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<mods:title id="FEF8667ECD5B5B5F31E7D7DECF87AFB7">Redescription of Lepidodactylus flaviocularis (Squamata: Gekkonidae), with the description of a new species from Makira Island, Solomon Islands</mods:title>
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<treatment id="03858789476CFFF1EDEF9CCFFD32F87D" ID-DOI="http://doi.org/10.5281/zenodo.8319962" ID-GBIF-Taxon="212783322" ID-Zenodo-Dep="8319962" LSID="urn:lsid:plazi:treatment:03858789476CFFF1EDEF9CCFFD32F87D" httpUri="http://treatment.plazi.org/id/03858789476CFFF1EDEF9CCFFD32F87D" lastPageId="10" lastPageNumber="572" pageId="4" pageNumber="566">
<subSubSection id="C3366514476CFFFFEDEF9CCFFDFFFF3F" box="[151,528,151,178]" pageId="4" pageNumber="566" type="nomenclature">
<paragraph id="8B93369F476CFFFFEDEF9CCFFDFFFF3F" blockId="4.[151,528,151,213]" box="[151,528,151,178]" pageId="4" pageNumber="566">
<heading id="D0DB81F3476CFFFFEDEF9CCFFDFFFF3F" bold="true" box="[151,528,151,178]" fontSize="11" level="1" pageId="4" pageNumber="566" reason="1">
<emphasis id="B958EA8D476CFFFFEDEF9CCFFDFFFF3F" bold="true" box="[151,528,151,178]" pageId="4" pageNumber="566">
<taxonomicName id="4C2C4D1C476CFFFFEDEF9CCFFE44FF3C" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[151,427,151,178]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D476CFFFFEDEF9CCFFE44FF3C" bold="true" box="[151,427,151,178]" italics="true" pageId="4" pageNumber="566">Lepidodactylus makira</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B57F6476CFFFFECCB9CCFFDFFFF3F" box="[435,528,151,177]" pageId="4" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<paragraph id="8B93369F476CFFFFEDEF9CE3FEFDFF5B" blockId="4.[151,528,151,213]" box="[151,274,187,213]" pageId="4" pageNumber="566">
<figureCitation id="13172A1A476CFFFFEDEF9CE3FF16FF5B" box="[151,249,187,213]" captionStart="FIGURE 2" captionStartId="7.[151,250,1886,1910]" captionTargetBox="[163,1425,181,1862]" captionTargetId="figure-26@7.[163,1425,181,1862]" captionTargetPageId="7" captionText="FIGURE 2. Portraitsin life of (A) an individualof Lepidodactylus flaviocularis from near the summit of Mt.Austen,Guadalcanal, Solomon Islands; and (B) the holotype of Lepidodactylus makira sp. nov. (KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. (A) taken by M. McCoy; (B) by I. Tigulu." figureDoi="http://doi.org/10.5281/zenodo.8313793" httpUri="https://zenodo.org/record/8313793/files/figure.png" pageId="4" pageNumber="566">Figs. 2B</figureCitation>
,
<figureCitation id="13172A1A476CFFFFEC7C9CE3FEFDFF5B" box="[260,274,187,213]" captionStart="FIGURE 3" captionStartId="8.[151,250,1757,1781]" captionTargetBox="[166,1421,188,1726]" captionTargetId="figure-19@8.[159,1428,181,1733]" captionTargetPageId="8" captionText="FIGURE 3. (A) Dorsum, (B) venter, (C) palmar view of left hand, and (D) plantar view of left foot of holotype of Lepidodactylus makira sp. nov. (KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. Scale bars are 1 cm." figureDoi="http://doi.org/10.5281/zenodo.8313795" httpUri="https://zenodo.org/record/8313795/files/figure.png" pageId="4" pageNumber="566">3</figureCitation>
</paragraph>
<paragraph id="8B93369F476CFFFFEDEF9D5BFECEFEEB" blockId="4.[151,1437,259,1978]" pageId="4" pageNumber="566">
<materialsCitation id="3B443CC2476CFFFFEDEF9D5BFECEFEEB" ID-GBIF-Occurrence="4404709304" collectingDate="2018-07-04" collectionCode="KU" collectorName="I. Tigulu" country="Solomon Islands" county="Makira Island" elevation="775" latitude="-10.5639" location="Na'ra Village" longLatPrecision="7" longitude="161.9049" municipality="Central Bauro Highlands" pageId="4" pageNumber="566" specimenCode="KU 350843" specimenCount="1" stateProvince="Makira Province" typeStatus="holotype">
<emphasis id="B958EA8D476CFFFFEDEF9D5BFF11FE93" box="[151,254,259,285]" italics="true" pageId="4" pageNumber="566">
<typeStatus id="5497883D476CFFFFEDEF9D5BFF11FE93" box="[151,254,259,285]" pageId="4" pageNumber="566" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCode id="DB8A9EE4476CFFFFEC749D5CFE7FFE90" box="[268,400,259,286]" collectionCode="KU" country="USA" name="Biodiversity Institute, University of Kansas" pageId="4" pageNumber="566" type="University or college">KU 350843</specimenCode>
(field tag SLT 1204), mature
<specimenCount id="9D2AFD16476CFFFFEF9A9D5BFCDEFE93" box="[738,817,259,285]" pageId="4" pageNumber="566" type="female">female</specimenCount>
, collected by
<collectorName id="26D95349476CFFFFEEB59D5CFBC1FE93" box="[973,1070,259,286]" pageId="4" pageNumber="566">I. Tigulu</collectorName>
at
<location id="8EF36044476CFFFFE9299D5BFB03FE93" LSID="urn:lsid:plazi:treatment:03858789476CFFF1EDEF9CCFFD32F87D:8EF36044476CFFFFE9299D5BFB03FE93" box="[1105,1260,259,285]" country="Solomon Islands" county="Makira Island" latitude="-10.5639" longLatPrecision="7" longitude="161.9049" municipality="Central Bauro Highlands" name="Na'ra Village" pageId="4" pageNumber="566" stateProvince="Makira Province">Nara Village</location>
,
<location id="8EF36044476CFFFFE9809D5CFA77FE90" LSID="urn:lsid:plazi:treatment:03858789476CFFF1EDEF9CCFFD32F87D:8EF36044476CFFFFE9809D5CFA77FE90" box="[1272,1432,260,286]" country="Solomon Islands" county="Makira Island" latitude="-10.5639" longLatPrecision="7" longitude="161.9049" municipality="Central Bauro Highlands" name="Naepaepa Mt." pageId="4" pageNumber="566" stateProvince="Makira Province">Naepaepa Mt.</location>
,
<collectingMunicipality id="6BF7ACE5476CFFFFEDEF9D7FFE5DFECF" box="[151,434,295,322]" pageId="4" pageNumber="566">Central Bauro Highlands</collectingMunicipality>
,
<geoCoordinate id="EE185058476CFFFFECC69D7FFDD9FECC" box="[446,566,295,322]" degrees="10.5639" direction="south" orientation="latitude" pageId="4" pageNumber="566" precision="5" value="-10.5639">10.5639°S</geoCoordinate>
,
<geoCoordinate id="EE185058476CFFFFEF399D7FFD25FECF" box="[577,714,295,321]" degrees="161.9049" direction="east" orientation="longitude" pageId="4" pageNumber="566" precision="5" value="161.9049">161.9049°E</geoCoordinate>
,
<elevation id="0001D1AC476CFFFFEFAD9D7FFCB6FECF" box="[725,857,295,322]" metricMagnitude="2" metricUnit="m" metricValue="7.75" pageId="4" pageNumber="566" unit="m" value="775.0">
<quantity id="4CD49B7A476CFFFFEFAD9D7FFCF3FECC" box="[725,796,295,322]" metricMagnitude="2" metricUnit="m" metricValue="7.75" pageId="4" pageNumber="566" unit="m" value="775.0">775 m</quantity>
a.s.l.
</elevation>
,
<collectingCounty id="62F24E13476CFFFFEE1D9D7FFBEBFECF" box="[869,1028,295,321]" pageId="4" pageNumber="566">Makira Island</collectingCounty>
,
<collectingRegion id="49E8F87D476CFFFFE9689D7FFB21FECF" box="[1040,1230,295,321]" country="Solomon Islands" name="Makira" pageId="4" pageNumber="566">Makira Province</collectingRegion>
,
<collectingCountry id="F33B760F476CFFFFE9A29D7FFA77FECF" box="[1242,1432,295,321]" name="Solomon Islands" pageId="4" pageNumber="566">Solomon Islands</collectingCountry>
,
<date id="FF92105F476CFFFFEDEF9D13FEF1FEEB" box="[151,286,331,357]" pageId="4" pageNumber="566" value="2018-07-04">
<collectingDate id="EFD6E9B7476CFFFFEDEF9D13FEF1FEEB" box="[151,286,331,357]" pageId="4" pageNumber="566" value="2018-07-04">4 July 2018</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8B93369F476CFFFFEDBF9D28FD06FE5F" blockId="4.[151,1437,259,1978]" pageId="4" pageNumber="566">
<materialsCitation id="3B443CC2476CFFFFEDBF9D28FB72FE23" ID-GBIF-Occurrence="4404709301" collectionCode="BMNH" country="Solomon Islands" county="The" location="Wainoni Bay" municipality="Makira Island" pageId="4" pageNumber="566" specimenCode="BMNH 1973.222" specimenCount="1" stateProvince="Makira Province" typeStatus="paratype">
<emphasis id="B958EA8D476CFFFFEDBF9D28FEC1FE07" box="[199,302,368,393]" italics="true" pageId="4" pageNumber="566">
<typeStatus id="5497883D476CFFFFEDBF9D28FEC1FE07" box="[199,302,368,393]" pageId="4" pageNumber="566" type="paratype">Paratype</typeStatus>
</emphasis>
.
<specimenCode id="DB8A9EE4476CFFFFEC459D28FDE9FE07" box="[317,518,367,394]" collectionCode="BMNH" pageId="4" pageNumber="566">BMNH 1973.222</specimenCode>
, mature
<specimenCount id="9D2AFD16476CFFFFEF119D37FD4DFE07" box="[617,674,367,393]" pageId="4" pageNumber="566" type="male">male</specimenCount>
, collected
<quantity id="4CD49B7A476CFFFFEE659D28FCBEFE07" box="[797,849,367,394]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="4" pageNumber="566" unit="mi" value="7.0">
<locationDeviation id="9A212FBD476CFFFFEE659D28FCBEFE07" box="[797,849,367,394]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="4" pageNumber="566" unit="mi" value="7.0">7 mi</locationDeviation>
</quantity>
(
<quantity id="4CD49B7A476CFFFFEE199D37FC2EFE07" box="[865,961,367,393]" metricMagnitude="4" metricUnit="m" metricValue="1.16" pageId="4" pageNumber="566" unit="km" value="11.6">
<locationDeviation id="9A212FBD476CFFFFEE199D37FC2EFE07" box="[865,961,367,393]" metricMagnitude="4" metricUnit="m" metricValue="1.16" pageId="4" pageNumber="566" unit="km" value="11.6">11.6 km</locationDeviation>
</quantity>
)
<locationDeviation id="9A212FBD476CFFFFEEA99D37FB89FE07" box="[977,1126,367,393]" location="Wainoni" pageId="4" pageNumber="566">S of Wainoni</locationDeviation>
(presumably
<location id="8EF36044476CFFFFE87B9D37FA77FE04" LSID="urn:lsid:plazi:treatment:03858789476CFFF1EDEF9CCFFD32F87D:8EF36044476CFFFFE87B9D37FA77FE04" box="[1283,1432,367,394]" country="Solomon Islands" county="The" municipality="Makira Island" name="Wainoni Bay" pageId="4" pageNumber="566" stateProvince="Makira Province">Wainoni Bay</location>
, since I can find no village of that name),
<collectingMunicipality id="6BF7ACE5476CFFFFEF1D9DCBFCEBFE23" box="[613,772,403,429]" pageId="4" pageNumber="566">Makira Island</collectingMunicipality>
,
<collectingRegion id="49E8F87D476CFFFFEE689DCBFC21FE23" box="[784,974,403,429]" country="Solomon Islands" name="Makira" pageId="4" pageNumber="566">Makira Province</collectingRegion>
,
<collectingCountry id="F33B760F476CFFFFEEA29DCBFB76FE23" box="[986,1177,403,429]" name="Solomon Islands" pageId="4" pageNumber="566">Solomon Islands</collectingCountry>
.
</materialsCitation>
<collectingCounty id="62F24E13476CFFFFE9DB9DCBFB20FE23" box="[1187,1231,403,429]" pageId="4" pageNumber="566">The</collectingCounty>
BMNH catalogue lists no collector or collection date for this specimen.
</paragraph>
</subSubSection>
<subSubSection id="C3366514476CFFFFEDBF9D83FCE8F837" pageId="4" pageNumber="566" type="diagnosis">
<paragraph id="8B93369F476CFFFFEDBF9D83FB65FCB7" blockId="4.[151,1437,259,1978]" pageId="4" pageNumber="566">
<emphasis id="B958EA8D476CFFFFEDBF9D83FEADFE7B" bold="true" box="[199,322,475,501]" pageId="4" pageNumber="566">Diagnosis.</emphasis>
An intermediately sized (adult female SVL
<quantity id="4CD49B7A476CFFFFEE5D9D83FC65FE78" box="[805,906,475,502]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" pageId="4" pageNumber="566" unit="mm" value="52.5">52.5 mm</quantity>
, adult male 45.0 mm) species of
<taxonomicName id="4C2C4D1C476CFFFFE9899D83FA73FE7B" box="[1265,1436,475,501]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="genus">
<emphasis id="B958EA8D476CFFFFE9899D83FA73FE7B" box="[1265,1436,475,501]" italics="true" pageId="4" pageNumber="566">Lepidodactylus</emphasis>
</taxonomicName>
having 4445 enlarged pore-bearing precloacal/femoral scales in a continous row extending to distal end of each thigh, 45 precloacal/femoral pores in sole male, enlarged scales of pore-bearing series entirely separated from patch of enlarged pubic scales by intervening row of tiny scales, entirely undivided subterminal lamellae on all toes though two on each toe may be grooved, 1719 T4 lamellae, 12 T1 lamellae, moderately long toes (T4L/ SVL = 0.100.11), lamellae occupying almost all of toes or extending onto palm (T4 lamellaeL/T4L = 0.941.04), toes fairly wide (T4W/T4L = 0.320.36) with extensive webbing (T3T4webL/T4L = 0.280.30, T4T5webL/T4L = 0.220.28), two internasals between supranasals along posterior margin of rostral, 810 supralabials to center of eye, circumorbital scales dark brown, foot webbing dark brown dorsally and ventrally, dorsum dark brown with darker-brown dorsolateral and lateral blotches, and pale ventral field ~15 scale rows wide.
</paragraph>
<paragraph id="8B93369F476CFFFFEDBF9F1BFD85FBBB" blockId="4.[151,1437,259,1978]" pageId="4" pageNumber="566">
<emphasis id="B958EA8D476CFFFFEDBF9F1BFDBCFCD3" bold="true" box="[199,595,835,861]" pageId="4" pageNumber="566">Comparisons with other species.</emphasis>
The undivided lamellae under the toes place
<taxonomicName id="4C2C4D1C476CFFFFE9139F1BFA9FFCD3" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[1131,1392,835,861]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D476CFFFFE9139F1BFA9FFCD3" box="[1131,1392,835,861]" italics="true" pageId="4" pageNumber="566">Lepidodactylus makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D476CFFFFE8039F1BFF27FC0F" bold="true" pageId="4" pageNumber="566">
<taxonomicNameLabel id="A26B57F6476CFFFFE8039F1BFF27FC0F" pageId="4" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in
<bibRefCitation id="EFBD4B6E476CFFFFED8A9F30FDF5FC0F" author="Brown, W. C. &amp; Parker, F." box="[242,538,871,898]" pageId="4" pageNumber="566" pagination="253 - 265" refId="ref9603" refString="Brown, W. C. &amp; Parker, F. (1977) Lizards of the genus Lepidodactylus (Gekkonidae) from the Indo-Australian Archipelago and the islands of the Pacific, with descriptions of new species. Proceedings of the California Academy of Sciences, 41, 253 - 265." type="journal article" year="1977">Brown &amp; Parkers (1977)</bibRefCitation>
phenetic Group I. Hence, it is readily distinguished from Melanesian species belonging to Group II (
<taxonomicName id="4C2C4D1C476CFFFFECE19FD4FDABFC2B" authority="Ineich" authorityName="Ineich" authorityYear="2008" box="[409,580,907,933]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="buleli">
<emphasis id="B958EA8D476CFFFFECE19FD4FE19FC2B" box="[409,502,907,933]" italics="true" pageId="4" pageNumber="566">L. buleli</emphasis>
Ineich
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEF369FD4FCCCFC28" authority="Kraus" authorityName="Kraus" authorityYear="2019" box="[590,803,907,934]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="dialeukos">
<emphasis id="B958EA8D476CFFFFEF369FD4FD39FC2B" box="[590,726,907,933]" italics="true" pageId="4" pageNumber="566">L. dialeukos</emphasis>
Kraus
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEE549FD4FBC1FC2B" authority="Boulenger" authorityName="Boulenger" authorityYear="1897" box="[812,1070,907,933]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="gardineri">
<emphasis id="B958EA8D476CFFFFEE549FD4FC5DFC2B" box="[812,946,907,933]" italics="true" pageId="4" pageNumber="566">L. gardineri</emphasis>
Boulenger
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFE9419FD4FB4EFC2B" authorityName="Boulenger" authorityYear="1884" box="[1081,1185,908,933]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="guppyi">
<emphasis id="B958EA8D476CFFFFE9419FD4FB4EFC2B" box="[1081,1185,908,933]" italics="true" pageId="4" pageNumber="566">L. guppyi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFE9D79FD4FA77FC28" authority="Kraus" authorityName="Kraus" authorityYear="2019" box="[1199,1432,907,934]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="kwasnickae">
<emphasis id="B958EA8D476CFFFFE9D79FD4FAA3FC2B" box="[1199,1356,907,933]" italics="true" pageId="4" pageNumber="566">L. kwasnickae</emphasis>
Kraus
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEDEF9FE8FEC6FC47" box="[151,297,943,969]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="laticinctus">
<emphasis id="B958EA8D476CFFFFEDEF9FE8FEC6FC47" box="[151,297,943,969]" italics="true" pageId="4" pageNumber="566">L. laticinctus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEC4E9FE8FDEEFC44" authority="Kraus" authorityName="Kraus" authorityYear="2019" box="[310,513,943,970]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="mitchelli">
<emphasis id="B958EA8D476CFFFFEC4E9FE8FE5AFC47" box="[310,437,943,969]" italics="true" pageId="4" pageNumber="566">L. mitchelli</emphasis>
Kraus
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEF739FE8FC6BFC47" authority="Brown &amp; Parker" authorityName="Brown &amp; Parker" authorityYear="1977" box="[523,900,943,970]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="B958EA8D476CFFFFEF739FE8FD2BFC47" box="[523,708,944,969]" italics="true" pageId="4" pageNumber="566">L. novaeguineae</emphasis>
Brown &amp; Parker
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEEF69FE8FBEEFC47" authorityName="Boulenger" authorityYear="1885" box="[910,1025,943,969]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="pulcher">
<emphasis id="B958EA8D476CFFFFEEF69FE8FBEEFC47" box="[910,1025,943,969]" italics="true" pageId="4" pageNumber="566">L. pulcher</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFE9769FE8FB97FC47" baseAuthorityName="Brown &amp; Tanner" baseAuthorityYear="1949" box="[1038,1144,943,969]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="shebae">
<emphasis id="B958EA8D476CFFFFE9769FE8FB97FC47" box="[1038,1144,943,969]" italics="true" pageId="4" pageNumber="566">L. shebae</emphasis>
</taxonomicName>
), which have two or more divided subterminal lamellae, and from Group III species (
<taxonomicName id="4C2C4D1C476CFFFFEE5C9F8CFC71FC63" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1836" box="[804,926,979,1005]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="B958EA8D476CFFFFEE5C9F8CFC71FC63" box="[804,926,979,1005]" italics="true" pageId="4" pageNumber="566">L. lugubris</emphasis>
</taxonomicName>
[Duméril &amp; Bibron],
<taxonomicName id="4C2C4D1C476CFFFFE9E99F8CFCC0FB9F" authority="Stubbs, Karin, Arifin, Iskandar, Arida, Reilly, Bloch, Kusnadi &amp; McGuire" authorityName="Stubbs, Karin, Arifin, Iskandar, Arida, Reilly, Bloch, Kusnadi &amp; McGuire" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="pantai">
<emphasis id="B958EA8D476CFFFFE9E99F8CFB19FC63" box="[1169,1270,980,1005]" italics="true" pageId="4" pageNumber="566">L. pantai</emphasis>
Stubbs, Karin, Arifin, Iskandar, Arida, Reilly, Bloch, Kusnadi &amp; McGuire
</taxonomicName>
, and
<taxonomicName id="4C2C4D1C476CFFFFEE129FA0FB94FB9F" authority="Boulenger" authorityName="Boulenger" authorityYear="1887" box="[874,1147,1015,1041]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="woodfordi">
<emphasis id="B958EA8D476CFFFFEE129FA0FC15FB9F" box="[874,1018,1015,1041]" italics="true" pageId="4" pageNumber="566">L. woodfordi</emphasis>
Boulenger
</taxonomicName>
), which have the terminal and several subterminal lamellae divided.
</paragraph>
<paragraph id="8B93369F476CFFFFEDBF9818FE92F9FB" blockId="4.[151,1437,259,1978]" pageId="4" pageNumber="566">
From other Group I species,
<taxonomicName id="4C2C4D1C476CFFFFEF779818FD93FBD7" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[527,636,1087,1113]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D476CFFFFEF779818FD93FBD7" box="[527,636,1087,1113]" italics="true" pageId="4" pageNumber="566">L. makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D476CFFFFEFFD9867FD30FBD7" bold="true" box="[645,735,1087,1113]" pageId="4" pageNumber="566">
<taxonomicNameLabel id="A26B57F6476CFFFFEFFD9867FD30FBD7" box="[645,735,1087,1113]" pageId="4" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from
<taxonomicName id="4C2C4D1C476CFFFFEEA69818FB8BFBD7" authorityName="Kraus" authorityYear="2019" box="[990,1124,1088,1113]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="aignanus">
<emphasis id="B958EA8D476CFFFFEEA69818FB8BFBD7" box="[990,1124,1088,1113]" italics="true" pageId="4" pageNumber="566">L. aignanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFE90A9818FB3DFBD7" baseAuthorityName="Boulenger" baseAuthorityYear="1889" box="[1138,1234,1087,1113]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="listeri">
<emphasis id="B958EA8D476CFFFFE90A9818FB3DFBD7" box="[1138,1234,1087,1113]" italics="true" pageId="4" pageNumber="566">L. listeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFE9999818FAA9FBD7" authorityName="Schmidt" authorityYear="1923" box="[1249,1350,1088,1113]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="manni">
<emphasis id="B958EA8D476CFFFFE9999818FAA9FBD7" box="[1249,1350,1088,1113]" italics="true" pageId="4" pageNumber="566">L. manni</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2C4D1C476CFFFFE8FE9818FEEDFBF3" authorityName="Brown &amp; Parker" authorityYear="1977" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="B958EA8D476CFFFFE8FE9818FEEDFBF3" italics="true" pageId="4" pageNumber="566">L. orientalis</emphasis>
</taxonomicName>
in having a continuous row of 45 precloacal/femoral pores in the sole male (vs. 31 or fewer precloacal/ femoral pores in
<taxonomicName id="4C2C4D1C476CFFFFEC2498D0FE0CFB2F" authorityName="Kraus" authorityYear="2019" box="[348,483,1160,1185]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="aignanus">
<emphasis id="B958EA8D476CFFFFEC2498D0FE0CFB2F" box="[348,483,1160,1185]" italics="true" pageId="4" pageNumber="566">L. aignanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEC8B98D0FDBBFB2F" baseAuthorityName="Boulenger" baseAuthorityYear="1889" box="[499,596,1159,1185]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="listeri">
<emphasis id="B958EA8D476CFFFFEC8B98D0FDBBFB2F" box="[499,596,1159,1185]" italics="true" pageId="4" pageNumber="566">L. listeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEF1C98D0FD25FB2F" authorityName="Schmidt" authorityYear="1923" box="[612,714,1160,1185]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="manni">
<emphasis id="B958EA8D476CFFFFEF1C98D0FD25FB2F" box="[612,714,1160,1185]" italics="true" pageId="4" pageNumber="566">L. manni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEFA298D0FC8AFB2F" authorityName="Brown &amp; Parker" authorityYear="1977" box="[730,869,1159,1185]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="B958EA8D476CFFFFEFA298D0FC8AFB2F" box="[730,869,1159,1185]" italics="true" pageId="4" pageNumber="566">L. orientalis</emphasis>
</taxonomicName>
) and far more T4 lamellae (1719 vs.
<quantity id="4CD49B7A476CFFFFE85E98DFFA92FB2F" box="[1318,1405,1159,1185]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="4" pageNumber="566" unit="in" value="10.5" valueMax="12.0" valueMin="9.0">912 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFE8FE98D0FF11FB4B" authorityName="Kraus" authorityYear="2019" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="aignanus">
<emphasis id="B958EA8D476CFFFFE8FE98D0FF11FB4B" italics="true" pageId="4" pageNumber="566">L. aignanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEC7398F4FE86FB4B" baseAuthorityName="Boulenger" baseAuthorityYear="1889" box="[267,361,1195,1221]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="listeri">
<emphasis id="B958EA8D476CFFFFEC7398F4FE86FB4B" box="[267,361,1195,1221]" italics="true" pageId="4" pageNumber="566">L. listeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEC0E98F4FE36FB4B" authorityName="Schmidt" authorityYear="1923" box="[374,473,1196,1221]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="manni">
<emphasis id="B958EA8D476CFFFFEC0E98F4FE36FB4B" box="[374,473,1196,1221]" italics="true" pageId="4" pageNumber="566">L. manni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEC9F98F4FD81FB4B" authorityName="Brown &amp; Parker" authorityYear="1977" box="[487,622,1195,1221]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="B958EA8D476CFFFFEC9F98F4FD81FB4B" box="[487,622,1195,1221]" italics="true" pageId="4" pageNumber="566">L. orientalis</emphasis>
</taxonomicName>
); from
<taxonomicName id="4C2C4D1C476CFFFFEFBB98F4FCD6FB4B" baseAuthorityName="Boulenger" baseAuthorityYear="1885" box="[707,825,1195,1221]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B958EA8D476CFFFFEFBB98F4FCD6FB4B" box="[707,825,1195,1221]" italics="true" pageId="4" pageNumber="566">L. pumilus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEE3D98F4FC17FB4B" authorityName="Kraus &amp; Oliver" authorityYear="2020" box="[837,1016,1195,1221]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="sacrolineatus">
<emphasis id="B958EA8D476CFFFFEE3D98F4FC17FB4B" box="[837,1016,1195,1221]" italics="true" pageId="4" pageNumber="566">L. sacrolineatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2C4D1C476CFFFFE94C98F4FB48FB4B" authorityName="Kraus" authorityYear="2019" box="[1076,1191,1195,1221]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="zweifeli">
<emphasis id="B958EA8D476CFFFFE94C98F4FB48FB4B" box="[1076,1191,1195,1221]" italics="true" pageId="4" pageNumber="566">L. zweifeli</emphasis>
</taxonomicName>
by its continuous row of precloacal/femoral pores and enlarged precloacal/femoral scales (vs. enlarged precloacal scales/pores divided from enlarged femoral scales/pores by intervening smaller scales lacking pores, producing three discrete pore series in
<taxonomicName id="4C2C4D1C476CFFFFEDCD9940FEC3FABF" baseAuthorityName="Boulenger" baseAuthorityYear="1885" box="[181,300,1303,1329]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="B958EA8D476CFFFFEDCD9940FEC3FABF" box="[181,300,1303,1329]" italics="true" pageId="4" pageNumber="566">L. pumilus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C4D1C476CFFFFEC429940FE01FABF" authorityName="Kraus &amp; Oliver" authorityYear="2020" box="[314,494,1303,1329]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="sacrolineatus">
<emphasis id="B958EA8D476CFFFFEC429940FE01FABF" box="[314,494,1303,1329]" italics="true" pageId="4" pageNumber="566">L. sacrolineatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2C4D1C476CFFFFEF559940FD4EFABF" authorityName="Kraus" authorityYear="2019" box="[557,673,1303,1329]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="zweifeli">
<emphasis id="B958EA8D476CFFFFEF559940FD4EFABF" box="[557,673,1303,1329]" italics="true" pageId="4" pageNumber="566">L. zweifeli</emphasis>
</taxonomicName>
); from
<taxonomicName id="4C2C4D1C476CFFFFEF8F9940FC83FABF" box="[759,876,1304,1329]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="magnus">
<emphasis id="B958EA8D476CFFFFEF8F9940FC83FABF" box="[759,876,1304,1329]" italics="true" pageId="4" pageNumber="566">L. magnus</emphasis>
</taxonomicName>
by its much smaller size (45.0
<quantity id="4CD49B7A476CFFFFE9AD994FFAD5FABC" box="[1237,1338,1303,1330]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" pageId="4" pageNumber="566" unit="mm" value="52.5">52.5 mm</quantity>
SVL vs.
<quantity id="4CD49B7A476CFFFFEDEF9963FEFEFAD8" box="[151,273,1339,1366]" metricMagnitude="-2" metricUnit="m" metricValue="6.05" metricValueMax="7.1" metricValueMin="5.0" pageId="4" pageNumber="566" unit="mm" value="60.5" valueMax="71.0" valueMin="50.0">5071 mm</quantity>
in
<taxonomicName id="4C2C4D1C476CFFFFEC409964FE42FADB" box="[312,429,1340,1365]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="magnus">
<emphasis id="B958EA8D476CFFFFEC409964FE42FADB" box="[312,429,1340,1365]" italics="true" pageId="4" pageNumber="566">L. magnus</emphasis>
</taxonomicName>
), greater number of T4 lamellae (1719 vs.
<quantity id="4CD49B7A476CFFFFEED09963FBE4FADB" box="[936,1035,1339,1365]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.556" metricValueMin="2.794" pageId="4" pageNumber="566" unit="in" value="12.5" valueMax="14.0" valueMin="11.0">1114 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFE96B9964FB66FADB" box="[1043,1161,1340,1365]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="magnus">
<emphasis id="B958EA8D476CFFFFE96B9964FB66FADB" box="[1043,1161,1340,1365]" italics="true" pageId="4" pageNumber="566">L. magnus</emphasis>
</taxonomicName>
), and greater amount of toe webbing (T3T4webL/T4L = 0.280.30 vs.
<quantity id="4CD49B7A476CFFFFEFD19907FCD8FAF7" box="[681,823,1375,1401]" metricMagnitude="-3" metricUnit="m" metricValue="3.937" metricValueMax="4.826" metricValueMin="3.048" pageId="4" pageNumber="566" unit="in" value="0.155" valueMax="0.19" valueMin="0.12">0.120.19 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFEE479938FC5AFAF7" box="[831,949,1376,1401]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="magnus">
<emphasis id="B958EA8D476CFFFFEE479938FC5AFAF7" box="[831,949,1376,1401]" italics="true" pageId="4" pageNumber="566">L. magnus</emphasis>
</taxonomicName>
); from
<taxonomicName id="4C2C4D1C476CFFFFE9759938FB8EFAF7" baseAuthorityName="Kopstein" baseAuthorityYear="1926" box="[1037,1121,1376,1401]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="oorti">
<emphasis id="B958EA8D476CFFFFE9759938FB8EFAF7" box="[1037,1121,1376,1401]" italics="true" pageId="4" pageNumber="566">L. oorti</emphasis>
</taxonomicName>
in its greater number of T4 lamellae (1719 vs.
<quantity id="4CD49B7A476CFFFFECF899DBFE0AFA13" box="[384,485,1411,1437]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.556" metricValueMin="3.048" pageId="4" pageNumber="566" unit="in" value="13.0" valueMax="14.0" valueMin="12.0">1214 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFEC9699DCFDABFA13" baseAuthorityName="Kopstein" baseAuthorityYear="1926" box="[494,580,1412,1437]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="oorti">
<emphasis id="B958EA8D476CFFFFEC9699DCFDABFA13" box="[494,580,1412,1437]" italics="true" pageId="4" pageNumber="566">L. oorti</emphasis>
</taxonomicName>
) and T1 lamellae (1112 vs.
<quantity id="4CD49B7A476CFFFFEEE099DBFC00FA13" box="[920,1007,1411,1437]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="4" pageNumber="566" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">910 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFEE8F99DCFBA2FA13" baseAuthorityName="Kopstein" baseAuthorityYear="1926" box="[1015,1101,1412,1437]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="oorti">
<emphasis id="B958EA8D476CFFFFEE8F99DCFBA2FA13" box="[1015,1101,1412,1437]" italics="true" pageId="4" pageNumber="566">L. oorti</emphasis>
</taxonomicName>
), greater number of enlarged precloacal/femoral scales (4445 vs.
<quantity id="4CD49B7A476CFFFFEF4B99FFFD7AFA4F" box="[563,661,1447,1473]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" metricValueMax="9.144" metricValueMin="8.128" pageId="4" pageNumber="566" unit="in" value="34.0" valueMax="36.0" valueMin="32.0">3236 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFEFE399F0FD01FA4F" baseAuthorityName="Kopstein" baseAuthorityYear="1926" box="[667,750,1448,1473]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="oorti">
<emphasis id="B958EA8D476CFFFFEFE399F0FD01FA4F" box="[667,750,1448,1473]" italics="true" pageId="4" pageNumber="566">L. oorti</emphasis>
</taxonomicName>
) and pores (45 vs.
<quantity id="4CD49B7A476CFFFFEEB899FFFBCDFA4F" box="[960,1058,1447,1473]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" metricValueMax="7.62" metricValueMin="7.112" pageId="4" pageNumber="566" unit="in" value="29.0" valueMax="30.0" valueMin="28.0">2830 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFE95099F0FB94FA4F" baseAuthorityName="Kopstein" baseAuthorityYear="1926" box="[1064,1147,1448,1473]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="oorti">
<emphasis id="B958EA8D476CFFFFE95099F0FB94FA4F" box="[1064,1147,1448,1473]" italics="true" pageId="4" pageNumber="566">L. oorti</emphasis>
</taxonomicName>
), and greater toe webbing (all toes webbed and T3T4webL/T4L = 0.280.30 vs. basal webbing only between T3 and T
<quantity id="4CD49B7A476CFFFFE9E39993FB2AFA6B" box="[1179,1221,1483,1509]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="4" pageNumber="566" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFE9B39994FAF0FA6B" baseAuthorityName="Kopstein" baseAuthorityYear="1926" box="[1227,1311,1484,1509]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="oorti">
<emphasis id="B958EA8D476CFFFFE9B39994FAF0FA6B" box="[1227,1311,1484,1509]" italics="true" pageId="4" pageNumber="566">L. oorti</emphasis>
</taxonomicName>
); and from
<taxonomicName id="4C2C4D1C476CFFFFEDEF99A8FEFBF987" authorityName="Gibbons &amp; Brown" authorityYear="1988" box="[151,276,1520,1545]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="euaensis">
<emphasis id="B958EA8D476CFFFFEDEF99A8FEFBF987" box="[151,276,1520,1545]" italics="true" pageId="4" pageNumber="566">L. euaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C4D1C476CFFFFEC3299A8FE5AF987" authorityName="Brown &amp; Parker" authorityYear="1977" box="[330,437,1519,1545]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="mutahi">
<emphasis id="B958EA8D476CFFFFEC3299A8FE5AF987" box="[330,437,1519,1545]" italics="true" pageId="4" pageNumber="566">L. mutahi</emphasis>
</taxonomicName>
in its greater number of T4 lamellae (1719 vs.
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<taxonomicName id="4C2C4D1C476CFFFFE94E99A8FB5CF987" authorityName="Gibbons &amp; Brown" authorityYear="1988" box="[1078,1203,1520,1545]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="euaensis">
<emphasis id="B958EA8D476CFFFFE94E99A8FB5CF987" box="[1078,1203,1520,1545]" italics="true" pageId="4" pageNumber="566">L. euaensis</emphasis>
</taxonomicName>
,
<quantity id="4CD49B7A476CFFFFE9B899B7FACDF987" box="[1216,1314,1519,1545]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="4" pageNumber="566" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">1011 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFE85099A8FA7CF987" authorityName="Brown &amp; Parker" authorityYear="1977" box="[1320,1427,1519,1545]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="mutahi">
<emphasis id="B958EA8D476CFFFFE85099A8FA7CF987" box="[1320,1427,1519,1545]" italics="true" pageId="4" pageNumber="566">L. mutahi</emphasis>
</taxonomicName>
) covering most of toe or more (T4 lamellaeL/T4L = 0.941.04 vs. ~
<quantity id="4CD49B7A476CFFFFEEFD9A4BFC3BF9A3" box="[901,980,1555,1582]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" pageId="4" pageNumber="566" unit="in" value="0.75">0.75 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFEEA39A4CFBB6F9A3" authorityName="Gibbons &amp; Brown" authorityYear="1988" box="[987,1113,1556,1581]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="euaensis">
<emphasis id="B958EA8D476CFFFFEEA39A4CFBB6F9A3" box="[987,1113,1556,1581]" italics="true" pageId="4" pageNumber="566">L. euaensis</emphasis>
</taxonomicName>
, ~
<quantity id="4CD49B7A476CFFFFE90D9A4BFB2AF9A3" box="[1141,1221,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.7018" pageId="4" pageNumber="566" unit="in" value="0.67">0.67 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFE9B49A4CFAD8F9A3" authorityName="Brown &amp; Parker" authorityYear="1977" box="[1228,1335,1555,1581]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="mutahi">
<emphasis id="B958EA8D476CFFFFE9B49A4CFAD8F9A3" box="[1228,1335,1555,1581]" italics="true" pageId="4" pageNumber="566">L. mutahi</emphasis>
</taxonomicName>
), greater number of enlarged precloacal/femoral scales (4445 vs.
<quantity id="4CD49B7A476CFFFFEE589A6FFC6BF9DF" box="[800,900,1591,1617]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" metricValueMax="9.144" metricValueMin="7.112" pageId="4" pageNumber="566" unit="in" value="32.0" valueMax="36.0" valueMin="28.0">2836 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFEEF49A60FC16F9DF" authorityName="Brown &amp; Parker" authorityYear="1977" box="[908,1017,1591,1617]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="mutahi">
<emphasis id="B958EA8D476CFFFFEEF49A60FC16F9DF" box="[908,1017,1591,1617]" italics="true" pageId="4" pageNumber="566">L. mutahi</emphasis>
</taxonomicName>
) and pores (45 vs.
<quantity id="4CD49B7A476CFFFFE9AC9A6FFAE1F9DF" box="[1236,1294,1591,1617]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" pageId="4" pageNumber="566" unit="in" value="33.0">33 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFE86E9A60FA7AF9DF" authorityName="Gibbons &amp; Brown" authorityYear="1988" box="[1302,1429,1592,1617]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="euaensis">
<emphasis id="B958EA8D476CFFFFE86E9A60FA7AF9DF" box="[1302,1429,1592,1617]" italics="true" pageId="4" pageNumber="566">L. euaensis</emphasis>
</taxonomicName>
,
<quantity id="4CD49B7A476CFFFFEDEF9A03FF15F9FB" box="[151,250,1627,1653]" metricMagnitude="-1" metricUnit="m" metricValue="7.747" metricValueMax="8.636" metricValueMin="6.858" pageId="4" pageNumber="566" unit="in" value="30.5" valueMax="34.0" valueMin="27.0">2734 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFEC799A04FE82F9FB" authorityName="Brown &amp; Parker" authorityYear="1977" box="[257,365,1627,1653]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="mutahi">
<emphasis id="B958EA8D476CFFFFEC799A04FE82F9FB" box="[257,365,1627,1653]" italics="true" pageId="4" pageNumber="566">L. mutahi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B93369F476CFFFFEDBF9A27FCE8F837" blockId="4.[151,1437,259,1978]" pageId="4" pageNumber="566">
<taxonomicName id="4C2C4D1C476CFFFFEDBF9A27FE27F917" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[199,456,1663,1689]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D476CFFFFEDBF9A27FE27F917" box="[199,456,1663,1689]" italics="true" pageId="4" pageNumber="566">Lepidodactylus makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D476CFFFFECB79A27FDC7F917" bold="true" box="[463,552,1663,1689]" pageId="4" pageNumber="566">
<taxonomicNameLabel id="A26B57F6476CFFFFECB79A27FDC7F917" box="[463,552,1663,1689]" pageId="4" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="4C2C4D1C476CFFFFEF8E9AD8FC4EF917" box="[758,929,1663,1689]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D476CFFFFEF8E9AD8FC4EF917" box="[758,929,1663,1689]" italics="true" pageId="4" pageNumber="566">L. flaviocularis</emphasis>
</taxonomicName>
from nearby
<collectingRegion id="49E8F87D476CFFFFE94F9A27FB28F917" box="[1079,1223,1663,1689]" country="Solomon Islands" name="Guadalcanal" pageId="4" pageNumber="566">Guadalcanal</collectingRegion>
. From that species,
<taxonomicName id="4C2C4D1C476CFFFFEDEF9AFCFEEAF933" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[151,261,1699,1725]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D476CFFFFEDEF9AFCFEEAF933" box="[151,261,1699,1725]" italics="true" pageId="4" pageNumber="566">L. makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D476CFFFFEC779AFBFE85F933" bold="true" box="[271,362,1699,1725]" pageId="4" pageNumber="566">
<taxonomicNameLabel id="A26B57F6476CFFFFEC779AFBFE85F933" box="[271,362,1699,1725]" pageId="4" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs in its larger size (female SVL
<quantity id="4CD49B7A476CFFFFEE669AFBFC6BF930" box="[798,900,1699,1726]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" pageId="4" pageNumber="566" unit="mm" value="52.5">52.5 mm</quantity>
vs. 46.0 mm in
<taxonomicName id="4C2C4D1C476CFFFFE9399AFCFB00F933" box="[1089,1263,1699,1725]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D476CFFFFE9399AFCFB00F933" box="[1089,1263,1699,1725]" italics="true" pageId="4" pageNumber="566">L. flaviocularis</emphasis>
</taxonomicName>
) and in having more extensive toe webbing (T3T4webL = 0.280.30 vs.
<quantity id="4CD49B7A476CFFFFEE4B9A9FFC2CF96F" box="[819,963,1735,1761]" metricMagnitude="-3" metricUnit="m" metricValue="6.731" metricValueMax="6.858" metricValueMin="6.604" pageId="4" pageNumber="566" unit="in" value="0.265" valueMax="0.27" valueMin="0.26">0.260.27 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFEEB59A90FB94F96F" box="[973,1147,1735,1761]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D476CFFFFEEB59A90FB94F96F" box="[973,1147,1735,1761]" italics="true" pageId="4" pageNumber="566">L. flaviocularis</emphasis>
</taxonomicName>
, T4T5webL = 0.220.28 vs.
<quantity id="4CD49B7A476CFFFFEDC79AB3FEA1F88B" box="[191,334,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="4.064" metricValueMax="4.572" metricValueMin="3.556" pageId="4" pageNumber="566" unit="in" value="0.16" valueMax="0.18" valueMin="0.14">0.140.18 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFEC2E9AB4FDEDF88B" box="[342,514,1771,1797]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D476CFFFFEC2E9AB4FDEDF88B" box="[342,514,1771,1797]" italics="true" pageId="4" pageNumber="566">L. flaviocularis</emphasis>
</taxonomicName>
), two internasal scales (vs. three in
<taxonomicName id="4C2C4D1C476CFFFFEEE49AB4FBA6F88B" box="[924,1097,1771,1797]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D476CFFFFEEE49AB4FBA6F88B" box="[924,1097,1771,1797]" italics="true" pageId="4" pageNumber="566">L. flaviocularis</emphasis>
</taxonomicName>
), 45 precloacal/femoral pores in the sole male (vs.
<quantity id="4CD49B7A476CFFFFECFD9B57FE50F8A7" box="[389,447,1807,1833]" metricMagnitude="-1" metricUnit="m" metricValue="9.144" pageId="4" pageNumber="566" unit="in" value="36.0">36 in</quantity>
the sole male
<taxonomicName id="4C2C4D1C476CFFFFEF1E9B48FCFCF8A7" box="[614,787,1807,1833]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D476CFFFFEF1E9B48FCFCF8A7" box="[614,787,1807,1833]" italics="true" pageId="4" pageNumber="566">L. flaviocularis</emphasis>
</taxonomicName>
), 4445 enlarged precloacal/femoral scales (vs.
<quantity id="4CD49B7A476CFFFFE8409B57FA73F8A7" box="[1336,1436,1807,1833]" metricMagnitude="-1" metricUnit="m" metricValue="9.271" metricValueMax="9.398" metricValueMin="9.144" pageId="4" pageNumber="566" unit="in" value="36.5" valueMax="37.0" valueMin="36.0">3637 in</quantity>
<taxonomicName id="4C2C4D1C476CFFFFEDEF9B6CFEABF8C3" box="[151,324,1843,1869]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D476CFFFFEDEF9B6CFEABF8C3" box="[151,324,1843,1869]" italics="true" pageId="4" pageNumber="566">L. flaviocularis</emphasis>
</taxonomicName>
), a continuous row of tiny scales entirely separating the enlarged pore-bearing precloacal/femoral scales from the patch of enlarged pubic scales (vs. tiny scales only intervening between these enlarged scale series laterally in
<taxonomicName id="4C2C4D1C476CFFFFEC629B24FE28F81B" box="[282,455,1915,1941]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D476CFFFFEC629B24FE28F81B" box="[282,455,1915,1941]" italics="true" pageId="4" pageNumber="566">L. flaviocularis</emphasis>
</taxonomicName>
), brown circumorbital scales (vs. yellow in
<taxonomicName id="4C2C4D1C476CFFFFEEB89B24FB82F81B" box="[960,1133,1915,1941]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D476CFFFFEEB89B24FB82F81B" box="[960,1133,1915,1941]" italics="true" pageId="4" pageNumber="566">L. flaviocularis</emphasis>
</taxonomicName>
), and pale ventral field 15 scale rows wide (vs. ~25 rows wide in
<taxonomicName id="4C2C4D1C476CFFFFEF339BF8FD18F837" box="[587,759,1951,1977]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="4" pageNumber="566" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D476CFFFFEF339BF8FD18F837" box="[587,759,1951,1977]" italics="true" pageId="4" pageNumber="566">L. flaviocularis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF536617476DFFFEEDEF9AE8FB49F962" ID-DOI="http://doi.org/10.5281/zenodo.8313791" ID-Zenodo-Dep="8313791" httpUri="https://zenodo.org/record/8313791/files/figure.png" pageId="5" pageNumber="567" startId="5.[151,250,1712,1736]" targetBox="[171,1417,186,1680]" targetPageId="5" targetType="figure">
<paragraph id="8B93369F476DFFFEEDEF9AE8FB49F962" blockId="5.[151,1436,1712,1773]" pageId="5" pageNumber="567">
<emphasis id="B958EA8D476DFFFEEDEF9AE8FEFDF947" bold="true" box="[151,274,1712,1737]" pageId="5" pageNumber="567">FIGURE 1.</emphasis>
(A) Dorsum, (B) venter, (C) palmar view of left hand, and (D) plantar view of left foot of holotype of
<taxonomicName id="4C2C4D1C476DFFFEE8799AE9FEF8F963" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="5" pageNumber="567" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D476DFFFEE8799AE9FEF8F963" italics="true" pageId="5" pageNumber="567">Lepidodactylus flaviocularis</emphasis>
</taxonomicName>
(USNM 313865), Mt. Austen, Guadalcanal Island, Solomon Islands. Scale bars are 1 cm.
</paragraph>
</caption>
<subSubSection id="C3366514476DFFFDEDBF9B43FE54F853" lastPageId="6" lastPageNumber="568" pageId="5" pageNumber="567" type="description">
<paragraph id="8B93369F476DFFFDEDBF9B43FAF1FE7B" blockId="5.[151,1437,1819,2025]" lastBlockId="6.[151,1437,151,2013]" lastPageId="6" lastPageNumber="568" pageId="5" pageNumber="567">
<emphasis id="B958EA8D476DFFFEEDBF9B43FE0DF8BB" bold="true" box="[199,482,1819,1845]" pageId="5" pageNumber="567">
Description of
<typeStatus id="5497883D476DFFFEEC0C9B43FE32F8BB" box="[372,477,1819,1845]" pageId="5" pageNumber="567" type="holotype">holotype</typeStatus>
.
</emphasis>
A mature female of medium size (SVL =
<quantity id="4CD49B7A476DFFFEEED69B43FBFDF8BB" box="[942,1042,1819,1846]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" pageId="5" pageNumber="567" unit="mm" value="52.5">52.5 mm</quantity>
) with a mid-ventral incision behind the pectoral region. Head relatively long (HL/SVL = 0.24) and wide (HW/HL = 0.82), distinct from neck (
<figureCitation id="13172A1A476DFFFEE84E9B67FA60F8D7" box="[1334,1423,1855,1881]" captionStart="FIGURE 3" captionStartId="8.[151,250,1757,1781]" captionTargetBox="[166,1421,188,1726]" captionTargetId="figure-19@8.[159,1428,181,1733]" captionTargetPageId="8" captionText="FIGURE 3. (A) Dorsum, (B) venter, (C) palmar view of left hand, and (D) plantar view of left foot of holotype of Lepidodactylus makira sp. nov. (KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. Scale bars are 1 cm." figureDoi="http://doi.org/10.5281/zenodo.8313795" httpUri="https://zenodo.org/record/8313795/files/figure.png" pageId="5" pageNumber="567">Fig. 3A</figureCitation>
). Loreal region slightly inflated; no distinct canthus rostralis. Top of snout, area between nares, and area posterior to nares shallowly concave. Snout tapered and rounded at tip, relatively long (SN/HL = 0.45), significantly longer than eye diameter (SN/EY = 1.9). Eye of modest size (EY/HL = 0.23, EY/EN = 0.64); pupil vertical, constricted into series of four lobes; anterior supraciliaries slightly larger than adjacent granules, posterior ones subequal to adjacent granules. Ear opening small (Ear/HL = 0.086), narrowly compressed, oriented obliquely; distance between ear and eye larger than eye diameter (EE/EY = 1.4). Rostral twice as wide (
<quantity id="4CD49B7A476EFFFDEE079CE3FC38FF5B" box="[895,983,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="568" unit="mm" value="2.4">2.4 mm</quantity>
) as high (
<quantity id="4CD49B7A476EFFFDE93A9CE3FB72FF5B" box="[1090,1181,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="568" unit="mm" value="1.2">1.2 mm</quantity>
), highest just medial to nares, lower between these points; length
<quantity id="4CD49B7A476EFFFDEF119C86FD20FF77" box="[617,719,222,249]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="568" unit="mm" value="0.55">0.55 mm</quantity>
. Supranasals separated by two internasals along posterior rostral margin. Rostral in contact with first supralabials, two supranasals, and two internasals. External nares circular; each bordered by rostral, two supranasals, first supralabial, and one postnasal. Mental triangular,
<quantity id="4CD49B7A476EFFFDE9E69D7EFAEDFECF" box="[1182,1282,294,321]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="6" pageNumber="568" unit="mm" value="0.95">0.95 mm</quantity>
wide. Mental bordered posteriorly by one small scale, and this bordered posteriorly by subequal scales that progressively decrease in size posteriorly to join granular chin scales. First five infralabials bordered below by enlarged scales; remaining scales below infralabials smaller, rapidly decreasing posteriorly to approximately same size as throat scales, which decrease in size medially. Supralabials to mid-orbital position eight on each side; three (R) or two (L) enlarged supralabials posterior to this; angle of jaw bordered with granular scales. Infralabials 12 (R) and 11 (L).
</paragraph>
<paragraph id="8B93369F476EFFFDEDBF9DA7FBF9FDEF" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="568">Body of rather narrow habitus (TrL/SVL = 0.52), slightly depressed. Dorsal scales on head, body, limbs, and throat tiny, juxtaposed granules, slightly larger on sides and snout; tubercles absent. Ventral scales larger, flat and smooth, subimbricate, gradually decreasing in size laterally to become granular.</paragraph>
<paragraph id="8B93369F476EFFFDEDBF9E33FC7BFC9B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="568">
Enlarged precloacal/femoral scales in single series of 44 scales extending to distal end of each thigh, 20 of these containing small, shallow pores or dimples; thigh scales anterior to this row larger than those posterior. Enlarged scales form a pubic patch between precloacal series and vent (
<figureCitation id="13172A1A476EFFFDEE0A9EEBFC22FD43" box="[882,973,691,717]" captionStart="FIGURE 3" captionStartId="8.[151,250,1757,1781]" captionTargetBox="[166,1421,188,1726]" captionTargetId="figure-19@8.[159,1428,181,1733]" captionTargetPageId="8" captionText="FIGURE 3. (A) Dorsum, (B) venter, (C) palmar view of left hand, and (D) plantar view of left foot of holotype of Lepidodactylus makira sp. nov. (KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. Scale bars are 1 cm." figureDoi="http://doi.org/10.5281/zenodo.8313795" httpUri="https://zenodo.org/record/8313795/files/figure.png" pageId="6" pageNumber="568">Fig. 3B</figureCitation>
); continuous row of tiny scales entirely separate precloacal series and pubic patch; nine scales in a row between apex of enlarged precloacal series and vent. Scales on palms and soles rounded, flattened, smooth, subimbricate.
</paragraph>
<paragraph id="8B93369F476EFFFDEDBF9F47FB30FBBB" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="568">
Fore- and hindlimbs relatively small but well-developed (FA/SVL = 0.10, CS/SVL = 0.13). Digits well-developed (
<figureCitation id="13172A1A476EFFFDEC629F1BFE7BFCD3" box="[282,404,835,861]" captionStart="FIGURE 3" captionStartId="8.[151,250,1757,1781]" captionTargetBox="[166,1421,188,1726]" captionTargetId="figure-19@8.[159,1428,181,1733]" captionTargetPageId="8" captionText="FIGURE 3. (A) Dorsum, (B) venter, (C) palmar view of left hand, and (D) plantar view of left foot of holotype of Lepidodactylus makira sp. nov. (KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. Scale bars are 1 cm." figureDoi="http://doi.org/10.5281/zenodo.8313795" httpUri="https://zenodo.org/record/8313795/files/figure.png" pageId="6" pageNumber="568">Fig. 3C, D</figureCitation>
), moderately dilated throughout their length (T4W/T4L = 0.36), all but first fingers and toes with recurved claws; clawed phalanges laterally compressed, free above and extending slightly beyond terminal lamellae. Subdigital lamellae narrow and smooth, all undivided (
<figureCitation id="13172A1A476EFFFDEE009FD3FC1BFC2B" box="[888,1012,907,933]" captionStart="FIGURE 3" captionStartId="8.[151,250,1757,1781]" captionTargetBox="[166,1421,188,1726]" captionTargetId="figure-19@8.[159,1428,181,1733]" captionTargetPageId="8" captionText="FIGURE 3. (A) Dorsum, (B) venter, (C) palmar view of left hand, and (D) plantar view of left foot of holotype of Lepidodactylus makira sp. nov. (KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. Scale bars are 1 cm." figureDoi="http://doi.org/10.5281/zenodo.8313795" httpUri="https://zenodo.org/record/8313795/files/figure.png" pageId="6" pageNumber="568">Fig. 3C, D</figureCitation>
); lamellae extend for entire length of each toe (T4lamellaeL/T4L = 1.04). Lamellae of manus 1112141612 on right, 913141511 on left; of pes 1215161812 on right, 1213161912 on left. Relative lengths of digits on manus and pes I &lt;II &lt;V &lt;III &lt;IV. Webbing present between all digits, most extensive between T3 and T4 (T3T4webL/T4L = 0.30, T4T5webL/ T4L = 0.28). A fringe of scales extends from base of fifth toe anteriorly along inner margin of leg.
</paragraph>
<paragraph id="8B93369F476EFFFDEDBF9867FC93FB2F" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="568">
Tail missing. Cloacal sacs not swollen (
<figureCitation id="13172A1A476EFFFDEFF49867FD0AFBD7" box="[652,741,1087,1113]" captionStart="FIGURE 3" captionStartId="8.[151,250,1757,1781]" captionTargetBox="[166,1421,188,1726]" captionTargetId="figure-19@8.[159,1428,181,1733]" captionTargetPageId="8" captionText="FIGURE 3. (A) Dorsum, (B) venter, (C) palmar view of left hand, and (D) plantar view of left foot of holotype of Lepidodactylus makira sp. nov. (KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. Scale bars are 1 cm." figureDoi="http://doi.org/10.5281/zenodo.8313795" httpUri="https://zenodo.org/record/8313795/files/figure.png" pageId="6" pageNumber="568">Fig. 3B</figureCitation>
), with small external orifices situated near lateral margins of vent; three (R) or two (L) slightly enlarged, blunt postcloacal spurs on each side of tailbase; midventral scales of sac hexagonal, subimbricate, slightly larger than those ventrolaterally.
</paragraph>
<paragraph id="8B93369F476EFFFDEDBF98F3FE9EFAF7" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="568">
<emphasis id="B958EA8D476EFFFDEDBF98F3FE54FB4A" box="[199,443,1195,1221]" italics="true" pageId="6" pageNumber="568">Color in preservative</emphasis>
: Dorsal ground color on body, head, and limbs dark brown with four dark chocolatebrown dorsolateral blotches between limbs on each side (
<figureCitation id="13172A1A476EFFFDEE609897FC9FFB67" box="[792,880,1231,1257]" captionStart="FIGURE 3" captionStartId="8.[151,250,1757,1781]" captionTargetBox="[166,1421,188,1726]" captionTargetId="figure-19@8.[159,1428,181,1733]" captionTargetPageId="8" captionText="FIGURE 3. (A) Dorsum, (B) venter, (C) palmar view of left hand, and (D) plantar view of left foot of holotype of Lepidodactylus makira sp. nov. (KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. Scale bars are 1 cm." figureDoi="http://doi.org/10.5281/zenodo.8313795" httpUri="https://zenodo.org/record/8313795/files/figure.png" pageId="6" pageNumber="568">Fig. 3A</figureCitation>
) below which are several smaller dark chocolatebrown lateral blotches. Approximately 15 mid-ventral rows of white scales; sides of venter brown like dorsum. Chin and throat white heavily spotted with brown, brown decreasing posteriorly (
<figureCitation id="13172A1A476EFFFDEE8C994FFBA2FABF" box="[1012,1101,1303,1329]" captionStart="FIGURE 3" captionStartId="8.[151,250,1757,1781]" captionTargetBox="[166,1421,188,1726]" captionTargetId="figure-19@8.[159,1428,181,1733]" captionTargetPageId="8" captionText="FIGURE 3. (A) Dorsum, (B) venter, (C) palmar view of left hand, and (D) plantar view of left foot of holotype of Lepidodactylus makira sp. nov. (KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. Scale bars are 1 cm." figureDoi="http://doi.org/10.5281/zenodo.8313795" httpUri="https://zenodo.org/record/8313795/files/figure.png" pageId="6" pageNumber="568">Fig. 3B</figureCitation>
). Palmar and plantar surfaces white punctated with some brown (
<figureCitation id="13172A1A476EFFFDEF649963FD7BFADB" box="[540,660,1339,1365]" captionStart="FIGURE 3" captionStartId="8.[151,250,1757,1781]" captionTargetBox="[166,1421,188,1726]" captionTargetId="figure-19@8.[159,1428,181,1733]" captionTargetPageId="8" captionText="FIGURE 3. (A) Dorsum, (B) venter, (C) palmar view of left hand, and (D) plantar view of left foot of holotype of Lepidodactylus makira sp. nov. (KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. Scale bars are 1 cm." figureDoi="http://doi.org/10.5281/zenodo.8313795" httpUri="https://zenodo.org/record/8313795/files/figure.png" pageId="6" pageNumber="568">Fig. 3C, D</figureCitation>
); tips of digits brown; webbing dark brown above and below. Iris gold veined with brown.
</paragraph>
<paragraph id="8B93369F476EFFFDEDBF99DBFF2EFA6B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="568">
<emphasis id="B958EA8D476EFFFDEDBF99DBFE29FA13" box="[199,454,1411,1437]" italics="true" pageId="6" pageNumber="568">Measurements (in mm)</emphasis>
. SVL = 52.5, TrL = 27.5, FA = 5.4, CS = 7.0, HL = 12.8, HW = 10.5, Ear = 1.1, EE = 4.3, EY = 3.0, SN = 5.8, EN = 4.7, IN = 2.6, T4L = 5.0, T4W = 1.8, T4lamellaeL = 5.2, T3T4webL = 1.5, T4T5webL = 1.4.
</paragraph>
<paragraph id="8B93369F476EFFFDEDBF99B7FE20F96F" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="568">
<emphasis id="B958EA8D476EFFFDEDBF99B7FEC0F986" box="[199,303,1519,1544]" italics="true" pageId="6" pageNumber="568">Variation</emphasis>
. The sole
<typeStatus id="5497883D476EFFFDECDD99B7FDEAF987" box="[421,517,1519,1545]" pageId="6" pageNumber="568" type="paratype">paratype</typeStatus>
is similar to the
<typeStatus id="5497883D476EFFFDEFBC99B7FCC9F987" box="[708,806,1519,1545]" pageId="6" pageNumber="568" type="holotype">holotype</typeStatus>
in most respects but is a male of smaller size (45.0 mm SVL), with 45 enlarged precloacal/femoral scales and 45 well-developed precloacal/femoral pores that extend to distal end of each thigh. It has nine (R) and ten (L) supralabials to center of eye, and 14 infralabials on each side. Lamellae of manus 913141714 on right, 1113171713 on left; of pes 1216171714 on right, 1214 191814 on left. It has somewhat less toe webbing (
<specimenCode id="DB8A9EE4476EFFFDEF9F9A27FCEBF917" box="[743,772,1663,1689]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="568">T3</specimenCode>
<specimenCode id="DB8A9EE4476EFFFDEE7C9A27FCCCF917" box="[772,803,1663,1689]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="568">T4</specimenCode>
webL/
<specimenCode id="DB8A9EE4476EFFFDEE159A27FC62F917" box="[877,909,1663,1689]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="568">T4</specimenCode>
L = 0.28,
<specimenCode id="DB8A9EE4476EFFFDEE819A26FBF7F917" box="[1017,1048,1662,1689]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="568">T4</specimenCode>
<specimenCode id="DB8A9EE4476EFFFDE96F9A26FBDBF917" box="[1047,1076,1662,1689]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="568">T5</specimenCode>
webL/
<specimenCode id="DB8A9EE4476EFFFDE9069A26FB72F917" box="[1150,1181,1662,1689]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="568">T4</specimenCode>
L = 0.22), but this may reflect a longer period of shrinkage in alcohol. The specimen is highly bleached such that reliable information on color pattern is unavailable.
</paragraph>
<paragraph id="8B93369F476EFFFDEDBF9AB3FEB3F8C3" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="568">
<emphasis id="B958EA8D476EFFFDEDBF9AB3FDA0F88B" box="[199,591,1771,1797]" italics="true" pageId="6" pageNumber="568">
Measurements of
<typeStatus id="5497883D476EFFFDECF59AB3FE1EF88A" box="[397,497,1771,1796]" pageId="6" pageNumber="568" type="paratype">paratype</typeStatus>
(in mm)
</emphasis>
. SVL = 45.0, TrL = 23.0, FA = 4.7, CS = 6.0, HL = 11.8, HW = 9.2, Ear = 1.0, EE = 4.0, EY = 2.9, SN = 5.2, EN = 4.3, IN = 2.4, T4L = 5.0, T4W = 1.6, T4lamellaeL = 4.7, T3T4webL = 1.4, T4T5webL = 1.1.
</paragraph>
<paragraph id="8B93369F476EFFFDEDBF9B0FFE54F853" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="568">
<emphasis id="B958EA8D476EFFFDEDBF9B0FFEBEF8FF" box="[199,337,1879,1905]" italics="true" pageId="6" pageNumber="568">Color in life</emphasis>
. A photo of the
<typeStatus id="5497883D476EFFFDEF7E9B0FFD87F8FF" box="[518,616,1879,1905]" pageId="6" pageNumber="568" type="holotype">holotype</typeStatus>
in life (
<figureCitation id="13172A1A476EFFFDEFB99B0FFCF8F8FF" box="[705,791,1879,1905]" captionStart="FIGURE 2" captionStartId="7.[151,250,1886,1910]" captionTargetBox="[163,1425,181,1862]" captionTargetId="figure-26@7.[163,1425,181,1862]" captionTargetPageId="7" captionText="FIGURE 2. Portraitsin life of (A) an individualof Lepidodactylus flaviocularis from near the summit of Mt.Austen,Guadalcanal, Solomon Islands; and (B) the holotype of Lepidodactylus makira sp. nov. (KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. (A) taken by M. McCoy; (B) by I. Tigulu." figureDoi="http://doi.org/10.5281/zenodo.8313793" httpUri="https://zenodo.org/record/8313793/files/figure.png" pageId="6" pageNumber="568">Fig. 2B</figureCitation>
) shows an animal that is yellow brown dorsally with five dark-brown dorsolateral blotches and smaller dark-brown lateral spots. It has two narrow brown lines behind the eye followed by three smaller brown lines arrayed behind those. The eye is golden, and the circumorbital scales are the same color as the dorsum.
</paragraph>
</subSubSection>
<caption id="DF536617476FFFFCEDEF9B06FCAEF830" ID-DOI="http://doi.org/10.5281/zenodo.8313793" ID-Zenodo-Dep="8313793" httpUri="https://zenodo.org/record/8313793/files/figure.png" pageId="7" pageNumber="569" startId="7.[151,250,1886,1910]" targetBox="[163,1425,181,1862]" targetPageId="7" targetType="figure">
<paragraph id="8B93369F476FFFFCEDEF9B06FCAEF830" blockId="7.[151,1437,1886,1982]" pageId="7" pageNumber="569">
<emphasis id="B958EA8D476FFFFCEDEF9B06FEFEF8F8" bold="true" box="[151,273,1886,1910]" pageId="7" pageNumber="569">FIGURE 2.</emphasis>
Portraits in life of (A) an individual of
<taxonomicName id="4C2C4D1C476FFFFCEFF39B06FC45F8F8" box="[651,938,1886,1910]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="7" pageNumber="569" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D476FFFFCEFF39B06FC45F8F8" box="[651,938,1886,1910]" italics="true" pageId="7" pageNumber="569">Lepidodactylus flaviocularis</emphasis>
</taxonomicName>
from near the summit of Mt.Austen, Guadalcanal, Solomon Islands; and (B) the holotype of
<taxonomicName id="4C2C4D1C476FFFFCEF3C9BDAFCC2F814" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[580,813,1922,1946]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="7" pageNumber="569" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D476FFFFCEF3C9BDAFCC2F814" box="[580,813,1922,1946]" italics="true" pageId="7" pageNumber="569">Lepidodactylus makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D476FFFFCEE4C9BDAFC6AF814" bold="true" box="[820,901,1922,1946]" pageId="7" pageNumber="569">
<taxonomicNameLabel id="A26B57F6476FFFFCEE4C9BDAFC6AF814" box="[820,901,1922,1946]" pageId="7" pageNumber="569" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. (A) taken by M. McCoy; (B) by I. Tigulu.
</paragraph>
</caption>
<caption id="DF5366174760FFF3EDEF9A85FAF1F897" ID-DOI="http://doi.org/10.5281/zenodo.8313795" ID-Zenodo-Dep="8313795" httpUri="https://zenodo.org/record/8313795/files/figure.png" pageId="8" pageNumber="570" startId="8.[151,250,1757,1781]" targetBox="[166,1421,188,1726]" targetPageId="8" targetType="figure">
<paragraph id="8B93369F4760FFF3EDEF9A85FAF1F897" blockId="8.[151,1436,1757,1817]" pageId="8" pageNumber="570">
<emphasis id="B958EA8D4760FFF3EDEF9A85FEFEF97B" bold="true" box="[151,273,1757,1781]" pageId="8" pageNumber="570">FIGURE 3.</emphasis>
(A) Dorsum, (B) venter, (C) palmar view of left hand, and (D) plantar view of left foot of holotype of
<taxonomicName id="4C2C4D1C4760FFF3E8799A85FF30F897" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="8" pageNumber="570" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D4760FFF3E8799A85FF30F897" italics="true" pageId="8" pageNumber="570">Lepidodactylus makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D4760FFF3ED9D9B59FED9F897" bold="true" box="[229,310,1793,1817]" pageId="8" pageNumber="570">
<taxonomicNameLabel id="A26B57F64760FFF3ED9D9B59FED9F897" box="[229,310,1793,1817]" pageId="8" pageNumber="570" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. Scale bars are 1 cm.
</paragraph>
</caption>
<subSubSection id="C33665144760FFF3EDBF9B1FFEA5F827" pageId="8" pageNumber="570" type="etymology">
<paragraph id="8B93369F4760FFF3EDBF9B1FFB0AF8EF" blockId="8.[151,1437,1863,2034]" box="[199,1253,1863,1890]" pageId="8" pageNumber="570">
<emphasis id="B958EA8D4760FFF3EDBF9B1FFEA1F8EF" bold="true" box="[199,334,1863,1889]" pageId="8" pageNumber="570">Etymology.</emphasis>
The species name is a noun in apposition and is named for its island of residence.
</paragraph>
<paragraph id="8B93369F4760FFF3EDBF9B33FEA5F827" blockId="8.[151,1437,1863,2034]" pageId="8" pageNumber="570">
<emphasis id="B958EA8D4760FFF3EDBF9B33FEF5F80B" bold="true" box="[199,282,1899,1925]" pageId="8" pageNumber="570">Range.</emphasis>
Known only from two localities that are within ~
<quantity id="4CD49B7A4760FFF3EE329B33FC7EF80B" box="[842,913,1899,1926]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="570" unit="km" value="15.0">15 km</quantity>
of each other although the
<typeStatus id="5497883D4760FFF3E9BE9B34FAC9F808" box="[1222,1318,1900,1926]" pageId="8" pageNumber="570" type="paratype">paratype</typeStatus>
locality is inexact (
<figureCitation id="13172A1A4760FFF3ED8F9BD7FED2F827" box="[247,317,1935,1961]" captionStart="FIGURE 4" captionStartId="9.[151,250,1843,1867]" captionTargetBox="[163,1424,181,1819]" captionTargetId="figure-26@9.[163,1424,181,1819]" captionTargetPageId="9" captionText="FIGURE 4. Map of (A) the Solomon Islands, with (B) Makira Island inset. Red dot is the type locality of Lepidodactylus makira sp. nov." figureDoi="http://doi.org/10.5281/zenodo.8313797" httpUri="https://zenodo.org/record/8313797/files/figure.png" pageId="8" pageNumber="570">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33665144760FFF3EDBF9BEBFD9BF87C" pageId="8" pageNumber="570" type="biology_ecology">
<paragraph id="8B93369F4760FFF3EDBF9BEBFD9BF87C" blockId="8.[151,1437,1863,2034]" pageId="8" pageNumber="570">
<emphasis id="B958EA8D4760FFF3EDBF9BEBFEC3F843" bold="true" box="[199,300,1971,1997]" pageId="8" pageNumber="570">Ecology.</emphasis>
The
<typeStatus id="5497883D4760FFF3EC1D9BEBFE28F843" box="[357,455,1971,1997]" pageId="8" pageNumber="570" type="holotype">holotype</typeStatus>
was collected on the leaf of a shrub ca.
<quantity id="4CD49B7A4760FFF3EEF09BEBFC5CF840" box="[904,947,1971,1998]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="570" unit="m" value="2.0">2 m</quantity>
above ground on a ridge in thickly forested lower-montane primary rainforest (
<figureCitation id="13172A1A4760FFF3EF599B8FFD89F87C" box="[545,614,2007,2034]" captionStart="FIGURE 5" captionStartId="10.[151,250,1146,1170]" captionTargetBox="[162,1424,181,1122]" captionTargetId="figure-19@10.[162,1424,181,1122]" captionTargetPageId="10" captionText="FIGURE 5. Location where the holotypeof Lepidodactylus makira sp. nov. was collected, on a ridge in lower-montane rainforest." figureDoi="http://doi.org/10.5281/zenodo.8313799" httpUri="https://zenodo.org/record/8313799/files/figure.png" pageId="8" pageNumber="570">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF5366174761FFF2EDEF9B6BFF07F8FE" ID-DOI="http://doi.org/10.5281/zenodo.8313797" ID-Zenodo-Dep="8313797" httpUri="https://zenodo.org/record/8313797/files/figure.png" pageId="9" pageNumber="571" startId="9.[151,250,1843,1867]" targetBox="[163,1424,181,1819]" targetPageId="9" targetType="figure">
<paragraph id="8B93369F4761FFF2EDEF9B6BFF07F8FE" blockId="9.[151,1436,1843,1904]" pageId="9" pageNumber="571">
<emphasis id="B958EA8D4761FFF2EDEF9B6BFEFDF8C5" bold="true" box="[151,274,1843,1867]" pageId="9" pageNumber="571">FIGURE 4.</emphasis>
Map of (A) the Solomon Islands, with (B) Makira Island inset. Red dot is the type locality of
<taxonomicName id="4C2C4D1C4761FFF2E9CD9B6BFA73F8C5" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[1205,1436,1843,1867]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="9" pageNumber="571" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D4761FFF2E9CD9B6BFA73F8C5" box="[1205,1436,1843,1867]" italics="true" pageId="9" pageNumber="571">Lepidodactylus makira</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B57F64761FFF2EDEF9B00FF07F8FE" box="[151,232,1880,1904]" pageId="9" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF5366174762FFF1EDEF9822FA72FB1C" ID-DOI="http://doi.org/10.5281/zenodo.8313799" ID-Zenodo-Dep="8313799" box="[151,1437,1146,1170]" httpUri="https://zenodo.org/record/8313799/files/figure.png" pageId="10" pageNumber="572" startId="10.[151,250,1146,1170]" targetBox="[162,1424,181,1122]" targetPageId="10" targetType="figure">
<paragraph id="8B93369F4762FFF1EDEF9822FA72FB1C" blockId="10.[151,1437,1146,1170]" box="[151,1437,1146,1170]" pageId="10" pageNumber="572">
<emphasis id="B958EA8D4762FFF1EDEF9822FEFEFB1C" bold="true" box="[151,273,1146,1170]" pageId="10" pageNumber="572">FIGURE 5.</emphasis>
Location where the holotype of
<taxonomicName id="4C2C4D1C4762FFF1EF359822FCDBFB1C" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[589,820,1146,1170]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D4762FFF1EF359822FCDBFB1C" box="[589,820,1146,1170]" italics="true" pageId="10" pageNumber="572">Lepidodactylus makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D4762FFF1EE409822FC69FB1C" bold="true" box="[824,902,1146,1170]" pageId="10" pageNumber="572">
<taxonomicNameLabel id="A26B57F64762FFF1EE409822FC69FB1C" box="[824,902,1146,1170]" pageId="10" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected, on a ridge in lower-montane rainforest.
</paragraph>
</caption>
<subSubSection id="C33665144762FFF1EDBF9898FD32F87D" pageId="10" pageNumber="572" type="discussion">
<paragraph id="8B93369F4762FFF1EDBF9898FCCEF9E8" blockId="10.[151,1437,1216,2035]" pageId="10" pageNumber="572">
<emphasis id="B958EA8D4762FFF1EDBF9898FED4FB54" bold="true" box="[199,315,1216,1242]" pageId="10" pageNumber="572">Remarks.</emphasis>
<taxonomicName id="4C2C4D1C4762FFF1EC389898FDADFB54" box="[320,578,1216,1242]" pageId="10" pageNumber="572">
<emphasis id="B958EA8D4762FFF1EC389898FE04FB54" box="[320,491,1216,1242]" italics="true" pageId="10" pageNumber="572">Lepidodactylus</emphasis>
species
</taxonomicName>
are often distinguished from each other in part by degree of toe webbing, with differences usually being estimated for the webbing between T3 and T4. I have found that distinction to generally work well, as it does for distinguishing
<taxonomicName id="4C2C4D1C4762FFF1EF359951FD57FAAC" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[589,696,1288,1314]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D4762FFF1EF359951FD57FAAC" box="[589,696,1288,1314]" italics="true" pageId="10" pageNumber="572">L. makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D4762FFF1EFC79950FCF7FAAC" bold="true" box="[703,792,1288,1314]" pageId="10" pageNumber="572">
<taxonomicNameLabel id="A26B57F64762FFF1EFC79950FCF7FAAC" box="[703,792,1288,1314]" pageId="10" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="4C2C4D1C4762FFF1EE229951FBEAFAAC" box="[858,1029,1288,1314]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D4762FFF1EE229951FBEAFAAC" box="[858,1029,1288,1314]" italics="true" pageId="10" pageNumber="572">L. flaviocularis</emphasis>
</taxonomicName>
. However, for this particular species pair, the difference in degree of webbing between T4 and T5 is even more striking, and I have included it in the diagnosis for both species. As well, differences in numbers of digital lamellae are generally summarized by comparing numbers under T4 and sometimes T1. Again, those numbers well serve to distinguish
<taxonomicName id="4C2C4D1C4762FFF1E9A8992DFAD3FA00" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[1232,1340,1396,1422]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D4762FFF1E9A8992DFAD3FA00" box="[1232,1340,1396,1422]" italics="true" pageId="10" pageNumber="572">L. makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D4762FFF1E83B992CFA73FA00" bold="true" box="[1347,1436,1396,1422]" pageId="10" pageNumber="572">
<taxonomicNameLabel id="A26B57F64762FFF1E83B992CFA73FA00" box="[1347,1436,1396,1422]" pageId="10" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from most other Melanesian
<taxonomicName id="4C2C4D1C4762FFF1ECAF99C0FD6DFA3C" box="[471,642,1432,1458]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="genus">
<emphasis id="B958EA8D4762FFF1ECAF99C0FD6DFA3C" box="[471,642,1432,1458]" italics="true" pageId="10" pageNumber="572">Lepidodactylus</emphasis>
</taxonomicName>
but not so well from
<taxonomicName id="4C2C4D1C4762FFF1EE1699C1FBF6FA3C" box="[878,1049,1432,1458]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D4762FFF1EE1699C1FBF6FA3C" box="[878,1049,1432,1458]" italics="true" pageId="10" pageNumber="572">L. flaviocularis</emphasis>
</taxonomicName>
. However,
<taxonomicName id="4C2C4D1C4762FFF1E9E099C1FAEBFA3C" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[1176,1284,1432,1458]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D4762FFF1E9E099C1FAEBFA3C" box="[1176,1284,1432,1458]" italics="true" pageId="10" pageNumber="572">L. makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D4762FFF1E87299C0FA8CFA3C" bold="true" box="[1290,1379,1432,1458]" pageId="10" pageNumber="572">
<taxonomicNameLabel id="A26B57F64762FFF1E87299C0FA8CFA3C" box="[1290,1379,1432,1458]" pageId="10" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
does have more digital lamellae overall than does
<taxonomicName id="4C2C4D1C4762FFF1EFF399E5FCD9FA58" box="[651,822,1468,1494]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D4762FFF1EFF399E5FCD9FA58" box="[651,822,1468,1494]" italics="true" pageId="10" pageNumber="572">L. flaviocularis</emphasis>
</taxonomicName>
, and this is best captured by summing total numbers of lamellae across all digits. For
<taxonomicName id="4C2C4D1C4762FFF1EC9D99B9FDBEFA74" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[485,593,1504,1530]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D4762FFF1EC9D99B9FDBEFA74" box="[485,593,1504,1530]" italics="true" pageId="10" pageNumber="572">L. makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D4762FFF1EF2F99B8FD5FFA74" bold="true" box="[599,688,1504,1530]" pageId="10" pageNumber="572">
<taxonomicNameLabel id="A26B57F64762FFF1EF2F99B8FD5FFA74" box="[599,688,1504,1530]" pageId="10" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, numbers of lamellae on each manus vary from 6271 (mean 66.2) and each pes from 7276 (mean 74.0), whereas those same figures for
<taxonomicName id="4C2C4D1C4762FFF1EEC09A5DFB8AF990" box="[952,1125,1540,1566]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D4762FFF1EEC09A5DFB8AF990" box="[952,1125,1540,1566]" italics="true" pageId="10" pageNumber="572">L. flaviocularis</emphasis>
</taxonomicName>
are 5759 (mean 58.0) and 6073 (mean 67.0). This better captures the impression of greater numbers of lamellae in
<taxonomicName id="4C2C4D1C4762FFF1E9F79A71FB12F9CC" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[1167,1277,1576,1602]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D4762FFF1E9F79A71FB12F9CC" box="[1167,1277,1576,1602]" italics="true" pageId="10" pageNumber="572">L. makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D4762FFF1E87D9A70FA8FF9CC" bold="true" box="[1285,1376,1576,1602]" pageId="10" pageNumber="572">
<taxonomicNameLabel id="A26B57F64762FFF1E87D9A70FA8FF9CC" box="[1285,1376,1576,1602]" pageId="10" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
even though comparisons for only T4 or T1 are less distinctive.
</paragraph>
<paragraph id="8B93369F4762FFF1EDBF9A28FD32F87D" blockId="10.[151,1437,1216,2035]" pageId="10" pageNumber="572">
There may be additional differences between
<taxonomicName id="4C2C4D1C4762FFF1EFBA9A29FCC1F904" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[706,814,1648,1674]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D4762FFF1EFBA9A29FCC1F904" box="[706,814,1648,1674]" italics="true" pageId="10" pageNumber="572">L. makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D4762FFF1EE4D9A28FC61F904" bold="true" box="[821,910,1648,1674]" pageId="10" pageNumber="572">
<taxonomicNameLabel id="A26B57F64762FFF1EE4D9A28FC61F904" box="[821,910,1648,1674]" pageId="10" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C2C4D1C4762FFF1EEBB9A29FB81F904" box="[963,1134,1648,1674]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D4762FFF1EEBB9A29FB81F904" box="[963,1134,1648,1674]" italics="true" pageId="10" pageNumber="572">L. flaviocularis</emphasis>
</taxonomicName>
, but these need to be better assessed when a broader series of each species becomes available. In preservative, the
<typeStatus id="5497883D4762FFF1E9019ACCFB34F920" box="[1145,1243,1684,1710]" pageId="10" pageNumber="572" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2C4D1C4762FFF1E87B9ACDFA9DF920" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[1283,1394,1684,1710]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D4762FFF1E87B9ACDFA9DF920" box="[1283,1394,1684,1710]" italics="true" pageId="10" pageNumber="572">L. makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D4762FFF1E8039ACCFF27F95C" bold="true" pageId="10" pageNumber="572">
<taxonomicNameLabel id="A26B57F64762FFF1E8039ACCFF27F95C" pageId="10" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has dark-brown webbing between the toes, which contrasts with the paler-brown color of the surrounding scales on the feet; this stands in contrast to what is seen in
<taxonomicName id="4C2C4D1C4762FFF1EE489A85FC33F978" box="[816,988,1756,1782]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D4762FFF1EE489A85FC33F978" box="[816,988,1756,1782]" italics="true" pageId="10" pageNumber="572">L. flaviocularis</emphasis>
</taxonomicName>
, in which the webbing and surrounding scales are the same color and do not contrast with each other. The bleached nature of the
<typeStatus id="5497883D4762FFF1E9F89B59FB0FF895" box="[1152,1248,1793,1819]" pageId="10" pageNumber="572" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C2C4D1C4762FFF1E87E9B59FA9CF894" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[1286,1395,1792,1818]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D4762FFF1E87E9B59FA9CF894" box="[1286,1395,1792,1818]" italics="true" pageId="10" pageNumber="572">L. makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D4762FFF1E8039B58FF27F8B0" bold="true" pageId="10" pageNumber="572">
<taxonomicNameLabel id="A26B57F64762FFF1E8039B58FF27F8B0" pageId="10" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
does not allow me to determine whether this feature is shared by both specimens. Similarly, the iris of the
<typeStatus id="5497883D4762FFF1EDEF9B10FF16F8EC" box="[151,249,1864,1890]" pageId="10" pageNumber="572" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2C4D1C4762FFF1EC669B11FE64F8EC" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[286,395,1864,1890]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D4762FFF1EC669B11FE64F8EC" box="[286,395,1864,1890]" italics="true" pageId="10" pageNumber="572">L. makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D4762FFF1ECEA9B10FE04F8EC" bold="true" box="[402,491,1864,1890]" pageId="10" pageNumber="572">
<taxonomicNameLabel id="A26B57F64762FFF1ECEA9B10FE04F8EC" box="[402,491,1864,1890]" pageId="10" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is gold veined with brown; that of both specimens of
<taxonomicName id="4C2C4D1C4762FFF1E93E9B11FB1DF8EC" box="[1094,1266,1864,1890]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D4762FFF1E93E9B11FB1DF8EC" box="[1094,1266,1864,1890]" italics="true" pageId="10" pageNumber="572">L. flaviocularis</emphasis>
</taxonomicName>
appears brown instead of gold, as does the photograph in life of that species (
<figureCitation id="13172A1A4762FFF1EE2B9B34FC41F808" box="[851,942,1900,1926]" captionStart="FIGURE 2" captionStartId="7.[151,250,1886,1910]" captionTargetBox="[163,1425,181,1862]" captionTargetId="figure-26@7.[163,1425,181,1862]" captionTargetPageId="7" captionText="FIGURE 2. Portraitsin life of (A) an individualof Lepidodactylus flaviocularis from near the summit of Mt.Austen,Guadalcanal, Solomon Islands; and (B) the holotype of Lepidodactylus makira sp. nov. (KU 350843), Nara Village, Naepaepa Mt., Makira Island, Solomon Islands. (A) taken by M. McCoy; (B) by I. Tigulu." figureDoi="http://doi.org/10.5281/zenodo.8313793" httpUri="https://zenodo.org/record/8313793/files/figure.png" pageId="10" pageNumber="572">Fig. 2A</figureCitation>
). Apparent differences in extent of webbing would also benefit from further evaluation as specimens become available, as would variation in the enlarged chin shields medial to the infralabials, which are smaller in the
<specimenCount id="9D2AFD164762FFF1EE659BEDFC2DF840" box="[797,962,1972,1999]" pageId="10" pageNumber="572" type="generic">two specimens</specimenCount>
of
<taxonomicName id="4C2C4D1C4762FFF1EE9D9BEDFB7FF840" box="[997,1168,1972,1998]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="B958EA8D4762FFF1EE9D9BEDFB7FF840" box="[997,1168,1972,1998]" italics="true" pageId="10" pageNumber="572">L. flaviocularis</emphasis>
</taxonomicName>
currently available than they are in the
<specimenCount id="9D2AFD164762FFF1EC469B81FE0BF87C" box="[318,484,2008,2035]" pageId="10" pageNumber="572" type="generic">two specimens</specimenCount>
of
<taxonomicName id="4C2C4D1C4762FFF1EF719B81FD9AF87C" authority="Kraus, 2023" authorityName="Kraus" authorityYear="2023" box="[521,629,2008,2034]" class="Squamata" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" pageId="10" pageNumber="572" phylum="Chordata" rank="species" species="makira" status="sp. nov.">
<emphasis id="B958EA8D4762FFF1EF719B81FD9AF87C" box="[521,629,2008,2034]" italics="true" pageId="10" pageNumber="572">L. makira</emphasis>
</taxonomicName>
<emphasis id="B958EA8D4762FFF1EF049B80FD39F87C" bold="true" box="[636,726,2008,2034]" pageId="10" pageNumber="572">
<taxonomicNameLabel id="A26B57F64762FFF1EF049B80FD39F87C" box="[636,726,2008,2034]" pageId="10" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>