168 lines
19 KiB
XML
168 lines
19 KiB
XML
<document id="01A844160E33E7BACCD78192601AB550" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE3C3FF57FF75FF3DBD08FCD8" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Madagaster quadricurvipes Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="199" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="197" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:titleInfo id="A9D5151B110351E004561DA7396EABE8">
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<mods:title id="DFE8EA0D9C41A53D6E7914B77C2FB3E6">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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<mods:name id="80E6AE36DBCBE7F2D7C7335A8405DBBD" type="personal">
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<mods:roleTerm id="FBA06518AC348D1F9701B59947970FF5">Author</mods:roleTerm>
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<mods:namePart id="133077AFAF052428C000F868C4DE4DBF">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="8631D4A441DD59EB01B767C44D6AA056">text</mods:typeOfResource>
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<mods:title id="6AFD345D2A40618A88F4FA057B1A3C31">Zootaxa</mods:title>
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<mods:part id="896F89E84E4C9CCDEA6D2AF50E3B87ED">
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<mods:date id="86B7C5BDC1E1F3093F8D6ECA95BC9D45">2017</mods:date>
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<mods:number id="1495F40B704316EE8B29B931A1620FFE">2017-11-03</mods:number>
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<mods:number id="340CCCFE4B146F72FC9BA4A93BFA84C0">4342</mods:number>
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<mods:start id="FCB41759EA45B8568C16B7E8FA89A957">1</mods:start>
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<mods:classification id="5213E9A8857FE8B906FCFD4D8BA3842F">journal article</mods:classification>
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<mods:identifier id="7A69F6B23ECED58620884E90230D59AF" type="CLB-Dataset">31639</mods:identifier>
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<mods:identifier id="2C15F54D3986BB60C4DFB27BB0301DDB" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<mods:identifier id="4D8CB29FD84E0ED05DCF833E728B290E" type="GBIF-Dataset">15799d2a-1968-4874-8100-452597d4e26b</mods:identifier>
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<mods:identifier id="E97BCE20558700EADE1B5E1CA81EEF4D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="52F6CEF7C8EC5021202F486D77D79808" type="Zenodo-Dep">1041066</mods:identifier>
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<treatment id="038587BBE3C3FF57FF75FF3DBD08FCD8" ID-DOI="http://doi.org/10.5281/zenodo.5323750" ID-GBIF-Taxon="137233380" ID-Zenodo-Dep="5323750" LSID="urn:lsid:plazi:treatment:038587BBE3C3FF57FF75FF3DBD08FCD8" httpUri="http://treatment.plazi.org/id/038587BBE3C3FF57FF75FF3DBD08FCD8" lastPageId="198" lastPageNumber="199" pageId="196" pageNumber="197">
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<subSubSection id="C3366526E3C3FF55FF75FF3DBB98FF56" pageId="196" pageNumber="197" type="nomenclature">
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<paragraph id="8B9336ADE3C3FF55FF75FF3DBBBDFF37" blockId="196.[151,676,152,211]" box="[151,641,152,178]" pageId="196" pageNumber="197">
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<heading id="D0DB81C1E3C3FF55FF75FF3DBBBDFF37" bold="true" box="[151,641,152,178]" fontSize="11" level="1" pageId="196" pageNumber="197" reason="1">
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<emphasis id="B958EABFE3C3FF55FF75FF3DBBBDFF37" bold="true" box="[151,641,152,178]" pageId="196" pageNumber="197">
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<taxonomicName id="4C2C4D2EE3C3FF55FF75FF3DB8DDFF37" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,481,152,178]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="quadricurvipes" status="sp. nov.">
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<emphasis id="B958EABFE3C3FF55FF75FF3DB8DDFF37" bold="true" box="[151,481,152,178]" italics="true" pageId="196" pageNumber="197">Madagaster quadricurvipes</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26B57C4E3C3FF55FE13FF3DBBBDFF37" box="[497,641,152,178]" pageId="196" pageNumber="197" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9336ADE3C3FF55FF75FF1FBB98FF56" blockId="196.[151,676,152,211]" box="[151,676,186,211]" pageId="196" pageNumber="197">
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<figureCitation id="13172A28E3C3FF55FF75FF1FB9C3FF56" box="[151,255,186,211]" captionStart="FIGURE 188" captionStartId="196.[151,250,1984,2006]" captionText="FIGURE 188. Madagaster quadricurvipes, holotype, habitus dorsal, lateral, and ventral." pageId="196" pageNumber="197">Figs. 188</figureCitation>
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(habitus), 190 (aedeagus), 300 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E3C3FF55FF75FEA7B876FEE6" pageId="196" pageNumber="197" type="materials_examined">
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<paragraph id="8B9336ADE3C3FF55FF75FEA7B876FEE6" blockId="196.[151,1437,258,679]" pageId="196" pageNumber="197">
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<materialsCitation id="3B443CF0E3C3FF55FF75FEA7B876FEE6" ID-GBIF-Occurrence="1697157761" collectingDate="2006-05-08" collectionCode="BMNH" collectorName="Andringitra NP & Riambavy R." elevation="1739" location="Fianarantsoa" pageId="196" pageNumber="197" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
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<emphasis id="B958EABFE3C3FF55FF75FEA7B8FAFE9E" bold="true" box="[151,454,258,283]" pageId="196" pageNumber="197">
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<typeStatus id="5497880FE3C3FF55FF75FEA7B9EEFE9E" box="[151,210,258,283]" pageId="196" pageNumber="197">Type</typeStatus>
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Material.
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<typeStatus id="5497880FE3C3FF55FEBBFEA7B8FAFE9E" box="[345,454,258,283]" pageId="196" pageNumber="197" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male):
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<collectingRegion id="49E8F84FE3C3FF55FDCDFEA7BBF2FE9E" box="[559,718,258,283]" country="Madagascar" name="Fianarantsoa" pageId="196" pageNumber="197">
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<emphasis id="B958EABFE3C3FF55FDCDFEA7BBF2FE9E" bold="true" box="[559,718,258,283]" pageId="196" pageNumber="197">Fianarantsoa</emphasis>
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</collectingRegion>
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,
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<collectorName id="26D9537BE3C3FF55FD03FEA7BAA4FE9F" box="[737,920,258,283]" pageId="196" pageNumber="197">Andringitra NP</collectorName>
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:
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<collectorName id="26D9537BE3C3FF55FC45FEA7BD7DFE9F" box="[935,1089,258,283]" pageId="196" pageNumber="197">Riambavy R.</collectorName>
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(female) cascade hygropetric habitat, N -22° 8.761, E:46° 53.4, wet rock surfaces next to high waterfall, elev.
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<elevation id="0001D19EE3C3FF55FBC1FE82BD47FEBB" box="[1059,1147,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.739" pageId="196" pageNumber="197" unit="m" value="1739.0">
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<quantity id="4CD49B48E3C3FF55FBC1FE82BD47FEBB" box="[1059,1147,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.739" pageId="196" pageNumber="197" unit="m" value="1739.0">1739 m</quantity>
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</elevation>
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,
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<collectingDate id="EFD6E985E3C3FF55FB65FE82BDD2FEBA" box="[1159,1262,294,319]" pageId="196" pageNumber="197" value="2006-05-08">
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<date id="FF92106DE3C3FF55FB65FE82BDD2FEBA" box="[1159,1262,294,319]" pageId="196" pageNumber="197" value="2006-05-08">8 v 2006</date>
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</collectingDate>
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, Bergsten
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<emphasis id="B958EABFE3C3FF55FA81FE82BCA0FEBA" box="[1379,1436,294,319]" italics="true" pageId="196" pageNumber="197">et al.</emphasis>
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(P39) (
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<collectionCode id="ED3DAE68E3C3FF55FF06FEEEB802FEE6" box="[228,318,331,355]" pageId="196" pageNumber="197">BMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E3C3FF55FF25FECBBD3EFDBE" pageId="196" pageNumber="197" type="diagnosis">
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<paragraph id="8B9336ADE3C3FF55FF25FECBBD3EFDBE" blockId="196.[151,1437,258,679]" pageId="196" pageNumber="197">
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<emphasis id="B958EABFE3C3FF55FF25FECBB8EEFE02" bold="true" box="[199,466,366,391]" pageId="196" pageNumber="197">Differential Diagnosis.</emphasis>
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Somewhat similar in habitus to
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<taxonomicName id="4C2C4D2EE3C3FF55FCDFFECABA8BFE03" box="[829,951,367,390]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="steineri">
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<emphasis id="B958EABFE3C3FF55FCDFFECABA8BFE03" box="[829,951,367,390]" italics="true" pageId="196" pageNumber="197">M. steineri</emphasis>
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</taxonomicName>
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; differing therefrom (males) in larger body size (ca. 3.38 vs.
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<quantity id="4CD49B48E3C3FF55FE8CFE36B8E7FE2E" box="[366,475,403,427]" metricMagnitude="-3" metricUnit="m" metricValue="2.99" pageId="196" pageNumber="197" unit="mm" value="2.99">2.99 mm</quantity>
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), having the interrupted areas of the elytral costae larger, and the costae shorter. Differing also in having arcuate meso- and metatibiae (
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<figureCitation id="13172A28E3C3FF55FCEBFE13BA4AFE4A" box="[777,886,438,463]" captionStart="FIGURE 184" captionStartId="192.[151,250,1367,1389]" captionText="FIGURE 184. Madagaster steineri, holotype, habitus dorsal, lateral, and ventral." pageId="196" pageNumber="197">Figs. 184</figureCitation>
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,
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<figureCitation id="13172A28E3C3FF55FC61FE12BA8DFE4A" box="[899,945,439,463]" captionStart="FIGURE 188" captionStartId="196.[151,250,1984,2006]" captionText="FIGURE 188. Madagaster quadricurvipes, holotype, habitus dorsal, lateral, and ventral." pageId="196" pageNumber="197">188</figureCitation>
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). The pronotal reliefs of
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<taxonomicName id="4C2C4D2EE3C3FF55FB2EFE12BCA0FE4A" authorityName="Perkins" authorityYear="2017" box="[1228,1436,438,463]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="quadricurvipes">
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<emphasis id="B958EABFE3C3FF55FB2EFE12BCA0FE4A" box="[1228,1436,438,463]" italics="true" pageId="196" pageNumber="197">M. quadricurvipes</emphasis>
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</taxonomicName>
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are more evidently tuberculate, whereas in the compared species the punctures are more evident than are the associated granules. The very complex aedeagi of the two species differ markedly in shapes of the parameres and main piece (
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<figureCitation id="13172A28E3C3FF55FEC2FD87B8B7FDBE" box="[288,395,546,571]" captionStart="FIGURES 185 – 186" captionStartId="193.[151,264,1869,1892]" captionText="FIGURES 185 – 186. Madagaster, aedeagi of holotypes." pageId="196" pageNumber="197">Figs. 186</figureCitation>
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,
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<figureCitation id="13172A28E3C3FF55FE75FD86B8F9FDBE" box="[407,453,547,571]" captionStart="FIGURES 189 – 190" captionStartId="197.[151,264,1704,1727]" captionText="FIGURES 189 – 190. Madagaster, aedeagi of holotypes." pageId="196" pageNumber="197">190</figureCitation>
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). Females of
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<taxonomicName id="4C2C4D2EE3C3FF55FDB9FD81BA17FDBE" authorityName="Perkins" authorityYear="2017" box="[603,811,546,571]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="quadricurvipes">
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<emphasis id="B958EABFE3C3FF55FDB9FD81BA17FDBE" box="[603,811,546,571]" italics="true" pageId="196" pageNumber="197">M. quadricurvipes</emphasis>
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</taxonomicName>
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are not yet known.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E3C3FF57FF25FDE3BCB8FCB2" lastPageId="198" lastPageNumber="199" pageId="196" pageNumber="197" type="description">
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<paragraph id="8B9336ADE3C3FF55FF25FDE3BA37FD06" blockId="196.[151,1437,258,679]" pageId="196" pageNumber="197">
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<emphasis id="B958EABFE3C3FF55FF25FDE3B86DFDDA" bold="true" box="[199,337,582,607]" pageId="196" pageNumber="197">Description</emphasis>
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. Size:
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<typeStatus id="5497880FE3C3FF55FE41FDE3BB39FDDA" box="[419,517,582,607]" pageId="196" pageNumber="197" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 3.38/1.55; head width 0.84; pronotum 0.74/0.86, PA 0.97, PB 1.04; elytra 2.09/1.55.
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</paragraph>
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<paragraph id="8B9336ADE3C3FF55FF25FD2ABA34FD22" blockId="196.[151,1437,258,679]" box="[199,776,654,679]" pageId="196" pageNumber="197">Dorsum and venter dark brown. Ocelli light brown.</paragraph>
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<caption id="DF536625E3C3FF55FF75F865BD20F853" box="[151,1052,1984,2006]" pageId="196" pageNumber="197">
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<paragraph id="8B9336ADE3C3FF55FF75F865BD20F853" blockId="196.[151,1052,1984,2006]" box="[151,1052,1984,2006]" pageId="196" pageNumber="197">
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<emphasis id="B958EABFE3C3FF55FF75F865B810F850" bold="true" box="[151,300,1984,2006]" pageId="196" pageNumber="197">FIGURE 188.</emphasis>
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<taxonomicName id="4C2C4D2EE3C3FF55FED0F865BB70F853" authorityName="Perkins" authorityYear="2017" box="[306,588,1984,2006]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="quadricurvipes">
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<emphasis id="B958EABFE3C3FF55FED0F865BB70F853" box="[306,588,1984,2006]" italics="true" pageId="196" pageNumber="197">Madagaster quadricurvipes</emphasis>
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</taxonomicName>
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, holotype, habitus dorsal, lateral, and ventral.
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</paragraph>
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</caption>
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<caption id="DF536625E3C2FF54FF75F90DBBEEF93A" box="[151,722,1704,1727]" pageId="197" pageNumber="198">
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<paragraph id="8B9336ADE3C2FF54FF75F90DBBEEF93A" blockId="197.[151,722,1704,1727]" box="[151,722,1704,1727]" pageId="197" pageNumber="198">
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<emphasis id="B958EABFE3C2FF54FF75F90DB851F93B" bold="true" box="[151,365,1704,1727]" pageId="197" pageNumber="198">FIGURES 189–190.</emphasis>
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<emphasis id="B958EABFE3C2FF54FE91F90CB8D0F93A" box="[371,492,1705,1727]" italics="true" pageId="197" pageNumber="198">Madagaster</emphasis>
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, aedeagi of holotypes.
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</paragraph>
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</caption>
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<paragraph id="8B9336ADE3C2FF54FF25F943BADCF80A" blockId="197.[151,1436,1766,2007]" pageId="197" pageNumber="198">Antennal club with last antennomere larger than penultimate. Maxillary palpi shorter than antenna, about as long as labrum is wide. Labrum deeply bilobed, pubescent, with longer setae along anterior margin. Labrum originating under slightly overhanging margin of clypeus. Labroclypeal suture very deep. Short, deep sulcus in front of each ocellus. Clypeus very weakly shining, surface slightly uneven and sparsely setose. Frons coarsely punctate, dull, slightly concave between ocelli. Eyes large and protuberant.</paragraph>
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<paragraph id="8B9336ADE3C2FF57FF25F83EBAF2FF50" blockId="197.[151,1436,1766,2007]" lastBlockId="198.[151,1437,151,861]" lastPageId="198" lastPageNumber="199" pageId="197" pageNumber="198">Pronotum with two narrowly separated midlongitudinal fovea, anterior three times as long as posterior; each side with two admedian and two sublateral foveae; anterior margin with deep postocular emarginations; lateral margin broadly rounded. Pronotal disc tuberculate, tubercles shining, each with short seta at posterior margin. Lateral foveae with very small and very sparse tubercles, weakly shining.</paragraph>
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<paragraph id="8B9336ADE3C1FF57FF25FF7AB846FEE0" blockId="198.[151,1437,151,861]" pageId="198" pageNumber="199">Elytral suture costate in posterior. First two elytral "interval" costae (from suture) interrupted in two places, third and fourth costa interrupted once. Interrupted areas often longer than costate areas in same line. Between costae usually three rows of setigerous punctures, in places with four rows, interpunctal surfaces sometimes smooth and shining.</paragraph>
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<paragraph id="8B9336ADE3C1FF57FF25FED5B9CBFE55" blockId="198.[151,1437,151,861]" pageId="198" pageNumber="199">Mentum and postmentum sparsely pubescent; Genae microreticulate, dull, deeply concave behind cardo, medial margin of concavity with short carina. Subocular groove for antenna deep and wide. Postocular margin not carinate.</paragraph>
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<paragraph id="8B9336ADE3C1FF57FF25FE79B8E0FDE5" blockId="198.[151,1437,151,861]" pageId="198" pageNumber="199">
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Mesoventrite without tubercle on intercoxal process. Metaventrite densely micropunctulate and with very short and very dense vestiture, except vestiture sparser in deep and wide midlongitudinal depression (
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<figureCitation id="13172A28E3C1FF57FB21FE5ABC19FD9D" box="[1219,1317,511,536]" captionStart="FIGURE 188" captionStartId="196.[151,250,1984,2006]" captionText="FIGURE 188. Madagaster quadricurvipes, holotype, habitus dorsal, lateral, and ventral." pageId="198" pageNumber="199">Fig. 188</figureCitation>
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). First two and apparently lateral area of third abdominal ventrites with hydrofuge pubescence, setae much sparser, nonhydrofuge, on other ventrites
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</paragraph>
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<paragraph id="8B9336ADE3C1FF57FF25FDC9BCB8FCB2" blockId="198.[151,1437,151,861]" pageId="198" pageNumber="199">
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Strongly sexually dimorphic (although females are not yet known, the following characters are thought to be male only, based on relationship to
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<taxonomicName id="4C2C4D2EE3C1FF57FDC0FD34BBA7FD2D" box="[546,667,657,680]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="steineri">
|
||
<emphasis id="B958EABFE3C1FF57FDC0FD34BBA7FD2D" box="[546,667,657,680]" italics="true" pageId="198" pageNumber="199">M. steineri</emphasis>
|
||
</taxonomicName>
|
||
: abdominal ventrite four with large tubercle on each side of midline; fifth ventrite with large posteromedian triangular process; sixth retractable (over median portion) within fifth, seventh large and completely exposed; profemur basally on lower margin produced in large pointed process; protibia widened at about midlength; mesotibia very slightly arcuate and apically slightly widened and densely setose; metafemur markedly arcuate in basal 1/3; metatibia arcuate, distal ½ widened and bearing brush of setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3366526E3C1FF57FF25FCE1BD08FCD8" box="[199,1076,836,861]" pageId="198" pageNumber="199" type="etymology">
|
||
<paragraph id="8B9336ADE3C1FF57FF25FCE1BD08FCD8" blockId="198.[151,1437,151,861]" box="[199,1076,836,861]" pageId="198" pageNumber="199">
|
||
<emphasis id="B958EABFE3C1FF57FF25FCE1B872FCD8" bold="true" box="[199,334,836,861]" pageId="198" pageNumber="199">Etymology.</emphasis>
|
||
Named in reference to the arcuate meso- and metatibiae of males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |