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<mods:title id="FB584E6E60181256BE1F7986370DD71D">Frasnian Atrypida (Brachiopoda) from Silesia (Poland) and the age of the eo-Variscan collision in the Sudetes</mods:title>
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<taxonomicName id="4C4E4CBEFFEDFFCFFF10FC18FE6D1ED9" authority="Rzhonsnitskaya, 1960" authorityName="Rzhonsnitskaya" authorityYear="1960" class="Rhynchonellata" family="Atrypidae" genus="Pseudogruenewaldtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="9" pageNumber="298" phylum="Brachiopoda" rank="species" species="tschernyschewi">
<emphasis id="B93AEB2FFFEDFFCFFF10FC18FDDE1EF8" bold="true" box="[183,576,944,971]" italics="true" pageId="9" pageNumber="298">Pseudogruenewaldtia tschernyschewi</emphasis>
<bibRefCitation id="EFDF4ACCFFEDFFCFFEA2FC78FE6D1ED9" author="RZHONSNITSKAYA M. A." box="[261,499,976,1002]" pageId="9" pageNumber="298" pagination="45 - 50" refId="ref11018" refString="RZHONSNITSKAYA M. A. 1960. - [The genus Gruenewaldtia Tschernyschew in the Devonian of the USSR] Rod Gruenewaldtia Cernysev v devone SSSR. Sbornik Stat'ej po Paleontologii i Biostratigrafii 20: 45 - 50 (in Russian)." type="journal article" year="1960">Rzhonsnitskaya, 1960</bibRefCitation>
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<subSubSection id="C35464B6FFEDFFCFFEE3FC58FE2A1938" box="[324,436,1008,1035]" pageId="9" pageNumber="298" type="description">
<paragraph id="8BF1373DFFEDFFCFFEE3FC58FE2A1938" blockId="9.[324,436,1008,1035]" box="[324,436,1008,1035]" pageId="9" pageNumber="298">
(
<figureCitation id="13752BB8FFEDFFCFFEEBFC58FE0E1938" box="[332,400,1008,1035]" captionStart="FIG" captionStartId="8.[114,125,1589,1607]" captionTargetBox="[134,1193,215,1485]" captionTargetId="figure-174@8.[134,1193,215,1485]" captionTargetPageId="8" captionText="FIG. 5. — Pseudogruenewaldtia tschernyschewi Rzhonsnitskaya,1960;Mokrzeszów (Oberkunzendorf),Late Frasnian:A-E, articulated shell MGUWr 5364s.1,specimen figured by Grocholski (1969;pl.5:2) as Atrypa ex gr.reticularis; F, V-Z, articulated shell MGUWr 5364s.1; G-K, articulated shell MB.B.2421.1; L-P, articulated shell MB.B.297.10; Q-U, articulated shell MB.B.2421.2; AA-EE, articulated shell MB.B.297.1. A, G, L, Q, V, AA, dorsal views; B, H, M, R, W, BB, ventral views; C, I, N, S, X, CC, lateral views; D, J, O, T, Y, DD, posterior views; E, K, P, U, Z, EE, anterior views; F, microornamentation. Scale bars: F, 5 mm; all others, 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4538049" httpUri="https://zenodo.org/record/4538049/files/figure.png" pageId="9" pageNumber="298">Figs 5</figureCitation>
;
<figureCitation id="13752BB8FFEDFFCFFE3BFC58FE341938" box="[412,426,1008,1035]" captionStart="FIG" captionStartId="9.[113,124,802,820]" captionTargetBox="[132,1191,215,762]" captionTargetId="figure-374@9.[132,1191,215,762]" captionTargetPageId="9" captionText="FIG. 6. — Pseudogruenewaldtia tschernyschewi Rzhonsnitskaya, 1960. The largest specimen in the collection, articulated shell MB.B.297.7. Mokrzeszów (Oberkunzendorf), Late Frasnian: A, dorsal view; B, ventral view; C, lateral view; D, posterior view; E, anterior view. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4538051" httpUri="https://zenodo.org/record/4538051/files/figure.png" pageId="9" pageNumber="298">6</figureCitation>
)
</paragraph>
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<subSubSection id="C35464B6FFEDFFCFFFD6FB81FF211809" pageId="9" pageNumber="298" type="reference_group">
<paragraph id="8BF1373DFFEDFFCFFFD6FB81FDC31945" blockId="9.[113,646,1065,1142]" pageId="9" pageNumber="298">
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<treatmentCitation id="0AEF112CFFEDFFCFFFD6FB81FF4E196E" author="RZHONSNITSKAYA M. A." page="46" pageId="9" pageNumber="298" year="1960">
<taxonomicName id="4C4E4CBEFFEDFFCFFFD6FB81FF4E196E" authority="Rzhonsnitskaya, 1960: 46" authorityName="Rzhonsnitskaya" authorityPageNumber="46" authorityYear="1960" class="Rhynchonellata" family="Atrypidae" genus="Pseudogruenewaldtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="9" pageNumber="298" phylum="Brachiopoda" rank="species" species="tschernyschewi">
<emphasis id="B93AEB2FFFEDFFCFFFD6FB81FE481972" box="[113,470,1065,1089]" italics="true" pageId="9" pageNumber="298">Pseudogruenewaldtia tschernyschewi</emphasis>
<bibRefCitation id="EFDF4ACCFFEDFFCFFE78FB82FF4E196E" author="RZHONSNITSKAYA M. A." pageId="9" pageNumber="298" pagination="45 - 50" refId="ref11018" refString="RZHONSNITSKAYA M. A. 1960. - [The genus Gruenewaldtia Tschernyschew in the Devonian of the USSR] Rod Gruenewaldtia Cernysev v devone SSSR. Sbornik Stat'ej po Paleontologii i Biostratigrafii 20: 45 - 50 (in Russian)." type="journal article" year="1960">Rzhonsnitskaya, 1960: 46</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 1: 7, pl. 2: 4; 1964: 107, pl. 2: 6, pl. 5: 4. —
<treatmentCitation id="0AEF112CFFEDFFCFFF0BFBF6FE7B1945" author="RZHONSNITSKAYA M. A. &amp; MARKOVSKII B. P. &amp; YUDINA Y. A. &amp; SOKIRAN E. V." box="[172,485,1118,1142]" page="328" pageId="9" pageNumber="298" year="1998">
<bibRefCitation id="EFDF4ACCFFEDFFCFFF0BFBF6FE7B1945" author="RZHONSNITSKAYA M. A. &amp; MARKOVSKII B. P. &amp; YUDINA Y. A. &amp; SOKIRAN E. V." box="[172,485,1118,1142]" pageId="9" pageNumber="298" pagination="305 - 344" refId="ref11163" refString="RZHONSNITSKAYA M. A., MARKOVSKII B. P., YUDINA Y. A. &amp; SOKIRAN E. V. 1998. - Late Frasnian Atrypida (Brachiopoda) from the South Urals, South Timan and Kuznetsk Basin (Russia). Acta Palaeontologica Polonica 43: 305 - 344." type="journal article" year="1998">
Rzhonsnitskaya
<emphasis id="B93AEB2FFFEDFFCFFEEEFBF6FEE61945" box="[329,376,1118,1142]" italics="true" pageId="9" pageNumber="298">et al.</emphasis>
1998: 328
</bibRefCitation>
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, figs 19, 20.
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</paragraph>
<paragraph id="8BF1373DFFEDFFCFFFD6FB26FDB61996" blockId="9.[113,552,1165,1191]" box="[113,552,1165,1191]" pageId="9" pageNumber="298">
<treatmentCitationGroup id="AB5E1013FFEDFFCFFFD6FB26FDB61996" box="[113,552,1165,1191]" pageId="9" pageNumber="298">
<taxonomicName id="4C4E4CBEFFEDFFCFFFD6FB26FF681996" box="[113,246,1166,1189]" class="Rhynchonellata" family="Atrypidae" genus="Atrypa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="9" pageNumber="298" phylum="Brachiopoda" rank="species" species="zonata">
<emphasis id="B93AEB2FFFEDFFCFFFD6FB26FF681996" box="[113,246,1166,1189]" italics="true" pageId="9" pageNumber="298">Atrypa zonata</emphasis>
</taxonomicName>
<treatmentCitation id="0AEF112CFFEDFFCFFEA8FB26FE5F1994" author="DAMES W. B." box="[271,449,1166,1191]" page="497" pageId="9" pageNumber="298" year="1868">
<bibRefCitation id="EFDF4ACCFFEDFFCFFEA8FB26FE5F1994" author="DAMES W. B." box="[271,449,1166,1191]" pageId="9" pageNumber="298" pagination="469 - 508" refId="ref8972" refString="DAMES W. B. 1868. - Ueber die in der Umgebung Freiburgs in Nieder-Schlesien auftretenden devonischer Ablagerungen. Zeitschrift der deutschen geologischen Gesellschaft 20: 469 - 508." type="journal article" year="1868">Dames 1868: 497</bibRefCitation>
</treatmentCitation>
, pl. 11a-c.
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</paragraph>
<paragraph id="8BF1373DFFEDFFCFFFD6FB15FE7519DC" blockId="9.[112,647,1213,1263]" pageId="9" pageNumber="298">
<emphasis id="B93AEB2FFFEDFFCFFFD6FB15FEC219E6" box="[113,348,1213,1237]" italics="true" pageId="9" pageNumber="298">Grunnewaldtia latilinguis</emphasis>
<bibRefCitation id="EFDF4ACCFFEDFFCFFED6FB16FDDC19E5" author="LJASCHENKO &amp; A. I" box="[369,578,1214,1238]" pageId="9" pageNumber="298" refId="ref10220" refString="LJASCHENKO (LAsENKO) A. I. (1959). - [Atlas of Brachiopods and Stratigraphy of Devonian Strata of the Central Regions of the Russian Platform] Atlas braxiopod i stratigrafia devonskix otlozenij central'nyx oblastej russkoj Platformy. Gostoltexizdat, Moskva, 451 p., 87 pls." type="book" year="1959">Ljaschenko 1959: 175</bibRefCitation>
, pl. 51: 5-7 [lapsus calami pro
<taxonomicName id="4C4E4CBEFFEDFFCFFEF7FB7FFE4319DC" authorityName="Tschernyschew" authorityYear="1885" box="[336,477,1239,1263]" class="Rhynchonellata" family="Palaferellidae" genus="Gruenewaldtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="9" pageNumber="298" phylum="Brachiopoda" rank="genus">
<emphasis id="B93AEB2FFFEDFFCFFEF7FB7FFE4319DC" box="[336,477,1239,1263]" italics="true" pageId="9" pageNumber="298">Gruenewaldtia</emphasis>
</taxonomicName>
].
</paragraph>
<paragraph id="8BF1373DFFEDFFCFFFD6FAAFFF211809" blockId="9.[113,646,1287,1338]" pageId="9" pageNumber="298">
<treatmentCitationGroup id="AB5E1013FFEDFFCFFFD6FAAFFF211809" pageId="9" pageNumber="298">
<taxonomicName id="4C4E4CBEFFEDFFCFFFD6FAAFFF2E182D" ID-CoL="3df4387b-1c40-49b9-880b-8059a37333d2" box="[113,176,1287,1310]" class="Rhynchonellata" family="Atrypidae" genus="Atrypa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="9" pageNumber="298" phylum="Brachiopoda" rank="genus">
<emphasis id="B93AEB2FFFEDFFCFFFD6FAAFFF2E182D" box="[113,176,1287,1310]" italics="true" pageId="9" pageNumber="298">Atrypa</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C4E4CBEFFEDFFCFFF52FAAFFECF182C" authorityName="Michael" authorityYear="1920" box="[245,337,1287,1311]" class="Rhynchonellata" family="Atrypidae" genus="Atrypa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="9" pageNumber="298" phylum="Brachiopoda" rank="species" species="reticularis">
<emphasis id="B93AEB2FFFEDFFCFFF52FAAFFECF182C" box="[245,337,1287,1311]" italics="true" pageId="9" pageNumber="298">reticularis</emphasis>
</taxonomicName>
<treatmentCitation id="0AEF112CFFEDFFCFFECBFAAFFDD9182C" author="GROCHOLSKI W." box="[364,583,1287,1311]" page="246" pageId="9" pageNumber="298" year="1969">
<bibRefCitation id="EFDF4ACCFFEDFFCFFECBFAAFFDD9182C" author="GROCHOLSKI W." box="[364,583,1287,1311]" pageId="9" pageNumber="298" refId="ref9327" refString="GROCHOLSKI W. (ed.). 1969. - [Geological guide book of the Sudetes] Przewodnik geologiczny po Sudetach. WydawnictwaGeologiczne, Warszawa, 535 p. (inPolish)." type="book" year="1969">Grocholski 1969: 246</bibRefCitation>
</treatmentCitation>
, pl. 5: 2 (e.p.).
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</paragraph>
</subSubSection>
<subSubSection id="C35464B6FFEDFFCFFFD6FAFAFE1918AD" pageId="9" pageNumber="298" type="materials_examined">
<paragraph id="8BF1373DFFEDFFCFFFD6FAFAFE1918AD" blockId="9.[113,648,1361,1438]" pageId="9" pageNumber="298">
<collectionCode id="ED5FAFF8FFEDFFCFFFD6FAFAFF46185B" box="[113,216,1362,1385]" pageId="9" pageNumber="298">MATERIAL</collectionCode>
EXAMINED. — 21 articulated shells from Mokrzeszów,
<collectionCode id="ED5FAFF8FFEDFFCFFEA7FAC4FEB218B7" box="[256,300,1388,1412]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="9" pageNumber="298">MB</collectionCode>
.B.297.1-10, 2419.1-2, 2420.1-2; MGUWr 5358s.1-
<geoCoordinate id="EE7A51FAFFEDFFCFFE8AFA2EFE1C18AD" box="[301,386,1414,1438]" degrees="6, 5360" direction="south" orientation="latitude" pageId="9" pageNumber="298" precision="5" value="-6.536">6, 5360s</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="C35464B6FFEDFFCFFFD6FA64FC0618D6" pageId="9" pageNumber="298" type="description">
<paragraph id="8BF1373DFFEDFFCFFFD6FA64FE9C18D7" blockId="9.[112,648,1484,1700]" box="[113,258,1484,1510]" pageId="9" pageNumber="298">DESCRIPTION</paragraph>
<paragraph id="8BF1373DFFEDFFCFFFD6FA43FC0618D6" blockId="9.[112,648,1484,1700]" lastBlockId="9.[675,1212,944,1510]" pageId="9" pageNumber="298">
Shell up to
<quantity id="4CB69AD8FFEDFFCFFF5CFA43FEFB1B36" box="[251,357,1515,1541]" metricMagnitude="-2" metricUnit="m" metricValue="4.9399999999999995" pageId="9" pageNumber="298" unit="mm" value="49.4">49.4 mm</quantity>
wide (mean value in the sample
<quantity id="4CB69AD8FFEDFFCFFF6DF9A3FEA91B16" box="[202,311,1547,1573]" metricMagnitude="-2" metricUnit="m" metricValue="2.98" pageId="9" pageNumber="298" unit="mm" value="29.8">29.8 mm</quantity>
), more often wider than long (width to length ratio from 0.88 to 1.44; mean value 1.07, N=10), rounded in outline, dorsibiconvex in young specimens, ventribiconvex (e.g.,
<figureCitation id="13752BB8FFEDFFCFFFD7F921FF301B97" box="[112,174,1673,1700]" captionStart="FIG" captionStartId="8.[114,125,1589,1607]" captionTargetBox="[134,1193,215,1485]" captionTargetId="figure-174@8.[134,1193,215,1485]" captionTargetPageId="8" captionText="FIG. 5. — Pseudogruenewaldtia tschernyschewi Rzhonsnitskaya,1960;Mokrzeszów (Oberkunzendorf),Late Frasnian:A-E, articulated shell MGUWr 5364s.1,specimen figured by Grocholski (1969;pl.5:2) as Atrypa ex gr.reticularis; F, V-Z, articulated shell MGUWr 5364s.1; G-K, articulated shell MB.B.2421.1; L-P, articulated shell MB.B.297.10; Q-U, articulated shell MB.B.2421.2; AA-EE, articulated shell MB.B.297.1. A, G, L, Q, V, AA, dorsal views; B, H, M, R, W, BB, ventral views; C, I, N, S, X, CC, lateral views; D, J, O, T, Y, DD, posterior views; E, K, P, U, Z, EE, anterior views; F, microornamentation. Scale bars: F, 5 mm; all others, 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4538049" httpUri="https://zenodo.org/record/4538049/files/figure.png" pageId="9" pageNumber="298">Fig. 5</figureCitation>
Q-U) to dorsibiconvex (e.g.,
<figureCitation id="13752BB8FFEDFFCFFE48F921FDB51B97" box="[495,555,1673,1700]" captionStart="FIG" captionStartId="8.[114,125,1589,1607]" captionTargetBox="[134,1193,215,1485]" captionTargetId="figure-174@8.[134,1193,215,1485]" captionTargetPageId="8" captionText="FIG. 5. — Pseudogruenewaldtia tschernyschewi Rzhonsnitskaya,1960;Mokrzeszów (Oberkunzendorf),Late Frasnian:A-E, articulated shell MGUWr 5364s.1,specimen figured by Grocholski (1969;pl.5:2) as Atrypa ex gr.reticularis; F, V-Z, articulated shell MGUWr 5364s.1; G-K, articulated shell MB.B.2421.1; L-P, articulated shell MB.B.297.10; Q-U, articulated shell MB.B.2421.2; AA-EE, articulated shell MB.B.297.1. A, G, L, Q, V, AA, dorsal views; B, H, M, R, W, BB, ventral views; C, I, N, S, X, CC, lateral views; D, J, O, T, Y, DD, posterior views; E, K, P, U, Z, EE, anterior views; F, microornamentation. Scale bars: F, 5 mm; all others, 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4538049" httpUri="https://zenodo.org/record/4538049/files/figure.png" pageId="9" pageNumber="298">Fig. 5</figureCitation>
AA-EE) in adults. Maximal width somewhat anteriorly to midlength (½ to ³⁄₅ of the shell length), maximal thickness posteriorly to midlength (⅓ to ½ of the shell length). Anterior commissure uniplicate, its deflexion rounded in outline, relatively low, occupying ⅓ to ³⁄₅ of the shell width. Ventral interarea concave, apsacline in young specimens; beak (foramen not observed) strongly incurved in older specimens, the interarea becoming thus anacline. Dorsal interarea not visible. A shallow flat-bottomed ventral sulcus is present in the largest specimens, its width at anterior commissure being 1₆ of that of the shell. Ribs undulose, fine, 8-10 per
<quantity id="4CB69AD8FFEDFFCFFCA0FAE4FCD91854" box="[775,839,1356,1383]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="298" unit="mm" value="5.0">5 mm</quantity>
at anterior commissure, new ones appearing (especially in posterior part) by bifurcation, seldom by intercalation. Growth lamellae (preserved only exceptionally) imbricate. Interior not studied.
</paragraph>
</subSubSection>
<subSubSection id="C35464B6FFEDFFCDFD02F9A3FE2E199B" lastPageId="11" lastPageNumber="300" pageId="9" pageNumber="298" type="discussion">
<paragraph id="8BF1373DFFEDFFCFFD02F9A3FCB81B10" blockId="9.[675,1209,1547,1700]" box="[677,806,1547,1573]" pageId="9" pageNumber="298">DISCUSSION</paragraph>
<paragraph id="8BF1373DFFEDFFCDFD02F982FE3C1CFC" blockId="9.[675,1209,1547,1700]" lastBlockId="11.[112,649,216,1192]" lastPageId="11" lastPageNumber="300" pageId="9" pageNumber="298">
The agreement of the described brachiopods from Mokrzeszów in external form (general shape and presence of a similar ventral furrow in the largest specimens), character and density of ornamenta- tion with the
<typeStatus id="54F5899FFFEFFFCDFEA0FF70FEA81DC1" box="[263,310,216,242]" pageId="11" pageNumber="300">type</typeStatus>
material of
<taxonomicName id="4C4E4CBEFFEFFFCDFE1EFF70FEAC1C22" authorityName="Rzhonsnitskaya" authorityYear="1960" class="Rhynchonellata" family="Atrypidae" genus="Pseudogruenewaldtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="11" pageNumber="300" phylum="Brachiopoda" rank="species" species="tschernyschewi">
<emphasis id="B93AEB2FFFEFFFCDFE1EFF70FEAC1C22" italics="true" pageId="11" pageNumber="300">Pseudogruenewaldtia tschernyschewi</emphasis>
</taxonomicName>
from Timan (northern Urals,
<collectingCountry id="F35977ADFFEFFFCDFFD3FEBFFF231C01" box="[116,189,279,306]" name="Russia" pageId="11" pageNumber="300">Russia</collectingCountry>
) make strong case for the conspecificity of both samples even if interior of the described material could not be studied because of scarcity of the collection.
<taxonomicName id="4C4E4CBEFFEFFFCDFEE6FEDEFEBA1C83" authority="Alekseeva, 1996" authorityName="Alekseeva" authorityYear="1996" class="Rhynchonellata" family="Atrypidae" genus="Pseudogruenewaldtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="11" pageNumber="300" phylum="Brachiopoda" rank="species" species="elongata">
<emphasis id="B93AEB2FFFEFFFCDFEE6FEDEFD171CA3" box="[321,649,374,400]" italics="true" pageId="11" pageNumber="300">Pseudogruenewaldtia elongata</emphasis>
Alekseeva, 1996
</taxonomicName>
from north-eastern Siberia, has much more elongate shells.
</paragraph>
<caption id="DF3167B5FFEEFFCCFFD5F9B7FDE51B91" ID-DOI="http://doi.org/10.5281/zenodo.4538053" ID-Zenodo-Dep="4538053" httpUri="https://zenodo.org/record/4538053/files/figure.png" pageId="10" pageNumber="299" startId="10.[114,125,1567,1585]" targetBox="[164,1155,215,1522]" targetPageId="10">
<paragraph id="8BF1373DFFEEFFCCFFD5F9B7FDE51B91" blockId="10.[114,1211,1566,1698]" pageId="10" pageNumber="299">
FIG. 7. —
<taxonomicName id="4C4E4CBEFFEEFFCCFF62F9B6FE391B02" authority="Halamski, 2013" authorityName="Halamski" authorityYear="2013" box="[197,423,1566,1585]" class="Rhynchonellata" family="Atrypidae" genus="Spinatrypa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="10" pageNumber="299" phylum="Brachiopoda" rank="species" species="mariaetheresiae" status="sp. nov.">
<emphasis id="B93AEB2FFFEEFFCCFF62F9B6FE391B02" box="[197,423,1566,1585]" italics="true" pageId="10" pageNumber="299">Spinatrypa mariaetheresiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2095654FFEEFFCCFE0BF9B7FE411B02" box="[428,479,1567,1585]" pageId="10" pageNumber="299" rank="species">n. sp.</taxonomicNameLabel>
, Mokrzeszów (Oberkunzendorf), Late Frasnian:
<emphasis id="B93AEB2FFFEEFFCCFCC9F9B7FC101B02" bold="true" box="[878,910,1567,1585]" pageId="10" pageNumber="299">
A
<emphasis id="B93AEB2FFFEEFFCCFCDDF9B7FC1F1B02" bold="true" box="[890,897,1567,1585]" italics="true" pageId="10" pageNumber="299">-</emphasis>
E
</emphasis>
, paratype, articulated shell MGUWr 5360s.1 figured by
<bibRefCitation id="EFDF4ACCFFEEFFCCFEB1F99DFE1E1B74" author="GUNIA T." box="[278,384,1589,1607]" pageId="10" pageNumber="299" pagination="115 - 220" refId="ref9426" refString="GUNIA T. 1968. - On the fauna, stratigraphy and conditions of sedimentation of the upper Devonian in the Swiebodzice depression (Middle Sudetes). Geologia Sudetica 4: 115 - 220." type="journal article" year="1968">Gunia (1968</bibRefCitation>
: pl. 7: 18, 19) as
<taxonomicName id="4C4E4CBEFFEEFFCCFDB4F99DFD081B74" authorityName="sensu Gunia" authorityYear="1968" box="[531,662,1589,1607]" class="Rhynchonellata" family="Atrypidae" genus="Spinatrypa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="10" pageNumber="299" phylum="Brachiopoda" rank="species" species="bifidaeformis">
<emphasis id="B93AEB2FFFEEFFCCFDB4F99DFD081B74" box="[531,662,1589,1607]" italics="true" pageId="10" pageNumber="299">S. bifidaeformis</emphasis>
</taxonomicName>
;
<emphasis id="B93AEB2FFFEEFFCCFD05F99DFD201B74" bold="true" box="[674,702,1589,1607]" pageId="10" pageNumber="299">
F
<emphasis id="B93AEB2FFFEEFFCCFD0AF99DFD2A1B74" bold="true" box="[685,692,1589,1607]" italics="true" pageId="10" pageNumber="299">-</emphasis>
J
</emphasis>
, paratype, articulated shell MGUWr 5361s.1, corresponding to
<taxonomicName id="4C4E4CBEFFEEFFCCFF2EF9E4FEE01B6D" authority="sensu Gunia, 1966" authorityName="sensu Gunia" authorityYear="1966" box="[137,382,1612,1630]" class="Rhynchonellata" family="Atrypidae" genus="Spinatrypa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="10" pageNumber="299" phylum="Brachiopoda" rank="species" species="aspera">
<emphasis id="B93AEB2FFFEEFFCCFF2EF9E4FE911B6D" box="[137,271,1612,1630]" italics="true" pageId="10" pageNumber="299">S. aspera sensu</emphasis>
<bibRefCitation id="EFDF4ACCFFEEFFCCFEB2F9E4FEE01B6D" author="GUNIA T." box="[277,382,1612,1630]" pageId="10" pageNumber="299" pagination="297 - 322" refId="ref9400" refString="GUNIA T. 1966. - Fauna and age of limestone pebbles from the Culm of Ksiaz. Geologia Sudetica 2: 297 - 322." type="journal article" year="1966">Gunia, 1966</bibRefCitation>
</taxonomicName>
;
<emphasis id="B93AEB2FFFEEFFCCFE20F9E4FE341B6D" bold="true" box="[391,426,1612,1630]" pageId="10" pageNumber="299">
K
<emphasis id="B93AEB2FFFEEFFCCFE33F9E4FE051B6D" bold="true" box="[404,411,1612,1630]" italics="true" pageId="10" pageNumber="299">-</emphasis>
O
</emphasis>
, paratype, articulated shell MB.B.2539;
<emphasis id="B93AEB2FFFEEFFCCFD59F9E4FC821B6D" bold="true" box="[766,796,1612,1630]" pageId="10" pageNumber="299">
P
<emphasis id="B93AEB2FFFEEFFCCFCADF9E4FC8F1B6D" bold="true" box="[778,785,1612,1630]" italics="true" pageId="10" pageNumber="299">-</emphasis>
T
</emphasis>
, paratype, articulated shell MGUWr 5360s.2 corresponding to
<taxonomicName id="4C4E4CBEFFEEFFCCFF4AF9CAFD8B1B46" authority="sensu Gunia, 1968" authorityName="sensu Gunia" authorityYear="1968" box="[237,533,1634,1653]" class="Rhynchonellata" family="Atrypidae" genus="Spinatrypa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="10" pageNumber="299" phylum="Brachiopoda" rank="species" species="bifidaeformis">
<emphasis id="B93AEB2FFFEEFFCCFF4AF9CAFE391B46" box="[237,423,1634,1653]" italics="true" pageId="10" pageNumber="299">S. bifidaeformis sensu</emphasis>
<bibRefCitation id="EFDF4ACCFFEEFFCCFE0AF9CAFD8B1B46" author="GUNIA T." box="[429,533,1634,1653]" pageId="10" pageNumber="299" pagination="115 - 220" refId="ref9426" refString="GUNIA T. 1968. - On the fauna, stratigraphy and conditions of sedimentation of the upper Devonian in the Swiebodzice depression (Middle Sudetes). Geologia Sudetica 4: 115 - 220." type="journal article" year="1968">Gunia, 1968</bibRefCitation>
</taxonomicName>
;
<emphasis id="B93AEB2FFFEEFFCCFDB9F9CBFDA11B46" bold="true" box="[542,575,1635,1653]" pageId="10" pageNumber="299">U -Y</emphasis>
, holotype, articulated shell MB.B. 2424.2;
<emphasis id="B93AEB2FFFEEFFCCFC00F9CBFC481B46" bold="true" box="[935,982,1635,1653]" pageId="10" pageNumber="299">Z -DD</emphasis>
, paratype, articulated shell MB.B.2424.1.
<emphasis id="B93AEB2FFFEEFFCCFF4DF9D1FF691BB8" bold="true" box="[234,247,1657,1675]" pageId="10" pageNumber="299">A</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFEA6F9D1FE921BB8" bold="true" box="[257,268,1657,1675]" pageId="10" pageNumber="299">F</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFEB1F9D1FEBD1BB8" bold="true" box="[278,291,1657,1675]" pageId="10" pageNumber="299">K</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFE8AF9D1FEA71BB8" bold="true" box="[301,313,1657,1675]" pageId="10" pageNumber="299">P</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFEE3F9D1FECC1BB8" bold="true" box="[324,338,1657,1675]" pageId="10" pageNumber="299">U</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFEFBF9D1FEF61BB8" bold="true" box="[348,360,1657,1675]" pageId="10" pageNumber="299">Z</emphasis>
, dorsal views;
<emphasis id="B93AEB2FFFEEFFCCFE44F9D1FE6E1BB8" bold="true" box="[483,496,1657,1675]" pageId="10" pageNumber="299">B</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFE5DF9D1FD961BB8" bold="true" box="[506,520,1657,1675]" pageId="10" pageNumber="299">G</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFDB4F9D1FD801BB8" bold="true" box="[531,542,1657,1675]" pageId="10" pageNumber="299">L</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFD8FF9D1FDA91BB8" bold="true" box="[552,567,1657,1675]" pageId="10" pageNumber="299">Q</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFDE7F9D1FDD21BB8" bold="true" box="[576,588,1657,1675]" pageId="10" pageNumber="299">V</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFDF1F9D1FDEE1BB8" bold="true" box="[598,624,1657,1675]" pageId="10" pageNumber="299">AA</emphasis>
, ventral views;
<emphasis id="B93AEB2FFFEEFFCCFD57F9D1FD601BB8" bold="true" box="[752,766,1657,1675]" pageId="10" pageNumber="299">C</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFCA0F9D1FC8B1BB8" bold="true" box="[775,789,1657,1675]" pageId="10" pageNumber="299">H</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFCB8F9D1FCAE1BB8" bold="true" box="[799,816,1657,1675]" pageId="10" pageNumber="299">M</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFC9DF9D1FCD91BB8" bold="true" box="[826,839,1657,1675]" pageId="10" pageNumber="299">R</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFCF5F9D1FCFA1BB8" bold="true" box="[850,868,1657,1675]" pageId="10" pageNumber="299">W</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFCC9F9D1FC161BB8" bold="true" box="[878,904,1657,1675]" pageId="10" pageNumber="299">BB</emphasis>
, lateral views;
<emphasis id="B93AEB2FFFEEFFCCFBA5F9D1FB8E1BB8" bold="true" box="[1026,1040,1657,1675]" pageId="10" pageNumber="299">D</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFBBDF9D1FBBE1BB8" bold="true" box="[1050,1056,1657,1675]" pageId="10" pageNumber="299">I</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFB8DF9D1FBA61BB8" bold="true" box="[1066,1080,1657,1675]" pageId="10" pageNumber="299">N</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFBE5F9D1FBD01BB8" bold="true" box="[1090,1102,1657,1675]" pageId="10" pageNumber="299">S</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFBFFF9D1FBFA1BB8" bold="true" box="[1112,1124,1657,1675]" pageId="10" pageNumber="299">X</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFBC9F9D1FB141BB8" bold="true" box="[1134,1162,1657,1675]" pageId="10" pageNumber="299">CC</emphasis>
, posterior views;
<emphasis id="B93AEB2FFFEEFFCCFF7BF938FF761B91" bold="true" box="[220,232,1680,1698]" pageId="10" pageNumber="299">E</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFF54F938FF631B91" bold="true" box="[243,253,1680,1698]" pageId="10" pageNumber="299">J</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFEA0F938FE881B91" bold="true" box="[263,278,1680,1698]" pageId="10" pageNumber="299">O</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFE87F938FEB51B91" bold="true" box="[288,299,1680,1698]" pageId="10" pageNumber="299">T</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFE91F938FEDC1B91" bold="true" box="[310,322,1680,1698]" pageId="10" pageNumber="299">Y</emphasis>
,
<emphasis id="B93AEB2FFFEEFFCCFEEAF938FEF71B91" bold="true" box="[333,361,1680,1698]" pageId="10" pageNumber="299">DD</emphasis>
, anterior views. Scale bar: 2 cm.
</paragraph>
</caption>
<paragraph id="8BF1373DFFEFFFCDFF2FFE7EFDA81F5D" blockId="11.[112,649,216,1192]" pageId="11" pageNumber="300">
For discussion of differences with
<taxonomicName id="4C4E4CBEFFEFFFCDFDB2FE7EFF571F3C" authority="Halamski, 2013" authorityName="Halamski" authorityYear="2013" class="Rhynchonellata" family="Atrypidae" genus="Kyrtatrypa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="11" pageNumber="300" phylum="Brachiopoda" rank="species" species="barnimi" status="sp. nov.">
<emphasis id="B93AEB2FFFEFFFCDFDB2FE7EFF571F3C" italics="true" pageId="11" pageNumber="300">Kyrtatrypa barnimi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2095654FFEFFFCDFF77FE5DFE951F3C" box="[208,267,501,527]" pageId="11" pageNumber="300" rank="species">n. sp.</taxonomicNameLabel>
co-occurring in the same outcrop, see above. The stratigraphic implications of the presence of
<taxonomicName id="4C4E4CBEFFEFFFCDFF48FD9CFDF71F7D" authorityName="Rzhonsnitskaya" authorityYear="1960" box="[239,617,564,590]" class="Rhynchonellata" family="Atrypidae" genus="Pseudogruenewaldtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="11" pageNumber="300" phylum="Brachiopoda" rank="species" species="tschernyschewi">
<emphasis id="B93AEB2FFFEFFFCDFF48FD9CFDF71F7D" box="[239,617,564,590]" italics="true" pageId="11" pageNumber="300">Pseudogruenewaldtia tschernyschewi</emphasis>
</taxonomicName>
in the studied material are discussed below.
</paragraph>
<paragraph id="8BF1373DFFEFFFCDFF2FFDDCFF251E3E" blockId="11.[112,649,216,1192]" pageId="11" pageNumber="300">
Reworked representatives of
<taxonomicName id="4C4E4CBEFFEFFFCDFE17FDDCFD1B1FBD" box="[432,645,628,654]" class="Rhynchonellata" family="Atrypidae" genus="Pseudogruenewaldtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="11" pageNumber="300" phylum="Brachiopoda" rank="genus">
<emphasis id="B93AEB2FFFEFFFCDFE17FDDCFD1B1FBD" box="[432,645,628,654]" italics="true" pageId="11" pageNumber="300">Pseudogruenewaldtia</emphasis>
</taxonomicName>
occur in Famennian volcanoclastics (Langenaubacher Tuffbrekzie) in the Dill Syncline (Rhenish Massif, Hessia,
<collectingCountry id="F35977ADFFEFFFCDFEBCFD7BFE1B1FDE" box="[283,389,723,749]" name="Germany" pageId="11" pageNumber="300">Germany</collectingCountry>
) (Copper, pers. comm. 2011).
</paragraph>
<paragraph id="8BF1373DFFEFFFCDFF2FFCBAFE2E199B" blockId="11.[112,649,216,1192]" pageId="11" pageNumber="300">
The peculiar palaeogeographic distribution of
<taxonomicName id="4C4E4CBEFFEFFFCDFF35FC9AFDB91E7F" authorityName="Rzhonsnitskaya" authorityYear="1960" box="[146,551,818,844]" class="Rhynchonellata" family="Atrypidae" genus="Pseudogruenewaldtia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Atrypida" pageId="11" pageNumber="300" phylum="Brachiopoda" rank="species" species="tschernyschewi">
<emphasis id="B93AEB2FFFEFFFCDFF35FC9AFDB91E7F" box="[146,551,818,844]" italics="true" pageId="11" pageNumber="300">Pseudogruenewaldtia tschernyschewi</emphasis>
</taxonomicName>
(Timan,
<collectingCountry id="F35977ADFFEFFFCDFFD6FCF9FF461E58" box="[113,216,849,875]" name="Czech Republic" pageId="11" pageNumber="300">Bohemia</collectingCountry>
, possibly Rhenish Massif; absent either in the Central Devonian Field or in the Holy Cross Mountains) would have been explained by a migration route along the southern margin of Fennoscandia through the Latvian Isthmus had it not been for brackish conditions in these regions in the Frasnian (
<bibRefCitation id="EFDF4ACCFFEFFFCDFE26FBA7FDFF1919" author="LEBEDEV O. A. &amp; ZAKHARENKO G. V." box="[385,609,1039,1066]" pageId="11" pageNumber="300" pagination="108 - 128" refId="ref10150" refString="LEBEDEV O. A., LUKsEVICs E. &amp; ZAKHARENKO G. V. 2010. - Palaeozoogeographical connections of the Devonian vertebrate communities of the Baltica Province. Part II. Late Devonian. Palaeoworld 19: 108 - 128." type="journal article" year="2010">
Lebedev
<emphasis id="B93AEB2FFFEFFFCDFE41FBB8FD85191A" box="[486,539,1039,1065]" italics="true" pageId="11" pageNumber="300">et al.</emphasis>
2010
</bibRefCitation>
). A straighter dispersion route is to be invoked and the absence of the discussed species in the abovementioned regions is more probably related to the patchiness of the record.
</paragraph>
</subSubSection>
</treatment>
</document>