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<document id="2E9E9F51D75C2784A42806817AF03D05" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE32BFFBCFF75FBB9BD48FE70" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena oscillata Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="46" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="45" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:title id="6DD2BD7244103F7334CA231B06269800">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
</mods:titleInfo>
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<mods:namePart id="92B387B85712CF503C057655C8769127">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="16EABD07B94D71058A30DAA443A0E8CF">text</mods:typeOfResource>
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<mods:title id="986E2020C25F979E4F6632C615C06FBB">Zootaxa</mods:title>
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<mods:part id="3C6102339B76F6B5C75EF34C8C38C944">
<mods:date id="8795A6A78D163D22DD4A043970017A56">2017</mods:date>
<mods:detail id="028F97B763C484940AE5DE9DAF21853D" type="pubDate">
<mods:number id="4421A57635C1944B45A43A8EE53C0850">2017-11-03</mods:number>
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<mods:number id="9137D31C3192B8EEEA8B01DB2BDA4B7B">4342</mods:number>
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<mods:identifier id="1BC7FE5F643F37E631D5DDD6011D1895" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<treatment id="038587BBE32BFFBCFF75FBB9BD48FE70" ID-DOI="http://doi.org/10.5281/zenodo.5323473" ID-GBIF-Taxon="137233456" ID-Zenodo-Dep="5323473" LSID="urn:lsid:plazi:treatment:038587BBE32BFFBCFF75FBB9BD48FE70" httpUri="http://treatment.plazi.org/id/038587BBE32BFFBCFF75FBB9BD48FE70" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<subSubSection id="C3366526E32BFFBDFF75FBB9BBB4FBD2" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="8B9336ADE32BFFBDFF75FBB9BB25FBB3" blockId="44.[151,648,1052,1111]" box="[151,537,1052,1078]" pageId="44" pageNumber="45">
<heading id="D0DB81C1E32BFFBDFF75FBB9BB25FBB3" bold="true" box="[151,537,1052,1078]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
<emphasis id="B958EABFE32BFFBDFF75FBB9BB25FBB3" bold="true" box="[151,537,1052,1078]" pageId="44" pageNumber="45">
<taxonomicName id="4C2C4D2EE32BFFBDFF75FBB9B846FBB3" ID-CoL="3N3M5" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,378,1052,1078]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="oscillata" status="sp. nov.">
<emphasis id="B958EABFE32BFFBDFF75FBB9B846FBB3" bold="true" box="[151,378,1052,1078]" italics="true" pageId="44" pageNumber="45">Hydraena oscillata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B57C4E32BFFBDFE6BFBB9BB25FBB3" box="[393,537,1052,1078]" pageId="44" pageNumber="45" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9336ADE32BFFBDFF75FB9BBBB4FBD2" blockId="44.[151,648,1052,1111]" box="[151,648,1086,1111]" pageId="44" pageNumber="45">
<figureCitation id="13172A28E32BFFBDFF75FB9BB9CDFBD2" box="[151,241,1086,1111]" captionStart="FIGURE 34" captionStartId="45.[151,250,1870,1892]" captionTargetBox="[298,1270,575,1840]" captionTargetId="figure@45.[298,1270,575,1840]" captionTargetPageId="45" captionText="FIGURE 34. Hydraena oscillata, holotype, habitus dorsal, lateral, and ventral." httpUri="https://zenodo.org/record/1041096/files/figure.png" pageId="44" pageNumber="45">Figs. 34</figureCitation>
(habitus), 36 (aedeagus), 243 (map)
</paragraph>
</subSubSection>
<subSubSection id="C3366526E32BFFBDFF75FB23B8A4FA8E" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="8B9336ADE32BFFBDFF75FB23B8A4FA8E" blockId="44.[151,1437,1158,2011]" pageId="44" pageNumber="45">
<emphasis id="B958EABFE32BFFBDFF75FB23B871FB1A" bold="true" box="[151,333,1158,1183]" pageId="44" pageNumber="45">
<typeStatus id="5497880FE32BFFBDFF75FB23B9EEFB1A" box="[151,210,1158,1183]" pageId="44" pageNumber="45">Type</typeStatus>
Material.
</emphasis>
<materialsCitation id="3B443CF0E32BFFBDFEBBFB23BDAAFB62" ID-GBIF-Occurrence="1697157519" collectedFrom="forest stream with small pools" collectingDate="2014-11-28" collectionCode="NHRS" collectorName="J. Bergsten &amp; R. Bukontaite &amp; J. H. Randriamihaja &amp; T. Ranarilalatiana &amp; S. Holmgren" elevation="930" latitude="-18.7697" location="Alaotra Mangoro Andasibe-Mantadia NP, Mantadia: 15 km N of park entrance" longLatPrecision="7" longitude="48.4372" pageId="44" pageNumber="45" specimenCode="MAD14-82" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
<emphasis id="B958EABFE32BFFBDFEBBFB23B8FAFB1A" bold="true" box="[345,454,1158,1183]" pageId="44" pageNumber="45">
<typeStatus id="5497880FE32BFFBDFEBBFB23B8FAFB1A" box="[345,454,1158,1183]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
</emphasis>
(
<specimenCount id="9D2AFD24E32BFFBDFE38FB23BB2BFB1A" box="[474,535,1158,1183]" pageId="44" pageNumber="45" type="male">male</specimenCount>
):
<collectingRegion id="49E8F84FE32BFFBDFDCCFB23BB8CFB1A" box="[558,688,1158,1183]" country="Madagascar" name="Toamasina" pageId="44" pageNumber="45">
<emphasis id="B958EABFE32BFFBDFDCCFB23BB8CFB1A" bold="true" box="[558,688,1158,1183]" pageId="44" pageNumber="45">Toamasina</emphasis>
</collectingRegion>
,
<geoCoordinate id="EE18506AE32BFFBDFD21FB23BA12FB1A" box="[707,814,1158,1183]" direction="south" orientation="latitude" pageId="44" pageNumber="45" precision="5" value="-18.7697">18.7697S</geoCoordinate>
<geoCoordinate id="EE18506AE32BFFBDFCD8FB22BA95FB1A" box="[826,937,1159,1183]" direction="east" orientation="longitude" pageId="44" pageNumber="45" precision="5" value="48.4372">48.4372E</geoCoordinate>
,
<location id="8EF36076E32BFFBDFC5BFB23BB03FB47" LSID="urn:lsid:plazi:treatment:038587BBE32BFFBCFF75FBB9BD48FE70:8EF36076E32BFFBDFC5BFB23BB03FB47" latitude="-18.7697" longLatPrecision="7" longitude="48.4372" name="Alaotra Mangoro Andasibe-Mantadia NP, Mantadia: 15 km N of park entrance" pageId="44" pageNumber="45" stateProvince="Toamasina">
Alaotra Mangoro Andasibe-Mantadia NP, Mantadia:
<quantity id="4CD49B48E32BFFBDFEF6FB0FB868FB46" box="[276,340,1194,1219]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="44" pageNumber="45" unit="km" value="15.0">15km</quantity>
N of park entrance
</location>
,
<collectedFrom id="38307550E32BFFBDFDACFB0FBA8EFB46" box="[590,946,1194,1219]" pageId="44" pageNumber="45">forest stream with small pools</collectedFrom>
, elev.
<quantity id="4CD49B48E32BFFBDFBE3FB0EBD72FB47" box="[1025,1102,1195,1219]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="44" pageNumber="45" unit="m" value="930.0">
<elevation id="0001D19EE32BFFBDFBE3FB0EBD72FB47" box="[1025,1102,1195,1219]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="44" pageNumber="45" unit="m" value="930.0">930 m</elevation>
</quantity>
,
<date id="FF92106DE32BFFBDFBBCFB0EBDDDFB46" box="[1118,1249,1194,1219]" pageId="44" pageNumber="45" value="2014-11-28">
<collectingDate id="EFD6E985E32BFFBDFBBCFB0EBDDDFB46" box="[1118,1249,1194,1219]" pageId="44" pageNumber="45" value="2014-11-28">28 xi 2014</collectingDate>
</date>
,
<collectorName id="26D9537BE32BFFBDFB12FB0EBC4FFB46" box="[1264,1395,1195,1219]" pageId="44" pageNumber="45">J. Bergsten</collectorName>
,
<collectorName id="26D9537BE32BFFBDFA61FB0EB829FB62" pageId="44" pageNumber="45">R. Bukontaite</collectorName>
,
<collectorName id="26D9537BE32BFFBDFEC3FB6AB8C3FB62" box="[289,511,1230,1255]" pageId="44" pageNumber="45">J.H. Randriamihaja</collectorName>
,
<collectorName id="26D9537BE32BFFBDFDE9FB6ABAB7FB62" box="[523,907,1230,1255]" pageId="44" pageNumber="45">T. Ranarilalatiana &amp; S. Holmgren</collectorName>
(
<specimenCode id="DB8A9ED6E32BFFBDFC7EFB6ABD1EFB62" box="[924,1058,1231,1255]" pageId="44" pageNumber="45">MAD14-82</specimenCode>
) (
<collectionCode id="ED3DAE68E32BFFBDFBD8FB6ABDB7FB62" box="[1082,1163,1231,1255]" collectionName="Sweden, Stockholm, Naturhistoriska riksmuseet" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="44" pageNumber="45">NHRS</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B443CF0E32BFFBDFB7CFB6BB8A8FA8E" ID-GBIF-Occurrence="1697157528" collectionCode="NHRS" pageId="44" pageNumber="45" specimenCount="1" typeStatus="paratype">
<typeStatus id="5497880FE32BFFBDFB7CFB6BBC2AFB62" box="[1182,1302,1230,1255]" pageId="44" pageNumber="45" type="paratype">
<emphasis id="B958EABFE32BFFBDFB7CFB6BBC2AFB62" bold="true" box="[1182,1302,1230,1255]" pageId="44" pageNumber="45">Paratypes</emphasis>
</typeStatus>
: Same data as holotype (2
<collectionCode id="ED3DAE68E32BFFBDFEDFFB56B8B1FA8E" box="[317,397,1267,1291]" collectionName="Sweden, Stockholm, Naturhistoriska riksmuseet" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="44" pageNumber="45">NHRS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3366526E32BFFBDFF25FAB3B88AFA3A" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="8B9336ADE32BFFBDFF25FAB3B88AFA3A" blockId="44.[151,1437,1158,2011]" pageId="44" pageNumber="45">
<emphasis id="B958EABFE32BFFBDFF25FAB3B8EEFAAA" bold="true" box="[199,466,1302,1327]" pageId="44" pageNumber="45">Differential Diagnosis.</emphasis>
Similar in color and size to
<taxonomicName id="4C2C4D2EE32BFFBDFCEBFAB2BAAEFAAA" authorityName="Perkins" authorityYear="2017" box="[777,914,1302,1327]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="bergsteni">
<emphasis id="B958EABFE32BFFBDFCEBFAB2BAAEFAAA" box="[777,914,1302,1327]" italics="true" pageId="44" pageNumber="45">H. bergsteni</emphasis>
</taxonomicName>
, but having a differently shaped pronotum that is more coarsely punctate, and elytra that are more ovate. The aedeagi of the two species differ markedly (
<figureCitation id="13172A28E32BFFBDFAD8FA9FBCABFAD6" box="[1338,1431,1338,1363]" captionStart="FIGURES 31 32" captionStartId="42.[151,264,1951,1974]" captionTargetBox="[366,1228,273,1907]" captionTargetId="figure@42.[290,1297,267,1930]" captionTargetPageId="42" captionText="FIGURES 31 32. Hydraena, aedeagi of H. impressicollis (lectotype) and H. bergsteni (holotype)." httpUri="https://zenodo.org/record/1041094/files/figure.png" pageId="44" pageNumber="45">Figs. 32</figureCitation>
,
<figureCitation id="13172A28E32BFFBDFF75FAFBB98BFAF2" box="[151,183,1374,1399]" captionStart="FIGURES 35 36" captionStartId="46.[151,264,1842,1865]" captionText="FIGURES 35 36. Hydraena, aedeagi of holotypes." pageId="44" pageNumber="45">36</figureCitation>
). Clearly related to
<taxonomicName id="4C2C4D2EE32BFFBDFE77FAFABB3BFAF2" authorityName="Perkins" authorityYear="2017" box="[405,519,1374,1399]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="furcula">
<emphasis id="B958EABFE32BFFBDFE77FAFABB3BFAF2" box="[405,519,1374,1399]" italics="true" pageId="44" pageNumber="45">H. furcula</emphasis>
</taxonomicName>
based on general structure of the aedeagus, but easily distinguished therefrom by the size and color differences, and the different shapes of the paramere and distal processes, and thickness of the main piece (
<figureCitation id="13172A28E32BFFBDFEC2FA03B895FA3A" box="[288,425,1446,1471]" captionStart="FIGURES 35 36" captionStartId="46.[151,264,1842,1865]" captionText="FIGURES 35 36. Hydraena, aedeagi of holotypes." pageId="44" pageNumber="45">Figs. 35, 36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3366526E32BFFBCFF25FA6FBAAFFE55" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="description">
<paragraph id="8B9336ADE32BFFBDFF25FA6FBA37F982" blockId="44.[151,1437,1158,2011]" pageId="44" pageNumber="45">
<emphasis id="B958EABFE32BFFBDFF25FA6FB86DFA66" bold="true" box="[199,337,1482,1507]" pageId="44" pageNumber="45">Description</emphasis>
. Size:
<typeStatus id="5497880FE32BFFBDFE41FA6FBB39FA66" box="[419,517,1482,1507]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.37/0.98; head width 0.54; pronotum 0.58/0.70, PA 0.61, PB 0.63; elytra 1.46/0.98.
</paragraph>
<paragraph id="8B9336ADE32BFFBDFF25F9B7BC33F9CA" blockId="44.[151,1437,1158,2011]" pageId="44" pageNumber="45">Head piceous, pronotum and elytra dark brown, anterior border of pronotum and humeral angles of elytra slightly lighter; legs dark brown; maxillary palpi reddish brown, distal ½ of last palpomere not darker.</paragraph>
<paragraph id="8B9336ADE32BFFBDFF25F9FFB82DF886" blockId="44.[151,1437,1158,2011]" pageId="44" pageNumber="45">Head with clypeus finely densely punctate; frons more coarsely punctate than clypeus, especially laterally where interstices narrow walls, medially interstices ca. 0.51xpd, each puncture with short seta. Pronotum densely coarsely punctate throughout, interstices ca. 0.51xpd, shining. Mentum and postmentum coarsely, densely punctate, rugulose, interstices shining. Genae raised, shining, with deeply demarcated posterior ridge, interrupted in midline.</paragraph>
<paragraph id="8B9336ADE32BFFBDFF25F8AABA4BF8CE" blockId="44.[151,1437,1158,2011]" pageId="44" pageNumber="45">Pronotum bisinuate; anterior margin straight behind eyes, emarginate behind frons; PF1 and PF2 absent; PF3 moderately deep; PF4 shallow; PF3 and PF4 shallowly confluent.</paragraph>
<paragraph id="8B9336ADE32BFFBDFF25F8F3BAF4F85E" blockId="44.[151,1437,1158,2011]" pageId="44" pageNumber="45">Elytra weakly arcuate laterally; summit of rather steep posterior declivity at ca. midlength; lateral explanate margins wide; serial punctures round, separated by ca. 0.51xpd, slightly smaller than largest pronotal punctures. Intervals ca. 0.51xpd; discal intervals weakly raised, shining. Apices in dorsal aspect rather sharply conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="8B9336ADE32AFFBCFF25FF32BAAFFE55" blockId="45.[151,1437,151,501]" pageId="45" pageNumber="46">Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/2/1. P1 width ca. 0.5x P2; median carina sinuate in profile. P2 l/w ca.2/1, sides parallel, apex blunt and on same level as anteromedian carina of metaventrite. Plaques roundly raised, converging and anteriorly contiguous with anteromedian carina, separated basally by slightly more than plaque width, located at sides of median depression. Metaventrite anteromedian carina shorter than plaques, as 10 to 13, joining anterior level of plaques, together forming inverted Y-shape; long and strong, slightly arcuate longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at arcuate posterior margin ca. 1.5x P2. Protibia straight, slender. Mesotibia very slightly arcuate, medial margin over ca. middle 1/3 with 5 or 6 short sharp spines. Metatibia very slightly arcuate in distal 1/3, slender. Metafemur moderately slender. Abdominal apex with small apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="C3366526E32AFFBCFF25FE79BD48FE70" box="[199,1140,476,501]" pageId="45" pageNumber="46" type="etymology">
<paragraph id="8B9336ADE32AFFBCFF25FE79BD48FE70" blockId="45.[151,1437,151,501]" box="[199,1140,476,501]" pageId="45" pageNumber="46">
<emphasis id="B958EABFE32AFFBCFF25FE79B872FE70" bold="true" box="[199,334,476,501]" pageId="45" pageNumber="46">Etymology.</emphasis>
Named in reference to the tuning fork shape of the metaventral carinae.
</paragraph>
</subSubSection>
</treatment>
</document>