treatments-xml/data/03/D0/1A/03D01A5F5E7DFF8E09AE56CAFAB32E62.xml

<document id="35CF693BA78E95A40A1FF2E8EF11B223" ID-DOI="10.11646/zootaxa.3955.1.3" ID-GBIF-Dataset="17d67da2-0873-4808-b1aa-f31f7cc90e4f" ID-ISSN="1175-5326" ID-Zenodo-Dep="288451" ID-ZooBank="6E10D6BD-0574-4EA8-A160-A00F9CB9346E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461049056181" checkinUser="plazi" docAuthor="Pimenta, Bruno V. S., Caramaschi, Ulisses &amp; Cruz, Carlos Alberto Gonçalves" docDate="2015" docId="03D01A5F5E7DFF8E09AE56CAFAB32E62" docLanguage="en" docName="zt03955p082.pdf" docOrigin="Zootaxa 3955 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Crossodactylus franciscanus Pimenta, Caramaschi &amp; Cruz, 2015, sp. nov." docType="treatment" docVersion="6" lastPageNumber="78" masterDocId="FFE962275E75FF8309395735FF8B2832" masterDocTitle="Synonymy of Crossodactylus bokermanni Caramaschi &amp; Sazima, 1985 with Crossodactylus trachystomus (Reinhardt &amp; Lütken, 1862) and description of a new species from Minas Gerais, Brazil (Anura: Hylodidae)" masterLastPageNumber="82" masterPageNumber="65" pageNumber="73" updateTime="1698578057939" updateUser="plazi">
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<mods:title id="A4CC51FC9DCC91E98C02055DA54F989D">Synonymy of Crossodactylus bokermanni Caramaschi &amp; Sazima, 1985 with Crossodactylus trachystomus (Reinhardt &amp; Lütken, 1862) and description of a new species from Minas Gerais, Brazil (Anura: Hylodidae)</mods:title>
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<mods:namePart id="610BA744EBCFD7AC2EAE6920FF15EE5C">Pimenta, Bruno V. S.</mods:namePart>
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<mods:namePart id="DF943E8F20A8C7EAA7417B6B099B1695">Caramaschi, Ulisses</mods:namePart>
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<mods:namePart id="76D7E8E725E98A09BA61975F01E6F273">Cruz, Carlos Alberto Gonçalves</mods:namePart>
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<mods:title id="48521F3C700B95D1B94576B105936F81">Zootaxa</mods:title>
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<mods:date id="DB59130056FF4205B25A87CA91996AEE">2015</mods:date>
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<mods:number id="6E442FC35D2A24A2A001932B633199F5">3955</mods:number>
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<treatment id="03D01A5F5E7DFF8E09AE56CAFAB32E62" ID-DOI="http://doi.org/10.5281/zenodo.5616022" ID-GBIF-Taxon="127691238" ID-Zenodo-Dep="5616022" LSID="urn:lsid:plazi:treatment:03D01A5F5E7DFF8E09AE56CAFAB32E62" httpUri="http://treatment.plazi.org/id/03D01A5F5E7DFF8E09AE56CAFAB32E62" lastPageId="13" lastPageNumber="78" pageId="8" pageNumber="73">
<subSubSection id="C363F8C25E7DFF8B09AE56CAFD4A2A6E" pageId="8" pageNumber="73" type="nomenclature">
<paragraph id="8BC6AB495E7DFF8B09AE56CAFDDD2A28" blockId="8.[151,598,511,538]" box="[151,598,511,538]" pageId="8" pageNumber="73">
<heading id="D08E1C255E7DFF8B09AE56CAFDDD2A28" bold="true" box="[151,598,511,538]" fontSize="11" level="1" pageId="8" pageNumber="73" reason="1">
<emphasis id="B90D775B5E7DFF8B09AE56CAFDDD2A28" bold="true" box="[151,598,511,538]" pageId="8" pageNumber="73">
<taxonomicName id="4C79D0CA5E7DFF8B09AE56CAFE7A2A2B" ID-CoL="ZMBY" box="[151,497,511,537]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="franciscanus" status="sp. nov.">
<emphasis id="B90D775B5E7DFF8B09AE56CAFE7A2A2B" bold="true" box="[151,497,511,537]" italics="true" pageId="8" pageNumber="73">Crossodactylus franciscanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23ECA205E7DFF8B08C15535FDDD2A28" box="[504,598,512,538]" pageId="8" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC6AB495E7DFF8B09AE5573FD4A2A6E" blockId="8.[151,705,582,605]" box="[151,705,582,605]" pageId="8" pageNumber="73">
<taxonomicName id="4C79D0CA5E7DFF8B09AE5573FE6B2A6E" box="[151,480,582,604]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="trachystomus">
<emphasis id="B90D775B5E7DFF8B09AE5573FEB92A6E" box="[151,306,582,604]" italics="true" pageId="8" pageNumber="73">Crossodactylus</emphasis>
cf. 
<emphasis id="B90D775B5E7DFF8B08605573FE6B2A6E" box="[345,480,582,604]" italics="true" pageId="8" pageNumber="73">trachystomus</emphasis>
</taxonomicName>
— 
<bibRefCitation id="EFE8D6B85E7DFF8B08C05573FD362A6E" author="Haddad" box="[505,701,582,605]" pageId="8" pageNumber="73" refString="Haddad, C. F. B., Andrade, G. V. &amp; Cardoso, A. J. (1988) Anfibios anuros do Parque Nacional da Serra da Canastra, Estado de Minas Gerais. Brasil Florestal, 64, 9 - 20." type="journal article" year="1988">
Haddad 
<emphasis id="B90D775B5E7DFF8B0B77557DFD0A2A6E" box="[590,641,582,605]" italics="true" pageId="8" pageNumber="73">et al.</emphasis>
1988
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C363F8C25E7DFF8B09AE55BCFCAF2B48" pageId="8" pageNumber="73" type="materials_examined">
<paragraph id="8BC6AB495E7DFF8B09AE55BCFBCA2AD8" blockId="8.[151,1437,649,2006]" pageId="8" pageNumber="73">
<emphasis id="B90D775B5E7DFF8B09AE55BCFE802A90" bold="true" box="[151,267,649,674]" pageId="8" pageNumber="73">
<typeStatus id="54C215EB5E7DFF8B09AE55BCFE8D2A90" box="[151,262,649,674]" pageId="8" pageNumber="73" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED68338C5E7DFF8B082155BFFEE82A93" box="[280,355,650,673]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="8" pageNumber="73">MNRJ</collectionCode>
40137, adult male, collected at Baixo da Casca d’Anta (aprox. 
<geoCoordinate id="EE4DCD8E5E7DFF8B0D5555BFFB432A90" box="[1132,1224,650,674]" direction="south" orientation="latitude" pageId="8" pageNumber="73" precision="925" value="-20.216667">20º13’S</geoCoordinate>
, 
<geoCoordinate id="EE4DCD8E5E7DFF8B0DE355BCFAB52A90" box="[1242,1342,649,674]" direction="west" orientation="longitude" pageId="8" pageNumber="73" precision="925" value="-46.366665">46º22’W</geoCoordinate>
, 
<quantity id="4C8106AC5E7DFF8B0C7655BFFA172A93" box="[1359,1436,650,674]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="8" pageNumber="73" unit="m" value="870.0">870 m</quantity>
elevation), Parque Nacional da Serra da Canastra, municipality of São Roque de Minas, Minas Gerais state, 
<collectingCountry id="F36EEBD95E7DFF8B0C685598FA132AF4" box="[1361,1432,685,710]" name="Brazil" pageId="8" pageNumber="73">Brazil</collectingCountry>
, in 
<date id="FFC78D895E7DFF8B098D55E7FEE42AD8" box="[180,367,722,746]" pageId="8" pageNumber="73" value="1981-01-28">28 January 1981</date>
, by A.J. Cardoso, G.V. Andrade, and C.F.B. Haddad (
<figureCitation id="1342B7CC5E7DFF8B0AFE55E4FBBF2AD8" box="[967,1076,721,746]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="9.[151,250,1617,1639]" captionStartId-1="10.[151,250,1844,1866]" captionTargetBox-0="[164,1423,697,1591]" captionTargetBox-1="[170,1418,383,1804]" captionTargetId-0="figure@9.[164,1424,697,1596]" captionTargetId-1="figure@10.[152,1435,373,1823]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 4. Crossodactylus franciscanus sp. nov. (MNRJ 40137, holotype, SVL 22.1 mm), (A) dorsal, (B) ventral, and (C) lateral views." captionText-1="FIGURE 5. Crossodactylus franciscanus sp. nov. (MNRJ 40137, holotype, SVL 22.1 mm). Dorsal and lateral views of head and ventral views of hand and foot. Scales = 5 mm." httpUri-0="https://zenodo.org/record/288455/files/figure.png" httpUri-1="https://zenodo.org/record/288456/files/figure.png" pageId="8" pageNumber="73">Figs. 4–5</figureCitation>
).
</paragraph>
<paragraph id="8BC6AB495E7DFF8B09FE55C0FCAF2B48" blockId="8.[151,1437,649,2006]" pageId="8" pageNumber="73">
<emphasis id="B90D775B5E7DFF8B09FE55C0FECD2B3C" bold="true" box="[199,326,757,782]" pageId="8" pageNumber="73">
<typeStatus id="54C215EB5E7DFF8B09FE55C0FECA2B3C" box="[199,321,757,782]" pageId="8" pageNumber="73" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED68338C5E7DFF8B087655C3FE112B3F" box="[335,410,758,781]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="8" pageNumber="73">MNRJ</collectionCode>
50784–9, four adult males and two adult females, collected at 
<collectingCountry id="F36EEBD95E7DFF8B0D4C55C3FB242B3F" box="[1141,1199,758,781]" name="Malta" pageId="8" pageNumber="73">Mata</collectingCountry>
do Batatal, Serra da Canastra, municipality of Passos (
<geoCoordinate id="EE4DCD8E5E7DFF8B0B11542FFD0F2B00" box="[552,644,794,818]" direction="south" orientation="latitude" pageId="8" pageNumber="73" precision="925" value="-20.716667">20º43’S</geoCoordinate>
, 
<geoCoordinate id="EE4DCD8E5E7DFF8B0BAC542CFD762B00" box="[661,765,793,818]" direction="west" orientation="longitude" pageId="8" pageNumber="73" precision="925" value="-46.616665">46º37’W</geoCoordinate>
), state of Minas Gerais, 
<collectingCountry id="F36EEBD95E7DFF8B0D1F542CFBE62B00" box="[1062,1133,793,818]" name="Brazil" pageId="8" pageNumber="73">Brazil</collectingCountry>
, in 
<date id="FFC78D895E7DFF8B0DA6542FFAC22B00" box="[1183,1353,793,818]" pageId="8" pageNumber="73" value="1943-04-23">23 April 1943</date>
, by H. Veloso; 
<collectionCode id="ED68338C5E7DFF8B09CA540BFEB52B67" box="[243,318,830,853]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="8" pageNumber="73">MNRJ</collectionCode>
50790–1, two adult males, collected at 
<collectingCountry id="F36EEBD95E7DFF8B0A3B540BFCB72B67" box="[770,828,830,853]" name="Malta" pageId="8" pageNumber="73">Mata</collectingCountry>
da 
<taxonomicName id="4C79D0CA5E7DFF8B0A5F5408FC712B64" authority="Serra" authorityName="Serra" box="[870,1018,829,854]" class="Gastropoda" family="Cerithiidae" genus="Colina" kingdom="Animalia" pageId="8" pageNumber="73" phylum="Mollusca" rank="genus">Colina, Serra</taxonomicName>
da Canastra, municipality of Passos, state of Minas Gerais, 
<collectingCountry id="F36EEBD95E7DFF8B08AD5454FE502B48" box="[404,475,865,890]" name="Brazil" pageId="8" pageNumber="73">Brazil</collectingCountry>
, in 
<date id="FFC78D895E7DFF8B0B3A5454FD0B2B48" box="[515,640,865,890]" pageId="8" pageNumber="73" value="1943-04">April 1943</date>
, by H. Veloso.
</paragraph>
</subSubSection>
<subSubSection id="C363F8C25E7DFF8A09FE54B0FE7D2AB6" lastPageId="9" lastPageNumber="74" pageId="8" pageNumber="73" type="diagnosis">
<paragraph id="8BC6AB495E7DFF8B09FE54B0FD512C44" blockId="8.[151,1437,649,2006]" pageId="8" pageNumber="73">
<emphasis id="B90D775B5E7DFF8B09FE54B0FECA2BAC" bold="true" box="[199,321,901,926]" pageId="8" pageNumber="73">Diagnosis.</emphasis>
(1) body slender; (2) head longer than wide; (3) snout rounded in dorsal view, protruding in lateral view; (4) 
<emphasis id="B90D775B5E7DFF8B0829549CFE592BF0" box="[272,466,937,962]" italics="true" pageId="8" pageNumber="73">canthus rostralis</emphasis>
sharp; (5) tympanum distinct; (6) vocal sac subgular, bilobate; (7) thumb spines developed; (11) toe and tarsal fringes broadly developed in males, reduced in females; (9) finger tips undilated; (10) toe tips truncate, undilated; (11) postrictal tubercle continuous; (13) a narrow glandular crest in the anterior surface of the arm, from arm insertion to the elbow; (14) presence of dorsal and dorsolateral folds; (15) a poorly delimited area marbled/dotted of brown over light-brown background from snout to shoulder; (16) presence of partial oblique lateral stripe; (17) belly immaculate.
</paragraph>
<paragraph id="8BC6AB495E7DFF8B09FE53B4FAEF2D40" blockId="8.[151,1437,649,2006]" pageId="8" pageNumber="73">
<emphasis id="B90D775B5E7DFF8B09FE53B4FDEA2CA8" bold="true" box="[199,609,1153,1178]" pageId="8" pageNumber="73">Comparison with other species.</emphasis>
Character states for the other species are shown in parenthesis. 
<taxonomicName id="4C79D0CA5E7DFF8B09AE5390FE512C8C" box="[151,474,1189,1214]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7DFF8B09AE5390FE512C8C" box="[151,474,1189,1214]" italics="true" pageId="8" pageNumber="73">Crossodactylus franciscanus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C79D0CA5E7DFF8B0B485393FD512C8C" box="[625,730,1189,1214]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B90D775B5E7DFF8B0B485393FD512C8C" box="[625,730,1189,1214]" italics="true" pageId="8" pageNumber="73">C. dispar</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C79D0CA5E7DFF8B0A2D5393FC012C8C" box="[788,906,1189,1214]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B90D775B5E7DFF8B0A2D5393FC012C8C" box="[788,906,1189,1214]" italics="true" pageId="8" pageNumber="73">C. grandis</emphasis>
</taxonomicName>
by the smaller size of males (males 20.4–22.1 SVL in 
<taxonomicName id="4C79D0CA5E7DFF8B09CB53FFFE282CD0" box="[242,419,1226,1250]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7DFF8B09CB53FFFE282CD0" box="[242,419,1226,1250]" italics="true" pageId="8" pageNumber="73">C. franciscanus</emphasis>
</taxonomicName>
; combined male SVL in 
<taxonomicName id="4C79D0CA5E7DFF8B0BFD53FFFCA52CD0" box="[708,814,1225,1250]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B90D775B5E7DFF8B0BFD53FFFCA52CD0" box="[708,814,1225,1250]" italics="true" pageId="8" pageNumber="73">C. dispar</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C79D0CA5E7DFF8B0A5F53FFFC552CD0" box="[870,990,1225,1250]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B90D775B5E7DFF8B0A5F53FFFC552CD0" box="[870,990,1225,1250]" italics="true" pageId="8" pageNumber="73">C. grandis</emphasis>
</taxonomicName>
23.6–42.0 mm), slender body (robust), head longer than wide (nearly as long as wide in 
<taxonomicName id="4C79D0CA5E7DFF8B0BE753DBFCC12D34" box="[734,842,1261,1286]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B90D775B5E7DFF8B0BE753DBFCC12D34" box="[734,842,1261,1286]" italics="true" pageId="8" pageNumber="73">C. dispar</emphasis>
</taxonomicName>
and wider than long in 
<taxonomicName id="4C79D0CA5E7DFF8B0D5053DBFB692D34" box="[1129,1250,1261,1286]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B90D775B5E7DFF8B0D5053DBFB692D34" box="[1129,1250,1261,1286]" italics="true" pageId="8" pageNumber="73">C. grandis</emphasis>
</taxonomicName>
), sharp 
<emphasis id="B90D775B5E7DFF8B0C7C53D8FF7E2D18" italics="true" pageId="8" pageNumber="73">canthus rostralis</emphasis>
(rounded), males with extensively fringed feet (moderate), truncate toe tips (rounded), and postrictal tubercle continuous (fragmented into small granules). 
<taxonomicName id="4C79D0CA5E7DFF8B0BCE5200FBB22D7C" box="[759,1081,1333,1358]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7DFF8B0BCE5200FBB22D7C" box="[759,1081,1333,1358]" italics="true" pageId="8" pageNumber="73">Crossodactylus franciscanus</emphasis>
</taxonomicName>
also differs from 
<taxonomicName id="4C79D0CA5E7DFF8B0C3B5203FAF32D7C" box="[1282,1400,1333,1358]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B90D775B5E7DFF8B0C3B5203FAF32D7C" box="[1282,1400,1333,1358]" italics="true" pageId="8" pageNumber="73">C. grandis</emphasis>
</taxonomicName>
by the smaller size in females (females 20.3–23.6 mm SVL in 
<taxonomicName id="4C79D0CA5E7DFF8B0A09526FFC6B2D40" box="[816,992,1370,1394]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7DFF8B0A09526FFC6B2D40" box="[816,992,1370,1394]" italics="true" pageId="8" pageNumber="73">C. franciscanus</emphasis>
</taxonomicName>
and 29.6–39.2 mm in 
<taxonomicName id="4C79D0CA5E7DFF8B0DE4526FFAD82D40" box="[1245,1363,1369,1394]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B90D775B5E7DFF8B0DE4526FFAD82D40" box="[1245,1363,1369,1394]" italics="true" pageId="8" pageNumber="73">C. grandis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BC6AB495E7DFF8B09FE5248FE592E78" blockId="8.[151,1437,649,2006]" pageId="8" pageNumber="73">
<taxonomicName id="4C79D0CA5E7DFF8B09FE5248FD812DA4" box="[199,522,1405,1430]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7DFF8B09FE5248FD812DA4" box="[199,522,1405,1430]" italics="true" pageId="8" pageNumber="73">Crossodactylus franciscanus</emphasis>
</taxonomicName>
could be distinguished from 
<taxonomicName id="4C79D0CA5E7DFF8B0A6E524BFC772DA4" box="[855,1020,1405,1430]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="cyclospinus">
<emphasis id="B90D775B5E7DFF8B0A6E524BFC772DA4" box="[855,1020,1405,1430]" italics="true" pageId="8" pageNumber="73">C. cyclospinus</emphasis>
</taxonomicName>
by its smaller size (males 20.4–22.1 mm and females 20.3–23.6 mm SVL in 
<taxonomicName id="4C79D0CA5E7DFF8B0BBE5297FCB62D88" box="[647,829,1442,1466]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7DFF8B0BBE5297FCB62D88" box="[647,829,1442,1466]" italics="true" pageId="8" pageNumber="73">C. franciscanus</emphasis>
</taxonomicName>
; males 22.8–24.5 mm and female 29.0 mm in 
<taxonomicName id="4C79D0CA5E7DFF8B0CBB5297FE912DEC" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="cyclospinus">
<emphasis id="B90D775B5E7DFF8B0CBB5297FE912DEC" italics="true" pageId="8" pageNumber="73">C. cyclospinus</emphasis>
</taxonomicName>
), from 
<taxonomicName id="4C79D0CA5E7DFF8B085752F3FD912DEC" box="[366,538,1477,1502]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="caramaschii">
<emphasis id="B90D775B5E7DFF8B085752F3FD912DEC" box="[366,538,1477,1502]" italics="true" pageId="8" pageNumber="73">C. caramaschii</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C79D0CA5E7DFF8B0B6852F3FC8D2DEC" box="[593,774,1477,1502]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="gaudichaudii">
<emphasis id="B90D775B5E7DFF8B0B6852F3FC8D2DEC" box="[593,774,1477,1502]" italics="true" pageId="8" pageNumber="73">C. gaudichaudii</emphasis>
</taxonomicName>
by the smaller size of males (combined male SVL in these species 22.5–30.4 mm), and from 
<taxonomicName id="4C79D0CA5E7DFF8B0B2A52DFFD0A2E33" box="[531,641,1514,1537]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="aeneus">
<emphasis id="B90D775B5E7DFF8B0B2A52DFFD0A2E33" box="[531,641,1514,1537]" italics="true" pageId="8" pageNumber="73">C. aeneus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0BB752DFFC992E30" box="[654,786,1513,1538]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="lutzorum">
<emphasis id="B90D775B5E7DFF8B0BB752DFFC992E30" box="[654,786,1513,1538]" italics="true" pageId="8" pageNumber="73">C. lutzorum</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C79D0CA5E7DFF8B0A7752DFFC462E30" box="[846,973,1513,1538]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B90D775B5E7DFF8B0A7752DFFC462E30" box="[846,973,1513,1538]" italics="true" pageId="8" pageNumber="73">C. schmidti</emphasis>
</taxonomicName>
by the smaller size of females (combined female SVL in these species 23.8–31.7 mm). It differs from 
<taxonomicName id="4C79D0CA5E7DFF8B0A07513BFC262E17" box="[830,941,1550,1573]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="aeneus">
<emphasis id="B90D775B5E7DFF8B0A07513BFC262E17" box="[830,941,1550,1573]" italics="true" pageId="8" pageNumber="73">C. aeneus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0A82513BFBA92E14" box="[955,1058,1549,1574]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="dantei">
<emphasis id="B90D775B5E7DFF8B0A82513BFBA92E14" box="[955,1058,1549,1574]" italics="true" pageId="8" pageNumber="73">C. dantei</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0D08513BFB6D2E14" box="[1073,1254,1549,1574]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="gaudichaudii">
<emphasis id="B90D775B5E7DFF8B0D08513BFB6D2E14" box="[1073,1254,1549,1574]" italics="true" pageId="8" pageNumber="73">C. gaudichaudii</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C79D0CA5E7DFF8B0C1D513BFA102E17" box="[1316,1435,1550,1573]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="werneri">
<emphasis id="B90D775B5E7DFF8B0C1D513BFA102E17" box="[1316,1435,1550,1573]" italics="true" pageId="8" pageNumber="73">C. werneri</emphasis>
</taxonomicName>
by its slender body (robust).
</paragraph>
<paragraph id="8BC6AB495E7DFF8B09FE5160FDCB2F10" blockId="8.[151,1437,649,2006]" pageId="8" pageNumber="73">
The rounded snout in dorsal view distinguishes 
<taxonomicName id="4C79D0CA5E7DFF8B0A1A5160FBE42E5C" box="[803,1135,1621,1646]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7DFF8B0A1A5160FBE42E5C" box="[803,1135,1621,1646]" italics="true" pageId="8" pageNumber="73">Crossodactylus franciscanus</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C79D0CA5E7DFF8B0DFE5163FAE12E5C" box="[1223,1386,1621,1646]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="B90D775B5E7DFF8B0DFE5163FAE12E5C" box="[1223,1386,1621,1646]" italics="true" pageId="8" pageNumber="73">C. boulengeri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0CBB5163FEAA2EA0" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="caramaschii">
<emphasis id="B90D775B5E7DFF8B0CBB5163FEAA2EA0" italics="true" pageId="8" pageNumber="73">C. caramaschii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0816514FFE4D2EA0" box="[303,454,1657,1682]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="cylospinus">
<emphasis id="B90D775B5E7DFF8B0816514FFE4D2EA0" box="[303,454,1657,1682]" italics="true" pageId="8" pageNumber="73">C. cylospinus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B08ED514FFDB02EA0" box="[468,571,1657,1682]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="dantei">
<emphasis id="B90D775B5E7DFF8B08ED514FFDB02EA0" box="[468,571,1657,1682]" italics="true" pageId="8" pageNumber="73">C. dantei</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0B70514FFD452EA0" box="[585,718,1657,1682]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="lutzorum">
<emphasis id="B90D775B5E7DFF8B0B70514FFD452EA0" box="[585,718,1657,1682]" italics="true" pageId="8" pageNumber="73">C. lutzorum</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C79D0CA5E7DFF8B0A32514FFC0E2EA0" box="[779,901,1657,1682]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="timbuhy">
<emphasis id="B90D775B5E7DFF8B0A32514FFC0E2EA0" box="[779,901,1657,1682]" italics="true" pageId="8" pageNumber="73">C. timbuhy</emphasis>
</taxonomicName>
(nearly pentagon-shaped; variable in 
<taxonomicName id="4C79D0CA5E7DFF8B0C14514FFA172EA3" box="[1325,1436,1658,1681]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="aeneus">
<emphasis id="B90D775B5E7DFF8B0C14514FFA172EA3" box="[1325,1436,1658,1681]" italics="true" pageId="8" pageNumber="73">C. aeneus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C79D0CA5E7DFF8B09F451ABFE032E84" box="[205,392,1693,1718]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="gaudichaudii">
<emphasis id="B90D775B5E7DFF8B09F451ABFE032E84" box="[205,392,1693,1718]" italics="true" pageId="8" pageNumber="73">C. gaudichaudii</emphasis>
</taxonomicName>
). It differs from 
<taxonomicName id="4C79D0CA5E7DFF8B0B6751ABFD6F2E84" box="[606,740,1693,1718]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B90D775B5E7DFF8B0B6751ABFD6F2E84" box="[606,740,1693,1718]" italics="true" pageId="8" pageNumber="73">C. schmidti</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C79D0CA5E7DFF8B0A1E51ABFC2D2E87" box="[807,934,1694,1717]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="werneri">
<emphasis id="B90D775B5E7DFF8B0A1E51ABFC2D2E87" box="[807,934,1694,1717]" italics="true" pageId="8" pageNumber="73">C. werneri</emphasis>
</taxonomicName>
by the sharp 
<emphasis id="B90D775B5E7DFF8B0D6F51A8FA922E84" box="[1110,1305,1693,1718]" italics="true" pageId="8" pageNumber="73">canthus rostralis</emphasis>
(rounded). 
<taxonomicName id="4C79D0CA5E7DFF8B09AE51F4FE502EE8" box="[151,475,1729,1754]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7DFF8B09AE51F4FE502EE8" box="[151,475,1729,1754]" italics="true" pageId="8" pageNumber="73">Crossodactylus franciscanus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C79D0CA5E7DFF8B0B4C51F7FD6D2EEB" box="[629,742,1730,1753]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="aeneus">
<emphasis id="B90D775B5E7DFF8B0B4C51F7FD6D2EEB" box="[629,742,1730,1753]" italics="true" pageId="8" pageNumber="73">C. aeneus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0BCF51F7FC172EE8" box="[758,924,1729,1754]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="cyclospinus">
<emphasis id="B90D775B5E7DFF8B0BCF51F7FC172EE8" box="[758,924,1729,1754]" italics="true" pageId="8" pageNumber="73">C. cyclospinus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0A9551F7FB9E2EE8" box="[940,1045,1729,1754]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="dantei">
<emphasis id="B90D775B5E7DFF8B0A9551F7FB9E2EE8" box="[940,1045,1729,1754]" italics="true" pageId="8" pageNumber="73">C. dantei</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0D1C51F7FB572EE8" box="[1061,1244,1729,1754]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="gaudichaudii">
<emphasis id="B90D775B5E7DFF8B0D1C51F7FB572EE8" box="[1061,1244,1729,1754]" italics="true" pageId="8" pageNumber="73">C. gaudichaudii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0DD551F7FAF92EE8" box="[1260,1394,1729,1754]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="lutzorum">
<emphasis id="B90D775B5E7DFF8B0DD551F7FAF92EE8" box="[1260,1394,1729,1754]" italics="true" pageId="8" pageNumber="73">C. lutzorum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0CBB51F7FF7D2ECC" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B90D775B5E7DFF8B0CBB51F7FF7D2ECC" italics="true" pageId="8" pageNumber="73">C. schmidti</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C79D0CA5E7DFF8B080251D3FE7E2ECC" box="[315,501,1765,1790]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="trachystomus">
<emphasis id="B90D775B5E7DFF8B080251D3FE7E2ECC" box="[315,501,1765,1790]" italics="true" pageId="8" pageNumber="73">C. trachystomus</emphasis>
</taxonomicName>
by its bilobate subgular vocal sac (not expanded in 
<taxonomicName id="4C79D0CA5E7DFF8B0D5A51D3FB462ECC" box="[1123,1229,1765,1790]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="dantei">
<emphasis id="B90D775B5E7DFF8B0D5A51D3FB462ECC" box="[1123,1229,1765,1790]" italics="true" pageId="8" pageNumber="73">C. dantei</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C79D0CA5E7DFF8B0C3551D3FA1E2ECC" box="[1292,1429,1765,1790]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="lutzorum">
<emphasis id="B90D775B5E7DFF8B0C3551D3FA1E2ECC" box="[1292,1429,1765,1790]" italics="true" pageId="8" pageNumber="73">C. lutzorum</emphasis>
</taxonomicName>
; median subgular in the other species).
</paragraph>
<paragraph id="8BC6AB495E7DFF8B09FE5018FA172FE7" blockId="8.[151,1437,649,2006]" pageId="8" pageNumber="73">
<taxonomicName id="4C79D0CA5E7DFF8B09FE5018FD822F74" box="[199,521,1837,1862]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7DFF8B09FE5018FD822F74" box="[199,521,1837,1862]" italics="true" pageId="8" pageNumber="73">Crossodactylus franciscanus</emphasis>
</taxonomicName>
could be distinguished from 
<taxonomicName id="4C79D0CA5E7DFF8B0A6D501BFC482F77" box="[852,963,1838,1861]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="aeneus">
<emphasis id="B90D775B5E7DFF8B0A6D501BFC482F77" box="[852,963,1838,1861]" italics="true" pageId="8" pageNumber="73">C. aeneus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0AEB501BFBE72F74" box="[978,1132,1837,1862]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="B90D775B5E7DFF8B0AEB501BFBE72F74" box="[978,1132,1837,1862]" italics="true" pageId="8" pageNumber="73">C. boulengeri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0D42501BFA942F74" box="[1147,1311,1837,1862]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="cyclospinus">
<emphasis id="B90D775B5E7DFF8B0D42501BFA942F74" box="[1147,1311,1837,1862]" italics="true" pageId="8" pageNumber="73">C. cyclospinus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0C14501BFA1E2F74" box="[1325,1429,1837,1862]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="dantei">
<emphasis id="B90D775B5E7DFF8B0C14501BFA1E2F74" box="[1325,1429,1837,1862]" italics="true" pageId="8" pageNumber="73">C. dantei</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B09AE5067FEC72F58" box="[151,332,1873,1898]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="gaudichaudii">
<emphasis id="B90D775B5E7DFF8B09AE5067FEC72F58" box="[151,332,1873,1898]" italics="true" pageId="8" pageNumber="73">C. gaudichaudii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B08635067FE542F58" box="[346,479,1873,1898]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="lutzorum">
<emphasis id="B90D775B5E7DFF8B08635067FE542F58" box="[346,479,1873,1898]" italics="true" pageId="8" pageNumber="73">C. lutzorum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B08D55067FD292F58" box="[492,674,1873,1898]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="trachystomus">
<emphasis id="B90D775B5E7DFF8B08D55067FD292F58" box="[492,674,1873,1898]" italics="true" pageId="8" pageNumber="73">C. trachystomus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0B885067FCBA2F58" box="[689,817,1873,1898]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B90D775B5E7DFF8B0B885067FCBA2F58" box="[689,817,1873,1898]" italics="true" pageId="8" pageNumber="73">C. schmidti</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0A065067FC332F58" box="[831,952,1873,1898]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="timbuhy">
<emphasis id="B90D775B5E7DFF8B0A065067FC332F58" box="[831,952,1873,1898]" italics="true" pageId="8" pageNumber="73">C. timbuhy</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C79D0CA5E7DFF8B0ACF5067FBE52F5B" box="[1014,1134,1874,1897]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="werneri">
<emphasis id="B90D775B5E7DFF8B0ACF5067FBE52F5B" box="[1014,1134,1874,1897]" italics="true" pageId="8" pageNumber="73">C. werneri</emphasis>
</taxonomicName>
by its undilated finger tips (dilated). 
<taxonomicName id="4C79D0CA5E7DFF8B08365040FDD02FBC" box="[271,603,1909,1934]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7DFF8B08365040FDD02FBC" box="[271,603,1909,1934]" italics="true" pageId="8" pageNumber="73">Crossodactylus franciscanus</emphasis>
</taxonomicName>
is separated from 
<taxonomicName id="4C79D0CA5E7DFF8B0A615043FC5A2FBF" box="[856,977,1910,1933]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="aeneus">
<emphasis id="B90D775B5E7DFF8B0A615043FC5A2FBF" box="[856,977,1910,1933]" italics="true" pageId="8" pageNumber="73">C. aeneus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0AD25043FB2B2FBC" box="[1003,1184,1909,1934]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="caramaschii">
<emphasis id="B90D775B5E7DFF8B0AD25043FB2B2FBC" box="[1003,1184,1909,1934]" italics="true" pageId="8" pageNumber="73">C. caramaschii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0D835043FAE22FBC" box="[1210,1385,1909,1934]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="cyclospinus">
<emphasis id="B90D775B5E7DFF8B0D835043FAE22FBC" box="[1210,1385,1909,1934]" italics="true" pageId="8" pageNumber="73">C. cyclospinus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B0CBB5043FEA02F80" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="gaudichaudii">
<emphasis id="B90D775B5E7DFF8B0CBB5043FEA02F80" italics="true" pageId="8" pageNumber="73">C. gaudichaudii</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C79D0CA5E7DFF8B085E50AFFE6A2F80" box="[359,481,1945,1970]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="timbuhy">
<emphasis id="B90D775B5E7DFF8B085E50AFFE6A2F80" box="[359,481,1945,1970]" italics="true" pageId="8" pageNumber="73">C. timbuhy</emphasis>
</taxonomicName>
by the developed thumb spines (small). 
<taxonomicName id="4C79D0CA5E7DFF8B0A9C50ACFB6D2F80" box="[933,1254,1945,1970]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7DFF8B0A9C50ACFB6D2F80" box="[933,1254,1945,1970]" italics="true" pageId="8" pageNumber="73">Crossodactylus franciscanus</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C79D0CA5E7DFF8B09EE508BFECA2FE4" box="[215,321,1981,2006]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="dantei">
<emphasis id="B90D775B5E7DFF8B09EE508BFECA2FE4" box="[215,321,1981,2006]" italics="true" pageId="8" pageNumber="73">C. dantei</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7DFF8B086B508BFE512FE4" box="[338,474,1981,2006]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="lutzorum">
<emphasis id="B90D775B5E7DFF8B086B508BFE512FE4" box="[338,474,1981,2006]" italics="true" pageId="8" pageNumber="73">C. lutzorum</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C79D0CA5E7DFF8B0B24508BFD2B2FE4" box="[541,672,1981,2006]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B90D775B5E7DFF8B0B24508BFD2B2FE4" box="[541,672,1981,2006]" italics="true" pageId="8" pageNumber="73">C. schmidti</emphasis>
</taxonomicName>
by the truncate toe tips (rounded), and from these species and 
<emphasis id="B90D775B5E7DFF8B0CBB508BFA172FE7" box="[1410,1436,1982,2005]" italics="true" pageId="8" pageNumber="73">C.</emphasis>
</paragraph>
<paragraph id="8BC6AB495E7CFF8A09AE57ACFDE428E7" blockId="9.[151,1436,151,645]" pageId="9" pageNumber="74">
<taxonomicName id="4C79D0CA5E7CFF8A09AE57ACFF6E2882" box="[151,229,153,176]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="aeneus">
<emphasis id="B90D775B5E7CFF8A09AE57ACFF6E2882" box="[151,229,153,176]" italics="true" pageId="9" pageNumber="74">aeneus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A09CE57ADFE1E2882" box="[247,405,151,176]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="B90D775B5E7CFF8A09CE57ADFE1E2882" box="[247,405,151,176]" italics="true" pageId="9" pageNumber="74">C. boulengeri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A089157ADFDD32882" box="[424,600,151,176]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="caramaschii">
<emphasis id="B90D775B5E7CFF8A089157ADFDD32882" box="[424,600,151,176]" italics="true" pageId="9" pageNumber="74">C. caramaschii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A0B5257ADFC982882" box="[619,787,151,176]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="cyclospinus">
<emphasis id="B90D775B5E7CFF8A0B5257ADFC982882" box="[619,787,151,176]" italics="true" pageId="9" pageNumber="74">C. cyclospinus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A0A1C57ADFC552882" box="[805,990,151,176]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="gaudichaudii">
<emphasis id="B90D775B5E7CFF8A0A1C57ADFC552882" box="[805,990,151,176]" italics="true" pageId="9" pageNumber="74">C. gaudichaudii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A0AC857ADFBE52882" box="[1009,1134,151,176]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="timbuhy">
<emphasis id="B90D775B5E7CFF8A0AC857ADFBE52882" box="[1009,1134,151,176]" italics="true" pageId="9" pageNumber="74">C. timbuhy</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A0DB857ADFAB02882" box="[1153,1339,151,176]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="trachystomus">
<emphasis id="B90D775B5E7CFF8A0DB857ADFAB02882" box="[1153,1339,151,176]" italics="true" pageId="9" pageNumber="74">C. trachystomus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C79D0CA5E7CFF8A0CBB57ADFF6528E6" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="werneri">
<emphasis id="B90D775B5E7CFF8A0CBB57ADFF6528E6" italics="true" pageId="9" pageNumber="74">C. werneri</emphasis>
</taxonomicName>
by the undilated toe tips (dilated).
</paragraph>
<paragraph id="8BC6AB495E7CFF8A09FE57EAFF7229BA" blockId="9.[151,1436,151,645]" pageId="9" pageNumber="74">
The developed postrictal gland distinguishes 
<taxonomicName id="4C79D0CA5E7CFF8A0BD357EAFBB828CA" box="[746,1075,223,248]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7CFF8A0BD357EAFBB828CA" box="[746,1075,223,248]" italics="true" pageId="9" pageNumber="74">Crossodactylus franciscanus</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C79D0CA5E7CFF8A0DB157D5FAB728CA" box="[1160,1340,223,248]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="caramaschii">
<emphasis id="B90D775B5E7CFF8A0DB157D5FAB728CA" box="[1160,1340,223,248]" italics="true" pageId="9" pageNumber="74">C. caramaschii</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C79D0CA5E7CFF8A0CBB57D5FF70292F" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="lutzorum">
<emphasis id="B90D775B5E7CFF8A0CBB57D5FF70292F" italics="true" pageId="9" pageNumber="74">C. lutzorum</emphasis>
</taxonomicName>
(a discrete ridge). 
<taxonomicName id="4C79D0CA5E7CFF8A08E05631FC95292E" box="[473,798,260,285]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7CFF8A08E05631FC95292E" box="[473,798,260,285]" italics="true" pageId="9" pageNumber="74">Crossodactylus franciscanus</emphasis>
</taxonomicName>
is different from 
<taxonomicName id="4C79D0CA5E7CFF8A0AD65630FBD2292F" box="[1007,1113,260,285]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="dantei">
<emphasis id="B90D775B5E7CFF8A0AD65630FBD2292F" box="[1007,1113,260,285]" italics="true" pageId="9" pageNumber="74">C. dantei</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C79D0CA5E7CFF8A0DAF5630FA95292F" box="[1174,1310,260,285]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="lutzorum">
<emphasis id="B90D775B5E7CFF8A0DAF5630FA95292F" box="[1174,1310,260,285]" italics="true" pageId="9" pageNumber="74">C. lutzorum</emphasis>
</taxonomicName>
due to the presence of dorsolateral folds (absent), and from these species and 
<taxonomicName id="4C79D0CA5E7CFF8A0AAA561DFB882972" box="[915,1027,296,320]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="aeneus">
<emphasis id="B90D775B5E7CFF8A0AAA561DFB882972" box="[915,1027,296,320]" italics="true" pageId="9" pageNumber="74">C. aeneus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A0D2B561DFB342972" box="[1042,1215,295,320]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="caramaschii">
<emphasis id="B90D775B5E7CFF8A0D2B561DFB342972" box="[1042,1215,295,320]" italics="true" pageId="9" pageNumber="74">C. caramaschii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A0DF7561DFAF82972" box="[1230,1395,295,320]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="cyclospinus">
<emphasis id="B90D775B5E7CFF8A0DF7561DFAF82972" box="[1230,1395,295,320]" italics="true" pageId="9" pageNumber="74">C. cyclospinus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A0CBB561DFEA02957" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="gaudichaudii">
<emphasis id="B90D775B5E7CFF8A0CBB561DFEA02957" italics="true" pageId="9" pageNumber="74">C. gaudichaudii</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C79D0CA5E7CFF8A08505678FE622957" box="[361,489,332,357]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B90D775B5E7CFF8A08505678FE622957" box="[361,489,332,357]" italics="true" pageId="9" pageNumber="74">C. schmidti</emphasis>
</taxonomicName>
by the presence of a narrow glandular crest on the anterior surface of the upper arm (absent).
</paragraph>
<paragraph id="8BC6AB495E7CFF8A09FE56A1FE7D2AB6" blockId="9.[151,1436,151,645]" pageId="9" pageNumber="74">
<taxonomicName id="4C79D0CA5E7CFF8A09FE56A1FD82299E" box="[199,521,404,429]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7CFF8A09FE56A1FD82299E" box="[199,521,404,429]" italics="true" pageId="9" pageNumber="74">Crossodactylus franciscanus</emphasis>
</taxonomicName>
differs from 
<emphasis id="B90D775B5E7CFF8A0BA656A0FAC1299F" box="[671,1354,404,429]" italics="true" pageId="9" pageNumber="74">
<taxonomicName id="4C79D0CA5E7CFF8A0BA656A0FCC1299F" box="[671,842,404,429]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="bokermanni">C. bokermanni</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A0A6C56A0FB8F299F" box="[853,1028,404,429]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="caramaschii">C. caramaschii</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A0D2956A0FB3D299F" box="[1040,1206,404,429]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="cyclospinus">C. cyclospinus</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A0DFA56A0FACD299F" box="[1219,1350,404,429]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="schmidti">C. schmidti</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C79D0CA5E7CFF8A0CBB56A0FEA729E2" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="trachystomus">
<emphasis id="B90D775B5E7CFF8A0CBB56A0FEA729E2" italics="true" pageId="9" pageNumber="74">C. trachystomus</emphasis>
</taxonomicName>
by having a poorly delimited area marbled/dotted of brown over light-brown background from snout to shoulder (a white or cream stripe from the snout to the shoulder; variable in 
<taxonomicName id="4C79D0CA5E7CFF8A0D1456E8FB2C29C6" box="[1069,1191,477,500]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="werneri">
<emphasis id="B90D775B5E7CFF8A0D1456E8FB2C29C6" box="[1069,1191,477,500]" italics="true" pageId="9" pageNumber="74">C. werneri</emphasis>
</taxonomicName>
). The presence of an oblique lateral stripe separates 
<taxonomicName id="4C79D0CA5E7CFF8A08C45535FD3B2A2A" box="[509,688,512,536]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7CFF8A08C45535FD3B2A2A" box="[509,688,512,536]" italics="true" pageId="9" pageNumber="74">C. franciscanus</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C79D0CA5E7CFF8A0BC25535FCEF2A2A" box="[763,868,511,536]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="dantei">
<emphasis id="B90D775B5E7CFF8A0BC25535FCEF2A2A" box="[763,868,511,536]" italics="true" pageId="9" pageNumber="74">C. dantei</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A0A4C5535FC542A2A" box="[885,991,511,536]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B90D775B5E7CFF8A0A4C5535FC542A2A" box="[885,991,511,536]" italics="true" pageId="9" pageNumber="74">C. dispar</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A0AC85535FBFC2A2A" box="[1009,1143,511,536]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="lutzorum">
<emphasis id="B90D775B5E7CFF8A0AC85535FBFC2A2A" box="[1009,1143,511,536]" italics="true" pageId="9" pageNumber="74">C. lutzorum</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C79D0CA5E7CFF8A0D825535FAB22A2A" box="[1211,1337,511,536]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="timbuhy">
<emphasis id="B90D775B5E7CFF8A0D825535FAB22A2A" box="[1211,1337,511,536]" italics="true" pageId="9" pageNumber="74">C. timbuhy</emphasis>
</taxonomicName>
(absent; variable in 
<taxonomicName id="4C79D0CA5E7CFF8A08195510FE4B2A0F" box="[288,448,548,573]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="B90D775B5E7CFF8A08195510FE4B2A0F" box="[288,448,548,573]" italics="true" pageId="9" pageNumber="74">C. boulengeri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C79D0CA5E7CFF8A0B3A5510FD382A0F" box="[515,691,548,573]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="caramaschii">
<emphasis id="B90D775B5E7CFF8A0B3A5510FD382A0F" box="[515,691,548,573]" italics="true" pageId="9" pageNumber="74">C. caramaschii</emphasis>
</taxonomicName>
). The immaculate belly distinguishes 
<taxonomicName id="4C79D0CA5E7CFF8A0D455510FAB92A0E" box="[1148,1330,548,572]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7CFF8A0D455510FAB92A0E" box="[1148,1330,548,572]" italics="true" pageId="9" pageNumber="74">C. franciscanus</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C79D0CA5E7CFF8A0CBB5510FE9B2A52" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="B90D775B5E7CFF8A0CBB5510FE9B2A52" italics="true" pageId="9" pageNumber="74">C. boulengeri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A0819557DFE462A52" box="[288,461,583,608]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="caramaschii">
<emphasis id="B90D775B5E7CFF8A0819557DFE462A52" box="[288,461,583,608]" italics="true" pageId="9" pageNumber="74">C. caramaschii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E7CFF8A08E4557DFD082A52" box="[477,643,583,608]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="cyclospinus">
<emphasis id="B90D775B5E7CFF8A08E4557DFD082A52" box="[477,643,583,608]" italics="true" pageId="9" pageNumber="74">C. cyclospinus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C79D0CA5E7CFF8A0BFD557DFCF72A52" box="[708,892,583,608]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="trachystomus">
<emphasis id="B90D775B5E7CFF8A0BFD557DFCF72A52" box="[708,892,583,608]" italics="true" pageId="9" pageNumber="74">C. trachystomus</emphasis>
</taxonomicName>
(reticulated; with brown scattered blotches and short stripes in 
<taxonomicName id="4C79D0CA5E7CFF8A087A5558FE6D2AB7" box="[323,486,620,645]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="cyclospinus">
<emphasis id="B90D775B5E7CFF8A087A5558FE6D2AB7" box="[323,486,620,645]" italics="true" pageId="9" pageNumber="74">C. cyclospinus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF06FBC15E7CFF8A09AE5164FE942EB4" httpUri="https://zenodo.org/record/288455/files/figure.png" pageId="9" pageNumber="74" targetBox="[164,1423,697,1591]" targetPageId="9">
<paragraph id="8BC6AB495E7CFF8A09AE5164FE942EB4" blockId="9.[151,1436,1617,1670]" pageId="9" pageNumber="74">
<emphasis id="B90D775B5E7CFF8A09AE5164FE852E55" bold="true" box="[151,270,1617,1639]" pageId="9" pageNumber="74">FIGURE 4</emphasis>
. 
<taxonomicName id="4C79D0CA5E7CFF8A08255164FDC92E55" box="[284,578,1617,1639]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="franciscanus" status="sp. nov.">
<emphasis id="B90D775B5E7CFF8A08255164FDC92E55" box="[284,578,1617,1639]" italics="true" pageId="9" pageNumber="74">Crossodactylus franciscanus</emphasis>
</taxonomicName>
<emphasis id="B90D775B5E7CFF8A0B705164FD102E55" bold="true" box="[585,667,1617,1639]" pageId="9" pageNumber="74">
<taxonomicNameLabel id="A23ECA205E7CFF8A0B705164FD102E55" box="[585,667,1617,1639]" pageId="9" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MNRJ 40137, holotype, SVL 22.1 mm), (A) dorsal, (B) ventral, and (C) lateral views.
</paragraph>
</caption>
<subSubSection id="C363F8C25E7CFF8E09FE5198FCAA2D02" lastPageId="13" lastPageNumber="78" pageId="9" pageNumber="74" type="description">
<paragraph id="8BC6AB495E7CFF8A09FE5198FB1D2FAC" blockId="9.[151,1437,1709,2022]" pageId="9" pageNumber="74">
<emphasis id="B90D775B5E7CFF8A09FE5198FD9E2EF4" bold="true" box="[199,533,1709,1734]" pageId="9" pageNumber="74">
Description of the 
<typeStatus id="54C215EB5E7CFF8A089E5198FD9B2EF4" box="[423,528,1709,1734]" pageId="9" pageNumber="74" type="holotype">holotype</typeStatus>
.
</emphasis>
Body slender. Head longer than wide; nostrils prominent, situated and directed dorsolaterally, closer to the tip of snout than to eye. Snout with approx. 30% of HL, rounded in dorsal view, protruding in lateral view (
<figureCitation id="1342B7CC5E7CFF8A08F251C0FD9A2F3C" box="[459,529,1781,1806]" captionStart="FIGURE 5" captionStartId="10.[151,250,1844,1866]" captionTargetBox="[170,1418,383,1804]" captionTargetId="figure@10.[152,1435,373,1823]" captionTargetPageId="10" captionText="FIGURE 5. Crossodactylus franciscanus sp. nov. (MNRJ 40137, holotype, SVL 22.1 mm). Dorsal and lateral views of head and ventral views of hand and foot. Scales = 5 mm." httpUri="https://zenodo.org/record/288456/files/figure.png" pageId="9" pageNumber="74">Fig. 5</figureCitation>
). 
<emphasis id="B90D775B5E7CFF8A0B1151C0FD672F3C" box="[552,748,1781,1806]" italics="true" pageId="9" pageNumber="74">Canthus rostralis</emphasis>
well marked, sharp; loreal region oblique, slightly concave. Eyes approx. 30% of HL. Tympanum distinct, approx. 70% of ED, rounded; supratympanic fold well marked, extending from the posterior corner of the eye to the shoulder. Vocal sac median, subgular. Upper lip spines developed, brown, concentrated on the anterior region. Tongue almost covering the entire mouth floor surface, with a discrete posterior notch. Choanae ovoid, very distant from each other. No vomerine teeth.
</paragraph>
<paragraph id="8BC6AB495E7CFF8909FE509CFC6F2972" blockId="9.[151,1437,1709,2022]" lastBlockId="10.[151,1436,151,320]" lastPageId="10" lastPageNumber="75" pageId="9" pageNumber="74">
Arms slender; forearms thicker than upper arms; fingers narrow, long, with undilated tips and slightly enlarged bases; finger lengths II~IV&lt;I&lt;III (
<figureCitation id="1342B7CC5E7CFF8A0B2550F8FDE92FD4" box="[540,610,1997,2022]" captionStart="FIGURE 5" captionStartId="10.[151,250,1844,1866]" captionTargetBox="[170,1418,383,1804]" captionTargetId="figure@10.[152,1435,373,1823]" captionTargetPageId="10" captionText="FIGURE 5. Crossodactylus franciscanus sp. nov. (MNRJ 40137, holotype, SVL 22.1 mm). Dorsal and lateral views of head and ventral views of hand and foot. Scales = 5 mm." httpUri="https://zenodo.org/record/288456/files/figure.png" pageId="9" pageNumber="74">Fig. 5</figureCitation>
); three developed spines on each thumb, arranged triangularly; the spine on the inner margin of thumb is smaller than the others; the region around their bases also cornified. Scutes poorly developed on upper surfaces of finger tips, more evident on fingers III e IV; small dermal folds with rounded margins on the joints of distal phalanges; fringes on fingers weakly developed. Carpal tubercle rounded, large; thenar tubercle elongated, as long as the diameter of carpal tubercle; subarticular tubercles rounded, protruding, more developed on finger I; supernumerary tubercles small, scarce (
<figureCitation id="1342B7CC5E7FFF890AA85612FC5D2972" box="[913,982,295,320]" captionStart="FIGURE 5" captionStartId="10.[151,250,1844,1866]" captionTargetBox="[170,1418,383,1804]" captionTargetId="figure@10.[152,1435,373,1823]" captionTargetPageId="10" captionText="FIGURE 5. Crossodactylus franciscanus sp. nov. (MNRJ 40137, holotype, SVL 22.1 mm). Dorsal and lateral views of head and ventral views of hand and foot. Scales = 5 mm." httpUri="https://zenodo.org/record/288456/files/figure.png" pageId="10" pageNumber="75">Fig. 5</figureCitation>
).
</paragraph>
<caption id="DF06FBC15E7FFF8909AE5001FD2B2F5B" httpUri="https://zenodo.org/record/288456/files/figure.png" pageId="10" pageNumber="75" targetBox="[170,1418,383,1804]" targetPageId="10">
<paragraph id="8BC6AB495E7FFF8909AE5001FD2B2F5B" blockId="10.[151,1436,1844,1897]" pageId="10" pageNumber="75">
<emphasis id="B90D775B5E7FFF8909AE5001FE852F78" bold="true" box="[151,270,1844,1866]" pageId="10" pageNumber="75">FIGURE 5</emphasis>
. 
<taxonomicName id="4C79D0CA5E7FFF8908225001FDB42F78" box="[283,575,1844,1866]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="75" phylum="Chordata" rank="species" species="franciscanus" status="sp. nov.">
<emphasis id="B90D775B5E7FFF8908225001FDB42F78" box="[283,575,1844,1866]" italics="true" pageId="10" pageNumber="75">Crossodactylus franciscanus</emphasis>
</taxonomicName>
<emphasis id="B90D775B5E7FFF890B7F5001FD132F78" bold="true" box="[582,664,1844,1866]" pageId="10" pageNumber="75">
<taxonomicNameLabel id="A23ECA205E7FFF890B7F5001FD132F78" box="[582,664,1844,1866]" pageId="10" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MNRJ 40137, holotype, SVL 22.1 mm). Dorsal and lateral views of head and ventral views of hand and foot. Scales = 5 mm.
</paragraph>
</caption>
<paragraph id="8BC6AB495E7FFF8809FE50A4FC87292F" blockId="10.[151,1436,1936,2033]" lastBlockId="11.[151,1437,151,968]" lastPageId="11" lastPageNumber="76" pageId="10" pageNumber="75">
Legs slender; the sum of tibia, thigh, and foot lengths 1.8 times the SVL; toes slender, extensively fringed, with truncate, undilated tips; toe lengths I&lt;II&lt;V&lt;III&lt;IV. Scutes on upper surfaces of toe tips more developed than on fingers, less evident on toe I; small dermal folds with rounded margins on the joints of distal phalanges. Inner metatarsal tubercle elongated, protruding; outer metatarsal tubercle small, rounded, protruding; subarticular tubercles rounded, protruding; no supernumerary tubercles. Fringes joined at base; tarsal fringe very developed, continuous distally with outer fringe of toe I, almost reaching the joint with the tibia; outer fringe of toe V reaches the posterior margin of the basal tubercle of toe (
<figureCitation id="1342B7CC5E7EFF880B805631FD75292F" box="[697,766,260,285]" captionStart="FIGURE 5" captionStartId="10.[151,250,1844,1866]" captionTargetBox="[170,1418,383,1804]" captionTargetId="figure@10.[152,1435,373,1823]" captionTargetPageId="10" captionText="FIGURE 5. Crossodactylus franciscanus sp. nov. (MNRJ 40137, holotype, SVL 22.1 mm). Dorsal and lateral views of head and ventral views of hand and foot. Scales = 5 mm." httpUri="https://zenodo.org/record/288456/files/figure.png" pageId="11" pageNumber="76">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BC6AB495E7EFF8809FE5612FDDC2A2A" blockId="11.[151,1437,151,968]" pageId="11" pageNumber="76">Dorsal skin posteriorly granular; discrete vertebral glandular ridge from interorbital region to the vent, interrupted on sacral region; dorsolateral folds from the posterior corner of eyes to the inguinal region, fragmented on a row of granules from midbody to groin; a pair of developed oblique glandular ridges between the dorsolateral folds and the vertebral ridge, from upper eyelids toward midbody. Ventral surfaces smooth. Upper eyelids and legs with small granules; flanks with sparse large granules; cloacal region with a few large granules. Postrictal tubercle elongated, continuous, touching the posterior margin of tympanum. A thin glandular crest on the anterior surface of upper arm, from arm insertion to elbow.</paragraph>
<paragraph id="8BC6AB495E7EFF8809FE5511FA0D2AC2" blockId="11.[151,1437,151,968]" pageId="11" pageNumber="76">Color in preservative (70% ethanol) uniformly light brown. Poorly delimited area marbled/dotted of brown over light brown background from snout to shoulder; maxillary region brown; tympanum, postrictal tubercle, and oblique lateral stripe cream. Two poorly evident brown transversal bars on forearm. Three brown transversal bars on thigh, three on tibia, and four on tarsus-foot. Ventral surfaces cream; small brown blotches on gular region, chest and anterior region of belly; narrow longitudinal dark brown line on chest. Ventral surfaces of hands and feet brown with cream tubercles; fringes cream but transparent, with small brown dots. Several scratches on dorsal surfaces.</paragraph>
<paragraph id="8BC6AB495E7EFF8809FE55C8FC932B0A" blockId="11.[151,1437,151,968]" pageId="11" pageNumber="76">
Measurements of 
<typeStatus id="54C215EB5E7EFF8808A855C9FE732B27" box="[401,504,764,789]" pageId="11" pageNumber="76" type="holotype">holotype</typeStatus>
: SVL = 22.1; HL = 8.6; HW = 8.0; TL = 10.6; THL = 11.6; FL = 17.9; TD = 1.8; ED = 2.6; 
<collectingCountry id="F36EEBD95E7EFF8808355415FECD2B05" box="[268,326,800,823]" name="India" pageId="11" pageNumber="76">END</collectingCountry>
= 1.7; NSD = 0.9; 
<collectingCountry id="F36EEBD95E7EFF880B245415FDC42B05" box="[541,591,800,823]" name="India" pageId="11" pageNumber="76">IND</collectingCountry>
= 2.9; IOD = 2.4.
</paragraph>
<paragraph id="8BC6AB495E7EFF8809FE5471FA862BFA" blockId="11.[151,1437,151,968]" pageId="11" pageNumber="76">
<emphasis id="B90D775B5E7EFF8809FE5471FEB02B6F" bold="true" box="[199,315,836,861]" pageId="11" pageNumber="76">Variation.</emphasis>
Toe and tarsal fringes are broadly developed in males and reduced in females, and forearms are thicker in males than in females. Upper lip spines are absent in females. Some specimens present a cream vertebral line from snout to vent. The number of transversal bars on legs varies from three to five. 
<typeStatus id="54C215EB5E7EFF880DEB54B8FAB42B97" box="[1234,1343,909,933]" pageId="11" pageNumber="76" type="paratype">Paratypes</typeStatus>
present reticulated gular region and chest, but immaculate belly. Morphometric variation is shown on 
<tableCitation id="C6FB9EF25E7EFF880D8A549AFA822BFA" box="[1203,1289,943,968]" captionStart="TABLE 2" captionStartId="11.[151,239,1012,1035]" captionTargetBox="[155,1234,1136,1654]" captionTargetPageId="11" captionText="TABLE 2. Means, standard deviations (SD), and ranges of some measurements (in mm) of Crossodactylus franciscanus sp. nov. (N = number of specimens)." httpUri="http://table.plazi.org/id/DF06FBC15E7EFF8809AE54C1FD952C19" pageId="11" pageNumber="76" tableUuid="DF06FBC15E7EFF8809AE54C1FD952C19">Table 2</tableCitation>
.
</paragraph>
<caption id="DF06FBC15E7EFF8809AE54C1FD952C19" ID-Table-UUID="DF06FBC15E7EFF8809AE54C1FD952C19" httpUri="http://table.plazi.org/id/DF06FBC15E7EFF8809AE54C1FD952C19" pageId="11" pageNumber="76" targetBox="[155,1234,1136,1654]" targetIsTable="true" targetPageId="11">
<paragraph id="8BC6AB495E7EFF8809AE54C1FD952C19" blockId="11.[151,1435,1011,1067]" pageId="11" pageNumber="76">
<emphasis id="B90D775B5E7EFF8809AE54C1FE822C39" bold="true" box="[151,265,1012,1035]" pageId="11" pageNumber="76">TABLE 2.</emphasis>
Means, standard deviations (SD), and ranges of some measurements (in mm) of 
<taxonomicName id="4C79D0CA5E7EFF880D5354C6FA102C39" box="[1130,1435,1011,1035]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="76" phylum="Chordata" rank="species" species="franciscanus" status="sp. nov.">
<emphasis id="B90D775B5E7EFF880D5354C6FA102C39" box="[1130,1435,1011,1035]" italics="true" pageId="11" pageNumber="76">Crossodactylus franciscanus</emphasis>
</taxonomicName>
<emphasis id="B90D775B5E7EFF8809AE5321FF662C19" bold="true" box="[151,237,1044,1067]" pageId="11" pageNumber="76">
<taxonomicNameLabel id="A23ECA205E7EFF8809AE5321FF662C19" box="[151,237,1044,1067]" pageId="11" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<emphasis id="B90D775B5E7EFF8809C55321FE852C19" box="[252,270,1044,1067]" italics="true" pageId="11" pageNumber="76">N</emphasis>
= number of specimens).
</paragraph>
</caption>
<paragraph id="8BC6AB495E7EFF8808435373FE852FDD" blockId="11.[378,1234,1094,1117]" lastBlockId="11.[151,1437,1718,2031]" pageId="11" pageNumber="76">
Males (
<emphasis id="B90D775B5E7EFF8808FF537DFE5C2C6F" box="[454,471,1096,1117]" italics="true" pageId="11" pageNumber="76">N</emphasis>
= 7) Females (
<emphasis id="B90D775B5E7EFF880DAA537DFB2F2C6F" box="[1171,1188,1096,1117]" italics="true" pageId="11" pageNumber="76">N</emphasis>
= 2) 
<emphasis id="B90D775B5E7EFF8809FE5183FEEA2EFD" bold="true" box="[199,353,1718,1743]" pageId="11" pageNumber="76">Vocalization.</emphasis>
Some parameters of the advertisement call of 
<taxonomicName id="4C79D0CA5E7EFF880ABB5183FB4D2EFD" box="[898,1222,1718,1743]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="76" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E7EFF880ABB5183FB4D2EFD" box="[898,1222,1718,1743]" italics="true" pageId="11" pageNumber="76">Crossodactylus franciscanus</emphasis>
</taxonomicName>
were described in 
<bibRefCitation id="EFE8D6B85E7EFF8809AE51EFFEF32EC1" author="Haddad" box="[151,376,1754,1779]" pageId="11" pageNumber="76" refString="Haddad, C. F. B., Andrade, G. V. &amp; Cardoso, A. J. (1988) Anfibios anuros do Parque Nacional da Serra da Canastra, Estado de Minas Gerais. Brasil Florestal, 64, 9 - 20." type="journal article" year="1988">
Haddad 
<emphasis id="B90D775B5E7EFF8809CC51E9FEA62EC1" box="[245,301,1754,1779]" italics="true" pageId="11" pageNumber="76">et al.</emphasis>
(1988
</bibRefCitation>
; referred as 
<taxonomicName id="4C79D0CA5E7EFF880B3B51EEFD502EC1" box="[514,731,1754,1779]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="76" phylum="Chordata" rank="species" species="trachystomus">
<emphasis id="B90D775B5E7EFF880B3B51EEFD972EC0" box="[514,540,1755,1778]" italics="true" pageId="11" pageNumber="76">C.</emphasis>
cf. 
<emphasis id="B90D775B5E7EFF880B7F51EFFD502EC1" box="[582,731,1754,1779]" italics="true" pageId="11" pageNumber="76">trachystomus</emphasis>
</taxonomicName>
) based on the recording of one male, but a better refinement is necessary in order to allow comparisons with calls known for the genus. The re-analysis of the recording AJC 010/ 1, the same used by 
<bibRefCitation id="EFE8D6B85E7EFF8808B75017FD082F09" box="[398,643,1826,1851]" pageId="11" pageNumber="76" refString="Haddad, C. F. B., Andrade, G. V. &amp; Cardoso, A. J. (1988) Anfibios anuros do Parque Nacional da Serra da Canastra, Estado de Minas Gerais. Brasil Florestal, 64, 9 - 20." type="journal article">
Haddad 
<emphasis id="B90D775B5E7EFF8808C95011FDA72F09" box="[496,556,1826,1851]" italics="true" pageId="11" pageNumber="76">et al.</emphasis>
(1988)
</bibRefCitation>
, shows an advertisement call composed of harmonic notes and less intense initial notes followed by high intensity notes, subsequently decreasing on last notes (
<figureCitation id="1342B7CC5E7EFF880D975073FB782F6D" box="[1198,1267,1862,1887]" captionStart="FIGURE 6" captionStartId="12.[151,250,607,629]" captionTargetBox="[158,1436,212,571]" captionTargetId="figure@12.[158,1443,193,579]" captionTargetPageId="12" captionText="FIGURE 6. (A) Waveform and (B) audiospectrogram of a section of the advertisement call of Crossodactylus franciscanus (recording AJC 010 / 1) showing an interrupted call (arrow) followed by additional low-intensity notes. Recorded at the type locality in 28 January 1981 at 02: 00 PM. Water temperature 20 ºC; substrate temperature 25 ºC. Unreferred specimen (see Remarks)." httpUri="https://zenodo.org/record/288457/files/figure.png" pageId="11" pageNumber="76">Fig. 6</figureCitation>
). Final portion of the call (i.e. the portion after peak intensity) contains most of the notes in some of the calls analyzed. Some calls seem to have been altered for unknown reasons (maybe the presence of the observer). We used only regular calls for description of the advertisement call and comparisons with other species whereas altered calls were analyzed separately.
</paragraph>
<paragraph id="8BC6AB495E7EFF8808435344FBF02E44" pageId="11" pageNumber="76">
<table id="F97959E95E7E007C09A25345FB592E44" box="[155,1234,1136,1654]" gridcols="5" gridrows="13" pageId="11" pageNumber="76">
<tr id="3549A90B5E7E007C09A25345FB592CB4" box="[155,1234,1136,1158]" gridrow="0" pageId="11" pageNumber="76" rowspan-0="1">
<th id="7698C0775E7E007C08435345FE3F2CB4" box="[378,436,1136,1158]" gridcol="1" gridrow="0" pageId="11" pageNumber="76">Mean</th>
<th id="7698C0775E7E007C0B685345FDF62CB4" box="[593,637,1136,1158]" gridcol="2" gridrow="0" pageId="11" pageNumber="76">SD</th>
<th id="7698C0775E7E007C0BC75345FCEF2CB4" box="[766,868,1136,1158]" gridcol="3" gridrow="0" pageId="11" pageNumber="76">Range</th>
<th id="7698C0775E7E007C0D165345FB592CB4" box="[1071,1234,1136,1158]" gridcol="4" gridrow="0" pageId="11" pageNumber="76">Range</th>
</tr>
<tr id="3549A90B5E7E007C09A253ACFB592C9D" box="[155,1234,1177,1199]" gridrow="1" pageId="11" pageNumber="76">
<th id="7698C0775E7E007C09A253ACFF442C9D" box="[155,207,1177,1199]" gridcol="0" gridrow="1" pageId="11" pageNumber="76">SVL</th>
<td id="7698C0775E7E007C084353ACFE3F2C9D" box="[378,436,1177,1199]" gridcol="1" gridrow="1" pageId="11" pageNumber="76">21.3</td>
<td id="7698C0775E7E007C0B6853ACFDF62C9D" box="[593,637,1177,1199]" gridcol="2" gridrow="1" pageId="11" pageNumber="76">0.61</td>
<td id="7698C0775E7E007C0BC753ACFCEF2C9D" box="[766,868,1177,1199]" gridcol="3" gridrow="1" pageId="11" pageNumber="76">20.4–22.1</td>
<td id="7698C0775E7E007C0D1653ACFB592C9D" box="[1071,1234,1177,1199]" gridcol="4" gridrow="1" pageId="11" pageNumber="76">20.3–23.6</td>
</tr>
<tr id="3549A90B5E7E007C09A253F7FB592CEA" box="[155,1234,1218,1240]" gridrow="2" pageId="11" pageNumber="76">
<th id="7698C0775E7E007C09A253F7FF442CEA" box="[155,207,1218,1240]" gridcol="0" gridrow="2" pageId="11" pageNumber="76">HL</th>
<td id="7698C0775E7E007C084353F7FE3F2CEA" box="[378,436,1218,1240]" gridcol="1" gridrow="2" pageId="11" pageNumber="76">7.9</td>
<td id="7698C0775E7E007C0B6853F7FDF62CEA" box="[593,637,1218,1240]" gridcol="2" gridrow="2" pageId="11" pageNumber="76">0.36</td>
<td id="7698C0775E7E007C0BC753F7FCEF2CEA" box="[766,868,1218,1240]" gridcol="3" gridrow="2" pageId="11" pageNumber="76">7.6–8.6</td>
<td id="7698C0775E7E007C0D1653F7FB592CEA" box="[1071,1234,1218,1240]" gridcol="4" gridrow="2" pageId="11" pageNumber="76">6.9–8.4</td>
</tr>
<tr id="3549A90B5E7E007C09A253D9FB592D30" box="[155,1234,1260,1282]" gridrow="3" pageId="11" pageNumber="76">
<th id="7698C0775E7E007C09A253D9FF442D30" box="[155,207,1260,1282]" gridcol="0" gridrow="3" pageId="11" pageNumber="76">HW</th>
<td id="7698C0775E7E007C084353D9FE3F2D30" box="[378,436,1260,1282]" gridcol="1" gridrow="3" pageId="11" pageNumber="76">7.3</td>
<td id="7698C0775E7E007C0B6853D9FDF62D30" box="[593,637,1260,1282]" gridcol="2" gridrow="3" pageId="11" pageNumber="76">0.35</td>
<td id="7698C0775E7E007C0BC753D9FCEF2D30" box="[766,868,1260,1282]" gridcol="3" gridrow="3" pageId="11" pageNumber="76">6.9–8.0</td>
<td id="7698C0775E7E007C0D1653D9FB592D30" box="[1071,1234,1260,1282]" gridcol="4" gridrow="3" pageId="11" pageNumber="76">6.4–7.3</td>
</tr>
<tr id="3549A90B5E7E007C09A25220FB592D19" box="[155,1234,1301,1323]" gridrow="4" pageId="11" pageNumber="76">
<th id="7698C0775E7E007C09A25220FF442D19" box="[155,207,1301,1323]" gridcol="0" gridrow="4" pageId="11" pageNumber="76">TL</th>
<td id="7698C0775E7E007C08435220FE3F2D19" box="[378,436,1301,1323]" gridcol="1" gridrow="4" pageId="11" pageNumber="76">10.2</td>
<td id="7698C0775E7E007C0B685220FDF62D19" box="[593,637,1301,1323]" gridcol="2" gridrow="4" pageId="11" pageNumber="76">0.53</td>
<td id="7698C0775E7E007C0BC75220FCEF2D19" box="[766,868,1301,1323]" gridcol="3" gridrow="4" pageId="11" pageNumber="76">9.4–10.8</td>
<td id="7698C0775E7E007C0D165220FB592D19" box="[1071,1234,1301,1323]" gridcol="4" gridrow="4" pageId="11" pageNumber="76">9.6–11.1</td>
</tr>
<tr id="3549A90B5E7E007C09A2520BFB592D66" box="[155,1234,1342,1364]" gridrow="5" pageId="11" pageNumber="76">
<th id="7698C0775E7E007C09A2520BFF442D66" box="[155,207,1342,1364]" gridcol="0" gridrow="5" pageId="11" pageNumber="76">THL</th>
<td id="7698C0775E7E007C0843520BFE3F2D66" box="[378,436,1342,1364]" gridcol="1" gridrow="5" pageId="11" pageNumber="76">10.3</td>
<td id="7698C0775E7E007C0B68520BFDF62D66" box="[593,637,1342,1364]" gridcol="2" gridrow="5" pageId="11" pageNumber="76">1.10</td>
<td id="7698C0775E7E007C0BC7520BFCEF2D66" box="[766,868,1342,1364]" gridcol="3" gridrow="5" pageId="11" pageNumber="76">8.4–11.6</td>
<td id="7698C0775E7E007C0D16520BFB592D66" box="[1071,1234,1342,1364]" gridcol="4" gridrow="5" pageId="11" pageNumber="76">10.3–11.3</td>
</tr>
<tr id="3549A90B5E7E007C09A2525DFB592D4C" box="[155,1234,1384,1406]" gridrow="6" pageId="11" pageNumber="76">
<th id="7698C0775E7E007C09A2525DFF442D4C" box="[155,207,1384,1406]" gridcol="0" gridrow="6" pageId="11" pageNumber="76">FL</th>
<td id="7698C0775E7E007C0843525DFE3F2D4C" box="[378,436,1384,1406]" gridcol="1" gridrow="6" pageId="11" pageNumber="76">16.4</td>
<td id="7698C0775E7E007C0B68525DFDF62D4C" box="[593,637,1384,1406]" gridcol="2" gridrow="6" pageId="11" pageNumber="76">0.98</td>
<td id="7698C0775E7E007C0BC7525DFCEF2D4C" box="[766,868,1384,1406]" gridcol="3" gridrow="6" pageId="11" pageNumber="76">15.3–17.9</td>
<td id="7698C0775E7E007C0D16525DFB592D4C" box="[1071,1234,1384,1406]" gridcol="4" gridrow="6" pageId="11" pageNumber="76">14.2–17.9</td>
</tr>
<tr id="3549A90B5E7E007C09A252A4FB592D95" box="[155,1234,1425,1447]" gridrow="7" pageId="11" pageNumber="76">
<th id="7698C0775E7E007C09A252A4FF442D95" box="[155,207,1425,1447]" gridcol="0" gridrow="7" pageId="11" pageNumber="76">TD</th>
<td id="7698C0775E7E007C084352A4FE3F2D95" box="[378,436,1425,1447]" gridcol="1" gridrow="7" pageId="11" pageNumber="76">1.5</td>
<td id="7698C0775E7E007C0B6852A4FDF62D95" box="[593,637,1425,1447]" gridcol="2" gridrow="7" pageId="11" pageNumber="76">0.28</td>
<td id="7698C0775E7E007C0BC752A4FCEF2D95" box="[766,868,1425,1447]" gridcol="3" gridrow="7" pageId="11" pageNumber="76">1.2–1.8</td>
<td id="7698C0775E7E007C0D1652A4FB592D95" box="[1071,1234,1425,1447]" gridcol="4" gridrow="7" pageId="11" pageNumber="76">1.3–1.6</td>
</tr>
<tr id="3549A90B5E7E007C09A2528FFB592DE2" box="[155,1234,1466,1488]" gridrow="8" pageId="11" pageNumber="76">
<th id="7698C0775E7E007C09A2528FFF442DE2" box="[155,207,1466,1488]" gridcol="0" gridrow="8" pageId="11" pageNumber="76">ED</th>
<td id="7698C0775E7E007C0843528FFE3F2DE2" box="[378,436,1466,1488]" gridcol="1" gridrow="8" pageId="11" pageNumber="76">2.8</td>
<td id="7698C0775E7E007C0B68528FFDF62DE2" box="[593,637,1466,1488]" gridcol="2" gridrow="8" pageId="11" pageNumber="76">0.24</td>
<td id="7698C0775E7E007C0BC7528FFCEF2DE2" box="[766,868,1466,1488]" gridcol="3" gridrow="8" pageId="11" pageNumber="76">2.5–3.1</td>
<td id="7698C0775E7E007C0D16528FFB592DE2" box="[1071,1234,1466,1488]" gridcol="4" gridrow="8" pageId="11" pageNumber="76">2.9–2.9</td>
</tr>
<tr id="3549A90B5E7E007C09A252D1FB592DC8" box="[155,1234,1508,1530]" gridrow="9" pageId="11" pageNumber="76">
<th id="7698C0775E7E007C09A252D1FF442DC8" box="[155,207,1508,1530]" gridcol="0" gridrow="9" pageId="11" pageNumber="76">END</th>
<td id="7698C0775E7E007C084352D1FE3F2DC8" box="[378,436,1508,1530]" gridcol="1" gridrow="9" pageId="11" pageNumber="76">1.6</td>
<td id="7698C0775E7E007C0B6852D1FDF62DC8" box="[593,637,1508,1530]" gridcol="2" gridrow="9" pageId="11" pageNumber="76">0.18</td>
<td id="7698C0775E7E007C0BC752D1FCEF2DC8" box="[766,868,1508,1530]" gridcol="3" gridrow="9" pageId="11" pageNumber="76">1.4–2.0</td>
<td id="7698C0775E7E007C0D1652D1FB592DC8" box="[1071,1234,1508,1530]" gridcol="4" gridrow="9" pageId="11" pageNumber="76">1.6–2.1</td>
</tr>
<tr id="3549A90B5E7E007C09A25138FB592E11" box="[155,1234,1549,1571]" gridrow="10" pageId="11" pageNumber="76">
<th id="7698C0775E7E007C09A25138FF442E11" box="[155,207,1549,1571]" gridcol="0" gridrow="10" pageId="11" pageNumber="76">NSD</th>
<td id="7698C0775E7E007C08435138FE3F2E11" box="[378,436,1549,1571]" gridcol="1" gridrow="10" pageId="11" pageNumber="76">0.9</td>
<td id="7698C0775E7E007C0B685138FDF62E11" box="[593,637,1549,1571]" gridcol="2" gridrow="10" pageId="11" pageNumber="76">0.10</td>
<td id="7698C0775E7E007C0BC75138FCEF2E11" box="[766,868,1549,1571]" gridcol="3" gridrow="10" pageId="11" pageNumber="76">0.8–1.0</td>
<td id="7698C0775E7E007C0D165138FB592E11" box="[1071,1234,1549,1571]" gridcol="4" gridrow="10" pageId="11" pageNumber="76">1.0–1.2</td>
</tr>
<tr id="3549A90B5E7E007C09A25103FB592E7E" box="[155,1234,1590,1612]" gridrow="11" pageId="11" pageNumber="76">
<th id="7698C0775E7E007C09A25103FF442E7E" box="[155,207,1590,1612]" gridcol="0" gridrow="11" pageId="11" pageNumber="76">IND</th>
<td id="7698C0775E7E007C08435103FE3F2E7E" box="[378,436,1590,1612]" gridcol="1" gridrow="11" pageId="11" pageNumber="76">2.5</td>
<td id="7698C0775E7E007C0B685103FDF62E7E" box="[593,637,1590,1612]" gridcol="2" gridrow="11" pageId="11" pageNumber="76">0.22</td>
<td id="7698C0775E7E007C0BC75103FCEF2E7E" box="[766,868,1590,1612]" gridcol="3" gridrow="11" pageId="11" pageNumber="76">2.2–2.9</td>
<td id="7698C0775E7E007C0D165103FB592E7E" box="[1071,1234,1590,1612]" gridcol="4" gridrow="11" pageId="11" pageNumber="76">2.3–2.5</td>
</tr>
<tr id="3549A90B5E7E007C09A25155FB592E44" box="[155,1234,1632,1654]" gridrow="12" pageId="11" pageNumber="76">
<th id="7698C0775E7E007C09A25155FF442E44" box="[155,207,1632,1654]" gridcol="0" gridrow="12" pageId="11" pageNumber="76">IOD</th>
<td id="7698C0775E7E007C08435155FE3F2E44" box="[378,436,1632,1654]" gridcol="1" gridrow="12" pageId="11" pageNumber="76">2.1</td>
<td id="7698C0775E7E007C0B685155FDF62E44" box="[593,637,1632,1654]" gridcol="2" gridrow="12" pageId="11" pageNumber="76">0.27</td>
<td id="7698C0775E7E007C0BC75155FCEF2E44" box="[766,868,1632,1654]" gridcol="3" gridrow="12" pageId="11" pageNumber="76">1.7–2.4</td>
<td id="7698C0775E7E007C0D165155FB592E44" box="[1071,1234,1632,1654]" gridcol="4" gridrow="12" pageId="11" pageNumber="76">1.7–2.1</td>
</tr>
</table>
</paragraph>
<caption id="DF06FBC15E79FF8F09AE556AFF752AE3" httpUri="https://zenodo.org/record/288457/files/figure.png" pageId="12" pageNumber="77" targetBox="[158,1436,212,571]" targetPageId="12">
<paragraph id="8BC6AB495E79FF8F09AE556AFF752AE3" blockId="12.[151,1436,607,721]" pageId="12" pageNumber="77">
<emphasis id="B90D775B5E79FF8F09AE556AFE842A47" bold="true" box="[151,271,607,629]" pageId="12" pageNumber="77">FIGURE 6</emphasis>
. (A) Waveform and (B) audiospectrogram of a section of the advertisement call of 
<taxonomicName id="4C79D0CA5E79FF8F0D4E556AFA172A47" box="[1143,1436,607,629]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E79FF8F0D4E556AFA172A47" box="[1143,1436,607,629]" italics="true" pageId="12" pageNumber="77">Crossodactylus franciscanus</emphasis>
</taxonomicName>
(recording AJC 010/1) showing an interrupted call (arrow) followed by additional low-intensity notes. Recorded at the type locality in 28 January 1981 at 02:00 PM. Water temperature 20ºC; substrate temperature 25ºC. Unreferred specimen (see 
<emphasis id="B90D775B5E79FF8F09AE558EFF7B2AE3" box="[151,240,699,721]" italics="true" pageId="12" pageNumber="77">Remarks</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BC6AB495E79FF8F09FE55CAFB592C21" blockId="12.[151,1437,765,1043]" pageId="12" pageNumber="77">
Mean advertisement call duration (
<emphasis id="B90D775B5E79FF8F0B6D55CAFDEC2B24" box="[596,615,767,790]" italics="true" pageId="12" pageNumber="77">N</emphasis>
= 6 calls) is 
<geoCoordinate id="EE4DCD8E5E79FF8F0BC455C8FCCA2B24" box="[765,833,765,790]" direction="south" orientation="latitude" pageId="12" pageNumber="77" precision="555" value="-6.42">6.42 s</geoCoordinate>
and mean number of notes per call is 48. Mean note duration (
<emphasis id="B90D775B5E79FF8F083C5416FE932B08" box="[261,280,803,826]" italics="true" pageId="12" pageNumber="77">N</emphasis>
= 284 notes) is 12 ms and mean internote interval is 123 ms. Mean dominant frequency is 3460.1 Hz and is always situated on the second harmonic (
<figureCitation id="1342B7CC5E79FF8F0B865470FC8E2B6C" box="[703,773,837,862]" captionStart="FIGURE 6" captionStartId="12.[151,250,607,629]" captionTargetBox="[158,1436,212,571]" captionTargetId="figure@12.[158,1443,193,579]" captionTargetPageId="12" captionText="FIGURE 6. (A) Waveform and (B) audiospectrogram of a section of the advertisement call of Crossodactylus franciscanus (recording AJC 010 / 1) showing an interrupted call (arrow) followed by additional low-intensity notes. Recorded at the type locality in 28 January 1981 at 02: 00 PM. Water temperature 20 ºC; substrate temperature 25 ºC. Unreferred specimen (see Remarks)." httpUri="https://zenodo.org/record/288457/files/figure.png" pageId="12" pageNumber="77">Fig. 6</figureCitation>
). 
<tableCitation id="C6FB9EF25E79FF8F0A245470FCF92B6C" box="[797,882,837,862]" captionStart="TABLE 3" captionStartId="12.[151,239,1090,1113]" captionTargetBox="[151,1288,1172,1613]" captionTargetPageId="12" captionText="TABLE 3. Comparison of acoustic parameters of known advertisement calls of Crossodactylus. Data presented as mean ± SD; range (N, number analyzed) when available. * = data not available." httpUri="http://table.plazi.org/id/DF06FBC15E79FF8F09AE5377FC2C2C4B" pageId="12" pageNumber="77" tableUuid="DF06FBC15E79FF8F09AE5377FC2C2C4B">Table 3</tableCitation>
presents standard deviations and ranges of each parameter analyzed and comparative data on acoustic parameters of the advertisement calls known for 
<taxonomicName id="4C79D0CA5E79FF8F09AE54B8FEC82B94" box="[151,323,909,934]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="B90D775B5E79FF8F09AE54B8FEC82B94" box="[151,323,909,934]" italics="true" pageId="12" pageNumber="77">Crossodactylus</emphasis>
</taxonomicName>
: 
<emphasis id="B90D775B5E79FF8F086554BBFD502B94" box="[348,731,909,934]" italics="true" pageId="12" pageNumber="77">
<taxonomicName id="4C79D0CA5E79FF8F086554BBFD9E2B94" box="[348,533,909,934]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="caramaschii">C. caramaschii</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E79FF8F0B1554BBFD502B94" box="[556,731,909,934]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="cyclospinus">C. cyclospinus</taxonomicName>
</emphasis>
, 
<taxonomicName id="4C79D0CA5E79FF8F0BCC54BBFC3B2B94" box="[757,944,910,934]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E79FF8F0BCC54BBFC3B2B94" box="[757,944,910,934]" italics="true" pageId="12" pageNumber="77">C. franciscanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E79FF8F0AF354BBFB012B94" box="[970,1162,909,934]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="gaudichaudii">
<emphasis id="B90D775B5E79FF8F0AF354BBFB012B94" box="[970,1162,909,934]" italics="true" pageId="12" pageNumber="77">C. gaudichaudii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E79FF8F0D9A54BBFAA52B94" box="[1187,1326,909,934]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B90D775B5E79FF8F0D9A54BBFAA52B94" box="[1187,1326,909,934]" italics="true" pageId="12" pageNumber="77">C. schmidti</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C79D0CA5E79FF8F0CBB54BBFEA72BF9" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="trachystomus">
<emphasis id="B90D775B5E79FF8F0CBB54BBFEA72BF9" italics="true" pageId="12" pageNumber="77">C. trachystomus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE8D6B85E79FF8F08035487FD402BF8" author="Weygoldt" box="[314,715,946,971]" pageId="12" pageNumber="77" refString="Weygoldt, P. &amp; Carvalho-e-Silva, S. P. (1992) Mating and oviposition in the hylodine frog Crossodactylus gaudichaudii (Anura: Leptodactylidae). Amphibia-Reptilia, 13, 35 - 45. http: // dx. doi. org / 10.1163 / 156853892 X 00210" type="journal article" year="1992">Weygoldt &amp; Carvalho-e-Silva 1992</bibRefCitation>
; 
<bibRefCitation id="EFE8D6B85E79FF8F0BEF5486FC502BF9" author="Bastos" box="[726,987,946,971]" pageId="12" pageNumber="77" refString="Bastos, R. P. &amp; Pombal Jr., J. P. (1995) New species of Crossodactylus (Anura: Leptodactylidae) from the Atlantic Rain Forest of southeastern Brazil. Copeia, 1995, 436 - 439. http: // dx. doi. org / 10.2307 / 1446907" type="journal article" year="1995">Bastos &amp; Pombal 1995</bibRefCitation>
; 
<bibRefCitation id="EFE8D6B85E79FF8F0ADF5487FB732BF9" author="Nascimento" box="[998,1272,946,971]" pageId="12" pageNumber="77" refString="Nascimento, L. B., Cruz, C. A. G. &amp; Feio, R. N. (2005 a) A new species of diurnal frog in the genus Crossodactylus Dumeril and Bibron, 1841 (Anura, Leptodactylidae) from Southeastern Brazil. Amphibia-Reptilia, 26, 497 - 505." type="journal article" year="2005" yearSuffix="a">
Nascimento 
<emphasis id="B90D775B5E79FF8F0D4B5486FB212BF9" box="[1138,1194,946,971]" italics="true" pageId="12" pageNumber="77">et al.</emphasis>
2005a
</bibRefCitation>
; 
<bibRefCitation id="EFE8D6B85E79FF8F0C3B5487FF582BDC" author="Pimenta" pageId="12" pageNumber="77" refString="Pimenta, B. V. S., Wachlevski, M. &amp; Cruz, C. A. G. (2008) Morphological and Acoustical Variation, Geographic Distribution, and Conservation Status of the Spinythumb Frog Crossodactylus bokermanni Caramaschi and Sazima, 1985 (Anura, Hylodidae). Journal of Herpetology, 42, 481 - 492. http: // dx. doi. org / 10.1670 / 07 - 164.1" type="journal article" year="2008">
Pimenta 
<emphasis id="B90D775B5E79FF8F0C5D5486FA172BF9" box="[1380,1436,946,971]" italics="true" pageId="12" pageNumber="77">et al.</emphasis>
2008
</bibRefCitation>
; Caldart 
<emphasis id="B90D775B5E79FF8F087054E2FE012BDC" box="[329,394,981,1006]" italics="true" pageId="12" pageNumber="77">et al.</emphasis>
2011; present study). The advertisement call of 
<taxonomicName id="4C79D0CA5E79FF8F0AD454E3FBFB2BDC" box="[1005,1136,981,1006]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="timbuhy">
<emphasis id="B90D775B5E79FF8F0AD454E3FBFB2BDC" box="[1005,1136,981,1006]" italics="true" pageId="12" pageNumber="77">C. timbuhy</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE8D6B85E79FF8F0DB154E0FE3E2C21" author="Weygoldt" pageId="12" pageNumber="77" refString="Weygoldt, P. (1986) Beobachtungen zur Okologie und Biologie von Froschen an einem neotropischen Bergbach. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 113, 429 - 454." type="journal article" year="1986">
Weygoldt, 1986; under 
<taxonomicName id="4C79D0CA5E79FF8F09AE54CFFE3E2C21" box="[151,437,1018,1043]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="dispar">
<emphasis id="B90D775B5E79FF8F09AE54CFFEC82C21" box="[151,323,1018,1043]" italics="true" pageId="12" pageNumber="77">Crossodactylus</emphasis>
cf. 
<emphasis id="B90D775B5E79FF8F085454CFFE3E2C21" box="[365,437,1018,1043]" italics="true" pageId="12" pageNumber="77">dispar</emphasis>
</taxonomicName>
</bibRefCitation>
) still requires a detailed description and was not used for comparisons.
</paragraph>
<caption id="DF06FBC15E79FF8F09AE5377FC2C2C4B" ID-Table-UUID="DF06FBC15E79FF8F09AE5377FC2C2C4B" httpUri="http://table.plazi.org/id/DF06FBC15E79FF8F09AE5377FC2C2C4B" pageId="12" pageNumber="77" targetBox="[151,1288,1172,1613]" targetIsTable="true" targetPageId="12">
<paragraph id="8BC6AB495E79FF8F09AE5377FC2C2C4B" blockId="12.[151,1436,1089,1145]" pageId="12" pageNumber="77">
<emphasis id="B90D775B5E79FF8F09AE5377FE832C6B" bold="true" box="[151,264,1090,1113]" pageId="12" pageNumber="77">TABLE 3.</emphasis>
Comparison of acoustic parameters of known advertisement calls of 
<taxonomicName id="4C79D0CA5E79FF8F0AD75374FB1A2C6B" box="[1006,1169,1089,1113]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="B90D775B5E79FF8F0AD75374FB1A2C6B" box="[1006,1169,1089,1113]" italics="true" pageId="12" pageNumber="77">Crossodactylus</emphasis>
</taxonomicName>
. Data presented as mean ± SD; range (
<emphasis id="B90D775B5E79FF8F081E5357FEB22C4B" box="[295,313,1122,1145]" italics="true" pageId="12" pageNumber="77">N</emphasis>
, number analyzed) when available. * = data not available.
</paragraph>
</caption>
<paragraph id="8BC6AB495E79FF8F0B2653A0FE832E7F" pageId="12" pageNumber="77">
<table id="F97959E95E79007C09AE53A1FA832E7F" box="[151,1288,1172,1613]" gridcols="4" gridrows="7" pageId="12" pageNumber="77">
<tr id="3549A90B5E79007C09AE53A1FA832C98" box="[151,1288,1172,1194]" gridrow="0" pageId="12" pageNumber="77" rowspan-0="1">
<th id="7698C0775E79007C0B2653A1FD312C98" box="[543,698,1172,1194]" gridcol="1" gridrow="0" pageId="12" pageNumber="77">
<taxonomicName id="4C79D0CA5E79FF8F0B2653A0FD312C98" box="[543,698,1172,1194]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="caramaschii">
<emphasis id="B90D775B5E79FF8F0B2653A0FD312C98" box="[543,698,1172,1194]" italics="true" pageId="12" pageNumber="77">C. caramaschii</emphasis>
</taxonomicName>
</th>
<th id="7698C0775E79007C0A0C53A1FC532C98" box="[821,984,1172,1194]" gridcol="2" gridrow="0" pageId="12" pageNumber="77">
<taxonomicName id="4C79D0CA5E79FF8F0A0C53A0FC432C98" box="[821,968,1172,1194]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="cyclospinus">
<emphasis id="B90D775B5E79FF8F0A0C53A0FC432C98" box="[821,968,1172,1194]" italics="true" pageId="12" pageNumber="77">C. cyclospinus</emphasis>
</taxonomicName>
</th>
<th id="7698C0775E79007C0D6053A1FA832C98" box="[1113,1288,1172,1194]" gridcol="3" gridrow="0" pageId="12" pageNumber="77">
<taxonomicName id="4C79D0CA5E79FF8F0D6053A0FB722C98" box="[1113,1273,1172,1194]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E79FF8F0D6053A0FB722C98" box="[1113,1273,1172,1194]" italics="true" pageId="12" pageNumber="77">C. franciscanus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3549A90B5E79007C09AE538BFA832CC0" box="[151,1288,1214,1266]" gridrow="1" pageId="12" pageNumber="77">
<th id="7698C0775E79007C09AE538BFE282CC0" box="[151,419,1214,1266]" gridcol="0" gridrow="1" pageId="12" pageNumber="77">Notes per call</th>
<td id="7698C0775E79007C0B26538BFD312CC0" box="[543,698,1214,1266]" gridcol="1" gridrow="1" pageId="12" pageNumber="77">56.83 ± 5.41 49–69 (6)</td>
<td id="7698C0775E79007C0A0C538BFC532CC0" box="[821,984,1214,1266]" gridcol="2" gridrow="1" pageId="12" pageNumber="77">63.3 ± 11.9 35–98 (25)</td>
<td id="7698C0775E79007C0D60538BFA832CC0" box="[1113,1288,1214,1266]" gridcol="3" gridrow="1" pageId="12" pageNumber="77">48 ± 18.2 36–84 (48)</td>
</tr>
<tr id="3549A90B5E79007C09AE5233FA832D08" box="[151,1288,1286,1338]" gridrow="2" pageId="12" pageNumber="77">
<th id="7698C0775E79007C09AE5233FE282D08" box="[151,419,1286,1338]" gridcol="0" gridrow="2" pageId="12" pageNumber="77">Call duration (s)</th>
<td id="7698C0775E79007C0B265233FD312D08" box="[543,698,1286,1338]" gridcol="1" gridrow="2" pageId="12" pageNumber="77">5.50 ± 0.54 4.71–6.09 (6)</td>
<td id="7698C0775E79007C0A0C5233FC532D08" box="[821,984,1286,1338]" gridcol="2" gridrow="2" pageId="12" pageNumber="77">4.33 ± 0.74 3.57–6.25 (25)</td>
<td id="7698C0775E79007C0D605233FA832D08" box="[1113,1288,1286,1338]" gridcol="3" gridrow="2" pageId="12" pageNumber="77">6.42 ± 2.5 4.8–11.6 (6)</td>
</tr>
<tr id="3549A90B5E79007C09AE527BFA832DB0" box="[151,1288,1358,1410]" gridrow="3" pageId="12" pageNumber="77">
<th id="7698C0775E79007C09AE527BFE282DB0" box="[151,419,1358,1410]" gridcol="0" gridrow="3" pageId="12" pageNumber="77">Note duration (ms)</th>
<td id="7698C0775E79007C0B26527BFD312DB0" box="[543,698,1358,1410]" gridcol="1" gridrow="3" pageId="12" pageNumber="77">*</td>
<td id="7698C0775E79007C0A0C527BFC532DB0" box="[821,984,1358,1410]" gridcol="2" gridrow="3" pageId="12" pageNumber="77">28 ± 4 3–40 (1601)</td>
<td id="7698C0775E79007C0D60527BFA832DB0" box="[1113,1288,1358,1410]" gridcol="3" gridrow="3" pageId="12" pageNumber="77">12 ± 2.6 6–20 (284)</td>
</tr>
<tr id="3549A90B5E79007C09AE52A3FA832DF8" box="[151,1288,1430,1482]" gridrow="4" pageId="12" pageNumber="77">
<th id="7698C0775E79007C09AE52A3FE282DF8" box="[151,419,1430,1482]" gridcol="0" gridrow="4" pageId="12" pageNumber="77">Internote interval (ms)</th>
<td id="7698C0775E79007C0B2652A3FD312DF8" box="[543,698,1430,1482]" gridcol="1" gridrow="4" pageId="12" pageNumber="77">*</td>
<td id="7698C0775E79007C0A0C52A3FC532DF8" box="[821,984,1430,1482]" gridcol="2" gridrow="4" pageId="12" pageNumber="77">40 ± 3 29–65 (1576)</td>
<td id="7698C0775E79007C0D6052A3FA832DF8" box="[1113,1288,1430,1482]" gridcol="3" gridrow="4" pageId="12" pageNumber="77">123 ± 13.4 90–184 (284)</td>
</tr>
<tr id="3549A90B5E79007C09AE52EBFA832E20" box="[151,1288,1502,1554]" gridrow="5" pageId="12" pageNumber="77">
<th id="7698C0775E79007C09AE52EBFE282E20" box="[151,419,1502,1554]" gridcol="0" gridrow="5" pageId="12" pageNumber="77">Dominant frequency (Hz)</th>
<td id="7698C0775E79007C0B2652EBFD312E20" box="[543,698,1502,1554]" gridcol="1" gridrow="5" pageId="12" pageNumber="77">~5000</td>
<td id="7698C0775E79007C0A0C52EBFC532E20" box="[821,984,1502,1554]" gridcol="2" gridrow="5" pageId="12" pageNumber="77">4981 ± 626.6 3488–5447 (25)</td>
<td id="7698C0775E79007C0D6052EBFA832E20" box="[1113,1288,1502,1554]" gridcol="3" gridrow="5" pageId="12" pageNumber="77">3460.1 ± 282.5 2344–4031 (284)</td>
</tr>
<tr id="3549A90B5E79007C09AE5103FA832E7F" box="[151,1288,1590,1613]" gridrow="6" pageId="12" pageNumber="77" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="7698C0775E79007C09AE5103FE282E7F" box="[151,419,1590,1613]" gridcol="0" gridrow="6" pageId="12" pageNumber="77">continued.</th>
</tr>
</table>
</paragraph>
<paragraph id="8BC6AB495E79FF8F08D7515FFA802FAD" pageId="12" pageNumber="77">
<table id="F97959E95E79007C09A6515CFABB2FAD" box="[159,1328,1641,1951]" gridcols="4" gridrows="5" pageId="12" pageNumber="77">
<tr id="3549A90B5E79007C09A6515CFABB2E4D" box="[159,1328,1641,1663]" gridrow="0" pageId="12" pageNumber="77" rowspan-0="1">
<th id="7698C0775E79007C08D7515CFD1A2E4D" box="[494,657,1641,1663]" gridcol="1" gridrow="0" pageId="12" pageNumber="77">
<taxonomicName id="4C79D0CA5E79FF8F08D7515FFD1A2E4D" box="[494,657,1641,1663]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="gaudichaudii">
<emphasis id="B90D775B5E79FF8F08D7515FFD1A2E4D" box="[494,657,1641,1663]" italics="true" pageId="12" pageNumber="77">C. gaudichaudii</emphasis>
</taxonomicName>
</th>
<th id="7698C0775E79007C0A1B515CFC3C2E4D" box="[802,951,1641,1663]" gridcol="2" gridrow="0" pageId="12" pageNumber="77">
<taxonomicName id="4C79D0CA5E79FF8F0A1B515FFC1D2E4D" box="[802,918,1641,1663]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B90D775B5E79FF8F0A1B515FFC1D2E4D" box="[802,918,1641,1663]" italics="true" pageId="12" pageNumber="77">C. schmidti</emphasis>
</taxonomicName>
</th>
<th id="7698C0775E79007C0D4F515CFABB2E4D" box="[1142,1328,1641,1663]" gridcol="3" gridrow="0" pageId="12" pageNumber="77">
<taxonomicName id="4C79D0CA5E79FF8F0D4F515FFA912E4D" box="[1142,1306,1641,1663]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="77" phylum="Chordata" rank="species" species="trachystomus">
<emphasis id="B90D775B5E79FF8F0D4F515FFA912E4D" box="[1142,1306,1641,1663]" italics="true" pageId="12" pageNumber="77">C. trachystomus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3549A90B5E79007C09A651A6FABB2EF5" box="[159,1328,1683,1735]" gridrow="1" pageId="12" pageNumber="77">
<th id="7698C0775E79007C09A651A6FE092EF5" box="[159,386,1683,1735]" gridcol="0" gridrow="1" pageId="12" pageNumber="77">Notes per call</th>
<td id="7698C0775E79007C08D751A6FD1A2EF5" box="[494,657,1683,1735]" gridcol="1" gridrow="1" pageId="12" pageNumber="77">25–130</td>
<td id="7698C0775E79007C0A1B51A6FC3C2EF5" box="[802,951,1683,1735]" gridcol="2" gridrow="1" pageId="12" pageNumber="77">35 ± 6.64 13–45 (11)</td>
<td id="7698C0775E79007C0D4F51A6FABB2EF5" box="[1142,1328,1683,1735]" gridcol="3" gridrow="1" pageId="12" pageNumber="77">58.8 ± 26.2 13–121 (86)</td>
</tr>
<tr id="3549A90B5E79007C09A651EEFABB2F3D" box="[159,1328,1755,1807]" gridrow="2" pageId="12" pageNumber="77">
<th id="7698C0775E79007C09A651EEFE092F3D" box="[159,386,1755,1807]" gridcol="0" gridrow="2" pageId="12" pageNumber="77">Call duration (s)</th>
<td id="7698C0775E79007C08D751EEFD1A2F3D" box="[494,657,1755,1807]" gridcol="1" gridrow="2" pageId="12" pageNumber="77">2–13</td>
<td id="7698C0775E79007C0A1B51EEFC3C2F3D" box="[802,951,1755,1807]" gridcol="2" gridrow="2" pageId="12" pageNumber="77">4.14 ± 1.06 2.23–5.75 (11)</td>
<td id="7698C0775E79007C0D4F51EEFABB2F3D" box="[1142,1328,1755,1807]" gridcol="3" gridrow="2" pageId="12" pageNumber="77">4.95 ± 1.9 1.4–10.1 (52)</td>
</tr>
<tr id="3549A90B5E79007C09A65016FABB2F65" box="[159,1328,1827,1879]" gridrow="3" pageId="12" pageNumber="77">
<th id="7698C0775E79007C09A65016FE092F65" box="[159,386,1827,1879]" gridcol="0" gridrow="3" pageId="12" pageNumber="77">Note duration (ms)</th>
<td id="7698C0775E79007C08D75016FD1A2F65" box="[494,657,1827,1879]" gridcol="1" gridrow="3" pageId="12" pageNumber="77">40–50</td>
<td id="7698C0775E79007C0A1B5016FC3C2F65" box="[802,951,1827,1879]" gridcol="2" gridrow="3" pageId="12" pageNumber="77">22 ± 7 9–61 (289)</td>
<td id="7698C0775E79007C0D4F5016FABB2F65" box="[1142,1328,1827,1879]" gridcol="3" gridrow="3" pageId="12" pageNumber="77">14 ± 5 1–33 (2478)</td>
</tr>
<tr id="3549A90B5E79007C09A6505EFABB2FAD" box="[159,1328,1899,1951]" gridrow="4" pageId="12" pageNumber="77">
<th id="7698C0775E79007C09A6505EFE092FAD" box="[159,386,1899,1951]" gridcol="0" gridrow="4" pageId="12" pageNumber="77">Internote interval (ms)</th>
<td id="7698C0775E79007C08D7505EFD1A2FAD" box="[494,657,1899,1951]" gridcol="1" gridrow="4" pageId="12" pageNumber="77">40–50</td>
<td id="7698C0775E79007C0A1B505EFC3C2FAD" box="[802,951,1899,1951]" gridcol="2" gridrow="4" pageId="12" pageNumber="77">100 ± 29 17–249 (279)</td>
<td id="7698C0775E79007C0D4F505EFABB2FAD" box="[1142,1328,1899,1951]" gridcol="3" gridrow="4" pageId="12" pageNumber="77">92 ± 21 56–265 (2392)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC6AB495E79FF8F09A65086FB622FFB" blockId="12.[159,1328,1971,2023]" box="[159,1257,1971,1993]" pageId="12" pageNumber="77">Dominant frequency (Hz) 2000–5500 3306 ± 649 3894 ± 525</paragraph>
<paragraph id="8BC6AB495E79FF8E0A1B50E4FB2A29BA" blockId="12.[159,1328,1971,2023]" lastBlockId="13.[151,1437,151,1616]" lastPageId="13" lastPageNumber="78" pageId="12" pageNumber="77">
2017–4280 (11 calls) 1802–4826 (1535) 
<collectingCountry id="F36EEBD95E78FF8E09FE57ADFF6D2882" box="[199,230,152,176]" name="American Samoa" pageId="13" pageNumber="78">As</collectingCountry>
observed for 
<taxonomicName id="4C79D0CA5E78FF8E08B057A2FD582882" box="[393,723,151,176]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="78" phylum="Chordata" rank="species" species="trachystomus">
<emphasis id="B90D775B5E78FF8E08B057A2FD582882" box="[393,723,151,176]" italics="true" pageId="13" pageNumber="78">Crossodactylus trachystomus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE8D6B85E78FF8E0BDC57A2FC462882" author="Pimenta" box="[741,973,151,176]" pageId="13" pageNumber="78" refString="Pimenta, B. V. S., Wachlevski, M. &amp; Cruz, C. A. G. (2008) Morphological and Acoustical Variation, Geographic Distribution, and Conservation Status of the Spinythumb Frog Crossodactylus bokermanni Caramaschi and Sazima, 1985 (Anura, Hylodidae). Journal of Herpetology, 42, 481 - 492. http: // dx. doi. org / 10.1670 / 07 - 164.1" type="journal article" year="2008">
Pimenta 
<emphasis id="B90D775B5E78FF8E0A7557ACFC032882" box="[844,904,151,176]" italics="true" pageId="13" pageNumber="78">et al.</emphasis>
2008
</bibRefCitation>
; under 
<taxonomicName id="4C79D0CA5E78FF8E0D1D57ADFB452882" box="[1060,1230,151,176]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="78" phylum="Chordata" rank="species" species="bokermanni">
<emphasis id="B90D775B5E78FF8E0D1D57ADFB452882" box="[1060,1230,151,176]" italics="true" pageId="13" pageNumber="78">C. bokermanni</emphasis>
</taxonomicName>
), the first note of each call (
<emphasis id="B90D775B5E78FF8E08335788FE9628E6" box="[266,285,189,212]" italics="true" pageId="13" pageNumber="78">N</emphasis>
= 6 first notes) is less intense, with a slightly smaller duration (mean = 10 ms; SD = 4.8; range = 5–18 ms) and lower dominant frequency (mean = 3125 Hz; SD = 363.87; range = 2531–3656 Hz). First notes with less intense and lower mean dominant frequencies are attributes probably found in others 
<taxonomicName id="4C79D0CA5E78FF8E0D655631FA83292F" box="[1116,1288,260,285]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="78" phylum="Chordata" rank="genus">
<emphasis id="B90D775B5E78FF8E0D655631FA83292F" box="[1116,1288,260,285]" italics="true" pageId="13" pageNumber="78">Crossodactylus</emphasis>
</taxonomicName>
but have not yet been described. Lower dominant frequencies were also observed in 
<taxonomicName id="4C79D0CA5E78FF8E0A8D5612FE452956" authority="Pombal, Feio &amp; Haddad, 2002 (Pombal et al. 2002)" authorityName="Pombal, Feio &amp; Haddad, 2002 (Pombal et al." authorityYear="2002" class="Amphibia" family="Hylodidae" genus="Hylodes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="78" phylum="Chordata" rank="species" species="amnicola">
<emphasis id="B90D775B5E78FF8E0A8D5612FBF42972" box="[948,1151,295,320]" italics="true" pageId="13" pageNumber="78">Hylodes amnicola</emphasis>
Pombal, Feio &amp; 
<bibRefCitation id="EFE8D6B85E78FF8E0C075612FF442956" author="Haddad" pageId="13" pageNumber="78" refString="Bastos, R. P. &amp; Haddad, C. F. B. (2002) Acoustic and aggressive interactions in Scinax rizibilis (Anura: Hylidae) during the reproductive activity in southeastern Brazil. Amphibia-Reptilia, 23, 97 - 104." type="journal article" year="2002">Haddad, 2002</bibRefCitation>
(Pombal 
<emphasis id="B90D775B5E78FF8E087D5678FE0A2957" box="[324,385,332,357]" italics="true" pageId="13" pageNumber="78">et al.</emphasis>
2002)
</taxonomicName>
and in other anurans (e.g., 
<bibRefCitation id="EFE8D6B85E78FF8E0A235678FBA42956" author="Bastos" box="[794,1071,332,357]" pageId="13" pageNumber="78" refString="Bastos, R. P. &amp; Haddad, C. F. B. (2002) Acoustic and aggressive interactions in Scinax rizibilis (Anura: Hylidae) during the reproductive activity in southeastern Brazil. Amphibia-Reptilia, 23, 97 - 104." type="journal article" year="2002">Bastos &amp; Haddad 2002</bibRefCitation>
; 
<bibRefCitation id="EFE8D6B85E78FF8E0D065679FAC82956" author="Castellano" box="[1087,1347,332,357]" pageId="13" pageNumber="78" refString="Castellano, S., Cuatto, B., Rinella, R., Rosso, A. &amp; Giacoma, C. (2002) The advertisement call of the European treefrogs (Hyla arborea): a multilevel study of variation. Ethology, 108, 75 - 89. http: // dx. doi. org / 10.1046 / j. 1439 - 0310.2002.00761. x" type="journal article" year="2002">
Castellano 
<emphasis id="B90D775B5E78FF8E0DF85678FB762957" box="[1217,1277,332,357]" italics="true" pageId="13" pageNumber="78">et al.</emphasis>
2002
</bibRefCitation>
) and is considered a common effect associated with the beginning of the call (
<bibRefCitation id="EFE8D6B85E78FF8E0A92565AFB1F29BA" author="Giacoma" box="[939,1172,367,392]" pageId="13" pageNumber="78" refString="Giacoma, C., Zugolaro, C. &amp; Beani, L. (1997) The advertisement calls of the green toad (Bufo viridis): variability and role in mate choice. Herpetologica, 53, 454 - 464." type="journal article" year="1997">
Giacoma 
<emphasis id="B90D775B5E78FF8E0D205644FBDA29BA" box="[1049,1105,367,392]" italics="true" pageId="13" pageNumber="78">et al.</emphasis>
1997
</bibRefCitation>
).
</paragraph>
<paragraph id="8BC6AB495E78FF8E09FE56A0FE562BB2" blockId="13.[151,1437,151,1616]" pageId="13" pageNumber="78">
We considered signs of alteration on calls analyzed the interruptions followed by less intense calls after long intervals, the addition of very low intensity notes at the end of call, and also the emission of isolated notes. These alterations were also observed in 
<taxonomicName id="4C79D0CA5E78FF8E0B1256E9FCB929C6" box="[555,818,476,501]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="78" phylum="Chordata" rank="species" species="aeneus">
<emphasis id="B90D775B5E78FF8E0B1256E9FCB929C6" box="[555,818,476,501]" italics="true" pageId="13" pageNumber="78">Crossodactylus aeneus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C79D0CA5E78FF8E0A7C56E8FB8B29C7" box="[837,1024,476,501]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="78" phylum="Chordata" rank="species" species="gaudichaudii">
<emphasis id="B90D775B5E78FF8E0A7C56E8FB8B29C7" box="[837,1024,476,501]" italics="true" pageId="13" pageNumber="78">C. gaudichaudii</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C79D0CA5E78FF8E0D7056E8FA8E29C7" box="[1097,1285,476,501]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="78" phylum="Chordata" rank="species" species="trachystomus">
<emphasis id="B90D775B5E78FF8E0D7056E8FA8E29C7" box="[1097,1285,476,501]" italics="true" pageId="13" pageNumber="78">C. trachystomus</emphasis>
</taxonomicName>
during field activities during this study (B.V.S. Pimenta, pers. obs.). They appear to be associated with the calling male noticing the presence of an observer, the moment when a male resumed call activity after escaping and returning to call site, or the presence of a second male emitting calls with short intervals. When call was interrupted and followed by low intensity calls, the notes of these altered calls presented mean duration of 11 ms (SD = 2.6; range = 5-16 ms; 
<emphasis id="B90D775B5E78FF8E0C695558FAE82AB6" box="[1360,1379,621,644]" italics="true" pageId="13" pageNumber="78">N</emphasis>
= 93 notes), mean internote interval of 124 ms (SD = 13.6; 99–175 ms; 
<emphasis id="B90D775B5E78FF8E0A4755A4FC1A2A9A" box="[894,913,657,680]" italics="true" pageId="13" pageNumber="78">N</emphasis>
= 93 intervals), and mean dominant frequency of 3228.6 Hz (SD = 231.41; range = 2531–3844 Hz; 
<emphasis id="B90D775B5E78FF8E0BD65580FC892AFE" box="[751,770,693,716]" italics="true" pageId="13" pageNumber="78">N</emphasis>
= 89 notes). Low intensity notes added to the end of a call present mean duration of 11 ms (DP = 2.5; range = 7–17 ms; 
<emphasis id="B90D775B5E78FF8E0A7F55ECFCD22AC2" box="[838,857,729,752]" italics="true" pageId="13" pageNumber="78">N</emphasis>
= 28 notes), mean internote interval of 141 ms (SD = 45.8; range = 98–282 ms), and mean dominant frequency of 3170.6 Hz (SD = 262.95; range = 2531–3844 Hz; 
<emphasis id="B90D775B5E78FF8E0CB055C8FA172B26" box="[1417,1436,765,788]" italics="true" pageId="13" pageNumber="78">N</emphasis>
= 22 notes). The only notes showing considerable differences from unaltered calls were the isolated notes, which present mean duration of 12 ms (
<emphasis id="B90D775B5E78FF8E0B375470FDAA2B6E" box="[526,545,837,860]" italics="true" pageId="13" pageNumber="78">N</emphasis>
= 3 notes) and mean dominant frequency of 2031.3 Hz (SD = 107.96; range = 
<date id="FFC78D895E78FF8E09AE5452FE9E2BB2" box="[151,277,871,896]" pageId="13" pageNumber="78" value="1969" valueMax="2156">1969–2156</date>
Hz; 
<emphasis id="B90D775B5E78FF8E0872545CFED52BB2" box="[331,350,873,896]" italics="true" pageId="13" pageNumber="78">N</emphasis>
= 3 notes).
</paragraph>
<paragraph id="8BC6AB495E78FF8E09FE54B9FD512C4F" blockId="13.[151,1437,151,1616]" pageId="13" pageNumber="78">
<emphasis id="B90D775B5E78FF8E09FE54B9FB932B97" bold="true" box="[199,1048,908,933]" pageId="13" pageNumber="78">Geographic distribution, natural history, and conservation status.</emphasis>
<taxonomicName id="4C79D0CA5E78FF8E0D1354B9FAFC2B96" box="[1066,1399,908,933]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="78" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E78FF8E0D1354B9FAFC2B96" box="[1066,1399,908,933]" italics="true" pageId="13" pageNumber="78">Crossodactylus franciscanus</emphasis>
</taxonomicName>
is currently known from two municipalities in the western region of the state of Minas Gerais, both included in the Cerrado biome (
<figureCitation id="1342B7CC5E78FF8E086854E1FE1C2BDE" box="[337,407,980,1005]" captionStart="FIGURE 3" captionStartId="7.[151,250,1621,1643]" captionTargetBox="[160,1427,668,1594]" captionTargetId="figure@7.[151,1435,661,1600]" captionTargetPageId="7" captionText="FIGURE 3. Geographic distributions of Crossodactylus trachystomus (circles, including the capital city of Belo Horizonte) and Crossodactylus franciscanus (squares) in the state of Minas Gerais, Southeastern Brazil." httpUri="https://zenodo.org/record/288454/files/figure.png" pageId="13" pageNumber="78">Fig. 3</figureCitation>
). The Parque Nacional da Serra da Canastra, on the municipality of São Roque de Minas, and the municipality of Passos are on opposite margins of the Rio Grande, which is dammed in this region, and distant approx. 
<quantity id="4C8106AC5E78FF8E086A5329FE2A2C07" box="[339,417,1052,1077]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="78" unit="km" value="60.0">60 km</quantity>
from each other. Passos is not included on the complex of elevations and plateaus collectively known as Serra da Canastra. The localities 
<collectingCountry id="F36EEBD95E78FF8E0A225375FCDE2C65" box="[795,853,1088,1111]" name="Malta" pageId="13" pageNumber="78">Mata</collectingCountry>
do Batatal and 
<collectingCountry id="F36EEBD95E78FF8E0D365375FBC22C65" box="[1039,1097,1088,1111]" name="Malta" pageId="13" pageNumber="78">Mata</collectingCountry>
da Colina, situated at Passos according to the original jar label, were not located.
</paragraph>
<paragraph id="8BC6AB495E78FF8E09FE53B2FCAA2D02" blockId="13.[151,1437,151,1616]" pageId="13" pageNumber="78">
According to 
<bibRefCitation id="EFE8D6B85E78FF8E085553B2FDE82C92" box="[364,611,1159,1184]" pageId="13" pageNumber="78" refString="Haddad, C. F. B., Andrade, G. V. &amp; Cardoso, A. J. (1988) Anfibios anuros do Parque Nacional da Serra da Canastra, Estado de Minas Gerais. Brasil Florestal, 64, 9 - 20." type="journal article">
Haddad 
<emphasis id="B90D775B5E78FF8E08F653BCFD862C92" box="[463,525,1159,1184]" italics="true" pageId="13" pageNumber="78">et al.</emphasis>
(1988)
</bibRefCitation>
males of 
<taxonomicName id="4C79D0CA5E78FF8E0BD853B2FBA32C92" box="[737,1064,1159,1184]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="78" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E78FF8E0BD853B2FBA32C92" box="[737,1064,1159,1184]" italics="true" pageId="13" pageNumber="78">Crossodactylus franciscanus</emphasis>
</taxonomicName>
(referred as 
<taxonomicName id="4C79D0CA5E78FF8E0DFE53B2FEA72CF7" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="78" phylum="Chordata" rank="species" species="trachystomus">
<emphasis id="B90D775B5E78FF8E0DFE53B2FAF82C92" box="[1223,1395,1159,1184]" italics="true" pageId="13" pageNumber="78">Crossodactylus</emphasis>
cf. 
<emphasis id="B90D775B5E78FF8E09AE5399FEA72CF7" box="[151,300,1196,1221]" italics="true" pageId="13" pageNumber="78">trachystomus</emphasis>
</taxonomicName>
) call during the day and shortly after sunset. Specimens were found in small ponds with shrubs and grass on margins or sitting on debris at the bottom of shallow rivulets, where tadpoles were also recorded (no specimens of tadpoles were found in collections). 
<collectingCountry id="F36EEBD95E78FF8E0BE553C1FD702D3E" box="[732,763,1268,1292]" name="American Samoa" pageId="13" pageNumber="78">As</collectingCountry>
far as we are aware, the last specimen of 
<taxonomicName id="4C79D0CA5E78FF8E0DD053C0FA172D3E" box="[1257,1436,1268,1292]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="78" phylum="Chordata" rank="species" species="franciscanus">
<emphasis id="B90D775B5E78FF8E0DD053C0FA172D3E" box="[1257,1436,1268,1292]" italics="true" pageId="13" pageNumber="78">C. franciscanus</emphasis>
</taxonomicName>
which entered collection was the 
<typeStatus id="54C215EB5E78FF8E0B345222FDF92D02" box="[525,626,1303,1328]" pageId="13" pageNumber="78" type="holotype">holotype</typeStatus>
, found in 1981.
</paragraph>
</subSubSection>
<subSubSection id="C363F8C25E78FF8E09FE5209FE4C2DAF" pageId="13" pageNumber="78" type="etymology">
<paragraph id="8BC6AB495E78FF8E09FE5209FE4C2DAF" blockId="13.[151,1437,151,1616]" pageId="13" pageNumber="78">
<emphasis id="B90D775B5E78FF8E09FE5209FEC52D67" bold="true" box="[199,334,1340,1365]" pageId="13" pageNumber="78">Etymology.</emphasis>
The specific epithet “ 
<taxonomicName id="4C79D0CA5E78FF8E0B7D5209FD5E2D67" box="[580,725,1340,1365]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="78" phylum="Chordata" rank="species" species="franciscanus">franciscanus</taxonomicName>
” aludes to the Rio São Francisco, which forms one of the most important Brazilian river basins. Its headwaters are located in the Parque Nacional da Serra da Canastra, 
<typeStatus id="54C215EB5E78FF8E0C555254FA172D4B" box="[1388,1436,1377,1401]" pageId="13" pageNumber="78">type</typeStatus>
locality of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C363F8C25E78FF8E09FE5292FAB32E62" pageId="13" pageNumber="78" type="discussion">
<paragraph id="8BC6AB495E78FF8E09FE5292FAB32E62" blockId="13.[151,1437,151,1616]" pageId="13" pageNumber="78">
<emphasis id="B90D775B5E78FF8E09FE5292FEB02DF2" bold="true" box="[199,315,1447,1472]" pageId="13" pageNumber="78">Remarks.</emphasis>
Specimen ZUEC 4343, which was not found in the corresponding collection during our study, was collected at Serra da Canastra and recorded by A.J. Cardoso. The holotype’s original jar label says it is also a recording voucher. There is only one recording of 
<taxonomicName id="4C79D0CA5E78FF8E0BF252DAFCFC2E3A" box="[715,887,1519,1544]" class="Amphibia" family="Hylodidae" genus="Crossodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="78" phylum="Chordata" rank="genus">
<emphasis id="B90D775B5E78FF8E0BF252DAFCFC2E3A" box="[715,887,1519,1544]" italics="true" pageId="13" pageNumber="78">Crossodactylus</emphasis>
</taxonomicName>
from Serra da Canastra deposited at the Arquivo Sonoro Neotropical da Universidade de Campinas (Neotropical Sound Archive, University of Campinas; AJC 010/ 1), so we could not determine which of these two specimens, ZUEC 4343 or MNRJ 40137, was recorded.
</paragraph>
</subSubSection>
</treatment>
</document>