151 lines
18 KiB
XML
151 lines
18 KiB
XML
<document id="7E7455F1FB7BC5791F1A3AC1EF8C8D01" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE3BDFF2BFF75FED5B8DCF816" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Madagaster bergsteni Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="187" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="187" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:titleInfo id="DF709446F48E4D20531BD1AE4D3ACDBE">
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<mods:title id="243D901992E0C91A83FA2B17FD98B2EE">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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<mods:name id="22A12CCF329E973687849E5004CE8E8B" type="personal">
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<mods:roleTerm id="DF8C58D16909086A564D00BE18043302">Author</mods:roleTerm>
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<mods:namePart id="9C58EA9B251BED344C585683F324E257">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="E707F40EC27B720BE3CDC2FE73207793">text</mods:typeOfResource>
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<mods:title id="5A413D5C92E6C582A743CA462ACE3E27">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="51B69687ED7A1EB17E2402FF7EEABDAF">
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<mods:date id="8F8252D457FA9C67E5A9F27DC83A2AE4">2017</mods:date>
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<mods:number id="62A2A1823E96CA8957F4AEDBF3F84CA4">2017-11-03</mods:number>
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<mods:number id="D7BF56D9E39846A464956D190900BFFF">4342</mods:number>
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<mods:detail id="3E4FF1CBA568708E2A38D6D6A2D653C8" type="issue">
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<mods:number id="D8313A34FB52176E1E889579D3354D32">1</mods:number>
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<mods:classification id="105B8995D786087F34C5E2A841DDF574">journal article</mods:classification>
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<mods:identifier id="4FA61825E157FF644B5FFA439DB5BF6B" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<mods:identifier id="435680CEB94ED109D9C60040C51AB308" type="GBIF-Dataset">15799d2a-1968-4874-8100-452597d4e26b</mods:identifier>
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<mods:identifier id="A0DBCABB0853E6BD53D98DE29E9E0B7E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5D8AF19E610D2EFDBD46739D72D91D55" type="Zenodo-Dep">1041066</mods:identifier>
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<treatment id="038587BBE3BDFF2BFF75FED5B8DCF816" ID-DOI="http://doi.org/10.5281/zenodo.5323734" ID-GBIF-Taxon="137233385" ID-Zenodo-Dep="5323734" LSID="urn:lsid:plazi:treatment:038587BBE3BDFF2BFF75FED5B8DCF816" httpUri="http://treatment.plazi.org/id/038587BBE3BDFF2BFF75FED5B8DCF816" lastPageNumber="187" pageId="186" pageNumber="187">
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<subSubSection id="C3366526E3BDFF2BFF75FED5BB98FE2E" pageId="186" pageNumber="187" type="nomenclature">
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<paragraph id="8B9336ADE3BDFF2BFF75FED5BB07FE0F" blockId="186.[151,676,368,427]" box="[151,571,368,394]" pageId="186" pageNumber="187">
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<heading id="D0DB81C1E3BDFF2BFF75FED5BB07FE0F" bold="true" box="[151,571,368,394]" fontSize="11" level="1" pageId="186" pageNumber="187" reason="1">
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<emphasis id="B958EABFE3BDFF2BFF75FED5BB07FE0F" bold="true" box="[151,571,368,394]" pageId="186" pageNumber="187">
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<emphasis id="B958EABFE3BDFF2BFF75FED5B89FFE0F" bold="true" box="[151,419,368,394]" italics="true" pageId="186" pageNumber="187">
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<taxonomicName id="4C2C4D2EE3BDFF2BFF75FED5B8A1FE0F" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,413,368,394]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="bergsteni" status="sp. nov.">Madagaster bergsteni</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A26B57C4E3BDFF2BFE49FED5BB07FE0F" box="[427,571,368,394]" pageId="186" pageNumber="187" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9336ADE3BDFF2BFF75FE37BB98FE2E" blockId="186.[151,676,368,427]" box="[151,676,402,427]" pageId="186" pageNumber="187">
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<figureCitation id="13172A28E3BDFF2BFF75FE37B9C3FE2E" box="[151,255,402,427]" captionStart="FIGURE 179" captionStartId="187.[151,250,1464,1486]" captionText="FIGURE 179. Madagaster bergsteni, holotype, habitus dorsal, lateral, and ventral." pageId="186" pageNumber="187">Figs. 179</figureCitation>
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(habitus), 181 (aedeagus), 301 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E3BDFF2BFF75FE7FBA27FDDA" pageId="186" pageNumber="187" type="materials_examined">
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<paragraph id="8B9336ADE3BDFF2BFF75FE7FBA27FDDA" blockId="186.[151,1437,474,1939]" pageId="186" pageNumber="187">
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<emphasis id="B958EABFE3BDFF2BFF75FE7FB8FCFE76" bold="true" box="[151,448,474,499]" pageId="186" pageNumber="187">
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<typeStatus id="5497880FE3BDFF2BFF75FE7FB9EEFE76" box="[151,210,474,499]" pageId="186" pageNumber="187">Type</typeStatus>
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Material.
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<typeStatus id="5497880FE3BDFF2BFEB1FE7FB8FCFE76" box="[339,448,474,499]" pageId="186" pageNumber="187" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male):
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<collectingRegion id="49E8F84FE3BDFF2BFDC1FE7FBBF5FE76" box="[547,713,474,499]" country="Madagascar" name="Antananarivo" pageId="186" pageNumber="187">
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<emphasis id="B958EABFE3BDFF2BFDC1FE7FBBF5FE76" bold="true" box="[547,713,474,499]" pageId="186" pageNumber="187">Antananarivo</emphasis>
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</collectingRegion>
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,
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<geoCoordinate id="EE18506AE3BDFF2BFD38FE7EBA78FE76" box="[730,836,475,499]" direction="south" orientation="latitude" pageId="186" pageNumber="187" precision="5" value="-18.8704">18.8704S</geoCoordinate>
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<geoCoordinate id="EE18506AE3BDFF2BFCAFFE7FBA86FE76" box="[845,954,474,499]" direction="east" orientation="longitude" pageId="186" pageNumber="187" precision="5" value="47.6708">47.6708E</geoCoordinate>
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Analamanga, Anjomara mountain next to Ambato- lampikely village, RN2, hygropetric rock, elev.
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<quantity id="4CD49B48E3BDFF2BFCD0FE5ABAB1FD93" box="[818,909,511,535]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="186" pageNumber="187" unit="m" value="1470.0">1470 m</quantity>
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,
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<date id="FF92106DE3BDFF2BFC7EFE5ABD22FD92" box="[924,1054,510,535]" pageId="186" pageNumber="187" value="2014-11-24">24 xi 2014</date>
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, J. Bergsten, R. Bukontaite, J.H. Randriamihaja, T. Ranarilalatiana & S. Holmgren (
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<collectionCode id="ED3DAE68E3BDFF2BFD3AFD86BA22FDBE" box="[728,798,547,571]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="186" pageNumber="187">MAD</collectionCode>
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14-79) (
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<collectionCode id="ED3DAE68E3BDFF2BFC97FD86BAFAFDBE" box="[885,966,547,571]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="186" pageNumber="187">NHRS</collectionCode>
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).
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<emphasis id="B958EABFE3BDFF2BFC3BFD87BD66FDBE" bold="true" box="[985,1114,546,571]" pageId="186" pageNumber="187">
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<typeStatus id="5497880FE3BDFF2BFC3BFD87BD69FDBE" box="[985,1109,546,571]" pageId="186" pageNumber="187" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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Same data as
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<typeStatus id="5497880FE3BDFF2BFB1DFD87BC5DFDBE" box="[1279,1377,546,571]" pageId="186" pageNumber="187" type="holotype">holotype</typeStatus>
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(526
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<collectionCode id="ED3DAE68E3BDFF2BFF75FDE2B9DBFDDA" box="[151,231,583,607]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="186" pageNumber="187">NHRS</collectionCode>
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); one male "
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<collectionCode id="ED3DAE68E3BDFF2BFE8DFDE2B8FAFDDA" box="[367,454,583,607]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="186" pageNumber="187">MNCN</collectionCode>
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<collectionCode id="ED3DAE68E3BDFF2BFE2FFDE2B8C6FDDB" box="[461,506,583,606]" country="Spain" httpUri="http://grbio.org/cool/qyi3-sxew" name="Institut de Biologia Evolutiva, (CSIC-UPF)" pageId="186" pageNumber="187">IBE</collectionCode>
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AN333
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<collectionCode id="ED3DAE68E3BDFF2BFDB8FDE2BBABFDDA" box="[602,663,583,607]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="186" pageNumber="187">DNA</collectionCode>
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specimen".
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E3BDFF2BFF25FDCFB9EDFCDE" pageId="186" pageNumber="187" type="diagnosis">
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<paragraph id="8B9336ADE3BDFF2BFF25FDCFB9EDFCDE" blockId="186.[151,1437,474,1939]" pageId="186" pageNumber="187">
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<emphasis id="B958EABFE3BDFF2BFF25FDCFB8E1FD06" bold="true" box="[199,477,618,643]" pageId="186" pageNumber="187">Differential Diagnosis.</emphasis>
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Similar in habitus to
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<taxonomicName id="4C2C4D2EE3BDFF2BFCE6FDC9BA9FFD06" authorityName="Perkins" authorityYear="2017" box="[772,931,620,643]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="procarina">
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<emphasis id="B958EABFE3BDFF2BFCE6FDC9BA9FFD06" box="[772,931,620,643]" italics="true" pageId="186" pageNumber="187">M. procarina</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C2C4D2EE3BDFF2BFC0DFDC9BD44FD06" authorityName="Perkins" authorityYear="2017" box="[1007,1144,618,643]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="barbata">
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<emphasis id="B958EABFE3BDFF2BFC0DFDC9BD44FD06" box="[1007,1144,618,643]" italics="true" pageId="186" pageNumber="187">M. barbata</emphasis>
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</taxonomicName>
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(
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<figureCitation id="13172A28E3BDFF2BFB70FDCFBC34FD06" box="[1170,1288,618,643]" captionStart="FIGURE 179" captionStartId="187.[151,250,1464,1486]" captionText="FIGURE 179. Madagaster bergsteni, holotype, habitus dorsal, lateral, and ventral." pageId="186" pageNumber="187">Figs. 179</figureCitation>
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,
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<figureCitation id="13172A28E3BDFF2BFAFDFDCEBC77FD06" box="[1311,1355,619,643]" captionStart="FIGURE 180" captionStartId="188.[151,250,1601,1623]" captionText="FIGURE 180. Madagaster procarina, holotype, habitus dorsal, lateral, and ventral." pageId="186" pageNumber="187">180</figureCitation>
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,
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<figureCitation id="13172A28E3BDFF2BFA80FDCEBCACFD06" box="[1378,1424,619,643]" captionStart="FIGURE 183" captionStartId="191.[151,250,1440,1462]" captionTargetBox="[340,1247,195,1408]" captionTargetId="figure@191.[340,1247,195,1408]" captionTargetPageId="191" captionText="FIGURE 183. Madagaster barbata, holotype, frontal view, habitus dorsal, lateral, and ventral." httpUri="https://zenodo.org/record/1041124/files/figure.png" pageId="186" pageNumber="187">183</figureCitation>
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). Differentiated from both species by the more transverse pronotum (W/L ca. 1.69) and, in males, by the dense and long setae on the mentum, the elevated and setose median area of the fifth abdominal ventrite, and the metatibiae, which are slightly arcuate away from the midline. The pronotal foveae are shallower than in the compared species. The very complex male genitalia of the three species share a hook-shaped process on the ventral surface of the main piece, but the parameres and processes of the distal part of the main piece differ markedly (
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<figureCitation id="13172A28E3BDFF2BFB12FCBBBCABFCB2" box="[1264,1431,798,823]" captionStart="FIGURES 181 – 182" captionStartId="189.[151,264,1764,1787]" captionText="FIGURES 181 – 182. Madagaster, aedeagi of holotypes." pageId="186" pageNumber="187">Figs. 181, 182</figureCitation>
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,
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<figureCitation id="13172A28E3BDFF2BFF75FCE7B9F9FCDE" box="[151,197,834,859]" captionStart="FIGURES 185 – 186" captionStartId="193.[151,264,1869,1892]" captionText="FIGURES 185 – 186. Madagaster, aedeagi of holotypes." pageId="186" pageNumber="187">185</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E3BDFF2BFF25FCC3BB9CF8CE" pageId="186" pageNumber="187" type="description">
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<paragraph id="8B9336ADE3BDFF2BFF25FCC3BDDFFBD2" blockId="186.[151,1437,474,1939]" pageId="186" pageNumber="187">
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<emphasis id="B958EABFE3BDFF2BFF25FCC3B865FCFA" bold="true" box="[199,345,870,895]" pageId="186" pageNumber="187">Description.</emphasis>
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Size (
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<typeStatus id="5497880FE3BDFF2BFE44FCC3BB30FCFA" box="[422,524,870,895]" pageId="186" pageNumber="187" type="holotype">holotype</typeStatus>
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, mm, length/width): body (length to elytral apices) 2.29/1.06, head 0.59/0.39, pronotum 0.52/0.88, elytra 1.41/1.06. Dorsum and venter black, legs dark brown, ocelli light brown to testaceous. Entire dorsum dull, with very small, closely spaced tubercles, each with a short recumbent seta. Labrum bilobed, anterior margin slightly reflexed, sparsely granulate and with sparse small setae except with longer setae along anterior margin. Labrum originating at anterior, slightly overhanging, margin of clypeus. Labroclypeal suture moderately deep. Short, deep sulcus in front of each ocellus. Frons with deep concavity between ocelli. Maxillary palpi slightly shorter than antenna. Antenna with last antennomere much shorter than penultimate.
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</paragraph>
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<paragraph id="8B9336ADE3BDFF2BFF25FBC6BB98FB46" blockId="186.[151,1437,474,1939]" pageId="186" pageNumber="187">Pronotum with two narrowly separated midlongitudinal fovea, anterior ca. twice as long as posterior; each side with two admedian and two sublateral foveae; transverse fovea along base, on each side of posterior midlongitudinal fovea; lateral margin rounded.</paragraph>
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<paragraph id="8B9336ADE3BDFF2BFF25FB6BBA73FAAB" blockId="186.[151,1437,474,1939]" pageId="186" pageNumber="187">Elytral suture slightly raised in posterior. Each elytron with four very weakly raised unilinear rows of granules; first row (from suture) slightly curved in ca. middle ½ of length. Areas between unilinear rows of granules with four or five irregular rows of granules. Elytral apices truncate.</paragraph>
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<paragraph id="8B9336ADE3BDFF2BFF25FA9EBDAEFA1E" blockId="186.[151,1437,474,1939]" pageId="186" pageNumber="187">Mentum and postmentum (males) punctate and with dense, long pubescence; Genae micropunctulate, dull, concave behind cardo, posterior margin gradual, not carinate. Genae at posterior with transverse arcuate carinate ridge. Subocular groove for antenna moderately deep. Posterior margin of eye not carinate.</paragraph>
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<paragraph id="8B9336ADE3BDFF2BFF25FA03B885F9CA" blockId="186.[151,1437,474,1939]" pageId="186" pageNumber="187">
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Prosternal intercoxal process with raised tip. Mesoventrite without tubercle on intercoxal process. Metaventrite with very short and very dense vestiture, including in median depression (
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<figureCitation id="13172A28E3BDFF2BFC35FA6FBD06FA66" box="[983,1082,1482,1507]" captionStart="FIGURE 179" captionStartId="187.[151,250,1464,1486]" captionText="FIGURE 179. Madagaster bergsteni, holotype, habitus dorsal, lateral, and ventral." pageId="186" pageNumber="187">Fig. 179</figureCitation>
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). First four abdominal ventrites and anterior 1/3 of fifth with hydrofuge pubescence; setae much sparser, non-hydrofuge, on other ventrites. Fifth ventrite of males with large tumid prominence on posterior ½ in midline, tumidity concave behind and with long golden setae in concavity.
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</paragraph>
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<paragraph id="8B9336ADE3BDFF2BFF25F9FEBB34F93E" blockId="186.[151,1437,474,1939]" pageId="186" pageNumber="187">Pro- and mesotibiae straight, stout. All tibiae apically with small cluster of moderately long shining setae. Metafemur markedly arcuate in basal 1/3. Metatibia slightly arcuate, toward midline. Mesocoxae (male) with ventral surface slightly flattened.</paragraph>
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<paragraph id="8B9336ADE3BDFF2BFF25F963B84CF8A3" blockId="186.[151,1437,474,1939]" pageId="186" pageNumber="187">Abdominal ventrites five and six enlarged in both sexes, in male combined lengths of two to four equal ca. one-half combined lengths of five and six. Seventh ventrite (male) asymmetrical, with concavity on each side of raised median area.</paragraph>
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<paragraph id="8B9336ADE3BDFF2BFF25F897BB9CF8CE" blockId="186.[151,1437,474,1939]" box="[199,672,1842,1867]" pageId="186" pageNumber="187">Female smaller than male (ca. 2.00–2.24).</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E3BDFF2BFF25F8F3B8DCF816" pageId="186" pageNumber="187" type="etymology">
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<paragraph id="8B9336ADE3BDFF2BFF25F8F3B8DCF816" blockId="186.[151,1437,474,1939]" pageId="186" pageNumber="187">
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<emphasis id="B958EABFE3BDFF2BFF25F8F3B872F8EA" bold="true" box="[199,334,1878,1903]" pageId="186" pageNumber="187">Etymology.</emphasis>
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Named in honor of Johannes Bergsten, in recognition of his expert leadership in the
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<collectingCountry id="F33B763DE3BDFF2BFAF7F8F3BCA0F8EA" box="[1301,1436,1878,1903]" name="Madagascar" pageId="186" pageNumber="187">Madagascar</collectingCountry>
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aquatic beetle survey project.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |