166 lines
21 KiB
XML
166 lines
21 KiB
XML
<document id="A566E85D39CD48D38F28AB703CF20E5B" ID-DOI="10.11646/zootaxa.4088.3.1" ID-GBIF-Dataset="c09a5f42-c1ac-4b6c-a98d-d836a56711db" ID-ISSN="1175-5326" ID-Zenodo-Dep="267464" ID-ZooBank="D055AB86-A0A2-4E30-8671-4B0990C478FB" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457597009995" checkinUser="plazi" docAuthor="Komai, Tomoyuki & Chan, Tin-Yam" docDate="2016" docId="03D78781E243FF87D4A4FA4A0F3A9A4D" docLanguage="en" docName="15575-5104-1-PB.pdf.imf" docOrigin="Zootaxa 4088 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trizocheles sakaii Forest 1987" docType="treatment" docVersion="11" lastPageNumber="322" masterDocId="FFEEFFF9E250FF92D433FFE20D6A982D" masterDocTitle="“ Symmetrical ” hermit crabs of the family Pylochelidae (Crustacea: Decapoda: Anomura) collected by the “ BIOPAPUA ” and “ PAPUA NIUGINI ” expeditions in the Papua New Guinea, with descriptions of two new species" masterLastPageNumber="328" masterPageNumber="301" pageNumber="320" updateTime="1698177513233" updateUser="plazi">
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<mods:title id="6D4F86EC4EFEDCF09AA10F828BFF8E7C">“ Symmetrical ” hermit crabs of the family Pylochelidae (Crustacea: Decapoda: Anomura) collected by the “ BIOPAPUA ” and “ PAPUA NIUGINI ” expeditions in the Papua New Guinea, with descriptions of two new species</mods:title>
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<mods:namePart id="8094E5B2E59A9C9AA72ED61776829C5E">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="0D37248B65EE66E0AD12F23257AA09DA">Chan, Tin-Yam</mods:namePart>
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<mods:typeOfResource id="107C14BF60FE37BFC8C00F11012B3B4B">text</mods:typeOfResource>
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<mods:title id="5F290A3E23E7F02D9D790BF642398476">Zootaxa</mods:title>
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<mods:date id="1FE59276E0F328F58D0CE5EEDA1130A2">2016</mods:date>
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<mods:detail id="928060B1B912A8FA6BD6ABB50C55BFC6" type="volume">
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<mods:number id="BB60EFEF584723960DF7691264615F45">4088</mods:number>
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<mods:identifier id="51F8DEADBC46B7385346EA221761EF56" type="DOI">10.11646/zootaxa.4088.3.1</mods:identifier>
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<treatment id="03D78781E243FF87D4A4FA4A0F3A9A4D" ID-DOI="http://doi.org/10.5281/zenodo.6076083" ID-GBIF-Taxon="125783248" ID-Zenodo-Dep="6076083" LSID="urn:lsid:plazi:treatment:03D78781E243FF87D4A4FA4A0F3A9A4D" httpUri="http://treatment.plazi.org/id/03D78781E243FF87D4A4FA4A0F3A9A4D" lastPageId="21" lastPageNumber="322" pageId="19" pageNumber="320">
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<subSubSection id="C364651CE243FF81D4A4FA4A0C929E4B" pageId="19" pageNumber="320" type="nomenclature">
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<paragraph id="8BC13697E243FF81D4A4FA4A0F499DEF" blockId="19.[151,547,1448,1507]" box="[151,547,1448,1474]" pageId="19" pageNumber="320">
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<heading id="D08981FBE243FF81D4A4FA4A0F499DEF" bold="true" box="[151,547,1448,1474]" fontSize="11" level="1" pageId="19" pageNumber="320" reason="1">
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<taxonomicName id="4C7E4D14E243FF81D4A4FA4A0F499DEF" ID-CoL="3d4e9cb7-857f-4b84-9f29-7c2c9e6e59d6" authority="Forest, 1987" authorityName="Forest" authorityYear="1987" box="[151,547,1448,1474]" class="Malacostraca" family="Pylochelidae" genus="Trizocheles" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="sakaii">
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<emphasis id="B90AEA85E243FF81D4A4FA4A0F499DEF" bold="true" box="[151,547,1448,1474]" pageId="19" pageNumber="320">
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<emphasis id="B90AEA85E243FF81D4A4FA4A0C049DEF" bold="true" box="[151,366,1448,1474]" italics="true" pageId="19" pageNumber="320">Trizocheles sakaii</emphasis>
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(Forest, 1987)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BC13697E243FF81D4A4FA280D979DCE" blockId="19.[151,547,1448,1507]" box="[151,253,1482,1507]" pageId="19" pageNumber="320">
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(
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<figureCitation id="13452A12E243FF81D4ACFA280D8B9DCE" box="[159,225,1482,1507]" captionStart="FIGURE 5. A" captionStartId="20.[151,250,1721,1742]" captionTargetBox="[151,1436,913,1698]" captionTargetId="figure@20.[151,1436,913,1699]" captionTargetPageId="20" captionText="FIGURE 5. A, Trizocheles manningi Forest, 1987, entire animal in dorsal view, BIOPAPUA, stn DW 3732, ovigerous female (sl 4.6 mm), MNHN-IU- 2011 - 1039; B, Trizocheles sakaii Forest, 1987, stn DW 3745, males (sl 4.1 mm), MNHN-IU- 2011 - 3490." httpUri="https://zenodo.org/record/267469/files/figure.png" pageId="19" pageNumber="320">Fig. 5</figureCitation>
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B)
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</paragraph>
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<paragraph id="8BC13697E243FF81D4A4F9F00C929E4B" blockId="19.[151,1436,1554,1638]" pageId="19" pageNumber="320">
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<taxonomicName id="4C7E4D14E243FF81D4A4F9F00F7B9E05" authority="Forest, 1987a: 189" authorityName="Forest" authorityPageNumber="189" authorityYear="1987" box="[151,529,1554,1576]" class="Malacostraca" family="Pylochelidae" genus="Trizocheles" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="sakaii">
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<emphasis id="B90AEA85E243FF81D4A4F9F00C279E05" box="[151,333,1554,1576]" italics="true" pageId="19" pageNumber="320">Trizocheles sakaii</emphasis>
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Forest, 1987a: 189
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</taxonomicName>
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, figs. 46a–h, 48d, e, 61b, 63d, 64, pls. 5A, B, F, G, 8A, B [type locality: Tosa Bay, Japan]; 1987b: 315, fig. 2.—McLaughlin
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<emphasis id="B90AEA85E243FF81D617F9D00F389E6A" box="[548,594,1585,1607]" italics="true" pageId="19" pageNumber="320">et al</emphasis>
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. 2007: 36, unnumbered figs.—Lemaitre
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<emphasis id="B90AEA85E243FF81D7C4F9D0094F9E6A" box="[1015,1061,1585,1607]" italics="true" pageId="19" pageNumber="320">et al</emphasis>
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. 2009: 5.—McLaughlin & Lemaitre 2009: 214.—Komai 2013: 92.
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</paragraph>
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</subSubSection>
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<subSubSection id="C364651CE243FF86D4A4F97009AC9AC2" lastPageId="20" lastPageNumber="321" pageId="19" pageNumber="320" type="materials_examined">
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<paragraph id="8BC13697E243FF81D4A4F97009EB9F8A" blockId="19.[151,1436,1682,2031]" pageId="19" pageNumber="320">
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<emphasis id="B90AEA85E243FF81D4A4F9700CEB9E86" bold="true" box="[151,385,1682,1707]" pageId="19" pageNumber="320">Material examined</emphasis>
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. BIOPAPUA, stn CP 3691, SE of Manus Island, 02°11’S, 147°18’E, 499–517 m, 29 September 2010, 1 female (sl 6.1 mm), MNHN-IU-2011-3586; stn CP 3692, SE of Manus Island, 02°10’S, 147°19’E, 408–448 m, 29 September 2010, 1 male (sl 6.0 mm), 2 females (sl 2.2, 2.9 mm), MNHN-IU-2011-2784; stn CP 3707, 04°59’S, 145°50’E, 460–466 m, 2 October 2010, 1 male (sl 10.2 mm), MNHN-IU-2011-1947; same data, 1 female (sl 5.6 mm), MNHN-IU-2011-1658; stn DW 3745, off Bougainville Island, 05°33’S, 154°00’E, 369–377 m, 12 October 2010, 2 males (sl 4.1, 4.1 mm), 3 ovigerous females (sl 4.5–5.0 mm), MNHN-IU-2011- 3490; stn DW 3748, off Bougainville Island, 05°37’S, 154°01’E, 398–399 m, 12 October 2010, 3 males (sl 4.9–5.6 mm), 1 female (sl 4.0 mm), 2 ovigerous females (sl 4.8, 5.0 mm), MNHN-IU-2011-5062.
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</paragraph>
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<paragraph id="8BC13697E243FF86D4F4F8510EB39981" blockId="19.[151,1436,1682,2031]" lastBlockId="20.[151,1436,151,861]" lastPageId="20" lastPageNumber="321" pageId="19" pageNumber="320">PAPUA NIUGINI, stn CP 3970, W of Karkar Island, Bismarck Sea, 04°39’S, 145°52’E, 573 m, 4 December 2012, 2 females (sl 4.3, 4.3 mm), MNHN-IU-2013-13210; same data, 1 male (sl 5.8 mm), MNHN-IU-2013-14436; stn CP 3978, N of Bagabag Island, Bismarck Sea, 04°49’S, 146°12’E, 456–582 m, 5 December 2012, 7 males (sl 4.6–10.0 mm), 3 females (sl 3.4–7.3 mm), MNHN-IU-2013-14434; stn CP 3979, N of Bagabag Island, 04°44’S, 148°11’E, 540–580 m, 5 December 2012, 2 males (sl 5.1, 9.2 mm), 1 ovigerous female (sl 7.3 mm), MNHN-IU- 2013-14430; same data, 1 female (sl 4.6 mm), MNHN-IU-2013-14438; stn CP 3994, Vitjaz Strait, 06°00’S, 147°35’E, 645–652 m, 9 December 2012, 1 male (sl 5.3 mm), 1 ovigerous female (sl 5.5 mm), MNHN-IU-2013- 14435; stn CP 4006, SE of Tuam Island, Solomon Sea, 06°03’N, 148°08’E, 440–475 m, 11 December 2012, 1 female (sl 5.1 mm), MNHN-IU-2013-14432; stn CP 4009, SE of Tuam Island, 06°04’S, 148°12’E, 550–575 m, 11 December 2012, 1 ovigerous female (sl 4.8 mm), MNHN-IU-2014-12695.</paragraph>
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<paragraph id="8BC13697E244FF86D4F4FE550CDC9A4D" blockId="20.[151,1436,151,861]" pageId="20" pageNumber="321">Comparative material. Off Shionomisaki, Kii Peninsula, Japan, 250 m, 15 January 1990, dredge, coll. S. Nagai, 1 male (sl 4.7 mm), CBM-ZC 1084; same locality, 200 m, 19 December 1992, dredge, coll. S. Nagai, 1 ovigerous female (sl 5.1 mm), CBM-ZC 1182; same locality, 250 m, 22 March 1992, dredge, coll. S. Nagai, 1 female (sl 3.9 mm), CBM-ZC 1184; same locality, 300 m, October 1996, dredge, coll. S. Nagai, 1 male (sl 3.7 mm), 1 female (sl 4.0 mm), CBM-ZC 3561.</paragraph>
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<paragraph id="8BC13697E244FF86D4F4FD8E0FFF9AE1" blockId="20.[151,1436,151,861]" pageId="20" pageNumber="321">TAIWAN 2000, stn DW 56, 24°29.8’N, 122°12.6’E, 438–539 m, 4 August 2000, 2 males (cl 4.3, 6.6 mm), 1 female (cl 4.2 mm), NTOU. TAIWAN 2004, stn CP 269, 24°30.55’N, 122º05.78’E, 2 September 2004, 397–399 m, 2 ovigerous females (sl 4.9, 5.7 mm), NTOU.</paragraph>
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<paragraph id="8BC13697E244FF86D4F4FD3509AC9AC2" blockId="20.[151,1436,151,861]" box="[199,1222,727,752]" pageId="20" pageNumber="321">
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<emphasis id="B90AEA85E244FF86D4F4FD350D9D9ADD" bold="true" box="[199,247,727,752]" pageId="20" pageNumber="321">Size</emphasis>
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. Largest male sl 10.2 mm, largest female sl 7.3 mm, ovigerous females sl 4.8–7.3 mm.
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</paragraph>
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</subSubSection>
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<subSubSection id="C364651CE244FF86D4F4FD1E0EED9B70" pageId="20" pageNumber="321" type="description">
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<paragraph id="8BC13697E244FF86D4F4FD1E0EED9B70" blockId="20.[151,1436,151,861]" pageId="20" pageNumber="321">
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<emphasis id="B90AEA85E244FF86D4F4FD1E0C1B9B38" bold="true" box="[199,369,764,789]" pageId="20" pageNumber="321">Colour in life</emphasis>
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. Shield, ocular peduncles, antennular peduncles, chelipeds and ambulatory legs generally orange, posterior carapace and pleon paler; ocular acicles and antennal peduncles whitish; meri of chelipeds and ambulatory legs each with white transverse band distally (
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<figureCitation id="13452A12E244FF86D711FCA60E0E9B70" box="[802,868,836,861]" captionStart="FIGURE 5. A" captionStartId="20.[151,250,1721,1742]" captionTargetBox="[151,1436,913,1698]" captionTargetId="figure@20.[151,1436,913,1699]" captionTargetPageId="20" captionText="FIGURE 5. A, Trizocheles manningi Forest, 1987, entire animal in dorsal view, BIOPAPUA, stn DW 3732, ovigerous female (sl 4.6 mm), MNHN-IU- 2011 - 1039; B, Trizocheles sakaii Forest, 1987, stn DW 3745, males (sl 4.1 mm), MNHN-IU- 2011 - 3490." httpUri="https://zenodo.org/record/267469/files/figure.png" pageId="20" pageNumber="321">Fig. 5</figureCitation>
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B).
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</paragraph>
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</subSubSection>
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<caption id="DF01661FE244FF86D4A4F95B0DBB9F26" httpUri="https://zenodo.org/record/267469/files/figure.png" pageId="20" pageNumber="321" targetBox="[151,1436,913,1698]" targetPageId="20">
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<paragraph id="8BC13697E244FF86D4A4F95B0DBB9F26" blockId="20.[151,1436,1720,1803]" pageId="20" pageNumber="321">
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<emphasis id="B90AEA85E244FF86D4A4F95B0C679EE3" bold="true" box="[151,269,1721,1742]" pageId="20" pageNumber="321">FIGURE 5</emphasis>
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. A,
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<taxonomicName id="4C7E4D14E244FF86D50AF95A0FF19EE3" authority="Forest, 1987" authorityName="Forest" authorityYear="1987" box="[313,667,1720,1742]" class="Malacostraca" family="Pylochelidae" genus="Trizocheles" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="manningi">
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<emphasis id="B90AEA85E244FF86D50AF95A0F799EE3" box="[313,531,1720,1742]" italics="true" pageId="20" pageNumber="321">Trizocheles manningi</emphasis>
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Forest, 1987
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</taxonomicName>
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, entire animal in dorsal view, BIOPAPUA, stn DW 3732, ovigerous female (sl 4.6 mm), MNHN-IU-2011-1039; B,
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<taxonomicName id="4C7E4D14E244FF86D607F9350E1D9EC0" authority="Forest, 1987" authorityName="Forest" authorityYear="1987" box="[564,887,1751,1773]" class="Malacostraca" family="Pylochelidae" genus="Trizocheles" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="sakaii">
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<emphasis id="B90AEA85E244FF86D607F9350F819EC0" box="[564,747,1751,1773]" italics="true" pageId="20" pageNumber="321">Trizocheles sakaii</emphasis>
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Forest, 1987
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</taxonomicName>
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, stn DW 3745, males (sl 4.1 mm), MNHN-IU-2011- 3490.
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</paragraph>
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</caption>
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<subSubSection id="C364651CE244FF86D4F4F8D10F2E9F95" pageId="20" pageNumber="321" type="distribution">
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<paragraph id="8BC13697E244FF86D4F4F8D10CCD9FBE" blockId="20.[151,1436,1843,2012]" pageId="20" pageNumber="321">
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<emphasis id="B90AEA85E244FF86D4F4F8D10C339F61" bold="true" box="[199,345,1843,1868]" pageId="20" pageNumber="321">Distribution</emphasis>
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. Previously known from Japan and Taiwan, at depths of 200–539 m. The present material greatly extends the geographical range of this species to the south; in waters around Papua New Guinea, this species occurs at depths of 369–652 m.
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</paragraph>
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<paragraph id="8BC13697E244FF86D4F4F87D0F2E9F95" blockId="20.[151,1436,1843,2012]" box="[199,580,1951,1976]" pageId="20" pageNumber="321">
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<emphasis id="B90AEA85E244FF86D4F4F87D0C499F95" bold="true" box="[199,291,1951,1976]" pageId="20" pageNumber="321">Habitat</emphasis>
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. Scaphopod (tusk) shells.
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</paragraph>
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</subSubSection>
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<subSubSection id="C364651CE244FF87D4F4F8260F3A9A4D" lastPageId="21" lastPageNumber="322" pageId="20" pageNumber="321" type="discussion">
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<paragraph id="8BC13697E244FF87D4F4F82609F89980" blockId="20.[151,1436,1843,2012]" lastBlockId="21.[151,1437,151,608]" lastPageId="21" lastPageNumber="322" pageId="20" pageNumber="321">
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<emphasis id="B90AEA85E244FF86D4F4F8260C5E9FF6" bold="true" box="[199,308,1988,2011]" pageId="20" pageNumber="321">Remarks</emphasis>
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. The present specimens are identified with
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<taxonomicName id="4C7E4D14E244FF86D744F82109279FF1" box="[887,1101,1987,2012]" class="Malacostraca" family="Pylochelidae" genus="Trizocheles" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="sakaii">
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<emphasis id="B90AEA85E244FF86D744F82109279FF1" box="[887,1101,1987,2012]" italics="true" pageId="20" pageNumber="321">Trizocheles sakaii</emphasis>
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</taxonomicName>
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because of the following characteristics: palms of chelipeds armed with two to four rows of spines; carpi of chelipeds with stridulating rods or tubercles; propodi of second pereopods moderately stout (3.6 times as long as wide or less), armed only with dorsodistal spine, otherwise unarmed. Comparison with the material from Japan and Taiwan failed to detect significant morphological differences from the Papua New Guinea specimens, although there is a wide geographical gap between Papua New Guinea and the Japanese Archipelago. With regard to the living coloration, minor differences are seen: the male specimen from stn DW 3745 (MNHN-IU-2011-3490) has white distal bands on the meri of the chelipeds and ambulatory legs (
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<figureCitation id="13452A12E245FF87D6E3FE8D0E7999A5" box="[720,787,367,392]" captionStart="FIGURE 5. A" captionStartId="20.[151,250,1721,1742]" captionTargetBox="[151,1436,913,1698]" captionTargetId="figure@20.[151,1436,913,1699]" captionTargetPageId="20" captionText="FIGURE 5. A, Trizocheles manningi Forest, 1987, entire animal in dorsal view, BIOPAPUA, stn DW 3732, ovigerous female (sl 4.6 mm), MNHN-IU- 2011 - 1039; B, Trizocheles sakaii Forest, 1987, stn DW 3745, males (sl 4.1 mm), MNHN-IU- 2011 - 3490." httpUri="https://zenodo.org/record/267469/files/figure.png" pageId="21" pageNumber="322">Fig. 5</figureCitation>
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B), whereas such white bands are not seen on the meri of the chelipeds in the specimen from Taiwan (McLaughlin
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<emphasis id="B90AEA85E245FF87D72AFE770E209980" box="[793,842,404,429]" italics="true" pageId="21" pageNumber="322">et al</emphasis>
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. 2007: 36, unnumbered fig.).
|
||
</paragraph>
|
||
<paragraph id="8BC13697E245FF87D4F4FE5A0F3A9A4D" blockId="21.[151,1437,151,608]" pageId="21" pageNumber="322">
|
||
Forest (1987a) attributed all records of
|
||
<taxonomicName id="4C7E4D14E245FF87D64DFE550E3599E2" box="[638,863,439,464]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="spinosus">
|
||
<emphasis id="B90AEA85E245FF87D64DFE550E3599E2" box="[638,863,439,464]" italics="true" pageId="21" pageNumber="322">Pylocheles spinosus</emphasis>
|
||
</taxonomicName>
|
||
from Japanese waters (Ortmann 1892; Balss 1913; Yokoya 1933; Miyake 1978, 1982) to
|
||
<taxonomicName id="4C7E4D14E245FF87D60CFE3E0E6299D8" box="[575,776,476,501]" class="Malacostraca" family="Pylochelidae" genus="Trizocheles" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="sakaii">
|
||
<emphasis id="B90AEA85E245FF87D60CFE3E0E6299D8" box="[575,776,476,501]" italics="true" pageId="21" pageNumber="322">Trizocheles sakaii</emphasis>
|
||
</taxonomicName>
|
||
, and this was followed by McLaughlin & Lemaitre (2009). Komai (2013) clarified this by noting that Ortmann’s (1892) specimen actually represented
|
||
<taxonomicName id="4C7E4D14E245FF87D0AFFDE2084C9A35" box="[1180,1318,511,536]" class="Malacostraca" family="Pylochelidae" genus="Trizocheles" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="albatrossi">
|
||
<emphasis id="B90AEA85E245FF87D0AFFDE2084C9A35" box="[1180,1318,511,536]" italics="true" pageId="21" pageNumber="322">T. albatrossi</emphasis>
|
||
</taxonomicName>
|
||
instead of
|
||
<taxonomicName id="4C7E4D14E245FF87D4A4FDC70D9F9A10" box="[151,245,548,573]" class="Malacostraca" family="Pylochelidae" genus="Trizocheles" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="sakaii">
|
||
<emphasis id="B90AEA85E245FF87D4A4FDC70D9F9A10" box="[151,245,548,573]" italics="true" pageId="21" pageNumber="322">T. sakaii</emphasis>
|
||
</taxonomicName>
|
||
, and that two species,
|
||
<taxonomicName id="4C7E4D14E245FF87D5C7FDC70F149A10" box="[500,638,548,573]" class="Malacostraca" family="Pylochelidae" genus="Trizocheles" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="albatrossi">
|
||
<emphasis id="B90AEA85E245FF87D5C7FDC70F149A10" box="[500,638,548,573]" italics="true" pageId="21" pageNumber="322">T. albatrossi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D14E245FF87D686FDC60EC79A10" box="[693,941,548,573]" class="Malacostraca" family="Pylochelidae" genus="Pomatocheles" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="jeffreisi">
|
||
<emphasis id="B90AEA85E245FF87D686FDC60EC79A10" box="[693,941,548,573]" italics="true" pageId="21" pageNumber="322">Pomatocheles jeffreisi</emphasis>
|
||
</taxonomicName>
|
||
, were mixed up in the material identified as
|
||
<taxonomicName id="4C7E4D14E245FF87D4A4FDA50C1D9A72" box="[151,375,583,608]" class="Malacostraca" family="Pylochelidae" genus="Pylocheles" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="spinosus">
|
||
<emphasis id="B90AEA85E245FF87D4A4FDA50C1D9A72" box="[151,375,583,608]" italics="true" pageId="21" pageNumber="322">Pylocheles spinosus</emphasis>
|
||
</taxonomicName>
|
||
by Miyake (1978).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |