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<document id="7938FBA0FBC9AF3B9999C7B63B5C15A6" ID-DOI="10.5281/zenodo.5301059" ID-ISSN="0374-1036" ID-Zenodo-Dep="5301059" ID-ZooBank="EA465CB7-1796-4656-9D14-EF0964899EDAD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630099098575" checkinUser="marcus" docAuthor="Makranczy, György" docDate="2014" docId="03AECB0EDC4FFFF6FE86356D52F495CE" docLanguage="en" docName="ActaEntMusNatPra.2014.54.2.539-554.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Thinodromus circulus" docType="treatment" docVersion="3" lastPageNumber="544" masterDocId="FF97B376DC4CFFF3FFF33756520A915D" masterDocTitle="Review of the Thinodromus circulus species group (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="554" masterPageNumber="539" pageNumber="542" updateTime="1699149139555" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7B28A57E52637BAA50F46C8EB8966C51">Review of the Thinodromus circulus species group (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:namePart id="BF231411087F83C95C83AB8DCE7C45FB">Makranczy, György</mods:namePart>
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<mods:title id="5B08146A546BEEC85387AEF4DF5109ED">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="5E6D45D373E009CA09850ADC2F359F0C">2014</mods:date>
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<mods:number id="282685BC6C8BE1CFFDFF88BAD09A9370">2014-12-15</mods:number>
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<mods:identifier id="D7EFF80D4DC1B1EAC020821A1F5EC28A" type="ISSN">0374-1036</mods:identifier>
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<treatment id="03AECB0EDC4FFFF6FE86356D52F495CE" LSID="urn:lsid:plazi:treatment:03AECB0EDC4FFFF6FE86356D52F495CE" httpUri="http://treatment.plazi.org/id/03AECB0EDC4FFFF6FE86356D52F495CE" lastPageId="5" lastPageNumber="544" pageId="3" pageNumber="542">
<subSubSection id="C31D2993DC4FFFF0FE86356D5137930F" box="[373,829,571,594]" pageId="3" pageNumber="542" type="nomenclature">
<paragraph id="8BB87A18DC4FFFF0FE86356D5137930F" blockId="3.[373,829,571,594]" box="[373,829,571,594]" pageId="3" pageNumber="542">
<heading id="D0F0CD74DC4FFFF0FE86356D5137930F" bold="true" box="[373,829,571,594]" centered="true" fontSize="8" level="2" pageId="3" pageNumber="542" reason="3">
<taxonomicName id="4C07019BDC4FFFF0FE86356D5137930F" ID-CoL="56H4Q" authority="(Bernhauer, 1922)" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[373,829,571,594]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="circulus">
<emphasis id="B973A60ADC4FFFF0FE86356D5137930F" bold="true" box="[373,829,571,594]" pageId="3" pageNumber="542">
<emphasis id="B973A60ADC4FFFF0FE86356D5060930F" bold="true" box="[373,618,571,594]" italics="true" pageId="3" pageNumber="542">Thinodromus circulus</emphasis>
(Bernhauer, 1922)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31D2993DC4FFFF0FE1E350F50CF9331" box="[493,709,601,620]" pageId="3" pageNumber="542" type="description">
<paragraph id="8BB87A18DC4FFFF0FE1E350F50CF9331" blockId="3.[113,891,601,708]" box="[493,709,601,620]" pageId="3" pageNumber="542">
(
<figureCitation id="133C669DDC4FFFF0FE00350F50239331" box="[499,553,601,620]" captionStart="Figs 14" captionStartId="2.[151,188,1526,1545]" captionTargetBox="[163,1115,218,1465]" captionTargetId="figure-10@2.[163,1116,217,1465]" captionTargetPageId="2" captionText="Figs 14. Forebodies of Thinodromus species. 1 T. circulus (Bernhauer, 1922); 2 T. chagosanus (Bernhauer, 1922); 3 T. palustris (Bernhauer, 1922); 4 T. arcitenens (Fauvel, 1905)." figureDoi="http://doi.org/10.5281/zenodo.5301061" httpUri="https://zenodo.org/record/5301061/files/figure.png" pageId="3" pageNumber="542">Figs 1</figureCitation>
,
<figureCitation id="133C669DDC4FFFF0FDC0350F506A9331" box="[563,608,601,620]" captionStart="Figs 514. 510" captionStartId="4.[151,188,1497,1516]" captionTargetBox="[151,1100,223,1466]" captionTargetId="figure-9@4.[151,1103,221,1468]" captionTargetPageId="4" captionText="Figs 514. 510 Thinodromus circulus (Bernhauer, 1922) (5 mesotarsus; 6 male sternite VIII; 7 male tergite IX; 8 male sternite IX; 9 male tergite X; 10 spermatheca); 1112 T. velutinus (Sharp, 1887) (11 tergite X; 12 spermatheca); 1314 T. chagosanus (Bernhauer, 1922) (13 mesotarsus; 14 platelike armature in hypopharynx). Scale bar: 0.06 mm for 14; 0.07 mm for 5, 13; 0.1 mm for 1012; 0.12 mm for 89; 0.14 mm for 67." figureDoi="http://doi.org/10.5281/zenodo.5301063" httpUri="https://zenodo.org/record/5301063/files/figure.png" pageId="3" pageNumber="542">510</figureCitation>
,
<figureCitation id="133C669DDC4FFFF0FD9A350F50AA9331" box="[617,672,601,620]" captionStart="Figs 2024. 20" captionStartId="7.[113,150,1480,1499]" captionTargetBox="[127,1077,222,1463]" captionTargetId="figure-9@7.[125,1078,219,1466]" captionTargetPageId="7" captionText="Figs 2024. 20 Thinodromus velutinus (Sharp, 1887), tergite VIII; 2122 T. circulus (Bernhauer, 1922): 21 tergite VIII, 22 aedeagus; 2324 T. chagosanus (Bernhauer, 1922): 23 male sternite IX, 24 aedeagus. Scale bar: 0.1 mm for 2021; 0.15 mm for 2224." figureDoi="http://doi.org/10.5281/zenodo.5301069" httpUri="https://zenodo.org/record/5301069/files/figure.png" pageId="3" pageNumber="542">2122</figureCitation>
,
<figureCitation id="133C669DDC4FFFF0FD5A350F50B59331" box="[681,703,601,620]" captionStart="Figs 3135. 31" captionStartId="11.[113,150,1527,1546]" captionTargetBox="[114,1064,219,1466]" captionTargetId="figure-9@11.[113,1065,219,1466]" captionTargetPageId="11" captionText="Figs 3135. 31 Thinodromus velutinus (Sharp, 1887), female ringstructure. 32 T. circulus (Bernhauer, 1922), female ringstructure. 3334 T. palustris (Bernhauer, 1922): 33 male sternite IX, 34 aedeagus. 35 T.arcitenens (Fauvel, 1905), aedeagus. Scale bar: 0.1 mm for 31; 0.12 mm for 32, 35; 0.15 mm for 33; 0.2 mm for 34." figureDoi="http://doi.org/10.5281/zenodo.5301075" httpUri="https://zenodo.org/record/5301075/files/figure.png" pageId="3" pageNumber="542">32</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31D2993DC4FFFF0FF82352A51719399" pageId="3" pageNumber="542" type="reference_group">
<paragraph id="8BB87A18DC4FFFF0FF82352A51719399" blockId="3.[113,891,601,708]" pageId="3" pageNumber="542">
<treatmentCitationGroup id="AB175D36DC4FFFF0FF82352A51719399" pageId="3" pageNumber="542">
<treatmentCitation id="0AA65C09DC4FFFF0FF82352A53E893D2" author="BERNHAUER M." box="[113,482,636,655]" page="3" pageId="3" pageNumber="542" year="1922">
<taxonomicName id="4C07019BDC4FFFF0FF82352A53E893D2" ID-CoL="599QB" authority="Bernhauer, 1922 a: 3" authorityName="Bernhauer" authorityPageNumber="3" authorityYear="1922" box="[113,482,636,655]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="circulus">
<emphasis id="B973A60ADC4FFFF0FF82352A532593D2" box="[113,303,636,655]" italics="true" pageId="3" pageNumber="542">Trogophloeus circulus</emphasis>
<bibRefCitation id="EF9607E9DC4FFFF0FEC6352A53E893D2" author="BERNHAUER M." box="[309,482,636,655]" pageId="3" pageNumber="542" pagination="1 - 23" refId="ref6980" refString="BERNHAUER M. 1922 a: Neue Staphyliniden aus Sudamerika (23. Beitrag). Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 71 (1921): 1 - 23." type="journal article" year="1922">Bernhauer, 1922a: 3</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<treatmentCitation id="0AA65C09DC4FFFF0FF8235C0509A93F4" author="SCHEERPELTZ O." box="[113,656,662,681]" page="1081" pageId="3" pageNumber="542" year="1933">
<taxonomicName id="4C07019BDC4FFFF0FF8235C0505793F4" ID-CoL="599QB" authority=": SCHEERPELTZ (1933)" authorityName="SCHEERPELTZ" authorityYear="1933" box="[113,605,662,681]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="circulus" subGenus="Carpalimus">
<emphasis id="B973A60ADC4FFFF0FF8235C052EC93F4" box="[113,230,662,681]" italics="true" pageId="3" pageNumber="542">Trogophloeus</emphasis>
(
<emphasis id="B973A60ADC4FFFF0FF0135C0535D93F4" box="[242,343,662,681]" italics="true" pageId="3" pageNumber="542">Carpalimus</emphasis>
)
<emphasis id="B973A60ADC4FFFF0FE9735C053A293F4" box="[356,424,662,681]" italics="true" pageId="3" pageNumber="542">circulus</emphasis>
:
<bibRefCitation id="EF9607E9DC4FFFF0FE4035C0505793F4" author="SCHEERPELTZ O." box="[435,605,662,681]" pageId="3" pageNumber="542" pagination="989 - 1500" refId="ref7358" refString="SCHEERPELTZ O. 1933: Staphylinidae VII. In: SCHENKLING S. (ed.): Coleopterorum Catalogus, Pars 129. [Supplementum I.]. W. Junk, Berlin, pp. 989 - 1500." type="book chapter" year="1933">SCHEERPELTZ (1933)</bibRefCitation>
</taxonomicName>
: 1081
</treatmentCitation>
(catalogue).
<treatmentCitation id="0AA65C09DC4FFFF0FF8235E753EC9399" author="HERMAN L. H." box="[113,486,689,708]" page="387" pageId="3" pageNumber="542" year="1970">
<taxonomicName id="4C07019BDC4FFFF0FF8235E753B79399" ID-CoL="56H4Q" authority=": HERMAN (1970)" authorityName="HERMAN" authorityYear="1970" box="[113,445,689,708]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="circulus">
<emphasis id="B973A60ADC4FFFF0FF8235E753269399" box="[113,300,689,708]" italics="true" pageId="3" pageNumber="542">Thinodromus circulus</emphasis>
:
<bibRefCitation id="EF9607E9DC4FFFF0FEC435E753B79399" author="HERMAN L. H." box="[311,445,689,708]" pageId="3" pageNumber="542" pagination="343 - 454" refId="ref7167" refString="HERMAN L. H. 1970: Phylogeny and reclassi? cation of the genera of the rove-beetle subfamily Oxytelinae of the World (Coleoptera, Staphylinidae). Bulletin of the American Museum of Natural History 142 (5): 343 - 454." type="journal article" year="1970">HERMAN (1970)</bibRefCitation>
</taxonomicName>
: 387
</treatmentCitation>
(catalogue),
<treatmentCitation id="0AA65C09DC4FFFF0FDA535E751059399" author="HERMAN L. H." box="[598,783,689,708]" page="1765" pageId="3" pageNumber="542" year="2001">
<bibRefCitation id="EF9607E9DC4FFFF0FDA535E750D69399" author="HERMAN L. H." box="[598,732,689,708]" pageId="3" pageNumber="542" pagination="1 - 4218" refId="ref7213" refString="HERMAN L. H. 2001: Catalog of the Staphylinidae (Insecta: Coleoptera). 1785 to the end of the second Millennium. Volume I - VII. Bulletin of the American Museum of Natural History 265: 1 - 4218." type="journal article" year="2001">HERMAN (2001)</bibRefCitation>
: 1765
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31D2993DC4FFFF0FF82358D50B692D3" pageId="3" pageNumber="542" type="materials_examined">
<paragraph id="8BB87A18DC4FFFF0FF82358D50FC93B3" blockId="3.[113,1090,731,910]" box="[113,758,731,750]" pageId="3" pageNumber="542">
<emphasis id="B973A60ADC4FFFF0FF82358D52E093B3" bold="true" box="[113,234,731,750]" pageId="3" pageNumber="542">
<typeStatus id="54BCC4BADC4FFFF0FF82358D529493B3" box="[113,158,731,750]" pageId="3" pageNumber="542">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3103A88DC4FFFF0FF1C358D533A93B3" box="[239,304,731,750]" name="Bolivia" pageId="3" pageNumber="542">Bolivia</collectingCountry>
,
<collectingRegion id="49C3B4FADC4FFFF0FECA358D53A193B3" box="[313,427,731,750]" country="Bolivia" name="Cochabamba" pageId="3" pageNumber="542">Cochabamba</collectingRegion>
, Yuracares, approx. 15°56ƍS, 65°30ƍW.
</paragraph>
<paragraph id="8BB87A18DC4FFFF0FF8235A350B692D3" blockId="3.[113,1090,731,910]" pageId="3" pageNumber="542">
<emphasis id="B973A60ADC4FFFF0FF8235A3535C9255" bold="true" box="[113,342,757,776]" pageId="3" pageNumber="542">Type material examined.</emphasis>
<materialsCitation id="3B6F7045DC4FFFF0FEAE35A3564B927E" collectorName="M. Bernhauer" country="Bolivia" determinerName="A. Fauvel" location="Bolivien" pageId="3" pageNumber="542" specimenCount="1" typeStatus="lectotype">
<typeStatus id="54BCC4BADC4FFFF0FEAE35A353BC925A" box="[349,438,757,776]" pageId="3" pageNumber="542" type="lectotype">LECTOTYPE</typeStatus>
(here designated): J, Yuracaris,
<collectingCountry id="F3103A88DC4FFFF0FD2D35A351209255" box="[734,810,757,776]" name="Bolivia" pageId="3" pageNumber="542">Bolivien</collectingCountry>
;
<determinerName id="7E6496DCDC4FFFF0FCC735A351839255" box="[820,905,757,776]" pageId="3" pageNumber="542">A. Fauvel</determinerName>
determ. [label cut in two] \ circulus; Bernh.; Typus. \
<taxonomicName id="4C07019BDC4FFFF0FE7E34465000927E" authority="Fauv." authorityName="Fauv." box="[397,522,784,803]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="circulus">circulus Fauv.</taxonomicName>
; i.l. Mus. Hamburg \ Chicago NHMus;
<collectorName id="26F21FCEDC4FFFF0FC97344651D4927E" box="[868,990,784,803]" pageId="3" pageNumber="542">M. Bernhauer</collectorName>
; Collection
</materialsCitation>
\
<typeStatus id="54BCC4BADC4FFFF0FF8E347D52D59263" box="[125,223,811,830]" pageId="3" pageNumber="542" type="lectotype">Lectotypus</typeStatus>
;
<taxonomicName id="4C07019BDC4FFFF0FF1B347D536B9263" box="[232,353,811,830]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
;
<taxonomicName id="4C07019BDC4FFFF0FE9A347D50059263" authority="Bernhauer" authorityName="Bernhauer" box="[361,527,811,830]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="circulus">circulus Bernhauer</taxonomicName>
; des.
<bibRefCitation id="EF9607E9DC4FFFF0FDB3347D50DF9263" author="MAKRANCZY GY" box="[576,725,811,830]" pageId="3" pageNumber="542" pagination="177 - 208" refId="ref7326" refString="MAKRANCZY GY. 2013: Review of the southern African species of Thinodromus (Coleoptera: Staphylinidae: Oxytelinae). Acta Entomologica Musei Nationalis Pragae 53: 177 - 208." type="journal article" year="2013">Makranczy, 2013</bibRefCitation>
\
<taxonomicName id="4C07019BDC4FFFF0FD15347D51569263" authorityName="Kraatz" authorityYear="1857" box="[742,860,811,830]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">Thinodromus</taxonomicName>
;
<taxonomicName id="4C07019BDC4FFFF0FC96347D56129263" authority="(Bernhauer)" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[869,1048,811,830]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="circulus">circulus (Bernhauer)</taxonomicName>
; det.
<bibRefCitation id="EF9607E9DC4FFFF0FF823413530D9205" author="MAKRANCZY GY" box="[113,263,837,856]" pageId="3" pageNumber="542" pagination="177 - 208" refId="ref7326" refString="MAKRANCZY GY. 2013: Review of the southern African species of Thinodromus (Coleoptera: Staphylinidae: Oxytelinae). Acta Entomologica Musei Nationalis Pragae 53: 177 - 208." type="journal article" year="2013">Makranczy, 2013</bibRefCitation>
(
<collectionCode id="ED16E2DDDC4FFFF0FEE5341353539205" box="[278,345,837,856]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="542" type="Museum">FMNH</collectionCode>
).
<typeStatus id="54BCC4BADC4FFFF0FE95341353E5920A" box="[358,495,837,856]" pageId="3" pageNumber="542" type="paralectotype">PARALECTOTYPES</typeStatus>
: 3 JJ
<specimenCount id="9D01B191DC4FFFF0FDDD341350469204" box="[558,588,834,857]" pageId="3" pageNumber="542" type="female">1 ♀</specimenCount>
Yuracaris;
<collectingCountry id="F3103A88DC4FFFF0FD46341350FE9205" box="[693,756,837,856]" name="Bolivia" pageId="3" pageNumber="542">Bolivie</collectingCountry>
\ circulus;
<materialsCitation id="3B6F7045DC4FFFF0FCA0341356349205" box="[851,1086,837,856]" collectorName="A. Fauvel" country="Bolivia" location="Fvl." municipality="Coll." pageId="3" pageNumber="542" specimenCount="1" typeStatus="paralectotype">
<location id="8ED82CC3DC4FFFF0FCA03413517C9205" LSID="urn:lsid:plazi:treatment:03AECB0EDC4FFFF6FE86356D52F495CE:8ED82CC3DC4FFFF0FCA03413517C9205" box="[851,886,837,856]" country="Bolivia" municipality="Coll." name="Fvl." pageId="3" pageNumber="542">Fvl.</location>
\
<collectingMunicipality id="6BDCE062DC4FFFF0FC76341351BA9205" box="[901,944,837,856]" pageId="3" pageNumber="542">Coll.</collectingMunicipality>
et det
<collectorName id="26F21FCEDC4FFFF0FC14341356349205" box="[999,1086,837,856]" pageId="3" pageNumber="542">A. Fauvel</collectorName>
</materialsCitation>
;
<collectionCode id="ED16E2DDDC4FFFF0FF8234365288922E" box="[113,130,864,883]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="542">R</collectionCode>
.I.Sc.N.B. 17.479 \
<typeStatus id="54BCC4BADC4FFFF0FEDD343653A5922E" box="[302,431,864,883]" pageId="3" pageNumber="542" type="paralectotype">Paralectotypus</typeStatus>
;
<taxonomicName id="4C07019BDC4FFFF0FE4934365039922E" box="[442,563,864,883]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
;
<taxonomicName id="4C07019BDC4FFFF0FDCD343650EC922E" authority="Bernhauer" authorityName="Bernhauer" box="[574,742,864,883]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="circulus">circulus Bernhauer</taxonomicName>
; des.
<bibRefCitation id="EF9607E9DC4FFFF0FCE8343651B8922E" author="MAKRANCZY GY" box="[795,946,864,883]" pageId="3" pageNumber="542" pagination="177 - 208" refId="ref7326" refString="MAKRANCZY GY. 2013: Review of the southern African species of Thinodromus (Coleoptera: Staphylinidae: Oxytelinae). Acta Entomologica Musei Nationalis Pragae 53: 177 - 208." type="journal article" year="2013">Makranczy, 2013</bibRefCitation>
\
<taxonomicName id="4C07019BDC4FFFF0FC3B34365634922E" authorityName="Kraatz" authorityYear="1857" box="[968,1086,864,883]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">Thinodromus</taxonomicName>
;
<taxonomicName id="4C07019BDC4FFFF0FF82342D532E92D3" authority="(Bernhauer)" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[113,292,891,910]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="circulus">circulus (Bernhauer)</taxonomicName>
; det.
<determinerName id="7E6496DCDC4FFFF0FEA2342D53B892D3" box="[337,434,891,910]" pageId="3" pageNumber="542">Makranczy</determinerName>
, 2001 (
<collectionCode id="ED16E2DDDC4FFFF0FE05342D502092D3" box="[502,554,891,910]" pageId="3" pageNumber="542">ISNB</collectionCode>
, 1 J in
<collectionCode id="ED16E2DDDC4FFFF0FD9E342D50B892D3" box="[621,690,891,910]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="3" pageNumber="542">HNHM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C31D2993DC4FFFF6FF8234F05051926E" lastPageId="5" lastPageNumber="544" pageId="3" pageNumber="542" type="description">
<paragraph id="8BB87A18DC4FFFF6FF8234F05051926E" blockId="3.[113,1089,934,1597]" lastBlockId="5.[113,1090,220,1171]" lastPageId="5" lastPageNumber="544" pageId="3" pageNumber="542">
<emphasis id="B973A60ADC4FFFF0FF8234F0531092E0" bold="true" box="[113,282,934,957]" pageId="3" pageNumber="542">Redescription.</emphasis>
<emphasis id="B973A60ADC4FFFF0FED334F053B792E0" box="[288,445,934,957]" italics="true" pageId="3" pageNumber="542">Measurements</emphasis>
(in mm, n = 5): HW = 0.40 (0.390.42); TW = 0.395 (0.39 0.41); PW = 0.44 (0.430.45); SW = 0.54 (0.520.56); AW = 0.59 (0.560.63); HL = 0.26 (0.240.27); EL = 0.12 (0.120.13); TL = 0.07 (0.060.07); PL = 0.33 (0.320.35); SL = 0.59 (0.570.60); SC = 0.56 (0.540.57); FB = 1.21 (1.161.26); BL = 2.24 (2.172.31).
<emphasis id="B973A60ADC4FFFF0FC0F335052E09560" italics="true" pageId="3" pageNumber="542">Lustre and colour</emphasis>
. Body medium dull, especially head and pronotum, due to surface sculpture, with only tiny glittery spots; rest of body more lustrous. Head, pronotum and abdomen very dark brown. Elytra just a little bit lighter, dark brown with some reddish tint. Antennae dark brown but basal 1/3 conspicuously lighter, light brown, even yellowish at base. Legs and mouthparts medium to dark brown but tarsi and tibiae (especially both ends) often lighter then femora.
<emphasis id="B973A60ADC4FFFF0FEF2339053D69580" box="[257,476,1222,1245]" italics="true" pageId="3" pageNumber="542">Shape and sculpture</emphasis>
. Forebody as in
<figureCitation id="133C669DDC4FFFF0FD63339050DB9580" box="[656,721,1222,1245]" captionStart="Figs 14" captionStartId="2.[151,188,1526,1545]" captionTargetBox="[163,1115,218,1465]" captionTargetId="figure-10@2.[163,1116,217,1465]" captionTargetPageId="2" captionText="Figs 14. Forebodies of Thinodromus species. 1 T. circulus (Bernhauer, 1922); 2 T. chagosanus (Bernhauer, 1922); 3 T. palustris (Bernhauer, 1922); 4 T. arcitenens (Fauvel, 1905)." figureDoi="http://doi.org/10.5281/zenodo.5301061" httpUri="https://zenodo.org/record/5301061/files/figure.png" pageId="3" pageNumber="542">Fig. 1</figureCitation>
. Head quite transverse, with large eyes and well developed, bulging temples nearing half the size of eyes. Neck delineated only by different, alveolate microsculpture, but without transversal groove. Antennae rather short, antennomeres slightly transverse: antennomeres 4 and 5 1.001.06× and 1.031.08× broader than long, respectively, antennomeres 9 and 10 1.301.38 and 1.601.70× broader than long, respectively. Pronotum rather transverse, strongly obtuse-angled anterior corners super¿cially appear somewhat rounded but still marked. Posterior half of pronotal sides quite straight, even feebly concave; posterior corners obtuse-angled and rounded. Horseshoe-shaped impression slightly marked except posterior/median part where rather impressed; slight lateral depressions connected by it. Middle of disc bearing pair of shallow depressions. Slight (thin) marginal bead (mostly lateral) observable only in slightly lateral view. Elytra combined imperceptably broader than long, gently dilated toward apex, with a pair of small, oval, slightly elongate impressions behind scutellum and extending posteriorly in longitudinal impressions, connected to somewhat depressed centre of elytral disc. Posterior elytral margin (slightly oblique) with very thin bead and in outer 1/3 with small membranous lobe protruding. Apex of abdominal tergite VII with palisade fringe.
<emphasis id="B973A60ADC49FFF6FC6A366A531E902E" italics="true" pageId="5" pageNumber="544">Punctation and microsculpture</emphasis>
. Head and pronotum densely but not too deeply punctured, yet appearance dominated by acinose microsculpture; puncture interspaces on average smaller than puncture diameters. Punctation denser near the edges than at centre of discs. Epistomal suture barely marked by transversal strigulate microsculpture. Elytral punctures only slightly larger in size but interspaces much larger and with more conspicuous microsculpture only apparent at the scutellar area, otherwise quite indistinct. Consequently, elytral surface more lustrous, even a little more than abdomen bearing strong coriaceous/imbricate microsculpture (with isodiametric or slightly transverse cells) giving a greasy lustre. Abdominal punctures tiny and scattered.
<emphasis id="B973A60ADC49FFF6FEFE350A5387932E" box="[269,397,604,627]" italics="true" pageId="5" pageNumber="544">Pubescence</emphasis>
. Body setation ¿ne, short and medium dense and equal sized on head and pronotum, a little longer and less dense on elytra. Longer hairs only on abdomen, especially apices of segments; otherwise elytral and abdominal setation with approximately the same density.
<emphasis id="B973A60ADC49FFF6FEC635EA50E9938E" box="[309,739,700,723]" italics="true" pageId="5" pageNumber="544">Primary and secondary sexual features</emphasis>
. Male sternite VIII as in
<figureCitation id="133C669DDC49FFF6FC0935EA5631938E" box="[1018,1083,700,723]" captionStart="Figs 514. 510" captionStartId="4.[151,188,1497,1516]" captionTargetBox="[151,1100,223,1466]" captionTargetId="figure-9@4.[151,1103,221,1468]" captionTargetPageId="4" captionText="Figs 514. 510 Thinodromus circulus (Bernhauer, 1922) (5 mesotarsus; 6 male sternite VIII; 7 male tergite IX; 8 male sternite IX; 9 male tergite X; 10 spermatheca); 1112 T. velutinus (Sharp, 1887) (11 tergite X; 12 spermatheca); 1314 T. chagosanus (Bernhauer, 1922) (13 mesotarsus; 14 platelike armature in hypopharynx). Scale bar: 0.06 mm for 14; 0.07 mm for 5, 13; 0.1 mm for 1012; 0.12 mm for 89; 0.14 mm for 67." figureDoi="http://doi.org/10.5281/zenodo.5301063" httpUri="https://zenodo.org/record/5301063/files/figure.png" pageId="5" pageNumber="544">Fig. 6</figureCitation>
, male tergite IX as in
<figureCitation id="133C669DDC49FFF6FEA4358A539393AE" box="[343,409,732,755]" captionStart="Figs 514. 510" captionStartId="4.[151,188,1497,1516]" captionTargetBox="[151,1100,223,1466]" captionTargetId="figure-9@4.[151,1103,221,1468]" captionTargetPageId="4" captionText="Figs 514. 510 Thinodromus circulus (Bernhauer, 1922) (5 mesotarsus; 6 male sternite VIII; 7 male tergite IX; 8 male sternite IX; 9 male tergite X; 10 spermatheca); 1112 T. velutinus (Sharp, 1887) (11 tergite X; 12 spermatheca); 1314 T. chagosanus (Bernhauer, 1922) (13 mesotarsus; 14 platelike armature in hypopharynx). Scale bar: 0.06 mm for 14; 0.07 mm for 5, 13; 0.1 mm for 1012; 0.12 mm for 89; 0.14 mm for 67." figureDoi="http://doi.org/10.5281/zenodo.5301063" httpUri="https://zenodo.org/record/5301063/files/figure.png" pageId="5" pageNumber="544">Fig. 7</figureCitation>
, male sternite IX as in
<figureCitation id="133C669DDC49FFF6FD65358A50DD93AE" box="[662,727,732,755]" captionStart="Figs 514. 510" captionStartId="4.[151,188,1497,1516]" captionTargetBox="[151,1100,223,1466]" captionTargetId="figure-9@4.[151,1103,221,1468]" captionTargetPageId="4" captionText="Figs 514. 510 Thinodromus circulus (Bernhauer, 1922) (5 mesotarsus; 6 male sternite VIII; 7 male tergite IX; 8 male sternite IX; 9 male tergite X; 10 spermatheca); 1112 T. velutinus (Sharp, 1887) (11 tergite X; 12 spermatheca); 1314 T. chagosanus (Bernhauer, 1922) (13 mesotarsus; 14 platelike armature in hypopharynx). Scale bar: 0.06 mm for 14; 0.07 mm for 5, 13; 0.1 mm for 1012; 0.12 mm for 89; 0.14 mm for 67." figureDoi="http://doi.org/10.5281/zenodo.5301063" httpUri="https://zenodo.org/record/5301063/files/figure.png" pageId="5" pageNumber="544">Fig. 8</figureCitation>
; tergite VIII (
<figureCitation id="133C669DDC49FFF6FC9C358A51B493AE" box="[879,958,732,755]" captionStart="Figs 2024. 20" captionStartId="7.[113,150,1480,1499]" captionTargetBox="[127,1077,222,1463]" captionTargetId="figure-9@7.[125,1078,219,1466]" captionTargetPageId="7" captionText="Figs 2024. 20 Thinodromus velutinus (Sharp, 1887), tergite VIII; 2122 T. circulus (Bernhauer, 1922): 21 tergite VIII, 22 aedeagus; 2324 T. chagosanus (Bernhauer, 1922): 23 male sternite IX, 24 aedeagus. Scale bar: 0.1 mm for 2021; 0.15 mm for 2224." figureDoi="http://doi.org/10.5281/zenodo.5301069" httpUri="https://zenodo.org/record/5301069/files/figure.png" pageId="5" pageNumber="544">Fig. 21</figureCitation>
) and tergite X (
<figureCitation id="133C669DDC49FFF6FF6735AA52DF924E" box="[148,213,764,787]" captionStart="Figs 514. 510" captionStartId="4.[151,188,1497,1516]" captionTargetBox="[151,1100,223,1466]" captionTargetId="figure-9@4.[151,1103,221,1468]" captionTargetPageId="4" captionText="Figs 514. 510 Thinodromus circulus (Bernhauer, 1922) (5 mesotarsus; 6 male sternite VIII; 7 male tergite IX; 8 male sternite IX; 9 male tergite X; 10 spermatheca); 1112 T. velutinus (Sharp, 1887) (11 tergite X; 12 spermatheca); 1314 T. chagosanus (Bernhauer, 1922) (13 mesotarsus; 14 platelike armature in hypopharynx). Scale bar: 0.06 mm for 14; 0.07 mm for 5, 13; 0.1 mm for 1012; 0.12 mm for 89; 0.14 mm for 67." figureDoi="http://doi.org/10.5281/zenodo.5301063" httpUri="https://zenodo.org/record/5301063/files/figure.png" pageId="5" pageNumber="544">Fig. 9</figureCitation>
) seemingly not different between sexes. Aedeagus as in
<figureCitation id="133C669DDC49FFF6FCCF35AA5186924E" box="[828,908,764,787]" captionStart="Figs 2024. 20" captionStartId="7.[113,150,1480,1499]" captionTargetBox="[127,1077,222,1463]" captionTargetId="figure-9@7.[125,1078,219,1466]" captionTargetPageId="7" captionText="Figs 2024. 20 Thinodromus velutinus (Sharp, 1887), tergite VIII; 2122 T. circulus (Bernhauer, 1922): 21 tergite VIII, 22 aedeagus; 2324 T. chagosanus (Bernhauer, 1922): 23 male sternite IX, 24 aedeagus. Scale bar: 0.1 mm for 2021; 0.15 mm for 2224." figureDoi="http://doi.org/10.5281/zenodo.5301069" httpUri="https://zenodo.org/record/5301069/files/figure.png" pageId="5" pageNumber="544">Fig. 22</figureCitation>
. Spermatheca as in
<figureCitation id="133C669DDC49FFF6FF7D344A52D7926E" box="[142,221,796,819]" captionStart="Figs 514. 510" captionStartId="4.[151,188,1497,1516]" captionTargetBox="[151,1100,223,1466]" captionTargetId="figure-9@4.[151,1103,221,1468]" captionTargetPageId="4" captionText="Figs 514. 510 Thinodromus circulus (Bernhauer, 1922) (5 mesotarsus; 6 male sternite VIII; 7 male tergite IX; 8 male sternite IX; 9 male tergite X; 10 spermatheca); 1112 T. velutinus (Sharp, 1887) (11 tergite X; 12 spermatheca); 1314 T. chagosanus (Bernhauer, 1922) (13 mesotarsus; 14 platelike armature in hypopharynx). Scale bar: 0.06 mm for 14; 0.07 mm for 5, 13; 0.1 mm for 1012; 0.12 mm for 89; 0.14 mm for 67." figureDoi="http://doi.org/10.5281/zenodo.5301063" httpUri="https://zenodo.org/record/5301063/files/figure.png" pageId="5" pageNumber="544">Fig. 10</figureCitation>
, female ringstructure as in
<figureCitation id="133C669DDC49FFF6FDF4344A505D926E" box="[519,599,796,819]" captionStart="Figs 3135. 31" captionStartId="11.[113,150,1527,1546]" captionTargetBox="[114,1064,219,1466]" captionTargetId="figure-9@11.[113,1065,219,1466]" captionTargetPageId="11" captionText="Figs 3135. 31 Thinodromus velutinus (Sharp, 1887), female ringstructure. 32 T. circulus (Bernhauer, 1922), female ringstructure. 3334 T. palustris (Bernhauer, 1922): 33 male sternite IX, 34 aedeagus. 35 T.arcitenens (Fauvel, 1905), aedeagus. Scale bar: 0.1 mm for 31; 0.12 mm for 32, 35; 0.15 mm for 33; 0.2 mm for 34." figureDoi="http://doi.org/10.5281/zenodo.5301075" httpUri="https://zenodo.org/record/5301075/files/figure.png" pageId="5" pageNumber="544">Fig. 32</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="DF782A90DC48FFF7FF64328F565E9761" ID-DOI="http://doi.org/10.5281/zenodo.5301063" ID-Zenodo-Dep="5301063" httpUri="https://zenodo.org/record/5301063/files/figure.png" pageId="4" pageNumber="543" startId="4.[151,188,1497,1516]" targetBox="[151,1100,223,1466]" targetPageId="4">
<paragraph id="8BB87A18DC48FFF7FF64328F565E9761" blockId="4.[151,1127,1497,1596]" pageId="4" pageNumber="543">
Figs 514. 510
<taxonomicName id="4C07019BDC48FFF7FEC6328F508494B1" authority="(Bernhauer, 1922)" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[309,654,1497,1516]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="circulus">
<emphasis id="B973A60ADC48FFF7FEC6328F53E594B1" box="[309,495,1497,1516]" italics="true" pageId="4" pageNumber="543">Thinodromus circulus</emphasis>
(Bernhauer, 1922)
</taxonomicName>
(5 mesotarsus; 6 male sternite VIII; 7 male tergite IX; 8 male sternite IX; 9 male tergite X; 10 spermatheca); 1112
<taxonomicName id="4C07019BDC48FFF7FCF032A251D5975A" authority="(Sharp, 1887)" authorityName="Makranczy" authorityYear="2014" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[771,991,1524,1543]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B973A60ADC48FFF7FCF032A2516F975A" box="[771,869,1524,1543]" italics="true" pageId="4" pageNumber="543">T. velutinus</emphasis>
(
<bibRefCitation id="EF9607E9DC48FFF7FC8332A251D0975A" author="SHARP D. S." box="[880,986,1524,1543]" pageId="4" pageNumber="543" pagination="673 - 824" refId="ref7488" refString="SHARP D. S. 1887: Staphylinidae. Pp. 673 - 824, pls. 17 - 19. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. I. Part 2. Taylor &amp; Francis, London, xvi + 824 pp., 19 pls." type="book chapter" year="1887">Sharp, 1887</bibRefCitation>
)
</taxonomicName>
(11 tergite X; 12 spermatheca); 1314
<taxonomicName id="4C07019BDC48FFF7FE7D315950A1977F" authority="(Bernhauer, 1922)" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[398,683,1551,1570]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="543" phylum="Arthropoda" rank="species" species="chagosanus">
<emphasis id="B973A60ADC48FFF7FE7D31595003977F" box="[398,521,1551,1570]" italics="true" pageId="4" pageNumber="543">T. chagosanus</emphasis>
(Bernhauer, 1922)
</taxonomicName>
(13 mesotarsus; 14 platelike armature in hypopharynx). Scale bar: 0.06 mm for 14; 0.07 mm for 5, 13; 0.1 mm for 1012; 0.12 mm for 89; 0.14 mm for 67.
</paragraph>
</caption>
<subSubSection id="C31D2993DC49FFF6FF82346A508092EE" pageId="5" pageNumber="544" type="diagnosis">
<paragraph id="8BB87A18DC49FFF6FF82346A508092EE" blockId="5.[113,1090,220,1171]" pageId="5" pageNumber="544">
<emphasis id="B973A60ADC49FFF6FF82346A5367920E" bold="true" box="[113,365,828,851]" pageId="5" pageNumber="544">Differential diagnosis.</emphasis>
This species is distinguishable from
<taxonomicName id="4C07019BDC49FFF6FD0E346A5172920E" authorityName="Makranczy" authorityYear="2014" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[765,888,828,851]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B973A60ADC49FFF6FD0E346A5172920E" box="[765,888,828,851]" italics="true" pageId="5" pageNumber="544">T. velutinus</emphasis>
</taxonomicName>
by its less conspicuously strong punctation of the head and pronotum. These two Neotropical species differ from the rest of the treated taxa by the less elongate antennae (mid-antennal segments more or less isodiametric or transverse, never elongate).
</paragraph>
</subSubSection>
<subSubSection id="C31D2993DC49FFF6FF8234EA5182928E" box="[113,904,956,979]" pageId="5" pageNumber="544" type="distribution">
<paragraph id="8BB87A18DC49FFF6FF8234EA5182928E" blockId="5.[113,1090,220,1171]" box="[113,904,956,979]" pageId="5" pageNumber="544">
<emphasis id="B973A60ADC49FFF6FF8234EA530E928E" bold="true" box="[113,260,956,979]" pageId="5" pageNumber="544">Distribution.</emphasis>
This species is known only from its
<typeStatus id="54BCC4BADC49FFF6FD7834EA50B3928E" box="[651,697,956,979]" pageId="5" pageNumber="544">type</typeStatus>
locality in
<collectingCountry id="F3103A88DC49FFF6FCC134EA518E928E" box="[818,900,956,979]" name="Bolivia" pageId="5" pageNumber="544">Bolivia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31D2993DC49FFF6FF82348A52F495CE" pageId="5" pageNumber="544" type="discussion">
<paragraph id="8BB87A18DC49FFF6FF82348A52F495CE" blockId="5.[113,1090,220,1171]" pageId="5" pageNumber="544">
<emphasis id="B973A60ADC49FFF6FF82348A52EA92AE" bold="true" box="[113,224,988,1011]" pageId="5" pageNumber="544">Remarks.</emphasis>
A short series of specimens are the only material available of this species and no further data survived. The series was ¿rst studied by Albert Fauvel, but had remained undescribed by him. Finally, a year after Fauvels death, Max Bernhauer published an article with Fauvels names, validating these taxon names.
<bibRefCitation id="EF9607E9DC49FFF6FD53336A517B950E" author="BERNHAUER M." box="[672,881,1084,1107]" pageId="5" pageNumber="544" pagination="1 - 23" refId="ref6980" refString="BERNHAUER M. 1922 a: Neue Staphyliniden aus Sudamerika (23. Beitrag). Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 71 (1921): 1 - 23." type="journal article" year="1922">BERNHAUER (1922a)</bibRefCitation>
gave no further information on how this material came to him. The
<typeStatus id="54BCC4BADC49FFF6FD8D330A50EA952E" box="[638,736,1116,1139]" pageId="5" pageNumber="544" type="lectotype">lectotype</typeStatus>
is chosen to ¿x the interpretation of this taxon.
</paragraph>
</subSubSection>
</treatment>
</document>