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<mods:title id="9988B85490F07ED7C148C6C5AB08C0B1">Review of unreported shore­fly genera of the tribe Scatellini from the New Zealand subregion (Diptera: Ephydridae) with description of three new species</mods:title>
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<emphasis id="B93A3A13DA32FFAE7C0FFE7E6150FE3E" bold="true" box="[264,579,473,499]" pageId="17" pageNumber="18">
Genus
<taxonomicName id="4C4E9D82DA32FFAE7C58FE7E6150FE3E" ID-CoL="62X6G" ID-ENA="92593" authority="Malloch" authorityName="Malloch" box="[351,579,473,499]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B93A3A13DA32FFAE7C58FE7E62C7FE3E" bold="true" box="[351,468,473,499]" italics="true" pageId="17" pageNumber="18">Limnellia</emphasis>
Malloch
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</emphasis>
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</paragraph>
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<subSubSection id="C354B58ADA32FFAE7C0FFD836154FDB6" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8BF1E601DA32FFAE7C0FFD836154FDB6" blockId="17.[264,1324,548,635]" pageId="17" pageNumber="18">
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<taxonomicName id="4C4E9D82DA32FFAE7C0FFD836156FDF6" ID-CoL="62X6G" authority="Malloch 1925: 331" authorityName="Malloch" authorityPageNumber="331" authorityYear="1925" box="[264,581,548,571]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B93A3A13DA32FFAE7C0FFD83627DFDF6" box="[264,366,548,571]" italics="true" pageId="17" pageNumber="18">Limnellia</emphasis>
<treatmentCitation id="0AEFC010DA32FFAE7C71FD836156FDF6" author="Malloch" box="[374,581,548,571]" page="331" pageId="17" pageNumber="18" year="1925">
<bibRefCitation id="EFDF9BF0DA32FFAE7C71FD83611CFDF6" author="Malloch" box="[374,527,548,571]" pageId="17" pageNumber="26" refString="Malloch, J. R. (1925) Notes on Australian Diptera. No. vii. Proceedings of the Linnean Society of New South Wales, 50 (4), 311 - 340." type="journal article" year="1925">Malloch 1925</bibRefCitation>
: 331
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</taxonomicName>
.
<typeStatus id="54F558A3DA32FFAE7F57FD836196FDF6" box="[592,645,548,571]" pageId="17" pageNumber="18">Type</typeStatus>
species:
<taxonomicName id="4C4E9D82DA32FFAE7FEFFD836790FDF6" ID-CoL="72BRZ" authority="Malloch 1925" authorityName="Malloch" authorityYear="1925" box="[744,1155,548,571]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="B93A3A13DA32FFAE7FEFFD8360F7FDF6" box="[744,996,548,571]" italics="true" pageId="17" pageNumber="18">Limnellia maculipennis</emphasis>
<bibRefCitation id="EFDF9BF0DA32FFAE7EECFD836790FDF6" author="Malloch" box="[1003,1155,548,571]" pageId="17" pageNumber="26" refString="Malloch, J. R. (1925) Notes on Australian Diptera. No. vii. Proceedings of the Linnean Society of New South Wales, 50 (4), 311 - 340." type="journal article" year="1925">Malloch 1925</bibRefCitation>
</taxonomicName>
, original designation.­
<treatmentCitation id="0AEFC010DA32FFAE7C8FFDE36146FD96" author="Mathis" box="[392,597,580,603]" page="648" pageId="17" pageNumber="18" year="1989">
<bibRefCitation id="EFDF9BF0DA32FFAE7C8FFDE36108FD96" author="Mathis" box="[392,539,580,603]" pageId="17" pageNumber="26" refString="Mathis, W. N. (1989) 66. Family Ephydridae. In: Evenhuis, N. L. (Ed.), Catalog of the Diptera of the Australasian and Oceanian Regions. Bishop Museum Press and E. J. Brill, Honolulu and Leiden, pp. 639 - 649." type="book chapter" year="1989">Mathis 1989</bibRefCitation>
: 648
</treatmentCitation>
[Australasian/Oceanian catalog].­
<treatmentCitation id="0AEFC010DA32FFAE7EC9FDE36273FDB6" author="Mathis" page="258" pageId="17" pageNumber="18" year="1995">
<bibRefCitation id="EFDF9BF0DA32FFAE7EC9FDE36634FD96" author="Mathis" box="[974,1319,580,603]" pageId="17" pageNumber="26" refString="Mathis, W. N. and Zatwarnicki, T. (1995) A world catalog of the shore flies (Diptera: Ephydridae). Memoirs on Entomology, International, 4, vi + 423 pp." type="book" year="1995">Mathis and Zatwarnicki 1995</bibRefCitation>
: 258
</treatmentCitation>
261 [world catalog].
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</paragraph>
</subSubSection>
<subSubSection id="C354B58ADA32FFAE7C0FFD0B61ABFBE3" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8BF1E601DA32FFAE7C0FFD0B674DFCF3" blockId="17.[264,1324,684,1870]" pageId="17" pageNumber="18">
<emphasis id="B93A3A13DA32FFAE7C0FFD0B629BFD0B" bold="true" box="[264,392,684,710]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
<taxonomicName id="4C4E9D82DA32FFAE7C96FD0B6112FD08" box="[401,513,684,709]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B93A3A13DA32FFAE7C96FD0B6112FD08" box="[401,513,684,709]" italics="true" pageId="17" pageNumber="18">Limnellia</emphasis>
</taxonomicName>
is distinguished from other genera of the tribe
<taxonomicName id="4C4E9D82DA32FFAE7932FD0B67B7FD0B" box="[1077,1188,684,710]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="tribe" tribe="Scatellini">Scatellini</taxonomicName>
by the following characters: Small to moderately small shore flies, body length
<quantity id="4CB64BE4DA32FFAE7973FD736636FD23" box="[1140,1317,724,750]" metricMagnitude="-3" metricUnit="m" metricValue="1.875" metricValueMax="2.5" metricValueMin="1.25" pageId="17" pageNumber="18" unit="mm" value="1.875" valueMax="2.5" valueMin="1.25">1.252.50 mm</quantity>
; mostly dark brown to black, microtomentose to bare, shiny, frequently with cinereous guttate and vittate maculae; wing maculate, generally dark with white spots.
</paragraph>
<paragraph id="8BF1E601DA32FFAE7C3FFCEB61ABFBE3" blockId="17.[264,1324,684,1870]" pageId="17" pageNumber="18">
<emphasis id="B93A3A13DA32FFAE7C3FFCEB6293FCA8" box="[312,384,844,869]" italics="true" pageId="17" pageNumber="18">Head:</emphasis>
Mesofrons distinct from duller parafrons, sometimes subshiny; lateroclinate fronto­orbital seta 1; inner and outer vertical setae both well developed; paravertical setae either reduced or absent; ocelli arranged in isosceles or equilateral triangle. Arista at most macropubescent. Face with shallow antennal grooves; interfoveal carina not projected or creased dorsally; facial setae mostly small, hair­like. Eye nearly round. Gena relatively short, bearing 1 well­developed seta.
</paragraph>
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<paragraph id="8BF1E601DA32FFAE7C3FFB9B618BFB03" blockId="17.[264,1324,684,1870]" pageId="17" pageNumber="18">
<emphasis id="B93A3A13DA32FFAE7C3FFB9B6284FB98" box="[312,407,1084,1109]" italics="true" pageId="17" pageNumber="18">Thorax:</emphasis>
Acrostichal setae uniform in size, small, arranged in 2 rows that extend to base of scutellum; dorsocentral setae 2 (0+2); supra­alar seta either reduced or lacking; disc of scutellum bare; lateral scutellar setae 2. Wing maculate, mostly dark brown with white, generally guttate markings.
</paragraph>
<paragraph id="8BF1E601DA32FFAE7C3FFB7B625CFA73" blockId="17.[264,1324,684,1870]" pageId="17" pageNumber="18">
<emphasis id="B93A3A13DA32FFAE7C3FFB7B62A3FB38" box="[312,432,1244,1269]" italics="true" pageId="17" pageNumber="18">Abdomen:</emphasis>
Tergites black, becoming shiny and polished posteriorly. Male terminalia: Epandrium a closed plate around cercal cavity, bearing articulated surstyli on anterior margin; gonites elongate with broad base, bearing setae on ventral portion; phallapodeme in lateral view nearly straight or deflected medially; dorsal aedeagal opening with a fold along ventral margin; aedeagus without distiphallus; ejaculatory apodeme usually present, small.
</paragraph>
</subSubSection>
<subSubSection id="C354B58ADA32FFAD7C3FFA6B67FDFD83" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BF1E601DA32FFAE7C3FFA6B6241F9FB" blockId="17.[264,1324,684,1870]" pageId="17" pageNumber="18">
<emphasis id="B93A3A13DA32FFAE7C3FFA6B62D0FA2B" bold="true" box="[312,451,1484,1510]" pageId="17" pageNumber="18">Discussion.</emphasis>
<taxonomicName id="4C4E9D82DA32FFAE7CCBFA6B612FFA28" box="[460,572,1484,1509]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B93A3A13DA32FFAE7CCBFA6B612FFA28" box="[460,572,1484,1509]" italics="true" pageId="17" pageNumber="18">Limnellia</emphasis>
</taxonomicName>
includes 18 species worldwide (
<bibRefCitation id="EFDF9BF0DA32FFAE7EC6FA6B6631FA2B" author="Mathis" box="[961,1314,1484,1510]" pageId="17" pageNumber="26" refString="Mathis, W. N. and Zatwarnicki, T. (1995) A world catalog of the shore flies (Diptera: Ephydridae). Memoirs on Entomology, International, 4, vi + 423 pp." type="book" year="1995">Mathis and Zatwarnicki 1995</bibRefCitation>
) that occur in most temperate regions or at higher elevation in subtropical and tropical zones.
</paragraph>
<paragraph id="8BF1E601DA32FFAD7C3FF9E367FDFD83" blockId="17.[264,1324,684,1870]" lastBlockId="18.[264,1323,284,590]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
Although now stable, the early nomenclatural history of
<taxonomicName id="4C4E9D82DA32FFAE7EE2F9E36745F990" box="[997,1110,1604,1629]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B93A3A13DA32FFAE7EE2F9E36745F990" box="[997,1110,1604,1629]" italics="true" pageId="17" pageNumber="18">Limnellia</emphasis>
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reflects a lack of communications and our very fragmentary knowledge about these shore flies on a world basis.
<bibRefCitation id="EFDF9BF0DA32FFAE7C55F9336118F963" author="Malloch" box="[338,523,1684,1710]" pageId="17" pageNumber="26" refString="Malloch, J. R. (1925) Notes on Australian Diptera. No. vii. Proceedings of the Linnean Society of New South Wales, 50 (4), 311 - 340." type="journal article" year="1925">Malloch (1925)</bibRefCitation>
first described
<taxonomicName id="4C4E9D82DA32FFAE7FC4F9336020F960" box="[707,819,1684,1709]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B93A3A13DA32FFAE7FC4F9336020F960" box="[707,819,1684,1709]" italics="true" pageId="17" pageNumber="18">Limnellia</emphasis>
</taxonomicName>
from a female specimen collected in Sydney,
<collectingCountry id="F359A691DA32FFAE7C38F91B62BCF91B" box="[319,431,1724,1750]" name="Australia" pageId="17" pageNumber="18">Australia</collectingCountry>
. Just five years later
<bibRefCitation id="EFDF9BF0DA32FFAE7FA4F91B6045F91B" author="Cresson" box="[675,854,1724,1750]" pageId="17" pageNumber="25" refString="Cresson, E. T., Jr. (1930) Studies in the dipterous family Ephydridae. Paper III. Transactions of the American Entomological Society, 56, 93 - 131." type="journal article" year="1930">Cresson (1930)</bibRefCitation>
and
<bibRefCitation id="EFDF9BF0DA32FFAE7E88F91B6722F91B" author="Collin" box="[911,1073,1724,1750]" pageId="17" pageNumber="25" refString="Collin, J. E. (1930) Some new species of the dipterous genus Scatella Dsv. and the differentiation of Stictoscatella gen. nov. (Ephydridae). The Entomologist's Monthly Magazine, 66, 133 - 139." type="journal article" year="1930">Collin (1930)</bibRefCitation>
, in separate but coincidental publications, redescribed the genus as
<emphasis id="B93A3A13DA32FFAE7E9DF9436721F930" box="[922,1074,1764,1789]" italics="true" pageId="17" pageNumber="18">Eustigoptera</emphasis>
and
<emphasis id="B93A3A13DA32FFAE798AF9436639F930" box="[1165,1322,1764,1789]" italics="true" pageId="17" pageNumber="18">Stictoscatella</emphasis>
respectively. Both Cresson and Collin based their genus­group names on the same generitype,
<taxonomicName id="4C4E9D82DA32FFAE7C4BF89361CDF883" authority="Fallen (1813)" authorityName="Fallen" authorityYear="1813" box="[332,734,1844,1870]" class="Insecta" family="Ephydridae" genus="Notiphila" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="B93A3A13DA32FFAE7C4BF893613CF880" box="[332,559,1844,1869]" italics="true" pageId="17" pageNumber="18">Notiphila quadrata</emphasis>
<bibRefCitation id="EFDF9BF0DA32FFAE7F3DF89361CDF883" author="Fallen" box="[570,734,1844,1870]" pageId="17" pageNumber="26" refString="Fallen, C. F. (1813) Beskrifning ofver nagra i Sverige funna Vattenflugor (Hydromyzides). Kongliga Vetenskaps-Academiens Handlingar series 3, 1813, 240 - 257." type="journal article" year="1813">Fallén (1813)</bibRefCitation>
</taxonomicName>
. Although Fallén's species was originally known only from Scandinavia, it is now known to be Holarctic in distribution (
<bibRefCitation id="EFDF9BF0DA31FFAD798DFEBB6635FEFB" author="Mathis" box="[1162,1318,284,310]" pageId="18" pageNumber="26" refString="Mathis, W. N. (1978) A revision of the Nearctic species of Limnellia Malloch (Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 91 (1): 250 - 293." type="journal article" year="1978">Mathis 1978</bibRefCitation>
,
<bibRefCitation id="EFDF9BF0DA31FFAD7C0FFEE36174FE93" author="Mathis" box="[264,615,324,350]" pageId="18" pageNumber="26" refString="Mathis, W. N. and Zatwarnicki, T. (1995) A world catalog of the shore flies (Diptera: Ephydridae). Memoirs on Entomology, International, 4, vi + 423 pp." type="book" year="1995">Mathis and Zatwarnicki 1995</bibRefCitation>
) but is obviously quite disjunct from the southeastern coast of
<collectingCountry id="F359A691DA31FFAD7C2AFECB628AFE4B" box="[301,409,364,390]" name="Australia" pageId="18" pageNumber="19">Australia</collectingCountry>
where its congener, which is the generitype of
<emphasis id="B93A3A13DA31FFAD7EE8FECB6774FE48" box="[1007,1127,364,389]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C4E9D82DA31FFAD7EE8FECB6770FE48" box="[1007,1123,364,389]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Limnellia</taxonomicName>
,
</emphasis>
is found.
<bibRefCitation id="EFDF9BF0DA31FFAD79ECFECB624AFE63" author="Duda" pageId="18" pageNumber="25" refString="Duda, O. (1942) Neue oder ungenugend bekannte Zweiflugler der palaarktischen Region aus meiner Sammlung. 2. Fortsetzung. Deutsche Entomologische Zeitschrift, 1942 (1 - 4), 1 - 39." type="journal article" year="1942">Duda (1942)</bibRefCitation>
, in an unusual paper from the standpoint of nomenclature, redescribed
<taxonomicName id="4C4E9D82DA31FFAD79BDFE336639FE60" box="[1210,1322,404,429]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B93A3A13DA31FFAD79BDFE336639FE60" box="[1210,1322,404,429]" italics="true" pageId="18" pageNumber="19">Limnellia</emphasis>
</taxonomicName>
yet again, this time as the subgenus
<emphasis id="B93A3A13DA31FFAD7FD7FE1B6049FE18" box="[720,858,444,469]" italics="true" pageId="18" pageNumber="19">Stranditella</emphasis>
of the genus
<taxonomicName id="4C4E9D82DA31FFAD790BFE1B6254FE33" authority="Hendel. Duda" authorityName="Hendel. Duda" class="Insecta" family="Ephydridae" genus="Lamproscatella" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B93A3A13DA31FFAD790BFE1B67D1FE18" box="[1036,1218,444,469]" italics="true" pageId="18" pageNumber="19">Lamproscatella</emphasis>
Hendel. Duda
</taxonomicName>
also based
<emphasis id="B93A3A13DA31FFAD7CE3FE43617EFE30" box="[484,621,484,509]" italics="true" pageId="18" pageNumber="19">Stranditella</emphasis>
on the generitype,
<emphasis id="B93A3A13DA31FFAD7E6CFE436748FE30" box="[875,1115,484,509]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C4E9D82DA31FFAD7E6CFE43674BFE30" box="[875,1112,484,509]" class="Insecta" family="Ephydridae" genus="Notiphila" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="quadrata">Notiphila quadrata</taxonomicName>
.
</emphasis>
Duda's paper is unconventional in having most of the new genus­group and several of the included new species names as patronyms, all based on his colleague, Embrik Strand (
<date id="FFF0C0C1DA31FFAD7950FD9367F2FD83" box="[1111,1249,564,590]" pageId="18" pageNumber="19" value="1876" valueMax="1953">18761953</date>
).
</paragraph>
</subSubSection>
</treatment>
</document>