lastPageId="4" lastPageNumber="653" pageId="3" pageNumber="652">
|
||
<emphasis id="B937EAC3E2239B7E2D56F19AFF16FA45" bold="true" box="[91,220,1452,1476]" pageId="3" pageNumber="652">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C434D52E2239B7E2DEBF19BFDCDFA44" authorityName="Barria & Quirós & Bayfield-Farrell" authorityYear="2023" box="[230,519,1453,1477]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="652" phylum="Arthropoda" rank="species" species="clavijobustosi" status="sp. nov.">
|
||
<emphasis id="B937EAC3E2239B7E2DEBF19BFDCDFA44" box="[230,519,1453,1477]" italics="true" pageId="3" pageNumber="652">Phyllophaga clavijobustosi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B937EAC3E2239B7E2F1CF19BFDA5FA44" bold="true" box="[529,623,1453,1477]" pageId="3" pageNumber="652">
|
||
<taxonomicNameLabel id="A20457B8E2239B7E2F1CF19BFDA5FA44" box="[529,623,1453,1477]" pageId="3" pageNumber="652" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be mostly recognized by the following characters: basal denticle of external protarsal claw not exceeding width of base; upper denticle of external mesotarsal claw exceeding length of basal denticle and base, upper margin straight; lobes of parameres curved ventrally, with a commissure on the ventral margin; posterior margin of parameres sinuated. In
|
||
<taxonomicName id="4C434D52E2239B7E2FD4F21BFC46F9C4" authorityName="Barria & Quirós & Bayfield-Farrell" authorityYear="2023" box="[729,908,1581,1605]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="652" phylum="Arthropoda" rank="species" species="clavijobustosi">
|
||
<emphasis id="B937EAC3E2239B7E2FD4F21BFD23F9C5" box="[729,745,1581,1604]" italics="true" pageId="3" pageNumber="652">P</emphasis>
|
||
.
|
||
<emphasis id="B937EAC3E2239B7E2FF7F21BFC46F9C4" box="[762,908,1581,1605]" italics="true" pageId="3" pageNumber="652">clavijobustosi</emphasis>
|
||
</taxonomicName>
|
||
the ventral process of the parameres is very short and does not separate clearly, while in
|
||
<taxonomicName id="4C434D52E2239B7E2E9AF27BFF61F904" authority="Moron" authorityName="Moron" authorityYear="2013" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="652" phylum="Arthropoda" rank="species" species="cocleana">
|
||
<emphasis id="B937EAC3E2239B7E2E9AF27BFBB7F9E4" box="[919,1149,1613,1637]" italics="true" pageId="3" pageNumber="652">Phyllophaga cocleana</emphasis>
|
||
Morón
|
||
</taxonomicName>
|
||
the process is almost as long as the parameres and clearly separates; in
|
||
<taxonomicName id="4C434D52E2239B7E2EF5F25BFEC7F924" authority="Saylor" authorityName="Saylor" authorityYear="1935" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="652" phylum="Arthropoda" rank="species" species="vandykei">
|
||
<emphasis id="B937EAC3E2239B7E2EF5F25BFF71F924" italics="true" pageId="3" pageNumber="652">Phyllophaga vandykei</emphasis>
|
||
Saylor
|
||
</taxonomicName>
|
||
the process is almost twice as long as the parameres and is very recurved. However, the genitalia are more similar to those of
|
||
<taxonomicName id="4C434D52E2239B7E2FA2F29BFC1FF944" authority="(Moser)" baseAuthorityName="Moser" baseAuthorityYear="1918" box="[687,981,1709,1733]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="652" phylum="Arthropoda" rank="species" species="nuda">
|
||
<emphasis id="B937EAC3E2239B7E2FA2F29BFCB8F944" box="[687,882,1709,1733]" italics="true" pageId="3" pageNumber="652">Phyllophaga nuda</emphasis>
|
||
(Moser)
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F3547641E2239B7E2EEEF29BFB88F944" box="[995,1090,1709,1733]" name="Guyana" pageId="3" pageNumber="652">Guyana</collectingCountry>
|
||
) and
|
||
<taxonomicName id="4C434D52E2249B792D9AF44BFE1EFF14" authority="(Moser)" baseAuthorityName="Moser" baseAuthorityYear="1918" box="[151,468,125,149]" class="Insecta" family="Scarabaeidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="653" phylum="Arthropoda" rank="species" species="orosina">
|
||
<emphasis id="B937EAC3E2249B792D9AF44BFEBBFF15" box="[151,369,125,149]" italics="true" pageId="4" pageNumber="653">Phyllophaga orosina</emphasis>
|
||
(Moser)
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F3547641E2249B792CEFF44BFDAAFF14" box="[482,608,125,149]" name="Nicaragua" pageId="4" pageNumber="653">Nicaragua</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3547641E2249B792F66F44BFD2FFF14" box="[619,741,125,149]" name="Honduras" pageId="4" pageNumber="653">Honduras</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3547641E2249B792FFDF44BFCA7FF14" box="[752,877,125,149]" name="Costa Rica" pageId="4" pageNumber="653">Costa Rica</collectingCountry>
|
||
) but differ in the proportion and curvature of parameres lobes, and ventral process (length and how separated they are from the lobes). Females can be identified only by association with males and given the similarity of the ventral setiferous cover.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359655AE2249B792D9AF52AFB50FBF6" pageId="4" pageNumber="653" type="distribution">
|
||
<paragraph id="8BFC36D1E2249B792D9AF52AFB50FBF6" blockId="4.[151,1055,284,309]" lastBlockId="4.[151,1178,1118,1143]" pageId="4" pageNumber="653">
|
||
<emphasis id="B937EAC3E2249B792D9AF52AFEFCFEB5" bold="true" box="[151,310,284,308]" pageId="4" pageNumber="653">Distribution.</emphasis>
|
||
Parque Nacional Santa Fe,
|
||
<collectingRegion id="4987F833E2249B792F75F52BFC9CFEB4" box="[632,854,285,309]" country="Panama" name="Veraguas" pageId="4" pageNumber="653">Veraguas province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3547641E2249B792E6CF52BFC0AFEB5" box="[865,960,285,308]" name="Panama" pageId="4" pageNumber="653">Panama</collectingCountry>
|
||
(
|
||
<figureCitation id="13782A54E2249B792EC2F52BFBDBFEB4" box="[975,1041,285,309]" captionStart="Figure 3" captionStartId="4.[151,222,1009,1031]" captionTargetBox="[151,1209,357,996]" captionTargetId="figure-94@4.[151,1209,357,999]" captionTargetPageId="4" captionText="Figure 3. Distribution map of the Panamanian species of the orosina species group. / Mapa de distribución de las especies panameñas del grupo de especies orosina." figureDoi="http://doi.org/10.5281/zenodo.13206451" httpUri="https://zenodo.org/record/13206451/files/figure.png" pageId="4" pageNumber="653">Fig. 3</figureCitation>
|
||
).
|
||
<emphasis id="B937EAC3E2249B792D9AF068FF30FBF7" bold="true" box="[151,250,1118,1142]" pageId="4" pageNumber="653">Habitat.</emphasis>
|
||
The specimens were collected in a tropical lower montane moist forest (
|
||
<figureCitation id="13782A54E2249B792947F069FB47FBF6" box="[1098,1165,1119,1143]" captionStart="Figure 4" captionStartId="5.[91,162,823,845]" captionTargetBox="[91,1148,182,816]" captionTargetId="figure-281@5.[90,1148,118,818]" captionTargetPageId="5" captionText="Figure 4. Habitat of Phyllophaga (Phyllophaga) clavijobustosi sp. nov. Tropical Lower Montane Moist Forest in P. N. Santa Fe (near to the continental divide), Veraguas prov., Panama. Photography by Manuel D. Barria. / Hábitat de Phyllophaga (Phyllophaga) clavijobustosi sp. nov. Bosque Húmedo Montano Bajo Tropical en el P. N. Santa Fe (cerca de la división continental), provincia de Veraguas, Panamá. FotografÍa de Manuel D. BarrÍa." figureDoi="http://doi.org/10.5281/zenodo.13206453" httpUri="https://zenodo.org/record/13206453/files/figure.png" pageId="4" pageNumber="653">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3C6659E2249B792D9AF7C7FCB7FBA5" ID-DOI="http://doi.org/10.5281/zenodo.13206451" ID-Zenodo-Dep="13206451" httpUri="https://zenodo.org/record/13206451/files/figure.png" pageId="4" pageNumber="653" startId="4.[151,222,1009,1031]" targetBox="[151,1209,357,996]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BFC36D1E2249B792D9AF7C7FCB7FBA5" blockId="4.[151,1209,1009,1060]" pageId="4" pageNumber="653">
|
||
<emphasis id="B937EAC3E2249B792D9AF7C7FF30FB87" bold="true" box="[151,250,1009,1031]" pageId="4" pageNumber="653">Figure 3.</emphasis>
|
||
Distribution map of the Panamanian species of the orosina species group. / Mapa de distribución de las especies panameñas del grupo de especies orosina.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C359655AE2249B792D9AF0A9FC7EFA96" pageId="4" pageNumber="653" type="etymology">
|
||
<paragraph id="8BFC36D1E2249B792D9AF0A9FC7EFA96" blockId="4.[151,1210,1183,1303]" pageId="4" pageNumber="653">
|
||
<emphasis id="B937EAC3E2249B792D9AF0A9FED5FB36" bold="true" box="[151,287,1183,1207]" pageId="4" pageNumber="653">Etymology.</emphasis>
|
||
The species is named in honor of MDB’s friend Julian Clavijo-Bustos (
|
||
<collectingCountry id="F3547641E2249B79293BF0A9FB67FB36" box="[1078,1197,1183,1207]" name="Colombia" pageId="4" pageNumber="653">Colombia</collectingCountry>
|
||
), as a thank you for his friendship all these years and in recognition of his great potential as a researcher and great knowledge in the
|
||
<taxonomicName id="4C434D52E2249B792F72F0E9FCE2FB76" authorityName="Latreille" authorityYear="1802" box="[639,808,1247,1271]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="654" phylum="Arthropoda" rank="superFamily" superFamily="Scarabaeoidea">Scarabaeoidea</taxonomicName>
|
||
superfamily, especially in groups such as Aphodiinae,
|
||
<taxonomicName id="4C434D52E2249B792C83F0C9FDE3FA96" authorityName="Erichson" authorityYear="1847" box="[398,553,1279,1303]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="653" phylum="Arthropoda" rank="family">Hybosoridae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C434D52E2249B792F39F0C9FD29FA96" box="[564,739,1279,1303]" class="Insecta" family="Bolboceratidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="653" phylum="Arthropoda" rank="family">Bolboceratidae</taxonomicName>
|
||
and Ochodaidae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |