treatments-xml/data/03/AE/EE/03AEEE34FFB8C800FF0AFF75FE37F895.xml
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<document id="78573F4DC1B674923955B3654CA66030" ID-CLB-Dataset="7372" ID-DOI="10.11646/zootaxa.4952.1.8" ID-GBIF-Dataset="d22e3fb7-eaed-40c7-8b5d-d6b3702492cf" ID-ISSN="1175-5326" ID-Zenodo-Dep="4671916" ID-ZooBank="6EC786D4-DF85-419A-B350-8CCC2C487CD1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617874823328" checkinUser="plazi" docAuthor="Freyhof, Jörg, Kaya, Cüneyt, Epitashvili, Giorgi &amp; Geiger, Matthias F." docDate="2021" docId="03AEEE34FFB8C800FF0AFF75FE37F895" docLanguage="en" docName="zootaxa.4952.1.8.pdf" docOrigin="Zootaxa 4952 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oxynoemacheilus phasicus Freyhof &amp; Kaya &amp; Epitashvili &amp; Geiger 2021, new species" docType="treatment" docVersion="9" lastPageNumber="146" masterDocId="FF97964CFFBCC80BFF9DFFE2FFE1FFC9" masterDocTitle="Oxynoemacheilus phasicus, a new nemacheilid loach from the eastern Black Sea basin with some remarks on other Caucasian Oxynoemacheilus (Teleostei: Nemacheilidae)" masterLastPageNumber="151" masterPageNumber="135" pageNumber="139" updateTime="1698936202155" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2755C54A81B3BE909AD57D92E0620BBE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C2B6F76E37CE01BC4401E636CB462869">
<mods:title id="1014BC85D037AC31FDA3324844A67E13">Oxynoemacheilus phasicus, a new nemacheilid loach from the eastern Black Sea basin with some remarks on other Caucasian Oxynoemacheilus (Teleostei: Nemacheilidae)</mods:title>
</mods:titleInfo>
<mods:name id="968458A7DD793B04F36CCDBDC729581D" type="personal">
<mods:role id="4F26ED2F8263152E4EFD27DBF6C7040A">
<mods:roleTerm id="66327A5628EDD0D678AD6A31EBC5F29D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2DD20B1691661437A9FA78EC89A02FCC">Freyhof, Jörg</mods:namePart>
<mods:nameIdentifier id="8352820516905BBC50C81D58AE2C425D" type="ORCID">0000-0002-7042-3127</mods:nameIdentifier>
<mods:affiliation id="AE6ED42E3C7BCF2DBDE3B5C392572806">Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, 10115 Berlin, Germany. joerg. freyhof @ mfn. berlin; https: // orcid. org / 0000 - 0002 - 7042 - 3127</mods:affiliation>
<mods:nameIdentifier id="31C55D2CC141F7EFBC584810AB530F41" type="email">joerg.freyhof@mfn.berlin</mods:nameIdentifier>
</mods:name>
<mods:name id="1AC65E046FFB9AB5FAB15EADB816A45E" type="personal">
<mods:role id="1E96A68CE77D2F30A21CA9C26C744368">
<mods:roleTerm id="38BADF4F5FCC709112FCF8C4584E24E4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8F97E83D3A4F49E88186C2C03E7EA78B">Kaya, Cüneyt</mods:namePart>
<mods:affiliation id="1A0746A9D220F1C2185B1E170C972D87">Recep Tayyip Erdogan University, Faculty of Fisheries and Aquatic Sciences, 53100 Rize, Turkey.</mods:affiliation>
</mods:name>
<mods:name id="E09EAB50931C60855A37BA7609979FCE" type="personal">
<mods:role id="BC4E2F6DD7826D68B73E2A389F1C8E97">
<mods:roleTerm id="238E06246D3FC61DAD6E8A9255BE1224">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EE4505B6ED44A56B573B642B3AA3033E">Epitashvili, Giorgi</mods:namePart>
<mods:affiliation id="F7FC9A54407AA55EFE717002E971DF2C">Institute of Zoology, Ilia State University, 0179, Tbilisi, Georgia.</mods:affiliation>
</mods:name>
<mods:name id="2F286A81C73549E024CAC6BEE322AF06" type="personal">
<mods:role id="EF9A334F993AE8B376E2E2DBAF73969A">
<mods:roleTerm id="7D5128F73DB464404977C83E83E84362">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="144D768F0BB2C6B694119627D09F5020">Geiger, Matthias F.</mods:namePart>
<mods:nameIdentifier id="49104DDA1D0D2B53D790C2982066D638" type="ORCID">0000-0003-2403-0805</mods:nameIdentifier>
<mods:affiliation id="64FB567D5EC38BE8023940932ED7D1D7">Zoological Research Museum Alexander Koenig, Leibniz Institute for Animal Biodiversity, Adenauerallee 160, 53113 Bonn, Germany. m. geiger @ leibniz-zfmk. de; https: // orcid. org / 0000 - 0003 - 2403 - 0805</mods:affiliation>
<mods:nameIdentifier id="00AC3213D334F8EC2E39DCD8F477B271" type="email">m.geiger@leibniz-zfmk.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="5E0938EC8BC47AA80AB92F00ADFC6D38">text</mods:typeOfResource>
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<mods:titleInfo id="E29002447010DA6D3A3BE1FCF8023B8B">
<mods:title id="430CC650F29D74C43DDF599E4400BD37">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1B47764B4D738F4A23D1371DA916A2E5">
<mods:date id="D15063F6280F35F574B2A8DA526DFFE9">2021</mods:date>
<mods:detail id="201D9F927664D2B06608DF4110DA4CA4" type="pubDate">
<mods:number id="6AE9E7528C700B595323DE2ADC0D3B81">2021-04-08</mods:number>
</mods:detail>
<mods:detail id="818A2E86C31AE67856ED22ACF5AEF670" type="volume">
<mods:number id="1879A873F728176C6C0C87746B12A0EE">4952</mods:number>
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<mods:classification id="69F637A9E65AFE9CA738C53A2D77E007">journal article</mods:classification>
<mods:identifier id="F2041EDDE1E1E4C9BDF1C8A1E720190E" type="CLB-Dataset">7372</mods:identifier>
<mods:identifier id="AC5FBE22DA4D24BB136D2A554DFCE940" type="DOI">10.11646/zootaxa.4952.1.8</mods:identifier>
<mods:identifier id="74D0C1AAFA5CB036C8F34DDC7068FB82" type="GBIF-Dataset">d22e3fb7-eaed-40c7-8b5d-d6b3702492cf</mods:identifier>
<mods:identifier id="2DB4B4F8B33DD3BDCFC326CB71303322" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EA74F297C068FAF54DA7459765B21B5C" type="Zenodo-Dep">4671916</mods:identifier>
<mods:identifier id="C4AC0F26014719DEF654FC0291B39630" type="ZooBank">6EC786D4-DF85-419A-B350-8CCC2C487CD1</mods:identifier>
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<treatment id="03AEEE34FFB8C800FF0AFF75FE37F895" ID-DOI="http://doi.org/10.5281/zenodo.4685572" ID-GBIF-Taxon="181003836" ID-Zenodo-Dep="4685572" LSID="urn:lsid:plazi:treatment:03AEEE34FFB8C800FF0AFF75FE37F895" httpUri="http://treatment.plazi.org/id/03AEEE34FFB8C800FF0AFF75FE37F895" lastPageId="11" lastPageNumber="146" pageId="4" pageNumber="139">
<subSubSection id="C31D0CA9FFB8C80FFF0AFF75FD61FF78" box="[151,640,151,178]" pageId="4" pageNumber="139" type="nomenclature">
<paragraph id="8BB85F22FFB8C80FFF0AFF75FD61FF78" blockId="4.[151,640,151,214]" box="[151,640,151,178]" pageId="4" pageNumber="139">
<heading id="D0F0E84EFFB8C80FFF0AFF75FD61FF78" bold="true" box="[151,640,151,178]" fontSize="11" level="1" pageId="4" pageNumber="139" reason="1">
<emphasis id="B9738330FFB8C80FFF0AFF75FD61FF78" bold="true" box="[151,640,151,178]" pageId="4" pageNumber="139">
<taxonomicName id="4C0724A1FFB8C80FFF0AFF75FE00FF7B" authority="Freyhof &amp; Kaya &amp; Epitashvili &amp; Geiger, 2021" authorityName="Freyhof &amp; Kaya &amp; Epitashvili &amp; Geiger" authorityYear="2021" box="[151,481,151,178]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="phasicus" status="sp. nov.">
<emphasis id="B9738330FFB8C80FFF0AFF75FE00FF7B" bold="true" box="[151,481,151,178]" italics="true" pageId="4" pageNumber="139">Oxynoemacheilus phasicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2403E4BFFB8C80FFE6DFF75FD61FF78" box="[496,640,151,177]" pageId="4" pageNumber="139" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31D0CA9FFB8C80FFF0AFF59FEEAFF1F" box="[151,267,187,214]" pageId="4" pageNumber="139" type="description">
<paragraph id="8BB85F22FFB8C80FFF0AFF59FEEAFF1F" blockId="4.[151,640,151,214]" box="[151,267,187,214]" pageId="4" pageNumber="139">
(
<figureCitation id="133C43A7FFB8C80FFF02FF59FEE2FF1F" box="[159,259,187,214]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="6.[151,250,1384,1408]" captionStartId-1="7.[151,250,1244,1268]" captionStartId-2="8.[151,250,1012,1036]" captionStartId-3="8.[151,250,1916,1940]" captionTargetBox-0="[152,1436,381,1361]" captionTargetBox-1="[151,1436,381,1221]" captionTargetBox-2="[151,1436,181,988]" captionTargetBox-3="[151,1436,1133,1893]" captionTargetId-0="figure-123@6.[152,1436,381,1361]" captionTargetId-1="figure-129@7.[151,1436,381,1221]" captionTargetId-2="figure-25@8.[151,1436,181,988]" captionTargetId-3="figure-82@8.[151,1436,1133,1893]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="8" captionText-0="FIGURE 2. Oxynoemacheilus phasicus, ZFMK ICH-124180, holotype, 74.1 mm SL; Georgia: Dzirula River." captionText-1="FIGURE 3. Oxynoemacheilus phasicus, paratypes, from top, left to right: FSJF 4061, 32 mm SL; 39 mm SL; Georgia: Gubistskali River; FSJF 3615, 44 mm SL, 54 mm SL; Georgia: Tskhenitskali River; FSJF 3576, 59 mm SL; Georgia: Enguri River; FSJF 3592, 77 mm SL; Georgia: Dzirula River." captionText-2="FIGURE 4. Oxynoemacheilus phasicus, paratypes, from top, left to right: FSJF 4061, 32 mm SL; 39 mm SL; Georgia: Gubistskali River; FSJF 3615, 44 mm SL, 54 mm SL; Georgia: Tskhenitskali River; FSJF 3576, 59 mm SL; Georgia: Enguri River; FSJF 3592, 77 mm SL; Georgia: Dzirula River." captionText-3="FIGURE 5. Oxynoemacheilus phasicus, paratypes, from top, left to right: FSJF 4061, 32 mm SL; 39 mm SL; Georgia: Gubistskali River; FSJF 3615, 44 mm SL, 54 mm SL; Georgia: Tskhenitskali River; FSJF 3576, 59 mm SL; Georgia: Enguri River; FSJF 3592, 77 mm SL; Georgia: Dzirula River." figureDoi-0="http://doi.org/10.5281/zenodo.4671920" figureDoi-1="http://doi.org/10.5281/zenodo.4671922" figureDoi-2="http://doi.org/10.5281/zenodo.4671926" figureDoi-3="http://doi.org/10.5281/zenodo.4671930" httpUri-0="https://zenodo.org/record/4671920/files/figure.png" httpUri-1="https://zenodo.org/record/4671922/files/figure.png" httpUri-2="https://zenodo.org/record/4671926/files/figure.png" httpUri-3="https://zenodo.org/record/4671930/files/figure.png" pageId="4" pageNumber="139">Figs 25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31D0CA9FFB8C80FFF0AFEE1FE2BFD3B" pageId="4" pageNumber="139" type="materials_examined">
<paragraph id="8BB85F22FFB8C80FFF0AFEE1FDE9FE88" blockId="4.[151,1437,259,2014]" pageId="4" pageNumber="139">
<materialsCitation id="3B6F557FFFB8C80FFF0AFEE1FDE9FE88" ID-GBIF-Occurrence="3071308302" collectionCode="ZFMK" country="Georgia" latitude="42.0975" location="Kutaisi" longLatPrecision="6" longitude="43.39" municipality="Dzirula River" pageId="4" pageNumber="139" specimenCode="ICH-124180, 74" specimenCount="1" stateProvince="Tbilisi" typeStatus="holotype">
<emphasis id="B9738330FFB8C80FFF0AFEE1FEEAFED4" bold="true" box="[151,267,259,285]" pageId="4" pageNumber="139">
<typeStatus id="54BCE180FFB8C80FFF0AFEE1FEE7FED4" box="[151,262,259,285]" pageId="4" pageNumber="139" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED16C7E7FFB8C80FFE8EFEE6FE80FED7" box="[275,353,260,286]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="139" type="Museum">ZFMK</collectionCode>
<specimenCode id="DBA1F759FFB8C80FFEF4FEE1FE18FED4" box="[361,505,259,285]" collectionCode="ICH-" pageId="4" pageNumber="139">ICH-124180</specimenCode>
,
<specimenCode id="DBA1F759FFB8C80FFD98FEE1FDC0FED4" box="[517,545,259,285]" collectionCode="ICH-" pageId="4" pageNumber="139">74</specimenCode>
mm SL; Georgia:
<collectingMunicipality id="6BDCC558FFB8C80FFD6BFEE1FC71FED4" box="[758,912,259,285]" pageId="4" pageNumber="139">Dzirula River</collectingMunicipality>
at road from
<collectingRegion id="49C391C0FFB8C80FFBB1FEE1FB94FED4" box="[1068,1141,259,285]" country="Georgia" name="Tbilisi" pageId="4" pageNumber="139">Tbilisi</collectingRegion>
to
<location id="8ED809F9FFB8C80FFB06FEE1FB0DFED4" LSID="urn:lsid:plazi:treatment:03AEEE34FFB8C800FF0AFF75FE37F895:8ED809F9FFB8C80FFB06FEE1FB0DFED4" box="[1179,1260,259,285]" country="Georgia" latitude="42.0975" longLatPrecision="6" longitude="43.39" municipality="Dzirula River" name="Kutaisi" pageId="4" pageNumber="139" stateProvince="Tbilisi">Kutaisi</location>
<locationDeviation id="9A0A4600FFB8C80FFB69FEE1FED9FE88" location="Khunevi" pageId="4" pageNumber="139">
<quantity id="4CFFF2C7FFB8C80FFB69FEE1FACFFED4" box="[1268,1326,259,285]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="139" unit="km" value="1.0">1 km</quantity>
upstream from Khunevi
</locationDeviation>
,
<geoCoordinate id="EE3339E5FFB8C80FFEDEFEC5FE7FFE8B" box="[323,414,295,322]" degrees="42.0975" direction="north" orientation="latitude" pageId="4" pageNumber="139" precision="5" value="42.0975">42.0975</geoCoordinate>
<geoCoordinate id="EE3339E5FFB8C80FFE38FEC5FDE5FE88" box="[421,516,295,321]" degrees="43.3900" direction="east" orientation="longitude" pageId="4" pageNumber="139" precision="5" value="43.39">43.3900</geoCoordinate>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB85F22FFB8C80FFF5AFEA9FCEBFE18" blockId="4.[151,1437,259,2014]" pageId="4" pageNumber="139">
<materialsCitation id="3B6F557FFFB8C80FFF5AFEA9FC90FEAC" ID-GBIF-Occurrence="3071308311" box="[199,881,331,358]" collectionCode="FSJF" country="Georgia" latitude="42.0975" location="Kutaisi" longLatPrecision="6" longitude="43.39" municipality="Dzirula River" pageId="4" pageNumber="139" specimenCode="FSJF 3592, 1" specimenCount="1" stateProvince="Tbilisi" typeStatus="paratype">
<emphasis id="B9738330FFB8C80FFF5AFEA9FEA7FEAC" bold="true" box="[199,326,331,357]" pageId="4" pageNumber="139">
<typeStatus id="54BCE180FFB8C80FFF5AFEA9FEA0FEAC" box="[199,321,331,357]" pageId="4" pageNumber="139" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="DBA1F759FFB8C80FFED0FEA9FE28FEAF" box="[333,457,331,358]" collectionCode="FSJF" pageId="4" pageNumber="139">FSJF 3592</specimenCode>
,
<specimenCode id="DBA1F759FFB8C80FFE48FEA9FE02FEAC" box="[469,483,331,357]" collectionCode="FSJF" pageId="4" pageNumber="139">1</specimenCode>
,
<quantity id="4CFFF2C7FFB8C80FFE6CFEAEFDA0FEAF" box="[497,577,332,358]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="4" pageNumber="139" unit="mm" value="77.0">77 mm</quantity>
SL; same data as holotype
</materialsCitation>
.—
<materialsCitation id="3B6F557FFFB8C80FFC0FFEA9FE05FE40" ID-GBIF-Occurrence="3071308305" collectionCode="FSJF" country="Georgia" latitude="42.6617" location="Enguri River" longLatPrecision="6" longitude="41.9911" municipality="Lia" pageId="4" pageNumber="139" specimenCode="FSJF 3576, 3" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBA1F759FFB8C80FFC0FFEA9FBEEFEAF" box="[914,1039,331,358]" collectionCode="FSJF" pageId="4" pageNumber="139">FSJF 3576</specimenCode>
,
<specimenCode id="DBA1F759FFB8C80FFB87FEA9FBC8FEAC" box="[1050,1065,331,357]" collectionCode="FSJF" pageId="4" pageNumber="139">3</specimenCode>
,
<quantity id="4CFFF2C7FFB8C80FFBAAFEA9FB51FEAF" box="[1079,1200,331,358]" metricMagnitude="-2" metricUnit="m" metricValue="5.45" metricValueMax="5.9" metricValueMin="5.0" pageId="4" pageNumber="139" unit="mm" value="54.5" valueMax="59.0" valueMin="50.0">5059 mm</quantity>
SL; Georgia:
<location id="8ED809F9FFB8C80FFACDFEA9FF34FE40" LSID="urn:lsid:plazi:treatment:03AEEE34FFB8C800FF0AFF75FE37F895:8ED809F9FFB8C80FFACDFEA9FF34FE40" country="Georgia" latitude="42.6617" longLatPrecision="6" longitude="41.9911" municipality="Lia" name="Enguri River" pageId="4" pageNumber="139">Enguri River</location>
at
<collectingMunicipality id="6BDCC558FFB8C80FFF6BFE8DFEFFFE40" box="[246,286,367,393]" pageId="4" pageNumber="139">Lia</collectingMunicipality>
,
<geoCoordinate id="EE3339E5FFB8C80FFEB5FE8DFE62FE40" box="[296,387,367,393]" degrees="42.6617" direction="north" orientation="latitude" pageId="4" pageNumber="139" precision="5" value="42.6617">42.6617</geoCoordinate>
<geoCoordinate id="EE3339E5FFB8C80FFE14FE8DFE05FE40" box="[393,484,367,393]" degrees="41.9911" direction="east" orientation="longitude" pageId="4" pageNumber="139" precision="5" value="41.9911">41.9911</geoCoordinate>
</materialsCitation>
.—
<materialsCitation id="3B6F557FFFB8C80FFD98FE8DFEB6FE64" ID-GBIF-Occurrence="3071308310" collectionCode="FSJF" country="Georgia" latitude="42.1694" location="Tskhenitskali River" longLatPrecision="6" longitude="42.4314" municipality="Samtredia" pageId="4" pageNumber="139" specimenCode="FSJF 3615, 2" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBA1F759FFB8C80FFD98FE8DFD60FE43" box="[517,641,367,394]" collectionCode="FSJF" pageId="4" pageNumber="139">FSJF 3615</specimenCode>
,
<specimenCode id="DBA1F759FFB8C80FFD16FE8DFD7AFE40" box="[651,667,367,393]" collectionCode="FSJF" pageId="4" pageNumber="139">2</specimenCode>
,
<quantity id="4CFFF2C7FFB8C80FFD3BFE8DFCFEFE43" box="[678,799,367,394]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" metricValueMax="5.4" metricValueMin="4.4" pageId="4" pageNumber="139" unit="mm" value="49.0" valueMax="54.0" valueMin="44.0">4454 mm</quantity>
SL; Georgia:
<location id="8ED809F9FFB8C80FFC27FE8DFB72FE40" LSID="urn:lsid:plazi:treatment:03AEEE34FFB8C800FF0AFF75FE37F895:8ED809F9FFB8C80FFC27FE8DFB72FE40" box="[954,1171,367,393]" country="Georgia" latitude="42.1694" longLatPrecision="6" longitude="42.4314" municipality="Samtredia" name="Tskhenitskali River" pageId="4" pageNumber="139">Tskhenitskali River</location>
upstream of
<collectingMunicipality id="6BDCC558FFB8C80FFAB9FE8DFA79FE40" box="[1316,1432,367,393]" pageId="4" pageNumber="139">Samtredia</collectingMunicipality>
,
<geoCoordinate id="EE3339E5FFB8C80FFF0AFE71FF13FE64" box="[151,242,403,429]" degrees="42.1694" direction="north" orientation="latitude" pageId="4" pageNumber="139" precision="5" value="42.1694">42.1694</geoCoordinate>
<geoCoordinate id="EE3339E5FFB8C80FFF67FE71FEB6FE64" box="[250,343,403,429]" degrees="42.4314" direction="east" orientation="longitude" pageId="4" pageNumber="139" precision="5" value="42.4314">42.4314</geoCoordinate>
</materialsCitation>
.—
<materialsCitation id="3B6F557FFFB8C80FFEEAFE71FCE7FE18" ID-GBIF-Occurrence="3071308313" collectionCode="FSJF" country="Georgia" county="Rioni" latitude="42.16251" location="Gubistskali River" longLatPrecision="1" longitude="42.41069" municipality="Rioni" pageId="4" pageNumber="139" specimenCode="FSJF 4061, 2" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBA1F759FFB8C80FFEEAFE71FE15FE64" box="[375,500,403,429]" collectionCode="FSJF" pageId="4" pageNumber="139">FSJF 4061</specimenCode>
,
<specimenCode id="DBA1F759FFB8C80FFE62FE71FDEEFE64" box="[511,527,403,429]" collectionCode="FSJF" pageId="4" pageNumber="139">2</specimenCode>
,
<quantity id="4CFFF2C7FFB8C80FFD81FE71FD74FE67" box="[540,661,403,430]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" metricValueMax="3.9" metricValueMin="3.2" pageId="4" pageNumber="139" unit="mm" value="35.5" valueMax="39.0" valueMin="32.0">3239 mm</quantity>
SL; Georgia:
<location id="8ED809F9FFB8C80FFCA8FE71FC1BFE64" LSID="urn:lsid:plazi:treatment:03AEEE34FFB8C800FF0AFF75FE37F895:8ED809F9FFB8C80FFCA8FE71FC1BFE64" box="[821,1018,403,429]" country="Georgia" county="Rioni" latitude="42.16251" longLatPrecision="1" longitude="42.41069" municipality="Rioni" name="Gubistskali River" pageId="4" pageNumber="139">Gubistskali River</location>
east of
<location id="8ED809F9FFB8C80FFBC9FE71FB29FE64" LSID="urn:lsid:plazi:treatment:03AEEE34FFB8C800FF0AFF75FE37F895:8ED809F9FFB8C80FFBC9FE71FB29FE64" box="[1108,1224,403,429]" country="Georgia" county="Rioni" latitude="42.16251" longLatPrecision="1" longitude="42.41069" municipality="Rioni" name="Samtredia" pageId="4" pageNumber="139">Samtredia</location>
, upriver of confluence with
<collectingMunicipality id="6BDCC558FFB8C80FFE97FE55FEA9FE18" box="[266,328,439,465]" pageId="4" pageNumber="139">Rioni</collectingMunicipality>
and
<collectingCounty id="62D927AEFFB8C80FFEE2FE55FE5CFE18" box="[383,445,439,465]" pageId="4" pageNumber="139">Rioni</collectingCounty>
channel,
<geoCoordinate id="EE3339E5FFB8C80FFDB4FE55FD73FE1B" box="[553,658,439,466]" degrees="42.16251" direction="north" orientation="latitude" pageId="4" pageNumber="139" precision="1" value="42.16251">42.16251</geoCoordinate>
<geoCoordinate id="EE3339E5FFB8C80FFD04FE55FCE7FE18" box="[665,774,439,465]" degrees="42.41069" direction="east" orientation="longitude" pageId="4" pageNumber="139" precision="1" value="42.41069">42.41069</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB85F22FFB8C80FFF5AFE39FE2BFD3B" blockId="4.[151,1437,259,2014]" pageId="4" pageNumber="139">
<emphasis id="B9738330FFB8C80FFF5AFE39FD2DFE3C" bold="true" box="[199,716,475,501]" pageId="4" pageNumber="139">Material used in molecular genetic analysis.</emphasis>
<materialsCitation id="3B6F557FFFB8C80FFD4FFE3EFC55FE3C" ID-GBIF-Occurrence="3071308306" box="[722,948,475,502]" collectionCode="ZFMK" pageId="4" pageNumber="139" specimenCode="ICH-NB184" specimenCount="1">
<collectionCode id="ED16C7E7FFB8C80FFD4FFE3EFCC1FE3F" box="[722,800,476,502]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="139" type="Museum">ZFMK</collectionCode>
<specimenCode id="DBA1F759FFB8C80FFCBBFE39FC55FE3C" box="[806,948,475,501]" collectionCode="ICH-NB" pageId="4" pageNumber="139">ICH-NB184</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B6F557FFFB8C80FFC23FE3EFC8DFDD3" ID-GBIF-Occurrence="3071308301" accessionNumber="MW564259" collectionCode="MW" country="Georgia" latitude="42.0975" location="GenBank" longLatPrecision="6" longitude="43.39" municipality="Dzirula River" pageId="4" pageNumber="139" specimenCount="2" stateProvince="Tbilisi" typeStatus="holotype">
same data as holotype (
<location id="8ED809F9FFB8C80FFB22FE39FACAFE3C" LSID="urn:lsid:plazi:treatment:03AEEE34FFB8C800FF0AFF75FE37F895:8ED809F9FFB8C80FFB22FE39FACAFE3C" box="[1215,1323,475,501]" name="GenBank" pageId="4" pageNumber="139">GenBank</location>
accession number:
<accessionNumber id="9454C2C1FFB8C80FFF60FE1DFE66FDD3" box="[253,391,511,538]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MW564259" pageId="4" pageNumber="139" type="GenBank">
<collectionCode id="ED16C7E7FFB8C80FFF60FE1DFED3FDD3" box="[253,306,511,538]" country="Netherlands" name="Museum Wasmann" pageId="4" pageNumber="139">MW</collectionCode>
564259
</accessionNumber>
,
<typeStatus id="54BCE180FFB8C80FFE0EFE1DFE14FDD0" box="[403,501,511,537]" pageId="4" pageNumber="139" type="holotype">holotype</typeStatus>
or
<typeStatus id="54BCE180FFB8C80FFD80FDE2FD61FDD3" box="[541,640,512,538]" pageId="4" pageNumber="139" type="paratype">paratype</typeStatus>
, association unclear)
</materialsCitation>
.—
<materialsCitation id="3B6F557FFFB8C80FFC10FE1DFB35FDD0" ID-GBIF-Occurrence="3071308309" box="[909,1236,511,538]" collectionCode="ZFMK" country="Georgia" pageId="4" pageNumber="139" specimenCode="ICH-106137-106138" specimenCount="1" stateProvince="Tbilisi" typeStatus="holotype">
<collectionCode id="ED16C7E7FFB8C80FFC10FE1DFC3CFDD3" box="[909,989,511,538]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="139" type="Museum">ZFMK</collectionCode>
<specimenCode id="DBA1F759FFB8C80FFC78FE1DFB35FDD0" box="[997,1236,511,537]" collectionCode="ICH-" pageId="4" pageNumber="139">ICH-106137-106138</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B6F557FFFB8C80FFB7DFE1DFACAFDF7" ID-GBIF-Occurrence="3071308312" accessionNumber="MW564307, MW564435" collectorName="GenBank" country="Georgia" latitude="42.07829" location="Swlana" longLatPrecision="1" longitude="41.91643" municipality="Jurukveti" pageId="4" pageNumber="139" specimenCount="1" typeStatus="holotype">
Georgia:
<location id="8ED809F9FFB8C80FFAD7FE1DFA7DFDD0" LSID="urn:lsid:plazi:treatment:03AEEE34FFB8C800FF0AFF75FE37F895:8ED809F9FFB8C80FFAD7FE1DFA7DFDD0" box="[1354,1436,511,537]" country="Georgia" latitude="42.07829" longLatPrecision="1" longitude="41.91643" municipality="Jurukveti" name="Swlana" pageId="4" pageNumber="139">Swlana</location>
stream west of
<collectingMunicipality id="6BDCC558FFB8C80FFEDAFDC1FE53FDF4" box="[327,434,547,573]" pageId="4" pageNumber="139">Jurukveti</collectingMunicipality>
,
<geoCoordinate id="EE3339E5FFB8C80FFE22FDC1FDC9FDF4" box="[447,552,547,573]" degrees="42.07829" direction="north" orientation="latitude" pageId="4" pageNumber="139" precision="1" value="42.07829">42.07829</geoCoordinate>
<geoCoordinate id="EE3339E5FFB8C80FFDACFDC1FD7BFDF4" box="[561,666,547,573]" degrees="41.91643" direction="east" orientation="longitude" pageId="4" pageNumber="139" precision="1" value="41.91643">41.91643</geoCoordinate>
(
<collectorName id="26F23AF4FFB8C80FFD37FDC1FCF7FDF4" box="[682,790,547,573]" pageId="4" pageNumber="139">GenBank</collectorName>
accession numbers:
<accessionNumber id="9454C2C1FFB8C80FFB99FDC1FB6CFDF7" box="[1028,1165,547,574]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MW564307" pageId="4" pageNumber="139" type="GenBank">MW564307</accessionNumber>
,
<accessionNumber id="9454C2C1FFB8C80FFB06FDC1FAC5FDF7" box="[1179,1316,547,574]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MW564435" pageId="4" pageNumber="139" type="GenBank">MW564435</accessionNumber>
)
</materialsCitation>
.—
<materialsCitation id="3B6F557FFFB8C80FFAD1FDC1FE67FDA8" ID-GBIF-Occurrence="3071308303" collectionCode="ZFMK" country="Georgia" pageId="4" pageNumber="139" specimenCode="ICH-106139-106140" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED16C7E7FFB8C80FFAD1FDC1FA7DFDF7" box="[1356,1436,547,574]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="139" type="Museum">ZFMK</collectionCode>
<specimenCode id="DBA1F759FFB8C80FFF0AFDA5FE67FDA8" box="[151,390,583,609]" collectionCode="ICH-" pageId="4" pageNumber="139">ICH-106139-106140</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B6F557FFFB8C80FFE13FDA5FBD6FD4F" ID-GBIF-Occurrence="3071308307" accessionNumber="MW564371, MW564381" collectorName="GenBank" country="Georgia" county="Rioni" latitude="42.16251" location="Gubistskali River" longLatPrecision="1" longitude="42.41069" municipality="Rioni" pageId="4" pageNumber="139" specimenCount="1" typeStatus="holotype">
Georgia:
<location id="8ED809F9FFB8C80FFE69FDA5FD56FDA8" LSID="urn:lsid:plazi:treatment:03AEEE34FFB8C800FF0AFF75FE37F895:8ED809F9FFB8C80FFE69FDA5FD56FDA8" box="[500,695,583,609]" country="Georgia" county="Rioni" latitude="42.16251" longLatPrecision="1" longitude="42.41069" municipality="Rioni" name="Gubistskali River" pageId="4" pageNumber="139">Gubistskali River</location>
east of
<location id="8ED809F9FFB8C80FFC9AFDA5FC9AFDA8" LSID="urn:lsid:plazi:treatment:03AEEE34FFB8C800FF0AFF75FE37F895:8ED809F9FFB8C80FFC9AFDA5FC9AFDA8" box="[775,891,583,609]" country="Georgia" county="Rioni" latitude="42.16251" longLatPrecision="1" longitude="42.41069" municipality="Rioni" name="Samtredia" pageId="4" pageNumber="139">Samtredia</location>
, upriver of confluence with
<collectingMunicipality id="6BDCC558FFB8C80FFB36FDA5FB08FDA8" box="[1195,1257,583,609]" pageId="4" pageNumber="139">Rioni</collectingMunicipality>
and
<collectingCounty id="62D927AEFFB8C80FFA86FDA5FAB8FDA8" box="[1307,1369,583,609]" pageId="4" pageNumber="139">Rioni</collectingCounty>
channel,
<geoCoordinate id="EE3339E5FFB8C80FFF55FD89FED0FD4F" box="[200,305,619,646]" degrees="42.16251" direction="north" orientation="latitude" pageId="4" pageNumber="139" precision="1" value="42.16251">42.16251</geoCoordinate>
<geoCoordinate id="EE3339E5FFB8C80FFEA4FD89FE47FD4C" box="[313,422,619,645]" degrees="42.41069" direction="east" orientation="longitude" pageId="4" pageNumber="139" precision="1" value="42.41069">42.41069</geoCoordinate>
. (
<collectorName id="26F23AF4FFB8C80FFE24FD89FDC4FD4C" box="[441,549,619,645]" pageId="4" pageNumber="139">GenBank</collectorName>
accession numbers:
<accessionNumber id="9454C2C1FFB8C80FFC8DFD89FC7BFD4F" box="[784,922,619,646]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MW564371" pageId="4" pageNumber="139" type="GenBank">MW564371</accessionNumber>
,
<accessionNumber id="9454C2C1FFB8C80FFC3BFD89FBCEFD4F" box="[934,1071,619,646]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MW564381" pageId="4" pageNumber="139" type="GenBank">MW564381</accessionNumber>
)
</materialsCitation>
.—
<materialsCitation id="3B6F557FFFB8C80FFBCAFD89FAA0FD4C" ID-GBIF-Occurrence="3071308314" box="[1111,1345,619,646]" collectionCode="ZFMK" country="Georgia" pageId="4" pageNumber="139" specimenCode="ICH-NB190" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED16C7E7FFB8C80FFBCAFD89FB49FD4F" box="[1111,1192,619,646]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="139" type="Museum">ZFMK</collectionCode>
<specimenCode id="DBA1F759FFB8C80FFB2DFD89FAA0FD4C" box="[1200,1345,619,645]" collectionCode="ICH-NB" pageId="4" pageNumber="139">ICH-NB190</specimenCode>
</materialsCitation>
-
<materialsCitation id="3B6F557FFFB8C80FFAD5FD89FE5CFD07" ID-GBIF-Occurrence="3071308304" accessionNumber="MW564373, MW564302, MW564481" collectorName="GenBank" country="Georgia" latitude="42.1694" location="Tskhenitskali River" longLatPrecision="6" longitude="42.4314" municipality="Samtredia" pageId="4" pageNumber="139" specimenCount="1" typeStatus="holotype">
NB191; Georgia:
<location id="8ED809F9FFB8C80FFE9DFD6DFE3AFD60" LSID="urn:lsid:plazi:treatment:03AEEE34FFB8C800FF0AFF75FE37F895:8ED809F9FFB8C80FFE9DFD6DFE3AFD60" box="[256,475,655,681]" country="Georgia" latitude="42.1694" longLatPrecision="6" longitude="42.4314" municipality="Samtredia" name="Tskhenitskali River" pageId="4" pageNumber="139">Tskhenitskali River</location>
upstream of
<collectingMunicipality id="6BDCC558FFB8C80FFDECFD6DFD04FD60" box="[625,741,655,681]" pageId="4" pageNumber="139">Samtredia</collectingMunicipality>
,
<geoCoordinate id="EE3339E5FFB8C80FFD6CFD6DFCADFD60" box="[753,844,655,681]" degrees="42.1694" direction="north" orientation="latitude" pageId="4" pageNumber="139" precision="5" value="42.1694">42.1694</geoCoordinate>
<geoCoordinate id="EE3339E5FFB8C80FFCC9FD6DFC4EFD60" box="[852,943,655,681]" degrees="42.4314" direction="east" orientation="longitude" pageId="4" pageNumber="139" precision="5" value="42.4314">42.4314</geoCoordinate>
(
<collectorName id="26F23AF4FFB8C80FFC22FD6DFBCAFD60" box="[959,1067,655,681]" pageId="4" pageNumber="139">GenBank</collectorName>
accession numbers:
<accessionNumber id="9454C2C1FFB8C80FFA90FD6DFA79FD63" box="[1293,1432,655,682]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MW564373" pageId="4" pageNumber="139" type="GenBank">MW564373</accessionNumber>
,
<accessionNumber id="9454C2C1FFB8C80FFF0AFD51FEC3FD07" box="[151,290,691,718]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MW564302" pageId="4" pageNumber="139" type="GenBank">MW564302</accessionNumber>
,
<accessionNumber id="9454C2C1FFB8C80FFEB1FD51FE54FD07" box="[300,437,691,718]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MW564481" pageId="4" pageNumber="139" type="GenBank">MW564481</accessionNumber>
)
</materialsCitation>
.—
<materialsCitation id="3B6F557FFFB8C80FFE40FD51FD23FD07" ID-GBIF-Occurrence="3071308308" box="[477,706,691,718]" collectionCode="ZFMK" country="Georgia" pageId="4" pageNumber="139" specimenCode="ICH-NB195" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED16C7E7FFB8C80FFE40FD51FDCFFD07" box="[477,558,691,718]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="139" type="Museum">ZFMK</collectionCode>
<specimenCode id="DBA1F759FFB8C80FFDA9FD51FD23FD07" box="[564,706,691,718]" collectionCode="ICH-NB" pageId="4" pageNumber="139">ICH-NB195</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B6F557FFFB8C80FFD50FD51FE27FD3B" ID-GBIF-Occurrence="3071308315" accessionNumber="MW564442" collectionCode="MW" collectorName="GenBank" country="Georgia" latitude="42.6617" location="Enguri River" longLatPrecision="6" longitude="41.9911" municipality="Lia" pageId="4" pageNumber="139" specimenCount="1" typeStatus="holotype">
Georgia:
<location id="8ED809F9FFB8C80FFCA8FD51FC27FD04" LSID="urn:lsid:plazi:treatment:03AEEE34FFB8C800FF0AFF75FE37F895:8ED809F9FFB8C80FFCA8FD51FC27FD04" box="[821,966,691,717]" country="Georgia" latitude="42.6617" longLatPrecision="6" longitude="41.9911" municipality="Lia" name="Enguri River" pageId="4" pageNumber="139">Enguri River</location>
at
<collectingMunicipality id="6BDCC558FFB8C80FFC7AFD51FBEEFD04" box="[999,1039,691,717]" pageId="4" pageNumber="139">Lia</collectingMunicipality>
,
<geoCoordinate id="EE3339E5FFB8C80FFB87FD51FB94FD04" box="[1050,1141,691,717]" degrees="42.6617" direction="north" orientation="latitude" pageId="4" pageNumber="139" precision="5" value="42.6617">42.6617</geoCoordinate>
<geoCoordinate id="EE3339E5FFB8C80FFBE1FD51FB37FD04" box="[1148,1238,691,717]" degrees="41.9911" direction="east" orientation="longitude" pageId="4" pageNumber="139" precision="5" value="41.9911">41.9911</geoCoordinate>
(
<collectorName id="26F23AF4FFB8C80FFB79FD51FAB1FD04" box="[1252,1360,691,717]" pageId="4" pageNumber="139">GenBank</collectorName>
accession number:
<accessionNumber id="9454C2C1FFB8C80FFEACFD35FE5FFD3B" box="[305,446,727,754]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MW564442" pageId="4" pageNumber="139" type="GenBank">
<collectionCode id="ED16C7E7FFB8C80FFEACFD35FE86FD3B" box="[305,359,727,754]" country="Netherlands" name="Museum Wasmann" pageId="4" pageNumber="139">MW</collectionCode>
564442
</accessionNumber>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31D0CA9FFB8C80FFF5AFD19FECCFAB3" pageId="4" pageNumber="139" type="diagnosis">
<paragraph id="8BB85F22FFB8C80FFF5AFD19FE9DFC27" blockId="4.[151,1437,259,2014]" pageId="4" pageNumber="139">
<emphasis id="B9738330FFB8C80FFF5AFD19FEA3FCDC" bold="true" box="[199,322,763,789]" pageId="4" pageNumber="139">Diagnosis.</emphasis>
<taxonomicName id="4C0724A1FFB8C80FFED7FD19FD9DFCDC" authorityName="Freyhof &amp; Kaya &amp; Epitashvili &amp; Geiger" authorityYear="2021" box="[330,636,763,789]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="phasicus">
<emphasis id="B9738330FFB8C80FFED7FD19FD9DFCDC" box="[330,636,763,789]" italics="true" pageId="4" pageNumber="139">Oxynoemacheilus phasicus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C0724A1FFB8C80FFCE1FD1EFC0AFCDC" box="[892,1003,763,790]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="cemali">
<emphasis id="B9738330FFB8C80FFCE1FD1EFC0AFCDC" box="[892,1003,763,790]" italics="true" pageId="4" pageNumber="139">O. cemali</emphasis>
</taxonomicName>
, the only other species of
<taxonomicName id="4C0724A1FFB8C80FFA86FD1EFF07FCF0" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="genus">
<emphasis id="B9738330FFB8C80FFA86FD1EFF07FCF0" italics="true" pageId="4" pageNumber="139">Oxynoemacheilus</emphasis>
</taxonomicName>
known from the eastern Black Sea basin, by a slightly emarginate caudal fin (vs. deeply emarginate), pelvic fin not reaching to origin of genital papillae (vs. reaching to or beyond posterior tip of genital papillae), pelvic-fin origin below first branched dorsal-fin ray (vs. below second or third ray), a wide band in posterior half of caudal fin (vs. absent), and a mottled or marbled flank pattern (juveniles with bars on flank behind dorsal-fin base) (vs. usually with 915 narrow, irregularly shaped and spaced, dark-grey bars or vertically elongate blotches on flank posterior to dorsal-fin origin).
</paragraph>
<paragraph id="8BB85F22FFB8C80FFF5AFC15FECCFAB3" blockId="4.[151,1437,259,2014]" pageId="4" pageNumber="139">
<taxonomicName id="4C0724A1FFB8C80FFF5AFC15FE1DFBD8" authorityName="Freyhof &amp; Kaya &amp; Epitashvili &amp; Geiger" authorityYear="2021" box="[199,508,1015,1041]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="phasicus">
<emphasis id="B9738330FFB8C80FFF5AFC15FE1DFBD8" box="[199,508,1015,1041]" italics="true" pageId="4" pageNumber="139">Oxynoemacheilus phasicus</emphasis>
</taxonomicName>
is distinguished from the other members of the
<taxonomicName id="4C0724A1FFB8C80FFBDCFC1AFB23FBD8" box="[1089,1218,1015,1042]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB8C80FFBDCFC1AFB23FBD8" box="[1089,1218,1015,1042]" italics="true" pageId="4" pageNumber="139">O. brandtii</emphasis>
</taxonomicName>
species group (
<taxonomicName id="4C0724A1FFB8C80FFA1CFC1AFF10FBFC" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB8C80FFA1CFC1AFF10FBFC" italics="true" pageId="4" pageNumber="139">O. brandtii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0724A1FFB8C80FFE9EFBFEFE83FBFC" box="[259,354,1051,1078]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB8C80FFE9EFBFEFE83FBFC" box="[259,354,1051,1078]" italics="true" pageId="4" pageNumber="139">O. elsae</emphasis>
</taxonomicName>
, and
<emphasis id="B9738330FFB8C80FFE3BFBFEFDFAFBFC" box="[422,539,1051,1078]" italics="true" pageId="4" pageNumber="139">
<taxonomicName id="4C0724A1FFB8C80FFE3BFBFEFDF2FBFC" authorityName="FSJF" authorityYear="1561" box="[422,531,1051,1078]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="merga">O. merga</taxonomicName>
)
</emphasis>
by having a slightly emarginate caudal fin (vs. forked in
<taxonomicName id="4C0724A1FFB8C80FFB5CFBFEFAA1FBFC" box="[1217,1344,1051,1078]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB8C80FFB5CFBFEFAA1FBFC" box="[1217,1344,1051,1078]" italics="true" pageId="4" pageNumber="139">O. brandtii</emphasis>
</taxonomicName>
, deeply emarginate in
<taxonomicName id="4C0724A1FFB8C80FFEA8FBA2FE70FB90" box="[309,401,1087,1114]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB8C80FFEA8FBA2FE70FB90" box="[309,401,1087,1114]" italics="true" pageId="4" pageNumber="139">O. elsae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0724A1FFB8C80FFE5BFBA2FDCFFB90" authorityName="FSJF" authorityYear="1561" box="[454,558,1088,1114]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="merga">
<emphasis id="B9738330FFB8C80FFE5BFBA2FDCFFB90" box="[454,558,1088,1114]" italics="true" pageId="4" pageNumber="139">O. merga</emphasis>
</taxonomicName>
), pelvic fin not reaching to origin of genital papillae (vs. reaching to or beyond anus in
<taxonomicName id="4C0724A1FFB8C80FFF72FB86FE8AFBB4" box="[239,363,1123,1150]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB8C80FFF72FB86FE8AFBB4" box="[239,363,1123,1150]" italics="true" pageId="4" pageNumber="139">O. brandtii</emphasis>
</taxonomicName>
and many
<taxonomicName id="4C0724A1FFB8C80FFE7AFB86FDA5FBB4" box="[487,580,1123,1150]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB8C80FFE7AFB86FDA5FBB4" box="[487,580,1123,1150]" italics="true" pageId="4" pageNumber="139">O. elsae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0724A1FFB8C80FFDE7FB86FD03FBB4" authorityName="FSJF" authorityYear="1561" box="[634,738,1124,1150]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="merga">
<emphasis id="B9738330FFB8C80FFDE7FB86FD03FBB4" box="[634,738,1124,1150]" italics="true" pageId="4" pageNumber="139">O. merga</emphasis>
</taxonomicName>
), caudal peduncle 1.52.1 times longer than deep (vs.
<quantity id="4CFFF2C7FFB8C80FFAD5FB81FF4CFB68" metricMagnitude="-2" metricUnit="m" metricValue="7.239" metricValueMax="8.128" metricValueMin="6.35" pageId="4" pageNumber="139" unit="in" value="2.85" valueMax="3.2" valueMin="2.5">2.53.2 in</quantity>
<taxonomicName id="4C0724A1FFB8C80FFF28FB6AFED3FB68" box="[181,306,1159,1186]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB8C80FFF28FB6AFED3FB68" box="[181,306,1159,1186]" italics="true" pageId="4" pageNumber="139">O. brandtii</emphasis>
</taxonomicName>
,
<quantity id="4CFFF2C7FFB8C80FFEDFFB65FE55FB68" box="[322,436,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="6.603999999999999" metricValueMax="7.366" metricValueMin="5.842" pageId="4" pageNumber="139" unit="in" value="2.5999999999999996" valueMax="2.9" valueMin="2.3">2.32.9 in</quantity>
<taxonomicName id="4C0724A1FFB8C80FFE26FB6AFDF8FB68" box="[443,537,1159,1186]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB8C80FFE26FB6AFDF8FB68" box="[443,537,1159,1186]" italics="true" pageId="4" pageNumber="139">O. elsae</emphasis>
</taxonomicName>
, 2.0
<quantity id="4CFFF2C7FFB8C80FFDC4FB65FD78FB68" box="[601,665,1159,1186]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" pageId="4" pageNumber="139" unit="in" value="2.5">2.5 in</quantity>
<taxonomicName id="4C0724A1FFB8C80FFD3CFB6AFCEAFB68" authorityName="FSJF" authorityYear="1561" box="[673,779,1160,1186]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="merga">
<emphasis id="B9738330FFB8C80FFD3CFB6AFCEAFB68" box="[673,779,1160,1186]" italics="true" pageId="4" pageNumber="139">O. merga</emphasis>
</taxonomicName>
), and a mottled or marbled flank pattern (vs. usually with irregularly shaped and spaced, dark-brown bars or vertically elongate blotches on flank posterior to dorsal-fin origin, much wider than interspaces, often fused into a plain brown pattern on flank in front of dorsal-fin base in
<taxonomicName id="4C0724A1FFB8C80FFA1CFB32FF10FAC4" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB8C80FFA1CFB32FF10FAC4" italics="true" pageId="4" pageNumber="139">O. brandtii</emphasis>
</taxonomicName>
; usually with irregularly shaped and spaced, rarely regularly shaped and spaced, dark-brown bars, often split in middle, or flank almost plain brown in
<taxonomicName id="4C0724A1FFB8C80FFD33FAFAFCEAFAF8" box="[686,779,1303,1330]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB8C80FFD33FAFAFCEAFAF8" box="[686,779,1303,1330]" italics="true" pageId="4" pageNumber="139">O. elsae</emphasis>
</taxonomicName>
; with many, narrow, pale-brown bars, usually much more narrow than interspaces, often dissociated in vertically elongate blotches on flank and dorsal saddles, rarely mottled in
<taxonomicName id="4C0724A1FFB8C80FFF29FA82FEFCFAB0" authorityName="FSJF" authorityYear="1561" box="[180,285,1376,1402]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="merga">
<emphasis id="B9738330FFB8C80FFF29FA82FEFCFAB0" box="[180,285,1376,1402]" italics="true" pageId="4" pageNumber="139">O. merga</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C31D0CA9FFB8C80DFF5AFA61FDC8FEAC" lastPageId="6" lastPageNumber="141" pageId="4" pageNumber="139" type="description">
<paragraph id="8BB85F22FFB8C80FFF5AFA61FBCCF928" blockId="4.[151,1437,259,2014]" pageId="4" pageNumber="139">
<emphasis id="B9738330FFB8C80FFF5AFA61FEB8FA54" bold="true" box="[199,345,1411,1437]" pageId="4" pageNumber="139">Description.</emphasis>
See
<figureCitation id="133C43A7FFB8C80FFE0FFA61FDF8FA57" box="[402,537,1411,1438]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="6.[151,250,1384,1408]" captionStartId-1="7.[151,250,1244,1268]" captionStartId-2="8.[151,250,1012,1036]" captionStartId-3="8.[151,250,1916,1940]" captionTargetBox-0="[152,1436,381,1361]" captionTargetBox-1="[151,1436,381,1221]" captionTargetBox-2="[151,1436,181,988]" captionTargetBox-3="[151,1436,1133,1893]" captionTargetId-0="figure-123@6.[152,1436,381,1361]" captionTargetId-1="figure-129@7.[151,1436,381,1221]" captionTargetId-2="figure-25@8.[151,1436,181,988]" captionTargetId-3="figure-82@8.[151,1436,1133,1893]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="8" captionText-0="FIGURE 2. Oxynoemacheilus phasicus, ZFMK ICH-124180, holotype, 74.1 mm SL; Georgia: Dzirula River." captionText-1="FIGURE 3. Oxynoemacheilus phasicus, paratypes, from top, left to right: FSJF 4061, 32 mm SL; 39 mm SL; Georgia: Gubistskali River; FSJF 3615, 44 mm SL, 54 mm SL; Georgia: Tskhenitskali River; FSJF 3576, 59 mm SL; Georgia: Enguri River; FSJF 3592, 77 mm SL; Georgia: Dzirula River." captionText-2="FIGURE 4. Oxynoemacheilus phasicus, paratypes, from top, left to right: FSJF 4061, 32 mm SL; 39 mm SL; Georgia: Gubistskali River; FSJF 3615, 44 mm SL, 54 mm SL; Georgia: Tskhenitskali River; FSJF 3576, 59 mm SL; Georgia: Enguri River; FSJF 3592, 77 mm SL; Georgia: Dzirula River." captionText-3="FIGURE 5. Oxynoemacheilus phasicus, paratypes, from top, left to right: FSJF 4061, 32 mm SL; 39 mm SL; Georgia: Gubistskali River; FSJF 3615, 44 mm SL, 54 mm SL; Georgia: Tskhenitskali River; FSJF 3576, 59 mm SL; Georgia: Enguri River; FSJF 3592, 77 mm SL; Georgia: Dzirula River." figureDoi-0="http://doi.org/10.5281/zenodo.4671920" figureDoi-1="http://doi.org/10.5281/zenodo.4671922" figureDoi-2="http://doi.org/10.5281/zenodo.4671926" figureDoi-3="http://doi.org/10.5281/zenodo.4671930" httpUri-0="https://zenodo.org/record/4671920/files/figure.png" httpUri-1="https://zenodo.org/record/4671922/files/figure.png" httpUri-2="https://zenodo.org/record/4671926/files/figure.png" httpUri-3="https://zenodo.org/record/4671930/files/figure.png" pageId="4" pageNumber="139">Figures 25</figureCitation>
for general appearance and
<tableCitation id="C6856A99FFB8C80FFCC0FA61FC53FA54" box="[861,946,1411,1437]" captionStart="TABLE 2" captionStartId="5.[151,244,474,500]" captionTargetPageId="5" captionText="TABLE 2. Morphometric data of Oxynoemacheilus phasicus (holotype ZFMK ICH-124180 and paratypes FSJF 3576, 3591, 3615, n=7). Holotype is included in calculations." httpUri="http://table.plazi.org/id/DF780FAAFFB9C80EFF0AFE38FCA6FDD0" pageId="4" pageNumber="139" tableUuid="DF780FAAFFB9C80EFF0AFE38FCA6FDD0">Table 2</tableCitation>
for morphometric data. Medium-sized and moderately slender species. Body deepest at about midline between nape and dorsal-fin origin. Body width greatest at pectoral-fin base, at cheek in largest individual. Section of head roundish, flattened on ventral surface, slightly convex in interorbital space, convex on snout. Snout blunt. Caudal peduncle compressed laterally, 1.52.1 times longer than deep. Pelvic axillary lobe present, its tip attached to flank. Pelvic-fin origin below first branched dorsalfin ray. Anal-fin origin at about vertical of midline between dorsal and caudal-fin origins. Pectoral fin reaching to approximately 5070% of distance from pectoral-fin origin to pelvic-fin origin. Pelvic fin not reaching anus; reaching slightly anterior to vertical of tip of last dorsal-fin ray; almost reaching anus in juveniles smaller than
<quantity id="4CFFF2C7FFB8C80FFABCF99DFA8EF953" box="[1313,1391,1663,1690]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="139" unit="mm" value="40.0">40 mm</quantity>
SL. Anus about 3070% of an eye diameter anterior to anal-fin origin. Anal fin not reaching caudal-fin base. No dorsal or ventral adipose crest on caudal peduncle. Largest known individual
<quantity id="4CFFF2C7FFB8C80FFC32F92AFC1FF92B" box="[943,1022,1736,1762]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="4" pageNumber="139" unit="mm" value="77.0">77 mm</quantity>
SL.
</paragraph>
<paragraph id="8BB85F22FFB8C80EFF5AF909FE68FE64" blockId="4.[151,1437,259,2014]" lastBlockId="5.[151,1437,151,429]" lastPageId="5" lastPageNumber="140" pageId="4" pageNumber="139">
Dorsal fin with 8½9½ branched rays, outer margin straight or convex. Anal fin with 5½ branched rays, outer margin straight or slightly concave. Pectoral fin with 911 branched rays, outer margin straight. Pelvic fin with 7 branched rays, outer margin straight or slightly convex. Caudal fin slightly emarginate with 8+8 branched rays, moderately emarginate in juveniles smaller than
<quantity id="4CFFF2C7FFB8C80FFD23F8B5FCEFF8BB" box="[702,782,1879,1906]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="139" unit="mm" value="40.0">40 mm</quantity>
SL. Flank, back and abdomen densely covered by scales. Lateral line complete, terminating at caudal-fin base. Anterior nostril opening at end of a low, ovoid and flap-like tube. Posterior tip of anterior nostril overlapping in small individuals, not overlapping posterior nostril when folded backwards in large individuals. One central pore, two in one individual, and one lateral pore on each side of supra- temporal head canal, 810 pores in anterior infraorbital canal, 34 pores in posterior infraorbital canal, 69 pores in supraorbital canal and 57 pores in mandibular canal. A short suborbital groove in male. Mouth small, arched. Lips thick without furrows, lower lip thicker than upper lip. A median interruption in lower lip. Upper lip with a small and short median incision in some individuals, absent in others. Processus dentiformis narrow, very small and rounded. Lower jaw rounded, without median notch. Barbels long; inner rostral barbel reaching to or beyond base of maxillary barbel, outer reaching beyond anterior margin of eye, to middle of eye in some individuals. Maxillary barbel reaching beyond vertical through middle of eye or to or beyond posterior margin of eye. Barbels very short in largest individuals.
</paragraph>
<caption id="DF780FAAFFB9C80EFF0AFE38FCA6FDD0" ID-Table-UUID="DF780FAAFFB9C80EFF0AFE38FCA6FDD0" httpUri="http://table.plazi.org/id/DF780FAAFFB9C80EFF0AFE38FCA6FDD0" pageId="5" pageNumber="140" startId="5.[151,244,474,500]" targetBox="[166,1394,552,1744]" targetIsTable="true" targetPageId="5">
<paragraph id="8BB85F22FFB9C80EFF0AFE38FCA6FDD0" blockId="5.[151,1437,474,537]" pageId="5" pageNumber="140">
<emphasis id="B9738330FFB9C80EFF0AFE38FEF2FE3D" bold="true" box="[151,275,474,500]" pageId="5" pageNumber="140">TABLE 2.</emphasis>
Morphometric data of
<taxonomicName id="4C0724A1FFB9C80EFD82FE39FCB0FE3C" authorityName="Freyhof &amp; Kaya &amp; Epitashvili &amp; Geiger" authorityYear="2021" box="[543,849,475,501]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="140" phylum="Chordata" rank="species" species="phasicus">
<emphasis id="B9738330FFB9C80EFD82FE39FCB0FE3C" box="[543,849,475,501]" italics="true" pageId="5" pageNumber="140">Oxynoemacheilus phasicus</emphasis>
</taxonomicName>
(holotype ZFMK ICH-124180 and paratypes FSJF 3576, 3591, 3615, n=7). Holotype is included in calculations.
</paragraph>
</caption>
<paragraph id="8BB85F22FFB9C80EFCF3FDCAFA93F919" pageId="5" pageNumber="140">
<table id="F907AD82FFB937F4FF3BFDCAFA93F919" box="[166,1394,552,1744]" gridcols="6" gridrows="30" pageId="5" pageNumber="140">
<tr id="35375D60FFB937F4FF3BFDCAFA93FD89" box="[166,1394,552,576]" gridrow="0" pageId="5" pageNumber="140" rowspan-0="1" rowspan-5="1">
<th id="76E6341CFFB937F4FCF3FDCAFC2AFD89" box="[878,971,552,576]" gridcol="1" gridrow="0" pageId="5" pageNumber="140">
<emphasis id="B9738330FFB9C80EFCF3FDCAFC2AFD89" bold="true" box="[878,971,552,576]" pageId="5" pageNumber="140">holotype</emphasis>
</th>
<th id="76E6341CFFB937F4FB93FDCAFAD6FD89" box="[1038,1335,552,576]" colspan="3" colspanRight="2" gridcol="2" gridrow="0" pageId="5" pageNumber="140">
<emphasis id="B9738330FFB9C80EFBD3FDCAFAD6FD89" bold="true" box="[1102,1335,552,576]" pageId="5" pageNumber="140">holotype &amp; paratypes</emphasis>
</th>
</tr>
<tr id="35375D60FFB937F4FF3BFDB3FA93FDA0" box="[166,1394,593,617]" gridrow="1" pageId="5" pageNumber="140" rowspan-0="1" rowspan-1="1">
<td id="76E6341CFFB937F4FB93FDB3FBA4FDA0" box="[1038,1093,593,617]" gridcol="2" gridrow="1" pageId="5" pageNumber="140">mean</td>
<td id="76E6341CFFB937F4FB1CFDB3FB4CFDA0" box="[1153,1197,593,617]" gridcol="3" gridrow="1" pageId="5" pageNumber="140">min</td>
<td id="76E6341CFFB937F4FB78FDB3FAD6FDA0" box="[1253,1335,593,617]" gridcol="4" gridrow="1" pageId="5" pageNumber="140">max</td>
<td id="76E6341CFFB937F4FACFFDB3FA93FDA0" box="[1362,1394,593,617]" gridcol="5" gridrow="1" pageId="5" pageNumber="140">SD</td>
</tr>
<tr id="35375D60FFB937F4FF3BFD9BFA93FD58" box="[166,1394,633,657]" gridrow="2" pageId="5" pageNumber="140" rowspan-2="1" rowspan-5="1">
<th id="76E6341CFFB937F4FF3BFD9BFD78FD58" box="[166,665,633,657]" gridcol="0" gridrow="2" pageId="5" pageNumber="140">Standard length (mm)</th>
<td id="76E6341CFFB937F4FCF3FD9BFC2AFD58" box="[878,971,633,657]" gridcol="1" gridrow="2" pageId="5" pageNumber="140">74.1</td>
<td id="76E6341CFFB937F4FB1CFD9BFB4CFD58" box="[1153,1197,633,657]" gridcol="3" gridrow="2" pageId="5" pageNumber="140">44.4</td>
<td id="76E6341CFFB937F4FB78FD9BFAD6FD58" box="[1253,1335,633,657]" gridcol="4" gridrow="2" pageId="5" pageNumber="140">77.0</td>
</tr>
<tr id="35375D60FFB937F4FF3BFD43FA93FD70" box="[166,1394,673,697]" gridrow="3" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFD43FA93FD70" box="[166,1394,673,697]" colspan="6" colspanRight="5" gridcol="0" gridrow="3" pageId="5" pageNumber="140">
<emphasis id="B9738330FFB9C80EFF3BFD43FE03FD70" bold="true" box="[166,482,673,697]" pageId="5" pageNumber="140">In percent of standard length</emphasis>
</th>
</tr>
<tr id="35375D60FFB937F4FF3BFD28FA93FD2B" box="[166,1394,714,738]" gridrow="4" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFD28FD78FD2B" box="[166,665,714,738]" gridcol="0" gridrow="4" pageId="5" pageNumber="140">Head length</th>
<td id="76E6341CFFB937F4FCF3FD28FC2AFD2B" box="[878,971,714,738]" gridcol="1" gridrow="4" pageId="5" pageNumber="140">22.3</td>
<td id="76E6341CFFB937F4FB93FD28FBA4FD2B" box="[1038,1093,714,738]" gridcol="2" gridrow="4" pageId="5" pageNumber="140">22.7</td>
<td id="76E6341CFFB937F4FB1CFD28FB4CFD2B" box="[1153,1197,714,738]" gridcol="3" gridrow="4" pageId="5" pageNumber="140">22.1</td>
<td id="76E6341CFFB937F4FB78FD28FAD6FD2B" box="[1253,1335,714,738]" gridcol="4" gridrow="4" pageId="5" pageNumber="140">23.8</td>
<td id="76E6341CFFB937F4FACFFD28FA93FD2B" box="[1362,1394,714,738]" gridcol="5" gridrow="4" pageId="5" pageNumber="140">0.6</td>
</tr>
<tr id="35375D60FFB937F4FF3BFD10FA93FCC3" box="[166,1394,754,778]" gridrow="5" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFD10FD78FCC3" box="[166,665,754,778]" gridcol="0" gridrow="5" pageId="5" pageNumber="140">Body depth at dorsal-fin origin</th>
<td id="76E6341CFFB937F4FCF3FD10FC2AFCC3" box="[878,971,754,778]" gridcol="1" gridrow="5" pageId="5" pageNumber="140">15.9</td>
<td id="76E6341CFFB937F4FB93FD10FBA4FCC3" box="[1038,1093,754,778]" gridcol="2" gridrow="5" pageId="5" pageNumber="140">15.6</td>
<td id="76E6341CFFB937F4FB1CFD10FB4CFCC3" box="[1153,1197,754,778]" gridcol="3" gridrow="5" pageId="5" pageNumber="140">13.8</td>
<td id="76E6341CFFB937F4FB78FD10FAD6FCC3" box="[1253,1335,754,778]" gridcol="4" gridrow="5" pageId="5" pageNumber="140">17.7</td>
<td id="76E6341CFFB937F4FACFFD10FA93FCC3" box="[1362,1394,754,778]" gridcol="5" gridrow="5" pageId="5" pageNumber="140">1.5</td>
</tr>
<tr id="35375D60FFB937F4FF3BFCF8FA93FCFB" box="[166,1394,794,818]" gridrow="6" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFCF8FD78FCFB" box="[166,665,794,818]" gridcol="0" gridrow="6" pageId="5" pageNumber="140">Body width at dorsal-fin origin</th>
<td id="76E6341CFFB937F4FCF3FCF8FC2AFCFB" box="[878,971,794,818]" gridcol="1" gridrow="6" pageId="5" pageNumber="140">12.9</td>
<td id="76E6341CFFB937F4FB93FCF8FBA4FCFB" box="[1038,1093,794,818]" gridcol="2" gridrow="6" pageId="5" pageNumber="140">12.5</td>
<td id="76E6341CFFB937F4FB1CFCF8FB4CFCFB" box="[1153,1197,794,818]" gridcol="3" gridrow="6" pageId="5" pageNumber="140">11.4</td>
<td id="76E6341CFFB937F4FB78FCF8FAD6FCFB" box="[1253,1335,794,818]" gridcol="4" gridrow="6" pageId="5" pageNumber="140">13.5</td>
<td id="76E6341CFFB937F4FACFFCF8FA93FCFB" box="[1362,1394,794,818]" gridcol="5" gridrow="6" pageId="5" pageNumber="140">0.9</td>
</tr>
<tr id="35375D60FFB937F4FF3BFCA0FA93FC93" box="[166,1394,834,858]" gridrow="7" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFCA0FD78FC93" box="[166,665,834,858]" gridcol="0" gridrow="7" pageId="5" pageNumber="140">Predorsal length</th>
<td id="76E6341CFFB937F4FCF3FCA0FC2AFC93" box="[878,971,834,858]" gridcol="1" gridrow="7" pageId="5" pageNumber="140">49.8</td>
<td id="76E6341CFFB937F4FB93FCA0FBA4FC93" box="[1038,1093,834,858]" gridcol="2" gridrow="7" pageId="5" pageNumber="140">50.0</td>
<td id="76E6341CFFB937F4FB1CFCA0FB4CFC93" box="[1153,1197,834,858]" gridcol="3" gridrow="7" pageId="5" pageNumber="140">48.8</td>
<td id="76E6341CFFB937F4FB78FCA0FAD6FC93" box="[1253,1335,834,858]" gridcol="4" gridrow="7" pageId="5" pageNumber="140">51.0</td>
<td id="76E6341CFFB937F4FACFFCA0FA93FC93" box="[1362,1394,834,858]" gridcol="5" gridrow="7" pageId="5" pageNumber="140">0.7</td>
</tr>
<tr id="35375D60FFB937F4FF3BFC89FA93FC4A" box="[166,1394,875,899]" gridrow="8" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFC89FD78FC4A" box="[166,665,875,899]" gridcol="0" gridrow="8" pageId="5" pageNumber="140">Postdorsal length</th>
<td id="76E6341CFFB937F4FCF3FC89FC2AFC4A" box="[878,971,875,899]" gridcol="1" gridrow="8" pageId="5" pageNumber="140">39.9</td>
<td id="76E6341CFFB937F4FB93FC89FBA4FC4A" box="[1038,1093,875,899]" gridcol="2" gridrow="8" pageId="5" pageNumber="140">40.1</td>
<td id="76E6341CFFB937F4FB1CFC89FB4CFC4A" box="[1153,1197,875,899]" gridcol="3" gridrow="8" pageId="5" pageNumber="140">39.1</td>
<td id="76E6341CFFB937F4FB78FC89FAD6FC4A" box="[1253,1335,875,899]" gridcol="4" gridrow="8" pageId="5" pageNumber="140">41.0</td>
<td id="76E6341CFFB937F4FACFFC89FA93FC4A" box="[1362,1394,875,899]" gridcol="5" gridrow="8" pageId="5" pageNumber="140">0.8</td>
</tr>
<tr id="35375D60FFB937F4FF3BFC71FA93FC62" box="[166,1394,915,939]" gridrow="9" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFC71FD78FC62" box="[166,665,915,939]" gridcol="0" gridrow="9" pageId="5" pageNumber="140">Preanal length</th>
<td id="76E6341CFFB937F4FCF3FC71FC2AFC62" box="[878,971,915,939]" gridcol="1" gridrow="9" pageId="5" pageNumber="140">74.1</td>
<td id="76E6341CFFB937F4FB93FC71FBA4FC62" box="[1038,1093,915,939]" gridcol="2" gridrow="9" pageId="5" pageNumber="140">74.0</td>
<td id="76E6341CFFB937F4FB1CFC71FB4CFC62" box="[1153,1197,915,939]" gridcol="3" gridrow="9" pageId="5" pageNumber="140">72.2</td>
<td id="76E6341CFFB937F4FB78FC71FAD6FC62" box="[1253,1335,915,939]" gridcol="4" gridrow="9" pageId="5" pageNumber="140">76.1</td>
<td id="76E6341CFFB937F4FACFFC71FA93FC62" box="[1362,1394,915,939]" gridcol="5" gridrow="9" pageId="5" pageNumber="140">1.4</td>
</tr>
<tr id="35375D60FFB937F4FF3BFC59FA93FC1A" box="[166,1394,955,979]" gridrow="10" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFC59FD78FC1A" box="[166,665,955,979]" gridcol="0" gridrow="10" pageId="5" pageNumber="140">Prepelvic length</th>
<td id="76E6341CFFB937F4FCF3FC59FC2AFC1A" box="[878,971,955,979]" gridcol="1" gridrow="10" pageId="5" pageNumber="140">50.9</td>
<td id="76E6341CFFB937F4FB93FC59FBA4FC1A" box="[1038,1093,955,979]" gridcol="2" gridrow="10" pageId="5" pageNumber="140">50.7</td>
<td id="76E6341CFFB937F4FB1CFC59FB4CFC1A" box="[1153,1197,955,979]" gridcol="3" gridrow="10" pageId="5" pageNumber="140">49.9</td>
<td id="76E6341CFFB937F4FB78FC59FAD6FC1A" box="[1253,1335,955,979]" gridcol="4" gridrow="10" pageId="5" pageNumber="140">51.4</td>
<td id="76E6341CFFB937F4FACFFC59FA93FC1A" box="[1362,1394,955,979]" gridcol="5" gridrow="10" pageId="5" pageNumber="140">0.6</td>
</tr>
<tr id="35375D60FFB937F4FF3BFC01FA93FC32" box="[166,1394,995,1019]" gridrow="11" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFC01FD78FC32" box="[166,665,995,1019]" gridcol="0" gridrow="11" pageId="5" pageNumber="140">Distance between pectoral- and pelvic-fin origins</th>
<td id="76E6341CFFB937F4FCF3FC01FC2AFC32" box="[878,971,995,1019]" gridcol="1" gridrow="11" pageId="5" pageNumber="140">29.0</td>
<td id="76E6341CFFB937F4FB93FC01FBA4FC32" box="[1038,1093,995,1019]" gridcol="2" gridrow="11" pageId="5" pageNumber="140">29.0</td>
<td id="76E6341CFFB937F4FB1CFC01FB4CFC32" box="[1153,1197,995,1019]" gridcol="3" gridrow="11" pageId="5" pageNumber="140">28.5</td>
<td id="76E6341CFFB937F4FB78FC01FAD6FC32" box="[1253,1335,995,1019]" gridcol="4" gridrow="11" pageId="5" pageNumber="140">29.4</td>
<td id="76E6341CFFB937F4FACFFC01FA93FC32" box="[1362,1394,995,1019]" gridcol="5" gridrow="11" pageId="5" pageNumber="140">0.3</td>
</tr>
<tr id="35375D60FFB937F4FF3BFBE9FA93FBED" box="[166,1394,1035,1060]" gridrow="12" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFBE9FD78FBED" box="[166,665,1035,1060]" gridcol="0" gridrow="12" pageId="5" pageNumber="140">Distance between pelvic- and anal-fin origins</th>
<td id="76E6341CFFB937F4FCF3FBE9FC2AFBED" box="[878,971,1035,1060]" gridcol="1" gridrow="12" pageId="5" pageNumber="140">23.4</td>
<td id="76E6341CFFB937F4FB93FBE9FBA4FBED" box="[1038,1093,1035,1060]" gridcol="2" gridrow="12" pageId="5" pageNumber="140">23.5</td>
<td id="76E6341CFFB937F4FB1CFBE9FB4CFBED" box="[1153,1197,1035,1060]" gridcol="3" gridrow="12" pageId="5" pageNumber="140">22.5</td>
<td id="76E6341CFFB937F4FB78FBE9FAD6FBED" box="[1253,1335,1035,1060]" gridcol="4" gridrow="12" pageId="5" pageNumber="140">25.5</td>
<td id="76E6341CFFB937F4FACFFBE9FA93FBED" box="[1362,1394,1035,1060]" gridcol="5" gridrow="12" pageId="5" pageNumber="140">1.0</td>
</tr>
<tr id="35375D60FFB937F4FF3BFBD6FA93FB85" box="[166,1394,1076,1100]" gridrow="13" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFBD6FD78FB85" box="[166,665,1076,1100]" gridcol="0" gridrow="13" pageId="5" pageNumber="140">Distance between vent and anal-fin origin</th>
<td id="76E6341CFFB937F4FCF3FBD6FC2AFB85" box="[878,971,1076,1100]" gridcol="1" gridrow="13" pageId="5" pageNumber="140">2.3</td>
<td id="76E6341CFFB937F4FB93FBD6FBA4FB85" box="[1038,1093,1076,1100]" gridcol="2" gridrow="13" pageId="5" pageNumber="140">1.5</td>
<td id="76E6341CFFB937F4FB1CFBD6FB4CFB85" box="[1153,1197,1076,1100]" gridcol="3" gridrow="13" pageId="5" pageNumber="140">0.9</td>
<td id="76E6341CFFB937F4FB78FBD6FAD6FB85" box="[1253,1335,1076,1100]" gridcol="4" gridrow="13" pageId="5" pageNumber="140">2.3</td>
<td id="76E6341CFFB937F4FACFFBD6FA93FB85" box="[1362,1394,1076,1100]" gridcol="5" gridrow="13" pageId="5" pageNumber="140">0.5</td>
</tr>
<tr id="35375D60FFB937F4FF3BFBBEFA93FBBD" box="[166,1394,1116,1140]" gridrow="14" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFBBEFD78FBBD" box="[166,665,1116,1140]" gridcol="0" gridrow="14" pageId="5" pageNumber="140">Depth of caudal peduncle</th>
<td id="76E6341CFFB937F4FCF3FBBEFC2AFBBD" box="[878,971,1116,1140]" gridcol="1" gridrow="14" pageId="5" pageNumber="140">12.3</td>
<td id="76E6341CFFB937F4FB93FBBEFBA4FBBD" box="[1038,1093,1116,1140]" gridcol="2" gridrow="14" pageId="5" pageNumber="140">11.2</td>
<td id="76E6341CFFB937F4FB1CFBBEFB4CFBBD" box="[1153,1197,1116,1140]" gridcol="3" gridrow="14" pageId="5" pageNumber="140">8.6</td>
<td id="76E6341CFFB937F4FB78FBBEFAD6FBBD" box="[1253,1335,1116,1140]" gridcol="4" gridrow="14" pageId="5" pageNumber="140">12.4</td>
<td id="76E6341CFFB937F4FACFFBBEFA93FBBD" box="[1362,1394,1116,1140]" gridcol="5" gridrow="14" pageId="5" pageNumber="140">1.3</td>
</tr>
<tr id="35375D60FFB937F4FF3BFB66FA93FB55" box="[166,1394,1156,1180]" gridrow="15" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFB66FD78FB55" box="[166,665,1156,1180]" gridcol="0" gridrow="15" pageId="5" pageNumber="140">Length of caudal peduncle</th>
<td id="76E6341CFFB937F4FCF3FB66FC2AFB55" box="[878,971,1156,1180]" gridcol="1" gridrow="15" pageId="5" pageNumber="140">19.3</td>
<td id="76E6341CFFB937F4FB93FB66FBA4FB55" box="[1038,1093,1156,1180]" gridcol="2" gridrow="15" pageId="5" pageNumber="140">19.0</td>
<td id="76E6341CFFB937F4FB1CFB66FB4CFB55" box="[1153,1197,1156,1180]" gridcol="3" gridrow="15" pageId="5" pageNumber="140">18.1</td>
<td id="76E6341CFFB937F4FB78FB66FAD6FB55" box="[1253,1335,1156,1180]" gridcol="4" gridrow="15" pageId="5" pageNumber="140">20.3</td>
<td id="76E6341CFFB937F4FACFFB66FA93FB55" box="[1362,1394,1156,1180]" gridcol="5" gridrow="15" pageId="5" pageNumber="140">0.9</td>
</tr>
<tr id="35375D60FFB937F4FF3BFB4EFA93FB0C" box="[166,1394,1196,1221]" gridrow="16" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFB4EFD78FB0C" box="[166,665,1196,1221]" gridcol="0" gridrow="16" pageId="5" pageNumber="140">Dorsal-fin depth</th>
<td id="76E6341CFFB937F4FCF3FB4EFC2AFB0C" box="[878,971,1196,1221]" gridcol="1" gridrow="16" pageId="5" pageNumber="140">15.6</td>
<td id="76E6341CFFB937F4FB93FB4EFBA4FB0C" box="[1038,1093,1196,1221]" gridcol="2" gridrow="16" pageId="5" pageNumber="140">17.2</td>
<td id="76E6341CFFB937F4FB1CFB4EFB4CFB0C" box="[1153,1197,1196,1221]" gridcol="3" gridrow="16" pageId="5" pageNumber="140">10.5</td>
<td id="76E6341CFFB937F4FB78FB4EFAD6FB0C" box="[1253,1335,1196,1221]" gridcol="4" gridrow="16" pageId="5" pageNumber="140">19.8</td>
<td id="76E6341CFFB937F4FACFFB4EFA93FB0C" box="[1362,1394,1196,1221]" gridcol="5" gridrow="16" pageId="5" pageNumber="140">3.3</td>
</tr>
<tr id="35375D60FFB937F4FF3BFB37FA93FB24" box="[166,1394,1237,1261]" gridrow="17" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFB37FD78FB24" box="[166,665,1237,1261]" gridcol="0" gridrow="17" pageId="5" pageNumber="140">Anal-fin height</th>
<td id="76E6341CFFB937F4FCF3FB37FC2AFB24" box="[878,971,1237,1261]" gridcol="1" gridrow="17" pageId="5" pageNumber="140">15.5</td>
<td id="76E6341CFFB937F4FB93FB37FBA4FB24" box="[1038,1093,1237,1261]" gridcol="2" gridrow="17" pageId="5" pageNumber="140">16.2</td>
<td id="76E6341CFFB937F4FB1CFB37FB4CFB24" box="[1153,1197,1237,1261]" gridcol="3" gridrow="17" pageId="5" pageNumber="140">14.3</td>
<td id="76E6341CFFB937F4FB78FB37FAD6FB24" box="[1253,1335,1237,1261]" gridcol="4" gridrow="17" pageId="5" pageNumber="140">19.2</td>
<td id="76E6341CFFB937F4FACFFB37FA93FB24" box="[1362,1394,1237,1261]" gridcol="5" gridrow="17" pageId="5" pageNumber="140">1.5</td>
</tr>
<tr id="35375D60FFB937F4FF3BFB1FFA93FADC" box="[166,1394,1277,1301]" gridrow="18" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFB1FFD78FADC" box="[166,665,1277,1301]" gridcol="0" gridrow="18" pageId="5" pageNumber="140">Pectoral-fin length</th>
<td id="76E6341CFFB937F4FCF3FB1FFC2AFADC" box="[878,971,1277,1301]" gridcol="1" gridrow="18" pageId="5" pageNumber="140">18.4</td>
<td id="76E6341CFFB937F4FB93FB1FFBA4FADC" box="[1038,1093,1277,1301]" gridcol="2" gridrow="18" pageId="5" pageNumber="140">20.5</td>
<td id="76E6341CFFB937F4FB1CFB1FFB4CFADC" box="[1153,1197,1277,1301]" gridcol="3" gridrow="18" pageId="5" pageNumber="140">18.4</td>
<td id="76E6341CFFB937F4FB78FB1FFAD6FADC" box="[1253,1335,1277,1301]" gridcol="4" gridrow="18" pageId="5" pageNumber="140">23.4</td>
<td id="76E6341CFFB937F4FACFFB1FFA93FADC" box="[1362,1394,1277,1301]" gridcol="5" gridrow="18" pageId="5" pageNumber="140">1.8</td>
</tr>
<tr id="35375D60FFB937F4FF3BFAC7FA93FAF4" box="[166,1394,1317,1341]" gridrow="19" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFAC7FD78FAF4" box="[166,665,1317,1341]" gridcol="0" gridrow="19" pageId="5" pageNumber="140">Pelvic-fin length</th>
<td id="76E6341CFFB937F4FCF3FAC7FC2AFAF4" box="[878,971,1317,1341]" gridcol="1" gridrow="19" pageId="5" pageNumber="140">16.4</td>
<td id="76E6341CFFB937F4FB93FAC7FBA4FAF4" box="[1038,1093,1317,1341]" gridcol="2" gridrow="19" pageId="5" pageNumber="140">15.9</td>
<td id="76E6341CFFB937F4FB1CFAC7FB4CFAF4" box="[1153,1197,1317,1341]" gridcol="3" gridrow="19" pageId="5" pageNumber="140">14.6</td>
<td id="76E6341CFFB937F4FB78FAC7FAD6FAF4" box="[1253,1335,1317,1341]" gridcol="4" gridrow="19" pageId="5" pageNumber="140">16.8</td>
<td id="76E6341CFFB937F4FACFFAC7FA93FAF4" box="[1362,1394,1317,1341]" gridcol="5" gridrow="19" pageId="5" pageNumber="140">0.9</td>
</tr>
<tr id="35375D60FFB937F4FF3BFAAFFA93FAAC" box="[166,1394,1357,1381]" gridrow="20" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFAAFFA93FAAC" box="[166,1394,1357,1381]" colspan="6" colspanRight="5" gridcol="0" gridrow="20" pageId="5" pageNumber="140">
<emphasis id="B9738330FFB9C80EFF3BFAAFFE54FAAC" bold="true" box="[166,437,1357,1381]" pageId="5" pageNumber="140">In percent of head length</emphasis>
</th>
</tr>
<tr id="35375D60FFB937F4FF3BFA94FA93FA47" box="[166,1394,1398,1422]" gridrow="21" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFA94FD78FA47" box="[166,665,1398,1422]" gridcol="0" gridrow="21" pageId="5" pageNumber="140">Head depth at eye</th>
<td id="76E6341CFFB937F4FCF3FA94FC2AFA47" box="[878,971,1398,1422]" gridcol="1" gridrow="21" pageId="5" pageNumber="140">46</td>
<td id="76E6341CFFB937F4FB93FA94FBA4FA47" box="[1038,1093,1398,1422]" gridcol="2" gridrow="21" pageId="5" pageNumber="140">47</td>
<td id="76E6341CFFB937F4FB1CFA94FB4CFA47" box="[1153,1197,1398,1422]" gridcol="3" gridrow="21" pageId="5" pageNumber="140">43</td>
<td id="76E6341CFFB937F4FB78FA94FAD6FA47" box="[1253,1335,1398,1422]" gridcol="4" gridrow="21" pageId="5" pageNumber="140">58</td>
<td id="76E6341CFFB937F4FACFFA94FA93FA47" box="[1362,1394,1398,1422]" gridcol="5" gridrow="21" pageId="5" pageNumber="140">4.9</td>
</tr>
<tr id="35375D60FFB937F4FF3BFA7CFA93FA7F" box="[166,1394,1438,1462]" gridrow="22" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFA7CFD78FA7F" box="[166,665,1438,1462]" gridcol="0" gridrow="22" pageId="5" pageNumber="140">Snout length</th>
<td id="76E6341CFFB937F4FCF3FA7CFC2AFA7F" box="[878,971,1438,1462]" gridcol="1" gridrow="22" pageId="5" pageNumber="140">43</td>
<td id="76E6341CFFB937F4FB93FA7CFBA4FA7F" box="[1038,1093,1438,1462]" gridcol="2" gridrow="22" pageId="5" pageNumber="140">42</td>
<td id="76E6341CFFB937F4FB1CFA7CFB4CFA7F" box="[1153,1197,1438,1462]" gridcol="3" gridrow="22" pageId="5" pageNumber="140">39</td>
<td id="76E6341CFFB937F4FB78FA7CFAD6FA7F" box="[1253,1335,1438,1462]" gridcol="4" gridrow="22" pageId="5" pageNumber="140">45</td>
<td id="76E6341CFFB937F4FACFFA7CFA93FA7F" box="[1362,1394,1438,1462]" gridcol="5" gridrow="22" pageId="5" pageNumber="140">1.6</td>
</tr>
<tr id="35375D60FFB937F4FF3BFA24FA93FA16" box="[166,1394,1478,1503]" gridrow="23" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFA24FD78FA16" box="[166,665,1478,1503]" gridcol="0" gridrow="23" pageId="5" pageNumber="140">Eye diameter</th>
<td id="76E6341CFFB937F4FCF3FA24FC2AFA16" box="[878,971,1478,1503]" gridcol="1" gridrow="23" pageId="5" pageNumber="140">15</td>
<td id="76E6341CFFB937F4FB93FA24FBA4FA16" box="[1038,1093,1478,1503]" gridcol="2" gridrow="23" pageId="5" pageNumber="140">17</td>
<td id="76E6341CFFB937F4FB1CFA24FB4CFA16" box="[1153,1197,1478,1503]" gridcol="3" gridrow="23" pageId="5" pageNumber="140">15</td>
<td id="76E6341CFFB937F4FB78FA24FAD6FA16" box="[1253,1335,1478,1503]" gridcol="4" gridrow="23" pageId="5" pageNumber="140">21</td>
<td id="76E6341CFFB937F4FACFFA24FA93FA16" box="[1362,1394,1478,1503]" gridcol="5" gridrow="23" pageId="5" pageNumber="140">2.2</td>
</tr>
<tr id="35375D60FFB937F4FF3BFA0CFA93F9CE" box="[166,1394,1518,1543]" gridrow="24" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BFA0CFD78F9CE" box="[166,665,1518,1543]" gridcol="0" gridrow="24" pageId="5" pageNumber="140">Postorbital distance</th>
<td id="76E6341CFFB937F4FCF3FA0CFC2AF9CE" box="[878,971,1518,1543]" gridcol="1" gridrow="24" pageId="5" pageNumber="140">47</td>
<td id="76E6341CFFB937F4FB93FA0CFBA4F9CE" box="[1038,1093,1518,1543]" gridcol="2" gridrow="24" pageId="5" pageNumber="140">46</td>
<td id="76E6341CFFB937F4FB1CFA0CFB4CF9CE" box="[1153,1197,1518,1543]" gridcol="3" gridrow="24" pageId="5" pageNumber="140">43</td>
<td id="76E6341CFFB937F4FB78FA0CFAD6F9CE" box="[1253,1335,1518,1543]" gridcol="4" gridrow="24" pageId="5" pageNumber="140">49</td>
<td id="76E6341CFFB937F4FACFFA0CFA93F9CE" box="[1362,1394,1518,1543]" gridcol="5" gridrow="24" pageId="5" pageNumber="140">1.8</td>
</tr>
<tr id="35375D60FFB937F4FF3BF9F5FA93F9E6" box="[166,1394,1559,1583]" gridrow="25" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BF9F5FD78F9E6" box="[166,665,1559,1583]" gridcol="0" gridrow="25" pageId="5" pageNumber="140">Maximum head width</th>
<td id="76E6341CFFB937F4FCF3F9F5FC2AF9E6" box="[878,971,1559,1583]" gridcol="1" gridrow="25" pageId="5" pageNumber="140">70</td>
<td id="76E6341CFFB937F4FB93F9F5FBA4F9E6" box="[1038,1093,1559,1583]" gridcol="2" gridrow="25" pageId="5" pageNumber="140">68</td>
<td id="76E6341CFFB937F4FB1CF9F5FB4CF9E6" box="[1153,1197,1559,1583]" gridcol="3" gridrow="25" pageId="5" pageNumber="140">61</td>
<td id="76E6341CFFB937F4FB78F9F5FAD6F9E6" box="[1253,1335,1559,1583]" gridcol="4" gridrow="25" pageId="5" pageNumber="140">75</td>
<td id="76E6341CFFB937F4FACFF9F5FA93F9E6" box="[1362,1394,1559,1583]" gridcol="5" gridrow="25" pageId="5" pageNumber="140">4.3</td>
</tr>
<tr id="35375D60FFB937F4FF3BF9DDFA93F99E" box="[166,1394,1599,1623]" gridrow="26" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BF9DDFD78F99E" box="[166,665,1599,1623]" gridcol="0" gridrow="26" pageId="5" pageNumber="140">Interorbital width</th>
<td id="76E6341CFFB937F4FCF3F9DDFC2AF99E" box="[878,971,1599,1623]" gridcol="1" gridrow="26" pageId="5" pageNumber="140">30</td>
<td id="76E6341CFFB937F4FB93F9DDFBA4F99E" box="[1038,1093,1599,1623]" gridcol="2" gridrow="26" pageId="5" pageNumber="140">29</td>
<td id="76E6341CFFB937F4FB1CF9DDFB4CF99E" box="[1153,1197,1599,1623]" gridcol="3" gridrow="26" pageId="5" pageNumber="140">25</td>
<td id="76E6341CFFB937F4FB78F9DDFAD6F99E" box="[1253,1335,1599,1623]" gridcol="4" gridrow="26" pageId="5" pageNumber="140">34</td>
<td id="76E6341CFFB937F4FACFF9DDFA93F99E" box="[1362,1394,1599,1623]" gridcol="5" gridrow="26" pageId="5" pageNumber="140">3.0</td>
</tr>
<tr id="35375D60FFB937F4FF3BF985FA93F9B6" box="[166,1394,1639,1663]" gridrow="27" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BF985FD78F9B6" box="[166,665,1639,1663]" gridcol="0" gridrow="27" pageId="5" pageNumber="140">Length of inner rostral barbel</th>
<td id="76E6341CFFB937F4FCF3F985FC2AF9B6" box="[878,971,1639,1663]" gridcol="1" gridrow="27" pageId="5" pageNumber="140">19</td>
<td id="76E6341CFFB937F4FB93F985FBA4F9B6" box="[1038,1093,1639,1663]" gridcol="2" gridrow="27" pageId="5" pageNumber="140">21</td>
<td id="76E6341CFFB937F4FB1CF985FB4CF9B6" box="[1153,1197,1639,1663]" gridcol="3" gridrow="27" pageId="5" pageNumber="140">14</td>
<td id="76E6341CFFB937F4FB78F985FAD6F9B6" box="[1253,1335,1639,1663]" gridcol="4" gridrow="27" pageId="5" pageNumber="140">26</td>
<td id="76E6341CFFB937F4FACFF985FA93F9B6" box="[1362,1394,1639,1663]" gridcol="5" gridrow="27" pageId="5" pageNumber="140">4.2</td>
</tr>
<tr id="35375D60FFB937F4FF3BF96DFA93F961" box="[166,1394,1679,1704]" gridrow="28" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BF96DFD78F961" box="[166,665,1679,1704]" gridcol="0" gridrow="28" pageId="5" pageNumber="140">Length of outer rostral barbel</th>
<td id="76E6341CFFB937F4FCF3F96DFC2AF961" box="[878,971,1679,1704]" gridcol="1" gridrow="28" pageId="5" pageNumber="140">19</td>
<td id="76E6341CFFB937F4FB93F96DFBA4F961" box="[1038,1093,1679,1704]" gridcol="2" gridrow="28" pageId="5" pageNumber="140">29</td>
<td id="76E6341CFFB937F4FB1CF96DFB4CF961" box="[1153,1197,1679,1704]" gridcol="3" gridrow="28" pageId="5" pageNumber="140">17</td>
<td id="76E6341CFFB937F4FB78F96DFAD6F961" box="[1253,1335,1679,1704]" gridcol="4" gridrow="28" pageId="5" pageNumber="140">42</td>
<td id="76E6341CFFB937F4FACFF96DFA93F961" box="[1362,1394,1679,1704]" gridcol="5" gridrow="28" pageId="5" pageNumber="140">9.3</td>
</tr>
<tr id="35375D60FFB937F4FF3BF95AFA93F919" box="[166,1394,1720,1744]" gridrow="29" pageId="5" pageNumber="140">
<th id="76E6341CFFB937F4FF3BF95AFD78F919" box="[166,665,1720,1744]" gridcol="0" gridrow="29" pageId="5" pageNumber="140">Length of maxillary barbel</th>
<td id="76E6341CFFB937F4FCF3F95AFC2AF919" box="[878,971,1720,1744]" gridcol="1" gridrow="29" pageId="5" pageNumber="140">23</td>
<td id="76E6341CFFB937F4FB93F95AFBA4F919" box="[1038,1093,1720,1744]" gridcol="2" gridrow="29" pageId="5" pageNumber="140">28</td>
<td id="76E6341CFFB937F4FB1CF95AFB4CF919" box="[1153,1197,1720,1744]" gridcol="3" gridrow="29" pageId="5" pageNumber="140">16</td>
<td id="76E6341CFFB937F4FB78F95AFAD6F919" box="[1253,1335,1720,1744]" gridcol="4" gridrow="29" pageId="5" pageNumber="140">33</td>
<td id="76E6341CFFB937F4FACFF95AFA93F919" box="[1362,1394,1720,1744]" gridcol="5" gridrow="29" pageId="5" pageNumber="140">6.3</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB85F22FFB9C80DFF5AF91DFDC8FEAC" blockId="5.[151,1437,1791,2034]" lastBlockId="6.[151,1437,151,357]" lastPageId="6" lastPageNumber="141" pageId="5" pageNumber="140">
<emphasis id="B9738330FFB9C80EFF5AF91DFEAEF8D0" bold="true" box="[199,335,1791,1817]" pageId="5" pageNumber="140">Coloration.</emphasis>
Body with yellowish or pale brown background and dark- or pale-brown pattern in live and preserved individuals. Preserved individuals with a wide inner-axial stripe, absent in life. Dorsal head and upper part of cheek brown, with a vermiculate pattern. Ventral surface of head yellowish without pattern. Flank densely mottled, without bars or blotches, marbled or with bars on flank behind dorsal-fin origin in individuals smaller than
<quantity id="4CFFF2C7FFB9C80EFAD0F889FA7CF84F" box="[1357,1437,1899,1926]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="140" unit="mm" value="50.0">50 mm</quantity>
SL, almost plain brown in some individuals. Flank pigmentation reaching down to horizontal of pectoral-fin base or to belly in some individuals. Back mottled, marbled or with 34 squarish or triangular predorsal blotches, confluent with flank pattern, one saddle at dorsal-fin origin, one or none in middle of dorsal-fin base and one at end of dorsal- fin base, and 36 saddles behind dorsal-fin base, all saddles wider than interspaces. A narrow, black bar at caudal-fin base. Posteriormost upper and lowermost caudal peduncle with a pale yellowish, unpigmented blotch. Dorsal and pectoral fins with small brown blotches on rays, forming 13 bands. Caudal fin with many small brown blotches on rays, forming 13 wide and bold bands, often band at second half of caudal fin very wide or caudal-fin rays almost completely brown. Pectoral, anal and pelvic fins with dark-brown blotches on rays; hyaline or with few blotches in individuals smaller than
<quantity id="4CFFF2C7FFBAC80DFE36FEA9FE1BFEAC" box="[427,506,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="141" unit="mm" value="60.0">60 mm</quantity>
SL.
</paragraph>
</subSubSection>
<caption id="DF780FAAFFBAC80DFF0AFA8AFB13FA49" ID-DOI="http://doi.org/10.5281/zenodo.4671920" ID-Zenodo-Dep="4671920" box="[151,1266,1384,1408]" httpUri="https://zenodo.org/record/4671920/files/figure.png" pageId="6" pageNumber="141" startId="6.[151,250,1384,1408]" targetBox="[152,1436,381,1361]" targetPageId="6">
<paragraph id="8BB85F22FFBAC80DFF0AFA8AFB13FA49" blockId="6.[151,1266,1384,1408]" box="[151,1266,1384,1408]" pageId="6" pageNumber="141">
<emphasis id="B9738330FFBAC80DFF0AFA8AFEF2FA49" bold="true" box="[151,275,1384,1408]" pageId="6" pageNumber="141">FIGURE 2.</emphasis>
<taxonomicName id="4C0724A1FFBAC80DFE84FA8AFD9EFAB6" authority=", ZFMK" authorityName="ZFMK" box="[281,639,1384,1408]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="phasicus">
<emphasis id="B9738330FFBAC80DFE84FA8AFDCCFA49" box="[281,557,1384,1408]" italics="true" pageId="6" pageNumber="141">Oxynoemacheilus phasicus</emphasis>
, ZFMK
</taxonomicName>
ICH-124180, holotype, 74.1 mm SL; Georgia: Dzirula River.
</paragraph>
</caption>
<subSubSection id="C31D0CA9FFBAC80DFF5AFA4CFE0DF9D8" pageId="6" pageNumber="141" type="distribution">
<paragraph id="8BB85F22FFBAC80DFF5AFA4CFE0DF9D8" blockId="6.[151,1437,1454,2021]" pageId="6" pageNumber="141">
<emphasis id="B9738330FFBAC80DFF5AFA4CFE80FA01" bold="true" box="[199,353,1454,1480]" pageId="6" pageNumber="141">Distribution.</emphasis>
<taxonomicName id="4C0724A1FFBAC80DFEFAFA4CFD76FA01" box="[359,663,1454,1480]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="phasicus">
<emphasis id="B9738330FFBAC80DFEFAFA4CFD76FA01" box="[359,663,1454,1480]" italics="true" pageId="6" pageNumber="141">Oxynoemacheilus phasicus</emphasis>
</taxonomicName>
was collected from the Enguri and Rioni River drainages in
<collectingCountry id="F3101FB2FFBAC80DFAA1FA4CFA78FA01" box="[1340,1433,1454,1480]" name="Georgia" pageId="6" pageNumber="141">Georgia</collectingCountry>
. It has not been found in the Chorokhi / Çoruh River in
<collectingCountry id="F3101FB2FFBAC80DFC98FA30FCBEFA25" box="[773,863,1490,1516]" name="Georgia" pageId="6" pageNumber="141">Georgia</collectingCountry>
or
<collectingCountry id="F3101FB2FFBAC80DFC19FA30FC35FA25" box="[900,980,1490,1516]" name="Turkey" pageId="6" pageNumber="141">Turkey</collectingCountry>
or in small rivers in-between despite intensive field work in that area.
</paragraph>
</subSubSection>
<subSubSection id="C31D0CA9FFBAC80DFF5AF9F8FAECF9FD" box="[199,1293,1562,1589]" pageId="6" pageNumber="141" type="etymology">
<paragraph id="8BB85F22FFBAC80DFF5AF9F8FAECF9FD" blockId="6.[151,1437,1454,2021]" box="[199,1293,1562,1589]" pageId="6" pageNumber="141">
<emphasis id="B9738330FFBAC80DFF5AF9F8FEAFF9FD" bold="true" box="[199,334,1562,1588]" pageId="6" pageNumber="141">Etymology.</emphasis>
The species name is derived from
<taxonomicName id="4C0724A1FFBAC80DFD49F9F8FCFEF9FD" box="[724,799,1562,1588]" class="Insecta" family="Lycaenidae" genus="Phasis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B9738330FFBAC80DFD49F9F8FCFEF9FD" box="[724,799,1562,1588]" italics="true" pageId="6" pageNumber="141">Phasis</emphasis>
</taxonomicName>
, the Greek name of the Rioni. An adjective.
</paragraph>
</subSubSection>
<subSubSection id="C31D0CA9FFBAC800FF5AF9DCFE37F895" lastPageId="11" lastPageNumber="146" pageId="6" pageNumber="141" type="discussion">
<paragraph id="8BB85F22FFBAC80CFF5AF9DCFCB4FEAC" blockId="6.[151,1437,1454,2021]" lastBlockId="7.[151,1437,151,357]" lastPageId="7" lastPageNumber="142" pageId="6" pageNumber="141">
<emphasis id="B9738330FFBAC80DFF5AF9DCFEDAF991" bold="true" box="[199,315,1598,1624]" pageId="6" pageNumber="141">Remarks.</emphasis>
<taxonomicName id="4C0724A1FFBAC80DFEDEF9DCFDF4F991" authorityName="Gratzianov" authorityYear="1907" box="[323,533,1598,1624]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="bergi">
<emphasis id="B9738330FFBAC80DFEDEF9DCFDF4F991" box="[323,533,1598,1624]" italics="true" pageId="6" pageNumber="141">Nemacheilus bergi</emphasis>
</taxonomicName>
was named by
<bibRefCitation id="EF9622D3FFBAC80DFD5BF9DCFC74F991" author="Gratzianov, V. I." box="[710,917,1598,1624]" pageId="6" pageNumber="141" pagination="1 - 567" refId="ref8818" refString="Gratzianov, V. I. (1907) Opyt' obzora ryby Rossiskoi Imperii v' sistematicheskom' i geograficescom' otnoshenii. Vol. 4. Trudy Otdela Ichtiologii Imperatorskago Russkago Obshchestva Akklimatizacii Zivotnych' i Rastenii, 1907, i - xxx + 1 - 567." type="book chapter" year="1907">Gratzianov (1907)</bibRefCitation>
based on a description by
<bibRefCitation id="EF9622D3FFBAC80DFB5EF9DDFAAAF991" author="Berg, L. S." box="[1219,1355,1598,1625]" pageId="6" pageNumber="141" pagination="1 - 80" refId="ref7980" refString="Berg, L. S. (1899) Beitrage zur Ichthyofauna des Kaukasus. Mitteilungen des Kaukasischen Museums [Bulletin du Musee du Caucase; Izvestija Kavkazskogo Muzeja], 1 (3), 1 - 80. [in Russian and German]." type="journal article" year="1899">Berg (1899)</bibRefCitation>
of two individuals (
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,
<quantity id="4CFFF2C7FFBAC80DFE19F980FE34F9B4" box="[388,469,1634,1661]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="141" unit="mm" value="35.0">35 mm</quantity>
) from the Akstapha [
<collectingRegion id="49C391C0FFBAC80DFD59F980FCC0F9B4" box="[708,801,1634,1661]" country="Azerbaijan" name="Agstafa" pageId="6" pageNumber="141">Agstafa</collectingRegion>
] River, a right tributary of the Kura in
<collectingCountry id="F3101FB2FFBAC80DFB41F980FABAF9B5" box="[1244,1371,1634,1660]" name="Azerbaijan" pageId="6" pageNumber="141">Azerbaijan</collectingCountry>
.
<bibRefCitation id="EF9622D3FFBAC80DFAFBF981FEE4F969" author="Berg, L. S." pageId="6" pageNumber="141" pagination="1 - 80" refId="ref7980" refString="Berg, L. S. (1899) Beitrage zur Ichthyofauna des Kaukasus. Mitteilungen des Kaukasischen Museums [Bulletin du Musee du Caucase; Izvestija Kavkazskogo Muzeja], 1 (3), 1 - 80. [in Russian and German]." type="journal article" year="1899">Berg (1899:73)</bibRefCitation>
wrote in his chapter on
<emphasis id="B9738330FFBAC80DFDBCF964FD35F969" box="[545,724,1670,1696]" italics="true" pageId="6" pageNumber="141">
<taxonomicName id="4C0724A1FFBAC80DFDBCF964FD53F969" box="[545,690,1670,1696]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="141" phylum="Chordata" rank="genus">Nemacheilus</taxonomicName>
sp
</emphasis>
.: „Die Schwanzflosse ist im Verhältnis zu N. Brandt schwach ausgeschnitten (bei dem Exemplar von
<quantity id="4CFFF2C7FFBAC80DFDC1F948FD51F90C" box="[604,688,1706,1733]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" pageId="6" pageNumber="141" unit="mm" value="61.0">61 mm</quantity>
. Länge ist der mittelste Strahl der Schwanzflosse
<quantity id="4CFFF2C7FFBAC80DFB73F948FAD5F90C" box="[1262,1332,1706,1733]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="141" unit="mm" value="8.0">8 mm</quantity>
. Und die seitlichen
<quantity id="4CFFF2C7FFBAC80DFE97F92CFEB9F920" box="[266,344,1742,1769]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="141" unit="mm" value="11.0">11 mm</quantity>
lang).” [The caudal fin is slightly emarginate in comparison with
<taxonomicName id="4C0724A1FFBAC80DFBDCF92DFB5DF921" box="[1089,1212,1742,1768]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFBAC80DFBDCF92DFB5DF921" box="[1089,1212,1742,1768]" italics="true" pageId="6" pageNumber="141">N. brandtii</emphasis>
</taxonomicName>
(in the specimen of
<quantity id="4CFFF2C7FFBAC80DFF0AF910FF06F8C4" box="[151,231,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" pageId="6" pageNumber="141" unit="mm" value="61.0">61 mm</quantity>
the length of the central caudal-fin ray is
<quantity id="4CFFF2C7FFBAC80DFD5BF910FCE9F8C4" box="[710,776,1778,1805]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="141" unit="mm" value="8.0">8 mm</quantity>
and of the lateral ones
<quantity id="4CFFF2C7FFBAC80DFB8EF910FB89F8C4" box="[1043,1128,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="141" unit="mm" value="11.0">11 mm</quantity>
)]. Also,
<bibRefCitation id="EF9622D3FFBAC80DFB5BF911FA7DF8C5" author="Berg, L. S." box="[1222,1436,1778,1805]" pageId="6" pageNumber="141" pagination="1 - 80" refId="ref7980" refString="Berg, L. S. (1899) Beitrage zur Ichthyofauna des Kaukasus. Mitteilungen des Kaukasischen Museums [Bulletin du Musee du Caucase; Izvestija Kavkazskogo Muzeja], 1 (3), 1 - 80. [in Russian and German]." type="journal article" year="1899">Berg (1899:7273)</bibRefCitation>
mentions that the pelvic fin reaches the anus, there is a bold bar at the caudal-fin base, a wide band in the middle of the caudal fin and incomplete bars on the flank. This species is further treated as a synonym of
<taxonomicName id="4C0724A1FFBAC80DFB20F8D9FADDF89D" baseAuthorityName="Steindachner" baseAuthorityYear="1897" box="[1213,1340,1851,1877]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="angorae">
<emphasis id="B9738330FFBAC80DFB20F8D9FADDF89D" box="[1213,1340,1851,1877]" italics="true" pageId="6" pageNumber="141">O. angorae</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF9622D3FFBAC80DFAFBF8D9FF05F8B1" author="Berg, L. S." pageId="6" pageNumber="141" pagination="927 - 1382" refId="ref8027" refString="Berg, L. S. (1949) The freshwater fishes of the USSR and adjacent countries. Part 3. 4 th Edition. Akademia Nauk USSR, Moscow &amp; Leningrad, pp. 927 - 1382. [in Russian]" type="book chapter" year="1949">Berg (1949)</bibRefCitation>
, who distinguished
<taxonomicName id="4C0724A1FFBAC80DFE58F8BDFDA4F8B1" baseAuthorityName="Steindachner" baseAuthorityYear="1897" box="[453,581,1887,1913]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="angorae">
<emphasis id="B9738330FFBAC80DFE58F8BDFDA4F8B1" box="[453,581,1887,1913]" italics="true" pageId="6" pageNumber="141">O. angorae</emphasis>
</taxonomicName>
from
<taxonomicName id="4C0724A1FFBAC80DFD16F8BDFCFCF8B1" box="[651,797,1886,1913]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="bergianus">
<emphasis id="B9738330FFBAC80DFD16F8BDFCFCF8B1" box="[651,797,1886,1913]" italics="true" pageId="6" pageNumber="141">O. bergianus</emphasis>
</taxonomicName>
by the caudal peduncle being less than two times longer than deep in
<taxonomicName id="4C0724A1FFBAC80DFEB6F861FE4DF855" baseAuthorityName="Steindachner" baseAuthorityYear="1897" box="[299,428,1923,1949]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="angorae">
<emphasis id="B9738330FFBAC80DFEB6F861FE4DF855" box="[299,428,1923,1949]" italics="true" pageId="6" pageNumber="141">O. angorae</emphasis>
</taxonomicName>
vs. more in
<taxonomicName id="4C0724A1FFBAC80DFDA0F861FD31F855" box="[573,720,1922,1949]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="bergianus">
<emphasis id="B9738330FFBAC80DFDA0F861FD31F855" box="[573,720,1922,1949]" italics="true" pageId="6" pageNumber="141">O. bergianus</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF9622D3FFBAC80DFD7DF860FC57F855" author="Cicek, E. &amp; Eagderi, S. &amp; Sungur, S." box="[736,950,1922,1949]" pageId="6" pageNumber="141" pagination="232 - 242" refId="ref8095" refString="Cicek, E., Eagderi, S. &amp; Sungur, S. (2018) Oxynoemacheilus veyseli, a new nemacheilid species from the upper Aras River drainage of Turkey (Teleostei: Nemacheilidae). Iranian Journal of Ichthyology, 5 (3), 232 - 242. https: // doi. org / 10.22034 / iji. v 5 i 3.302" type="journal article" year="2018">
Çiçek
<emphasis id="B9738330FFBAC80DFCB4F861FCBDF855" box="[809,860,1922,1948]" italics="true" pageId="6" pageNumber="141">et al</emphasis>
. (2018)
</bibRefCitation>
treat
<taxonomicName id="4C0724A1FFBAC80DFC64F861FBB4F855" authorityName="Gratzianov" authorityYear="1907" box="[1017,1109,1922,1948]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="bergi">
<emphasis id="B9738330FFBAC80DFC64F861FBB4F855" box="[1017,1109,1922,1948]" italics="true" pageId="6" pageNumber="141">N. bergi</emphasis>
</taxonomicName>
as a valid species in
<taxonomicName id="4C0724A1FFBAC80DFAD7F861FEF4F809" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="141" phylum="Chordata" rank="genus">
<emphasis id="B9738330FFBAC80DFAD7F861FEF4F809" italics="true" pageId="6" pageNumber="141">Oxynoemacheilus</emphasis>
</taxonomicName>
without giving further details. Soheil Eagderi (Tehran) sent JF a picture (
<figureCitation id="133C43A7FFBAC80DFBCCF844FB75F808" box="[1105,1172,1958,1985]" captionStart="FIGURE 7" captionStartId="9.[151,250,1288,1312]" captionTargetBox="[194,1361,993,1241]" captionTargetId="figure-42@9.[151,1437,971,1264]" captionTargetPageId="9" captionText="FIGURE 7. Oxynoemacheilus bergi; IMNRF-1030-2, 50.4 mm SL; Azerbaijan: Kura River, 41°1425N 45°2557E. Photo: Soheil Eagderi" figureDoi="http://doi.org/10.5281/zenodo.4671934" httpUri="https://zenodo.org/record/4671934/files/figure.png" pageId="6" pageNumber="141">Fig. 7</figureCitation>
) of one individual they identified as
<taxonomicName id="4C0724A1FFBAC80DFEB5F829FE65F82D" authorityName="Gratzianov" authorityYear="1907" box="[296,388,1994,2021]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="bergi">
<emphasis id="B9738330FFBAC80DFEB5F829FE65F82D" box="[296,388,1994,2021]" italics="true" pageId="6" pageNumber="141">O. bergi</emphasis>
</taxonomicName>
collected in the Kura, very close to the inflow of the
<collectingRegion id="49C391C0FFBAC80DFC4AF828FBD3F82D" box="[983,1074,1994,2020]" country="Azerbaijan" name="Agstafa" pageId="6" pageNumber="141">Agstafa</collectingRegion>
. Measured from the picture, this individual has the caudal-peduncle depth two times in the caudal-peduncle length, the caudal fin is emarginate and the middle caudal-fin ray is 75% of the length of the longest principal ray in the lower caudal-fin lobe (72% in the description of
<bibRefCitation id="EF9622D3FFBBC80CFEA1FF3DFE54FF30" author="Berg, L. S." box="[316,437,223,249]" pageId="7" pageNumber="142" pagination="1 - 80" refId="ref7980" refString="Berg, L. S. (1899) Beitrage zur Ichthyofauna des Kaukasus. Mitteilungen des Kaukasischen Museums [Bulletin du Musee du Caucase; Izvestija Kavkazskogo Muzeja], 1 (3), 1 - 80. [in Russian and German]." type="journal article" year="1899">Berg 1899</bibRefCitation>
, but unclear which lobe was measured). These two character states as well as its colour pattern distinguish this species clearly from
<taxonomicName id="4C0724A1FFBBC80CFD1FFEE1FD1FFED4" box="[642,766,259,285]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFBBC80CFD1FFEE1FD1FFED4" box="[642,766,259,285]" italics="true" pageId="7" pageNumber="142">O. brandtii</emphasis>
</taxonomicName>
(caudal peduncle 2.53.2 times longer than deep, caudal fin forked). Indeed the fish on
<figureCitation id="133C43A7FFBBC80CFE53FEC5FDF0FE88" box="[462,529,295,321]" captionStart="FIGURE 7" captionStartId="9.[151,250,1288,1312]" captionTargetBox="[194,1361,993,1241]" captionTargetId="figure-42@9.[151,1437,971,1264]" captionTargetPageId="9" captionText="FIGURE 7. Oxynoemacheilus bergi; IMNRF-1030-2, 50.4 mm SL; Azerbaijan: Kura River, 41°1425N 45°2557E. Photo: Soheil Eagderi" figureDoi="http://doi.org/10.5281/zenodo.4671934" httpUri="https://zenodo.org/record/4671934/files/figure.png" pageId="7" pageNumber="142">Fig. 7</figureCitation>
agree with the characters given for
<taxonomicName id="4C0724A1FFBBC80CFC36FEC5FBE6FE88" authorityName="Gratzianov" authorityYear="1907" box="[939,1031,295,321]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="bergi">
<emphasis id="B9738330FFBBC80CFC36FEC5FBE6FE88" box="[939,1031,295,321]" italics="true" pageId="7" pageNumber="142">N. bergi</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF9622D3FFBBC80CFBAEFEC5FB5DFE88" author="Berg, L. S." box="[1075,1212,295,321]" pageId="7" pageNumber="142" pagination="1 - 80" refId="ref7980" refString="Berg, L. S. (1899) Beitrage zur Ichthyofauna des Kaukasus. Mitteilungen des Kaukasischen Museums [Bulletin du Musee du Caucase; Izvestija Kavkazskogo Muzeja], 1 (3), 1 - 80. [in Russian and German]." type="journal article" year="1899">Berg (1899)</bibRefCitation>
and we support the view of
<bibRefCitation id="EF9622D3FFBBC80CFF6EFEA9FE24FEAC" author="Cicek, E. &amp; Eagderi, S. &amp; Sungur, S." box="[243,453,331,357]" pageId="7" pageNumber="142" pagination="232 - 242" refId="ref8095" refString="Cicek, E., Eagderi, S. &amp; Sungur, S. (2018) Oxynoemacheilus veyseli, a new nemacheilid species from the upper Aras River drainage of Turkey (Teleostei: Nemacheilidae). Iranian Journal of Ichthyology, 5 (3), 232 - 242. https: // doi. org / 10.22034 / iji. v 5 i 3.302" type="journal article" year="2018">
Çiçek
<emphasis id="B9738330FFBBC80CFEA6FEA9FE95FEAC" box="[315,372,331,357]" italics="true" pageId="7" pageNumber="142">et al.</emphasis>
(2018)
</bibRefCitation>
treating
<taxonomicName id="4C0724A1FFBBC80CFDB5FEA9FD64FEAC" authorityName="Gratzianov" authorityYear="1907" box="[552,645,331,357]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="bergi">
<emphasis id="B9738330FFBBC80CFDB5FEA9FD64FEAC" box="[552,645,331,357]" italics="true" pageId="7" pageNumber="142">O. bergi</emphasis>
</taxonomicName>
as a valid species.
</paragraph>
<caption id="DF780FAAFFBBC80CFF0AFB3EFD99FAF5" ID-DOI="http://doi.org/10.5281/zenodo.4671922" ID-Zenodo-Dep="4671922" httpUri="https://zenodo.org/record/4671922/files/figure.png" pageId="7" pageNumber="142" startId="7.[151,250,1244,1268]" targetBox="[151,1436,381,1221]" targetPageId="7">
<paragraph id="8BB85F22FFBBC80CFF0AFB3EFD99FAF5" blockId="7.[151,1436,1244,1340]" pageId="7" pageNumber="142">
<emphasis id="B9738330FFBBC80CFF0AFB3EFEF2FB3D" bold="true" box="[151,275,1244,1268]" pageId="7" pageNumber="142">FIGURE 3.</emphasis>
<taxonomicName id="4C0724A1FFBBC80CFE84FB3EFDCDFB3D" box="[281,556,1244,1268]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="phasicus">
<emphasis id="B9738330FFBBC80CFE84FB3EFDCDFB3D" box="[281,556,1244,1268]" italics="true" pageId="7" pageNumber="142">Oxynoemacheilus phasicus</emphasis>
</taxonomicName>
, paratypes, from top, left to right: FSJF 4061, 32 mm SL; 39 mm SL; Georgia: Gubistskali River; FSJF 3615, 44 mm SL, 54 mm SL; Georgia: Tskhenitskali River; FSJF 3576, 59 mm SL; Georgia: Enguri River; FSJF 3592, 77 mm SL; Georgia: Dzirula River.
</paragraph>
</caption>
<paragraph id="8BB85F22FFBBC80CFF5AFA88FC54F9DC" blockId="7.[151,1437,1386,1989]" pageId="7" pageNumber="142">
<taxonomicName id="4C0724A1FFBBC80CFF5AFA88FE2EFA4D" box="[199,463,1386,1412]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="bergi">
<emphasis id="B9738330FFBBC80CFF5AFA88FE2EFA4D" box="[199,463,1386,1412]" italics="true" pageId="7" pageNumber="142">Oxynoemacheilus bergi</emphasis>
</taxonomicName>
is clearly distinguished from
<taxonomicName id="4C0724A1FFBBC80CFC86FA89FC7EFA4D" box="[795,927,1386,1413]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="phasicus">
<emphasis id="B9738330FFBBC80CFC86FA89FC7EFA4D" box="[795,927,1386,1413]" italics="true" pageId="7" pageNumber="142">O. phasicus</emphasis>
</taxonomicName>
by possessing a deeply emarginate caudal fin (vs. slightly emarginate in
<taxonomicName id="4C0724A1FFBBC80CFE5CFA6DFDA4FA61" box="[449,581,1422,1449]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="phasicus">
<emphasis id="B9738330FFBBC80CFE5CFA6DFDA4FA61" box="[449,581,1422,1449]" italics="true" pageId="7" pageNumber="142">O. phasicus</emphasis>
</taxonomicName>
), and the pelvic fin reaching the anus (vs. not reaching). Potentially
<taxonomicName id="4C0724A1FFBBC80CFADDFA6DFA7DFA61" box="[1344,1436,1422,1449]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="bergi">
<emphasis id="B9738330FFBBC80CFADDFA6DFA7DFA61" box="[1344,1436,1422,1449]" italics="true" pageId="7" pageNumber="142">O. bergi</emphasis>
</taxonomicName>
is conspecific with
<taxonomicName id="4C0724A1FFBBC80CFEEEFA51FDE4FA05" box="[371,517,1458,1485]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="bergianus">
<emphasis id="B9738330FFBBC80CFEEEFA51FDE4FA05" box="[371,517,1458,1485]" italics="true" pageId="7" pageNumber="142">O. bergianus</emphasis>
</taxonomicName>
, described from the Sefid River in
<collectingCountry id="F3101FB2FFBBC80CFC0AFA51FC25FA04" box="[919,964,1459,1485]" name="Iran" pageId="7" pageNumber="142">Iran</collectingCountry>
and known to occur in the adjacent Aras, a major tributary of the Kura. Future research should clarify if
<taxonomicName id="4C0724A1FFBBC80CFCC2FA35FC5CFA39" box="[863,957,1494,1521]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="bergi">
<emphasis id="B9738330FFBBC80CFCC2FA35FC5CFA39" box="[863,957,1494,1521]" italics="true" pageId="7" pageNumber="142">O. bergi</emphasis>
</taxonomicName>
is different from
<taxonomicName id="4C0724A1FFBBC80CFB18FA35FAF6FA39" box="[1157,1303,1494,1521]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="bergianus">
<emphasis id="B9738330FFBBC80CFB18FA35FAF6FA39" box="[1157,1303,1494,1521]" italics="true" pageId="7" pageNumber="142">O. bergianus</emphasis>
</taxonomicName>
. If both are conspecific,
<taxonomicName id="4C0724A1FFBBC80CFEB8FA19FE57F9DD" box="[293,438,1530,1557]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="bergianus">
<emphasis id="B9738330FFBBC80CFEB8FA19FE57F9DD" box="[293,438,1530,1557]" italics="true" pageId="7" pageNumber="142">O. bergianus</emphasis>
</taxonomicName>
would become a junior synonym of
<taxonomicName id="4C0724A1FFBBC80CFCCCFA19FC4CF9DD" box="[849,941,1530,1557]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="bergi">
<emphasis id="B9738330FFBBC80CFCCCFA19FC4CF9DD" box="[849,941,1530,1557]" italics="true" pageId="7" pageNumber="142">O. bergi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB85F22FFBBC80CFF5AF9FDFD95F80C" blockId="7.[151,1437,1386,1989]" pageId="7" pageNumber="142">
We also follow
<bibRefCitation id="EF9622D3FFBBC80CFEE5F9FCFDABF9F0" author="Cicek, E. &amp; Eagderi, S. &amp; Sungur, S." box="[376,586,1566,1593]" pageId="7" pageNumber="142" pagination="232 - 242" refId="ref8095" refString="Cicek, E., Eagderi, S. &amp; Sungur, S. (2018) Oxynoemacheilus veyseli, a new nemacheilid species from the upper Aras River drainage of Turkey (Teleostei: Nemacheilidae). Iranian Journal of Ichthyology, 5 (3), 232 - 242. https: // doi. org / 10.22034 / iji. v 5 i 3.302" type="journal article" year="2018">
Çiçek
<emphasis id="B9738330FFBBC80CFE5DF9FDFE18F9F1" box="[448,505,1566,1592]" italics="true" pageId="7" pageNumber="142">et al.</emphasis>
(2018)
</bibRefCitation>
in treating
<taxonomicName id="4C0724A1FFBBC80CFD56F9FDFC29F9F1" box="[715,968,1566,1592]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="subSpecies" species="angorae" subSpecies="alasanicus">
<emphasis id="B9738330FFBBC80CFD56F9FDFC29F9F1" box="[715,968,1566,1592]" italics="true" pageId="7" pageNumber="142">N. angorae alasanicus</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="4C0724A1FFBBC80CFB13F9FDFB0BF9F1" box="[1166,1258,1566,1593]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="bergi">
<emphasis id="B9738330FFBBC80CFB13F9FDFB0BF9F1" box="[1166,1258,1566,1593]" italics="true" pageId="7" pageNumber="142">O. bergi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0724A1FFBBC80CFABCF9FDFEC0F995" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="subSpecies" species="brandtii" subSpecies="gibbusnazus">
<emphasis id="B9738330FFBBC80CFABCF9FDFEC0F995" italics="true" pageId="7" pageNumber="142">N. brandtii gibbusnazus</emphasis>
</taxonomicName>
as a synonym of
<emphasis id="B9738330FFBBC80CFE74F9A1FD8CF995" box="[489,621,1602,1629]" italics="true" pageId="7" pageNumber="142">
<taxonomicName id="4C0724A1FFBBC80CFE74F9A1FD88F995" box="[489,617,1602,1629]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="brandtii">O. brandtii</taxonomicName>
.
</emphasis>
<bibRefCitation id="EF9622D3FFBBC80CFDE9F9A0FCA6F994" author="Cicek, E. &amp; Eagderi, S. &amp; Sungur, S." box="[628,839,1602,1629]" pageId="7" pageNumber="142" pagination="232 - 242" refId="ref8095" refString="Cicek, E., Eagderi, S. &amp; Sungur, S. (2018) Oxynoemacheilus veyseli, a new nemacheilid species from the upper Aras River drainage of Turkey (Teleostei: Nemacheilidae). Iranian Journal of Ichthyology, 5 (3), 232 - 242. https: // doi. org / 10.22034 / iji. v 5 i 3.302" type="journal article" year="2018">
Çiçek
<emphasis id="B9738330FFBBC80CFD20F9A1FD14F995" box="[701,757,1602,1628]" italics="true" pageId="7" pageNumber="142">et al.</emphasis>
(2018)
</bibRefCitation>
did not examine any materials of
<emphasis id="B9738330FFBBC80CFB55F9A1FA9FF995" box="[1224,1406,1602,1628]" italics="true" pageId="7" pageNumber="142">N. a. alasanicus</emphasis>
or
<emphasis id="B9738330FFBBC80CFF0AF985FE83F949" box="[151,354,1638,1664]" italics="true" pageId="7" pageNumber="142">N. b. gibbusnazus</emphasis>
, both described by
<bibRefCitation id="EF9622D3FFBBC80CFDDBF984FD1AF948" author="Elanidze, R. F." box="[582,763,1638,1665]" pageId="7" pageNumber="142" refId="ref8253" refString="Elanidze, R. F. (1983) Ikhtiofauna rek i ozer Gruzii. Akademiya Nauk Gruzinskoi SSR, Institut Zoologii, Metsniereba, Tbilisi, Georgia, 319 pp. [in Russian]" type="book" year="1983">Elanidze (1983)</bibRefCitation>
from the upper Alazani in the Georgian Kura drainage. G. Epitashvili and L. Mumladze (Ilia State University,
<collectingRegion id="49C391C0FFBBC80CFD7FF968FCCFF96D" box="[738,814,1674,1700]" country="Georgia" name="Tbilisi" pageId="7" pageNumber="142">Tbilisi</collectingRegion>
,
<collectingCountry id="F3101FB2FFBBC80CFCA7F968FC77F96C" box="[826,918,1674,1701]" name="Georgia" pageId="7" pageNumber="142">Georgia</collectingCountry>
) visited the upper Alazani though yet only
<taxonomicName id="4C0724A1FFBBC80CFA1CF969FF10F901" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFBBC80CFA1CF969FF10F901" italics="true" pageId="7" pageNumber="142">O. brandtii</emphasis>
</taxonomicName>
was collected.
<bibRefCitation id="EF9622D3FFBBC80CFE07F94CFD89F900" author="Cicek, E. &amp; Eagderi, S. &amp; Sungur, S." box="[410,616,1710,1737]" pageId="7" pageNumber="142" pagination="232 - 242" refId="ref8095" refString="Cicek, E., Eagderi, S. &amp; Sungur, S. (2018) Oxynoemacheilus veyseli, a new nemacheilid species from the upper Aras River drainage of Turkey (Teleostei: Nemacheilidae). Iranian Journal of Ichthyology, 5 (3), 232 - 242. https: // doi. org / 10.22034 / iji. v 5 i 3.302" type="journal article" year="2018">
Çiçek
<emphasis id="B9738330FFBBC80CFE7CF94DFDF9F901" box="[481,536,1710,1736]" italics="true" pageId="7" pageNumber="142">et al.</emphasis>
(2018)
</bibRefCitation>
give no arguments for their conclusion and did not analyse the description of both species. However, we also have no arguments to reject the hypothesis of
<bibRefCitation id="EF9622D3FFBBC80CFB81F930FB11F924" author="Cicek, E. &amp; Eagderi, S. &amp; Sungur, S." box="[1052,1264,1746,1773]" pageId="7" pageNumber="142" pagination="232 - 242" refId="ref8095" refString="Cicek, E., Eagderi, S. &amp; Sungur, S. (2018) Oxynoemacheilus veyseli, a new nemacheilid species from the upper Aras River drainage of Turkey (Teleostei: Nemacheilidae). Iranian Journal of Ichthyology, 5 (3), 232 - 242. https: // doi. org / 10.22034 / iji. v 5 i 3.302" type="journal article" year="2018">
Çiçek
<emphasis id="B9738330FFBBC80CFBF9F931FB7DF925" box="[1124,1180,1746,1772]" italics="true" pageId="7" pageNumber="142">et al.</emphasis>
(2018)
</bibRefCitation>
, as the descriptions are not very conclusive, without figures and we also did not examine enough materials by our own. Only four loaches from the
<typeStatus id="54BCE180FFBBC80CFEC5F8F9FE69F8FC" box="[344,392,1819,1845]" pageId="7" pageNumber="142">type</typeStatus>
locality of both species were available for this study and these are identified as
<taxonomicName id="4C0724A1FFBBC80CFA9CF8F9FA9CF8FD" box="[1281,1405,1818,1845]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFBBC80CFA9CF8F9FA9CF8FD" box="[1281,1405,1818,1845]" italics="true" pageId="7" pageNumber="142">O. brandtii</emphasis>
</taxonomicName>
. It is likely that
<bibRefCitation id="EF9622D3FFBBC80CFEB7F8DCFE3FF890" author="Elanidze, R. F." box="[298,478,1854,1881]" pageId="7" pageNumber="142" refId="ref8253" refString="Elanidze, R. F. (1983) Ikhtiofauna rek i ozer Gruzii. Akademiya Nauk Gruzinskoi SSR, Institut Zoologii, Metsniereba, Tbilisi, Georgia, 319 pp. [in Russian]" type="book" year="1983">Elanidze (1983)</bibRefCitation>
had identified his materials by the key provided by
<bibRefCitation id="EF9622D3FFBBC80CFBB4F8DDFB50F890" author="Berg, L. S." box="[1065,1201,1855,1881]" pageId="7" pageNumber="142" pagination="927 - 1382" refId="ref8027" refString="Berg, L. S. (1949) The freshwater fishes of the USSR and adjacent countries. Part 3. 4 th Edition. Akademia Nauk USSR, Moscow &amp; Leningrad, pp. 927 - 1382. [in Russian]" type="book chapter" year="1949">Berg (1949)</bibRefCitation>
and correctly identified one species as
<taxonomicName id="4C0724A1FFBBC80CFEECF881FE0CF8B5" box="[369,493,1890,1917]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFBBC80CFEECF881FE0CF8B5" box="[369,493,1890,1917]" italics="true" pageId="7" pageNumber="142">O. brandtii</emphasis>
</taxonomicName>
and the second species, which has a deeper caudal peduncle and an emarginate (vs. furcate) fin, was identified as
<taxonomicName id="4C0724A1FFBBC80CFE7BF865FD84F869" baseAuthorityName="Steindachner" baseAuthorityYear="1897" box="[486,613,1927,1953]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="angorae">
<emphasis id="B9738330FFBBC80CFE7BF865FD84F869" box="[486,613,1927,1953]" italics="true" pageId="7" pageNumber="142">O. angorae</emphasis>
</taxonomicName>
. He then described new subspecies for both species, and thus
<taxonomicName id="4C0724A1FFBBC80CFA83F865FEEEF80D" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="subSpecies" species="angorae" subSpecies="alasanicus">
<emphasis id="B9738330FFBBC80CFA83F865FEEEF80D" italics="true" pageId="7" pageNumber="142">N. angorae alasanicus</emphasis>
</taxonomicName>
might be identified as
<taxonomicName id="4C0724A1FFBBC80CFD8DF849FD8CF80D" box="[528,621,1962,1989]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="142" phylum="Chordata" rank="species" species="bergi">
<emphasis id="B9738330FFBBC80CFD8DF849FD8CF80D" box="[528,621,1962,1989]" italics="true" pageId="7" pageNumber="142">O. bergi</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF780FAAFFB4C803FF0AFC16FD99FB9D" ID-DOI="http://doi.org/10.5281/zenodo.4671926" ID-Zenodo-Dep="4671926" httpUri="https://zenodo.org/record/4671926/files/figure.png" pageId="8" pageNumber="143" startId="8.[151,250,1012,1036]" targetBox="[151,1436,181,988]" targetPageId="8">
<paragraph id="8BB85F22FFB4C803FF0AFC16FD99FB9D" blockId="8.[151,1436,1012,1109]" pageId="8" pageNumber="143">
<emphasis id="B9738330FFB4C803FF0AFC16FEF2FBC5" bold="true" box="[151,275,1012,1036]" pageId="8" pageNumber="143">FIGURE 4.</emphasis>
<taxonomicName id="4C0724A1FFB4C803FE84FC16FDCDFBC5" box="[281,556,1012,1036]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="phasicus">
<emphasis id="B9738330FFB4C803FE84FC16FDCDFBC5" box="[281,556,1012,1036]" italics="true" pageId="8" pageNumber="143">Oxynoemacheilus phasicus</emphasis>
</taxonomicName>
, paratypes, from top, left to right: FSJF 4061, 32 mm SL; 39 mm SL; Georgia: Gubistskali River; FSJF 3615, 44 mm SL, 54 mm SL; Georgia: Tskhenitskali River; FSJF 3576, 59 mm SL; Georgia: Enguri River; FSJF 3592, 77 mm SL; Georgia: Dzirula River.
</paragraph>
</caption>
<caption id="DF780FAAFFB4C803FF0AF89EFD99F815" ID-DOI="http://doi.org/10.5281/zenodo.4671930" ID-Zenodo-Dep="4671930" httpUri="https://zenodo.org/record/4671930/files/figure.png" pageId="8" pageNumber="143" startId="8.[151,250,1916,1940]" targetBox="[151,1436,1133,1893]" targetPageId="8">
<paragraph id="8BB85F22FFB4C803FF0AF89EFD99F815" blockId="8.[151,1436,1916,2012]" pageId="8" pageNumber="143">
<emphasis id="B9738330FFB4C803FF0AF89EFEF2F85D" bold="true" box="[151,275,1916,1940]" pageId="8" pageNumber="143">FIGURE 5.</emphasis>
<taxonomicName id="4C0724A1FFB4C803FE84F89EFDCDF85D" box="[281,556,1916,1940]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="phasicus">
<emphasis id="B9738330FFB4C803FE84F89EFDCDF85D" box="[281,556,1916,1940]" italics="true" pageId="8" pageNumber="143">Oxynoemacheilus phasicus</emphasis>
</taxonomicName>
, paratypes, from top, left to right: FSJF 4061, 32 mm SL; 39 mm SL; Georgia: Gubistskali River; FSJF 3615, 44 mm SL, 54 mm SL; Georgia: Tskhenitskali River; FSJF 3576, 59 mm SL; Georgia: Enguri River; FSJF 3592, 77 mm SL; Georgia: Dzirula River.
</paragraph>
</caption>
<caption id="DF780FAAFFB5C802FF0AFC79FA64FC7A" ID-DOI="http://doi.org/10.5281/zenodo.4671932" ID-Zenodo-Dep="4671932" box="[151,1413,923,947]" httpUri="https://zenodo.org/record/4671932/files/figure.png" pageId="9" pageNumber="144" startId="9.[151,250,923,947]" targetBox="[181,1406,219,885]" targetPageId="9">
<paragraph id="8BB85F22FFB5C802FF0AFC79FA64FC7A" blockId="9.[151,1413,923,947]" box="[151,1413,923,947]" pageId="9" pageNumber="144">
<emphasis id="B9738330FFB5C802FF0AFC79FEF2FC7A" bold="true" box="[151,275,923,947]" pageId="9" pageNumber="144">FIGURE 6.</emphasis>
Records of
<taxonomicName id="4C0724A1FFB5C802FE13FC79FD7AFC7A" box="[398,667,923,947]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB5C802FE13FC79FD7AFC7A" box="[398,667,923,947]" italics="true" pageId="9" pageNumber="144">Oxynoemacheilus brandtii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0724A1FFB5C802FD56FC7EFCA2FC7A" box="[715,835,923,947]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="phasicus">
<emphasis id="B9738330FFB5C802FD56FC7EFCA2FC7A" box="[715,835,923,947]" italics="true" pageId="9" pageNumber="144">O. phasicus</emphasis>
</taxonomicName>
in Georgia based on site-scale records mentioned above.
</paragraph>
</caption>
<caption id="DF780FAAFFB5C802FF0AFAEAFECCFA8D" ID-DOI="http://doi.org/10.5281/zenodo.4671934" ID-Zenodo-Dep="4671934" httpUri="https://zenodo.org/record/4671934/files/figure.png" pageId="9" pageNumber="144" startId="9.[151,250,1288,1312]" targetBox="[194,1361,993,1241]" targetPageId="9">
<paragraph id="8BB85F22FFB5C802FF0AFAEAFECCFA8D" blockId="9.[151,1437,1288,1348]" pageId="9" pageNumber="144">
<emphasis id="B9738330FFB5C802FF0AFAEAFEF2FAE9" bold="true" box="[151,275,1288,1312]" pageId="9" pageNumber="144">FIGURE 7.</emphasis>
<taxonomicName id="4C0724A1FFB5C802FE87FAEAFDEBFAE9" box="[282,522,1288,1312]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="bergi">
<emphasis id="B9738330FFB5C802FE87FAEAFDEBFAE9" box="[282,522,1288,1312]" italics="true" pageId="9" pageNumber="144">Oxynoemacheilus bergi</emphasis>
</taxonomicName>
; IMNRF-1030-2, 50.4 mm SL; Azerbaijan: Kura River, 41°1425N 45°2557E. Photo: Soheil Eagderi
</paragraph>
</caption>
<paragraph id="8BB85F22FFB5C801FF5AFA90FCDEFF30" blockId="9.[151,1437,1394,2033]" lastBlockId="10.[151,1436,151,249]" lastPageId="10" lastPageNumber="145" pageId="9" pageNumber="144">
<bibRefCitation id="EF9622D3FFB5C802FF5AFA90FE5BFA45" author="Eagderi, S. &amp; Jalili, P. &amp; Cicek, E." box="[199,442,1394,1421]" pageId="9" pageNumber="144" pagination="453 - 459" refId="ref8162" refString="Eagderi, S., Jalili, P. &amp; Cicek, E. (2018) Oxynoemacheilus elsae, a new species from the Urmia Lake basin of Iran (Teleostei: Nemacheilidae). FishTaxa, 3 (2), 453 - 459." type="journal article" year="2018">
Eagderi
<emphasis id="B9738330FFB5C802FEB4FA91FEBFFA45" box="[297,350,1394,1420]" italics="true" pageId="9" pageNumber="144">et al</emphasis>
. (2018)
</bibRefCitation>
distinguish
<taxonomicName id="4C0724A1FFB5C802FDD7FA91FD4BFA45" box="[586,682,1394,1421]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB5C802FDD7FA91FD4BFA45" box="[586,682,1394,1421]" italics="true" pageId="9" pageNumber="144">O. elsae</emphasis>
</taxonomicName>
(
<figureCitation id="133C43A7FFB5C802FD21FA90FCE2FA45" box="[700,771,1394,1420]" captionStart="FIGURE 9" captionStartId="11.[151,250,1320,1344]" captionTargetBox="[151,1436,181,1296]" captionTargetId="figure-19@11.[151,1436,181,1296]" captionTargetPageId="11" captionText="FIGURE 9. Oxynoemacheilus elsae FSJF 4109, from top: 55 mm SL; 60 mm SL; 53 mm SL; Iran: Zarrineh River." figureDoi="http://doi.org/10.5281/zenodo.4671938" httpUri="https://zenodo.org/record/4671938/files/figure.png" pageId="9" pageNumber="144">Fig. 9</figureCitation>
) from
<taxonomicName id="4C0724A1FFB5C802FCCAFA91FC36FA45" box="[855,983,1394,1421]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB5C802FCCAFA91FC36FA45" box="[855,983,1394,1421]" italics="true" pageId="9" pageNumber="144">O. brandtii</emphasis>
</taxonomicName>
(
<figureCitation id="133C43A7FFB5C802FC77FA90FBD1FA45" box="[1002,1072,1394,1420]" captionStart="FIGURE 8" captionStartId="10.[151,250,1832,1856]" captionTargetBox="[151,1436,273,1809]" captionTargetId="figure-71@10.[151,1436,273,1809]" captionTargetPageId="10" captionText="FIGURE 8. Oxynoemacheilus brandtii, Kura drainage, from top: Not preserved, about 70 mm SL; Georgia: Alazani River; FSJF 3430, 50 mm SL; 48 mm SL; Turkey: stream between Göle and Ardahan; Not preserved; about 65 mm SL; Kura River at Yalnızçam." figureDoi="http://doi.org/10.5281/zenodo.4671936" httpUri="https://zenodo.org/record/4671936/files/figure.png" pageId="9" pageNumber="144">Fig. 8</figureCitation>
) based on 10, non-overlapping morphometric characters (dorsal-fin base length, anal-fin base length, anal-fin depth, pelvic-fin length, caudal-fin length, caudal-peduncle width, snout length, eye diameter, interorbital distance, maxillary barbel length). They compared 20
<taxonomicName id="4C0724A1FFB5C802FEB2FA3DFE6AFA31" box="[303,395,1502,1529]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB5C802FEB2FA3DFE6AFA31" box="[303,395,1502,1529]" italics="true" pageId="9" pageNumber="144">O. elsae</emphasis>
</taxonomicName>
with five
<taxonomicName id="4C0724A1FFB5C802FE66FA3DFD96FA31" box="[507,631,1502,1529]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB5C802FE66FA3DFD96FA31" box="[507,631,1502,1529]" italics="true" pageId="9" pageNumber="144">O. brandtii</emphasis>
</taxonomicName>
from the Kura drainage. We examined 20 individuals from each species and found all the 10 morphometric characters overlapping. In seven out of 10 characters no significant difference from equality of their mean values were detected. Only the means of the pelvic-fin length, length of the lower caudal-fin lobe, and the eye diameter were significantly different (
<figureCitation id="133C43A7FFB5C802FCCCF9A8FC45F9AD" box="[849,932,1610,1636]" captionStart="FIGURE 10" captionStartId="12.[151,250,1864,1888]" captionTargetBox="[156,1427,195,1827]" captionTargetId="figure-25@12.[156,1432,181,1838]" captionTargetPageId="12" captionText="FIGURE 10. Morphometric characters of O. brandtii from Aras (FSJF 3441 n=9, FFR 1427 n=11, FFR 1351 n=9), and Kura (FSJF 3544 n=7, FSJF 3571 n=5, FSJF 3630 n=1, FSJF 3627 n=1, FSJF 4023 n=6) and O. elsae from Zarrineh (FSJF 2264 n=9, FSJF 4109 (n=11)." figureDoi="http://doi.org/10.5281/zenodo.4671940" httpUri="https://zenodo.org/record/4671940/files/figure.png" pageId="9" pageNumber="144">Fig. 10</figureCitation>
). Also these morphometric characters largely overlap: pelvic-fin length 1420, mean 18% SL in
<taxonomicName id="4C0724A1FFB5C802FD42F98DFCBCF941" box="[735,861,1646,1673]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB5C802FD42F98DFCBCF941" box="[735,861,1646,1673]" italics="true" pageId="9" pageNumber="144">O. brandtii</emphasis>
</taxonomicName>
(vs. 1417, mean
<quantity id="4CFFF2C7FFB5C802FBAEF98CFB8FF941" box="[1075,1134,1646,1672]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="9" pageNumber="144" unit="in" value="16.0">16 in</quantity>
<taxonomicName id="4C0724A1FFB5C802FBEAF98DFB34F941" box="[1143,1237,1646,1673]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB5C802FBEAF98DFB34F941" box="[1143,1237,1646,1673]" italics="true" pageId="9" pageNumber="144">O. elsae</emphasis>
</taxonomicName>
), length of lower caudal-fin lobe 2027, mean 24% SL (vs. 1923, mean 21), and the eye diameter 1321, mean 17% HL (vs. 1724, mean 20). The overlap in the pelvic-fin length is due to two individuals only and from 49
<taxonomicName id="4C0724A1FFB5C802FBE0F955FB18F919" box="[1149,1273,1718,1745]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB5C802FBE0F955FB18F919" box="[1149,1273,1718,1745]" italics="true" pageId="9" pageNumber="144">O. brandtii</emphasis>
</taxonomicName>
all except two individuals, have a pelvic-fin length 1720 % SL, one has 14% SL and one has 15% SL. This character allows to distinguish
<taxonomicName id="4C0724A1FFB5C802FE87F91DFE76F8D1" box="[282,407,1790,1817]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB5C802FE87F91DFE76F8D1" box="[282,407,1790,1817]" italics="true" pageId="9" pageNumber="144">O. brandtii</emphasis>
</taxonomicName>
from
<taxonomicName id="4C0724A1FFB5C802FE43F91DFDDAF8D1" box="[478,571,1790,1817]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB5C802FE43F91DFDDAF8D1" box="[478,571,1790,1817]" italics="true" pageId="9" pageNumber="144">O. elsae</emphasis>
</taxonomicName>
if a larger number of individuals is available, while the other characters can just support their diagnosis.
<taxonomicName id="4C0724A1FFB5C802FE53F8C0FD34F8F5" box="[462,725,1826,1852]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB5C802FE53F8C0FD34F8F5" box="[462,725,1826,1852]" italics="true" pageId="9" pageNumber="144">Oxynoemacheilus elsae</emphasis>
</taxonomicName>
is well distinguished from
<taxonomicName id="4C0724A1FFB5C802FC62F8C1FB9BF8F5" box="[1023,1146,1826,1853]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB5C802FC62F8C1FB9BF8F5" box="[1023,1146,1826,1853]" italics="true" pageId="9" pageNumber="144">O. brandtii</emphasis>
</taxonomicName>
by the shape of the caudal fin, which is emarginate in
<taxonomicName id="4C0724A1FFB5C802FE52F8A5FDCCF8A9" box="[463,557,1862,1889]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB5C802FE52F8A5FDCCF8A9" box="[463,557,1862,1889]" italics="true" pageId="9" pageNumber="144">O. elsae</emphasis>
</taxonomicName>
(vs. forked in
<taxonomicName id="4C0724A1FFB5C802FD48F8A5FCB2F8A9" box="[725,851,1862,1889]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB5C802FD48F8A5FCB2F8A9" box="[725,851,1862,1889]" italics="true" pageId="9" pageNumber="144">O. brandtii</emphasis>
</taxonomicName>
) and by details in colour pattern.
<taxonomicName id="4C0724A1FFB5C802FB48F8A4FF30F84D" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="144" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB5C802FB48F8A4FF30F84D" italics="true" pageId="9" pageNumber="144">Oxynoemacheilus elsae</emphasis>
</taxonomicName>
usually possess a flank pattern of irregularly shaped and spaced, dark-brown bars, often split in middle, more narrow than interspaces, or flank almost plain brown (vs. flank with brown bars or vertically elongate blotches, much wider than interspaces, usually regularly set) and the caudal fin possess many small brown blotches on rays arranged in 36, narrow, pale-brown band (vs. caudal fin with 13, wide and very bold bands). However, the flank pattern is variable in both species and occasionally almost plain brown individuals or individuals with many, very narrow bars occur.
<taxonomicName id="4C0724A1FFB6C801FEF7FF59FD91FF1C" box="[362,624,187,213]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="145" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB6C801FEF7FF59FD91FF1C" box="[362,624,187,213]" italics="true" pageId="10" pageNumber="145">Oxynoemacheilus elsae</emphasis>
</taxonomicName>
is separated from
<taxonomicName id="4C0724A1FFB6C801FCA4FF59FC55FF1C" box="[825,948,187,213]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="145" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB6C801FCA4FF59FC55FF1C" box="[825,948,187,213]" italics="true" pageId="10" pageNumber="145">O. brandtii</emphasis>
</taxonomicName>
by a minimum K2P distance of 1.5% (Aras) and 4.9% (Kura) in the mtDNA COI barcode region (
<figureCitation id="133C43A7FFB6C801FD70FF3DFCCFFF30" box="[749,814,223,249]" captionStart="FIGURE 1" captionStartId="3.[151,250,1849,1873]" captionTargetBox="[327,1235,190,1820]" captionTargetId="figure-19@3.[319,1268,181,1825]" captionTargetPageId="3" captionText="FIGURE 1. Maximum likelihood estimation of the phylogenetic relationships based on the mitochondrial DNA barcode region (Tamura-Nei model, discrete Gamma distribution for rate differences with 3 categories (+G, parameter = 0.3477). Nucleotide positions with less than 98% site coverage were eliminated, resulting in 557 analysed positions. Numbers of major nodes indicate bootstrap values from 1000 pseudoreplicates from the ML, NJ and MP methods. The tree is drawn to scale with branch lengths depicting number of substitutions per site." figureDoi="http://doi.org/10.5281/zenodo.4671918" httpUri="https://zenodo.org/record/4671918/files/figure.png" pageId="10" pageNumber="145">Fig. 1</figureCitation>
).
</paragraph>
<caption id="DF780FAAFFB6C801FF0AF8CAFEEBF841" ID-DOI="http://doi.org/10.5281/zenodo.4671936" ID-Zenodo-Dep="4671936" httpUri="https://zenodo.org/record/4671936/files/figure.png" pageId="10" pageNumber="145" startId="10.[151,250,1832,1856]" targetBox="[151,1436,273,1809]" targetPageId="10">
<paragraph id="8BB85F22FFB6C801FF0AF8CAFEEBF841" blockId="10.[151,1436,1832,1928]" pageId="10" pageNumber="145">
<emphasis id="B9738330FFB6C801FF0AF8CAFEF4F889" bold="true" box="[151,277,1832,1856]" pageId="10" pageNumber="145">FIGURE 8.</emphasis>
<taxonomicName id="4C0724A1FFB6C801FE83F8CAFD8FF889" authority=", Kura" authorityName="Kura" box="[286,622,1832,1856]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="145" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB6C801FE83F8CAFDCCF889" box="[286,557,1832,1856]" italics="true" pageId="10" pageNumber="145">Oxynoemacheilus brandtii</emphasis>
, Kura
</taxonomicName>
drainage, from top: Not preserved, about 70 mm SL; Georgia: Alazani River; FSJF 3430, 50 mm SL; 48 mm SL; Turkey: stream between Göle and Ardahan; Not preserved; about 65 mm SL; Kura River at Yalnızçam.
</paragraph>
</caption>
<caption id="DF780FAAFFB7C800FF0AFACAFAC9FA89" ID-DOI="http://doi.org/10.5281/zenodo.4671938" ID-Zenodo-Dep="4671938" box="[151,1320,1320,1344]" httpUri="https://zenodo.org/record/4671938/files/figure.png" pageId="11" pageNumber="146" startId="11.[151,250,1320,1344]" targetBox="[151,1436,181,1296]" targetPageId="11">
<paragraph id="8BB85F22FFB7C800FF0AFACAFAC9FA89" blockId="11.[151,1320,1320,1344]" box="[151,1320,1320,1344]" pageId="11" pageNumber="146">
<emphasis id="B9738330FFB7C800FF0AFACAFEF2FA89" bold="true" box="[151,275,1320,1344]" pageId="11" pageNumber="146">FIGURE 9.</emphasis>
<taxonomicName id="4C0724A1FFB7C800FE84FACAFDA3FA89" authority="FSJF" authorityName="FSJF" box="[281,578,1320,1344]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB7C800FE84FACAFDE9FA89" box="[281,520,1320,1344]" italics="true" pageId="11" pageNumber="146">Oxynoemacheilus elsae</emphasis>
FSJF
</taxonomicName>
4109, from top: 55 mm SL; 60 mm SL; 53 mm SL; Iran: Zarrineh River.
</paragraph>
</caption>
<paragraph id="8BB85F22FFB7C800FF5AFA8CFE37F895" blockId="11.[151,1437,1390,1885]" pageId="11" pageNumber="146">
<taxonomicName id="4C0724A1FFB7C800FF5AFA8CFE6FFA41" box="[199,398,1390,1416]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="genus">
<emphasis id="B9738330FFB7C800FF5AFA8CFE6FFA41" box="[199,398,1390,1416]" italics="true" pageId="11" pageNumber="146">Oxynoemacheilus</emphasis>
</taxonomicName>
from the Aras River (
<figureCitation id="133C43A7FFB7C800FDE2FA8CFD31FA41" box="[639,720,1390,1416]" captionStart="FIGURE 11" captionStartId="13.[151,250,1892,1916]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-19@13.[151,1436,181,1868]" captionTargetPageId="13" captionText="FIGURE 11. Oxynoemacheilus brandtii,Aras drainage,Turkey:from top:FFR 15549, 54 mm SL; 52 mm SL, stream Büyükçayır; FFR 15551, 54 mm SL, stream Fehmiharabesi; FSJF 3441, 65 mm SL; Turkey: stream Derinöz." figureDoi="http://doi.org/10.5281/zenodo.4671942" httpUri="https://zenodo.org/record/4671942/files/figure.png" pageId="11" pageNumber="146">Fig. 11</figureCitation>
), a major tributary to the Kura, cluster as sister to
<taxonomicName id="4C0724A1FFB7C800FB67FA8DFAB4FA41" box="[1274,1365,1390,1417]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB7C800FB67FA8DFAB4FA41" box="[1274,1365,1390,1417]" italics="true" pageId="11" pageNumber="146">O. elsae</emphasis>
</taxonomicName>
in our molecular analysis based on all three approaches (ML, NJ, MP). In all morphological characters distinguishing
<taxonomicName id="4C0724A1FFB7C800FA1CFA71FF30FA19" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB7C800FA1CFA71FF30FA19" italics="true" pageId="11" pageNumber="146">O. elsae</emphasis>
</taxonomicName>
from
<taxonomicName id="4C0724A1FFB7C800FE8EFA55FE6EFA19" box="[275,399,1462,1489]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB7C800FE8EFA55FE6EFA19" box="[275,399,1462,1489]" italics="true" pageId="11" pageNumber="146">O. brandtii</emphasis>
</taxonomicName>
, the population from the Aras agrees with
<taxonomicName id="4C0724A1FFB7C800FCFBFA55FC03FA19" box="[870,994,1462,1489]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB7C800FCFBFA55FC03FA19" box="[870,994,1462,1489]" italics="true" pageId="11" pageNumber="146">O. brandtii</emphasis>
</taxonomicName>
and not with
<taxonomicName id="4C0724A1FFB7C800FBE5FA55FB35FA19" box="[1144,1236,1462,1489]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB7C800FBE5FA55FB35FA19" box="[1144,1236,1462,1489]" italics="true" pageId="11" pageNumber="146">O. elsae</emphasis>
</taxonomicName>
(
<figureCitation id="133C43A7FFB7C800FB7FFA54FAC5FA19" box="[1250,1316,1462,1488]" captionStart="FIGURE 9" captionStartId="11.[151,250,1320,1344]" captionTargetBox="[151,1436,181,1296]" captionTargetId="figure-19@11.[151,1436,181,1296]" captionTargetPageId="11" captionText="FIGURE 9. Oxynoemacheilus elsae FSJF 4109, from top: 55 mm SL; 60 mm SL; 53 mm SL; Iran: Zarrineh River." figureDoi="http://doi.org/10.5281/zenodo.4671938" httpUri="https://zenodo.org/record/4671938/files/figure.png" pageId="11" pageNumber="146">Fig. 9</figureCitation>
) including the characteristic colour pattern of
<taxonomicName id="4C0724A1FFB7C800FD82FA39FD7DFA3D" box="[543,668,1498,1525]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB7C800FD82FA39FD7DFA3D" box="[543,668,1498,1525]" italics="true" pageId="11" pageNumber="146">O. brandtii</emphasis>
</taxonomicName>
. These fish are significantly different from
<taxonomicName id="4C0724A1FFB7C800FB1BFA39FB03FA3D" box="[1158,1250,1498,1525]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB7C800FB1BFA39FB03FA3D" box="[1158,1250,1498,1525]" italics="true" pageId="11" pageNumber="146">O. elsae</emphasis>
</taxonomicName>
in the pelvic-fin length mentioned above, but not different from
<taxonomicName id="4C0724A1FFB7C800FD30FA1DFCC8F9D1" box="[685,809,1534,1561]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB7C800FD30FA1DFCC8F9D1" box="[685,809,1534,1561]" italics="true" pageId="11" pageNumber="146">O. brandtii</emphasis>
</taxonomicName>
in any of these or other characters examined. The obvious mismatch between morphological and molecular characters, which suggests that the fish from the Aras should be identified as
<taxonomicName id="4C0724A1FFB7C800FED5F9A5FE45F9A9" box="[328,420,1606,1633]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB7C800FED5F9A5FE45F9A9" box="[328,420,1606,1633]" italics="true" pageId="11" pageNumber="146">O. elsae</emphasis>
</taxonomicName>
cannot be resolved here. It could be speculated that past river-captures might have resulted in an invasion of a mitochondrial haplotype of
<taxonomicName id="4C0724A1FFB7C800FD36F989FCE8F94D" box="[683,777,1642,1669]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB7C800FD36F989FCE8F94D" box="[683,777,1642,1669]" italics="true" pageId="11" pageNumber="146">O. elsae</emphasis>
</taxonomicName>
from Urmia into the adjacent Aras. While they hybridised with native
<emphasis id="B9738330FFB7C800FE81F96DFE7EF961" box="[284,415,1678,1705]" italics="true" pageId="11" pageNumber="146">
<taxonomicName id="4C0724A1FFB7C800FE81F96DFE7AF961" box="[284,411,1678,1705]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="brandtii">O. brandtii</taxonomicName>
,
</emphasis>
their mitochondria might have performed better and gave the carriers an evolutionary benefit which spread in the Aras. If so, what cannot be excluded, we would find
<taxonomicName id="4C0724A1FFB7C800FC61F951FB98F905" box="[1020,1145,1714,1741]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB7C800FC61F951FB98F905" box="[1020,1145,1714,1741]" italics="true" pageId="11" pageNumber="146">O. brandtii</emphasis>
</taxonomicName>
with mtDNA of
<taxonomicName id="4C0724A1FFB7C800FAA4F951FA77F905" box="[1337,1430,1714,1741]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB7C800FAA4F951FA77F905" box="[1337,1430,1714,1741]" italics="true" pageId="11" pageNumber="146">O. elsae</emphasis>
</taxonomicName>
. The small molecular distance between
<taxonomicName id="4C0724A1FFB7C800FDC9F935FD53F939" box="[596,690,1750,1777]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="elsae">
<emphasis id="B9738330FFB7C800FDC9F935FD53F939" box="[596,690,1750,1777]" italics="true" pageId="11" pageNumber="146">O. elsae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0724A1FFB7C800FD76F935FC88F939" box="[747,873,1750,1777]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB7C800FD76F935FC88F939" box="[747,873,1750,1777]" italics="true" pageId="11" pageNumber="146">O. brandtii</emphasis>
</taxonomicName>
from the Aras in their mtDNA indicates the time passed since the connection of the Aras with the Urmia basin. Further nuclear molecular research is needed in the future aiming to clarify the case. We suggest to identify the fish from the Aras as
<taxonomicName id="4C0724A1FFB7C800FBBDF8FDFB7DF8F1" box="[1056,1180,1822,1849]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="146" phylum="Chordata" rank="species" species="brandtii">
<emphasis id="B9738330FFB7C800FBBDF8FDFB7DF8F1" box="[1056,1180,1822,1849]" italics="true" pageId="11" pageNumber="146">O. brandtii</emphasis>
</taxonomicName>
due to its morphological similarity to this species.
</paragraph>
</subSubSection>
</treatment>
</document>