638 lines
87 KiB
XML
638 lines
87 KiB
XML
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<mods:title id="0C5F987D45F99F5C2B8653A53B08EF70">Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species</mods:title>
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<taxonomicName id="4C3CA096A3407F50FF67F9D48845DFA4" ID-CoL="7D4HJ" authority="Andrewes, 1930" authorityName="Andrewes" authorityYear="1930" box="[151,793,1658,1684]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
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<emphasis id="B9480707A3407F50FF67F9D48845DFA4" bold="true" box="[151,793,1658,1684]" pageId="45" pageNumber="346">
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<emphasis id="B9480707A3407F50FF67F9D48A1DDFA4" bold="true" box="[151,321,1658,1684]" italics="true" pageId="45" pageNumber="346">Trichotichnus</emphasis>
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(
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<emphasis id="B9480707A3407F50FEA2F9D48A95DFA4" bold="true" box="[338,457,1658,1684]" italics="true" pageId="45" pageNumber="346">Bottchrus</emphasis>
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)
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<emphasis id="B9480707A3407F50FE2AF9D48910DFA4" bold="true" box="[474,588,1658,1684]" italics="true" pageId="45" pageNumber="346">hingstoni</emphasis>
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Andrewes, 1930
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</emphasis>
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<paragraph id="8B83DB15A3407F50FF67F9328A92DF85" blockId="45.[151,793,1658,1717]" box="[151,462,1692,1717]" pageId="45" pageNumber="346">
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(
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<figureCitation id="1307C790A3407F50FF6FF9328A5FDF85" box="[159,259,1692,1717]" captionStart="FIGURES 120 – 123" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[229,1356,202,1919]" captionTargetId="figure@41.[227,1360,193,1921]" captionTargetPageId="41" captionText="FIGURES 120 – 123. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 120, T. sikkimensis sp. n., holotype; 121, T. martensi sp. n., holotype; 122, T. hingstoni Andrewes, 1930, male from Kechu Pheri Lake; 123, T. birmanicus Bates, 1892, male from the Kali Gandaki, Tatopani." httpUri="https://zenodo.org/record/919672/files/figure.png" pageId="45" pageNumber="346">Figs 122</figureCitation>
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,
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<figureCitation id="1307C790A3407F50FEFFF9328A28DF85" box="[271,372,1692,1717]" captionStart-0="FIGURES 135 – 136" captionStart-1="FIGURES 137 – 145" captionStartId-0="46.[151,264,1244,1267]" captionStartId-1="47.[151,264,1705,1728]" captionTargetBox-0="[301,1277,456,1212]" captionTargetBox-1="[194,1406,318,1664]" captionTargetId-0="figure@46.[271,1316,445,1223]" captionTargetId-1="figure@47.[151,1436,303,1684]" captionTargetPageId-0="46" captionTargetPageId-1="47" captionText-0="FIGURES 135 – 136. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930, median lobe of aedeagus, lateral view (135, lectotype; 136, holotype of T. curvatus Ito, 1996). Scale bar = 1.0 mm." captionText-1="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri-0="https://zenodo.org/record/919684/files/figure.png" httpUri-1="https://zenodo.org/record/919688/files/figure.png" pageId="45" pageNumber="346">135–145</figureCitation>
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,
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<figureCitation id="1307C790A3407F50FE8FF9328AF7DF85" box="[383,427,1692,1717]" captionStart="FIGURE 152" captionStartId="53.[151,250,1975,1997]" captionTargetBox="[151,1436,1234,1954]" captionTargetId="figure@53.[151,1436,1234,1954]" captionTargetPageId="53" captionText="FIGURE 152. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. gupchiensis sp. n.; 2, T. brancuccii sp. n.; 3, T. manasluensis sp. n.; 4, T. loebli Ito, 1998; 5, T. sikkimensis sp. n.; 6, T. hingstoni Andrewes, 1930." httpUri="https://zenodo.org/record/919698/files/figure.png" pageId="45" pageNumber="346">152</figureCitation>
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,
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<figureCitation id="1307C790A3407F50FE47F9328A99DF85" box="[439,453,1692,1717]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="45" pageNumber="346">
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<emphasis id="B9480707A3407F50FE47F9328A99DF85" box="[439,453,1692,1717]" italics="true" pageId="45" pageNumber="346">6</emphasis>
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</figureCitation>
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)
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</paragraph>
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<subSubSection id="C326889EA3407F50FF67F94A8E7ADE29" pageId="45" pageNumber="346" type="reference_group">
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<taxonomicName id="4C3CA096A3407F50FF67F94A893BDFCA" ID-CoL="7D4HJ" authority="Andrewes, 1930 a: 19" authorityName="Andrewes" authorityPageNumber="19" authorityYear="1930" box="[151,615,1764,1786]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="hingstoni">
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<emphasis id="B9480707A3407F50FF67F94A8ADADFCA" box="[151,390,1764,1786]" italics="true" pageId="45" pageNumber="346">Trichotichnus hingstoni</emphasis>
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<treatmentCitation id="0A9DFD04A3407F50FE7CF94A893BDFCA" author="Andrewes" box="[396,615,1764,1786]" page="19" pageId="45" pageNumber="346" year="1930">
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: 19
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</taxonomicName>
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. Type locality: Darjeeling,
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[ca
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<taxonomicName id="4C3CA096A3407F50FF67F8AD89A2DE29" ID-CoL="8LPCF" authority="Ito, 1996: 205" authorityName="N. Ito" authorityPageNumber="205" authorityYear="1996" box="[151,766,1795,1817]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="curvatus" subGenus="Pseudotrichotichnus">
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<emphasis id="B9480707A3407F50FF67F8AD8A7FDE29" box="[151,291,1795,1817]" italics="true" pageId="45" pageNumber="346">Trichotichnus</emphasis>
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(
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<emphasis id="B9480707A3407F50FEC1F8AD895DDE29" box="[305,513,1795,1817]" italics="true" pageId="45" pageNumber="346">Pseudotrichotichnus</emphasis>
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)
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<emphasis id="B9480707A3407F50FDE0F8AA893BDE29" box="[528,615,1796,1817]" italics="true" pageId="45" pageNumber="346">curvatus</emphasis>
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<treatmentCitation id="0A9DFD04A3407F50FD9DF8AA89A2DE29" author="Ito" box="[621,766,1795,1817]" page="205" pageId="45" pageNumber="346" year="1996">
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: 205
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</taxonomicName>
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,
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<emphasis id="B9480707A3407F50FCF8F8AD8812DE28" bold="true" box="[776,846,1795,1816]" pageId="45" pageNumber="346">syn. n.</emphasis>
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</taxonomicNameLabel>
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<typeStatus id="548765B7A3407F50FCA4F8AA88DADE29" box="[852,902,1796,1817]" pageId="45" pageNumber="346">Type</typeStatus>
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locality:
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<location id="8EE38DCEA3407F50FC16F8AD8F0FDE29" LSID="urn:lsid:plazi:treatment:03956A03A3407F4CFF67F9D48F0AD8E0:8EE38DCEA3407F50FC16F8AD8F0FDE29" box="[998,1107,1795,1817]" country="India" name="Darjeeling" pageId="45" pageNumber="346" stateProvince="West Bengal">Darjeeling</location>
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,
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<collectingRegion id="49F815F7A3407F50FBAEF8AA8FBCDE29" box="[1118,1248,1795,1817]" country="India" name="West Bengal" pageId="45" pageNumber="346">West Bengal</collectingRegion>
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,
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<collectingCountry id="F32B9B85A3407F50FB1AF8AD8E7DDE29" box="[1258,1313,1795,1817]" name="India" pageId="45" pageNumber="346">India</collectingCountry>
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</materialsCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B83DB15A3407F50FF67F8EE8F67DEF5" blockId="45.[151,1436,1856,2025]" pageId="45" pageNumber="346">
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<emphasis id="B9480707A3407F50FF67F8EE8AE7DE69" bold="true" box="[151,443,1856,1881]" pageId="45" pageNumber="346">Type material examined</emphasis>
|
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. Lectotype (present designation) of
|
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<taxonomicName id="4C3CA096A3407F50FCABF8EF88BCDE69" authorityName="Andrewes" authorityYear="1930" box="[859,992,1856,1881]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="hingstoni">
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<emphasis id="B9480707A3407F50FCABF8EF88BCDE69" box="[859,992,1856,1881]" italics="true" pageId="45" pageNumber="346">T. hingstoni</emphasis>
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</taxonomicName>
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: ♂, “N. India: Darjeeling,
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.,
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<date id="FF82FDD5A3407F50FA70F8EF8A4BDE4D" pageId="45" pageNumber="346" value="1924-03-01" valueMax="1924-03-10" valueMin="1924-03-01">1– 10.III.1924</date>
|
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, Maj. R.W.G. Hingston”, “Under stones”, “Everest Exp., Brit. Mus., 1924–386”, “Type” [round label with red margin], “
|
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<taxonomicName id="4C3CA096A3407F50FE44F826886BDE91" authority="Andr." authorityName="Andrewes" authorityYear="1930" box="[436,823,1928,1953]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="hingstoni">
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<emphasis id="B9480707A3407F50FE44F8268989DE91" box="[436,725,1928,1953]" italics="true" pageId="45" pageNumber="346">Trichotichnus hingstoni</emphasis>
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Andr.
|
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</taxonomicName>
|
||
, Type, H.E. Andrewes det.”, “
|
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<taxonomicName id="4C3CA096A3407F50FAF2F8268927DEF5" authority="Andrewes" authorityName="Andrewes" authorityYear="1930" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Pseudotrichotichnus">
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<emphasis id="B9480707A3407F50FAF2F8268EC7DE91" box="[1282,1435,1928,1953]" italics="true" pageId="45" pageNumber="346">Trichotichnus</emphasis>
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(
|
||
<emphasis id="B9480707A3407F50FF50F8028ADADEF5" box="[160,390,1964,1989]" italics="true" pageId="45" pageNumber="346">Pseudotrichotichnus</emphasis>
|
||
)
|
||
<emphasis id="B9480707A3407F50FE67F8028AA3DEF5" box="[407,511,1964,1989]" italics="true" pageId="45" pageNumber="346">hingstoni</emphasis>
|
||
Andrewes
|
||
</taxonomicName>
|
||
, Holotype, Det. N. Ito, 2000” (BMNH).
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3407F50FF37F87E8ED2DED9" blockId="45.[151,1436,1856,2025]" box="[199,1422,2000,2025]" pageId="45" pageNumber="346">
|
||
<typeStatus id="548765B7A3407F50FF37F87E8A71DED9" box="[199,301,2000,2025]" pageId="45" pageNumber="346" type="holotype">Holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C3CA096A3407F50FEA1F87C8A94DED9" authorityName="N. Ito" authorityYear="1996" box="[337,456,2002,2025]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="curvatus">
|
||
<emphasis id="B9480707A3407F50FEA1F87C8A94DED9" box="[337,456,2002,2025]" italics="true" pageId="45" pageNumber="346">T. curvatus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<specimenCount id="9D3A109CA3407F50FE28F87F8AB3DED8" box="[472,495,2001,2024]" pageId="45" pageNumber="346" type="male">♂</specimenCount>
|
||
, “Himalaya, O. Thieme, Darj.”, “
|
||
<typeStatus id="548765B7A3407F50FCACF87E889BDED9" box="[860,967,2000,2025]" pageId="45" pageNumber="346" type="holotype">Holotype</typeStatus>
|
||
,
|
||
<taxonomicName id="4C3CA096A3407F50FC22F87E8E49DED9" authority="N. Ito" authorityName="N. Ito" authorityYear="1996" box="[978,1301,2000,2025]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="curvatus">
|
||
<emphasis id="B9480707A3407F50FC22F87E8F91DED9" box="[978,1229,2000,2025]" italics="true" pageId="45" pageNumber="346">Trichotichnus curvatus</emphasis>
|
||
N. Ito
|
||
</taxonomicName>
|
||
” (
|
||
<collectionCode id="ED2D43D0A3407F50FADEF87F8EDEDED9" box="[1326,1410,2001,2025]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="45" pageNumber="346">MFNB</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3437F53FF37FF398958D8B8" blockId="46.[151,1437,151,392]" pageId="46" pageNumber="347">
|
||
<materialsCitation id="3B54D148A3437F53FF37FF3988D4D9E4" ID-GBIF-Occurrence="1655986736" collectingDate="1998-05-19" collectorName="Ahrens & Fabrizi" country="India" elevation="2315" location="Maney Bhanjyang" longitude="88.183334" municipality="Darjeeling Distr." pageId="46" pageNumber="347" specimenCount="1" specimenCount-male="1" stateProvince="West Bengal">
|
||
<emphasis id="B9480707A3437F53FF37FF3989DDD980" bold="true" box="[199,641,151,176]" pageId="46" pageNumber="347">
|
||
Additional material examined.
|
||
<collectingCountry id="F32B9B85A3437F53FDB1FF3989DDD980" box="[577,641,151,176]" name="India" pageId="46" pageNumber="347">India</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<specimenCount id="9D3A109CA3437F53FD61FF3689E3D99F" box="[657,703,152,176]" pageId="46" pageNumber="347" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingRegion id="49F815F7A3437F53FD3DFF36883CD980" box="[717,864,151,176]" country="India" name="West Bengal" pageId="46" pageNumber="347">West Bengal</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BE7416FA3437F53FC9EFF398F72D980" box="[878,1070,151,176]" pageId="46" pageNumber="347">Darjeeling Distr.</collectingMunicipality>
|
||
,
|
||
<location id="8EE38DCEA3437F53FBCBFF368E57D980" LSID="urn:lsid:plazi:treatment:03956A03A3407F4CFF67F9D48F0AD8E0:8EE38DCEA3437F53FBCBFF368E57D980" box="[1083,1291,151,176]" country="India" longitude="88.183334" municipality="Darjeeling Distr." name="Maney Bhanjyang" pageId="46" pageNumber="347" stateProvince="West Bengal">Maney Bhanjyang</location>
|
||
[ca 26°98'N
|
||
<geoCoordinate id="EE08BDD2A3437F53FF67FF128BAFD9E4" box="[151,243,188,212]" direction="east" orientation="longitude" pageId="46" pageNumber="347" precision="925" value="88.183334">88°11'E</geoCoordinate>
|
||
],
|
||
<elevation id="00113C26A3437F53FEFBFF128AF0D9E4" box="[267,428,188,213]" metricMagnitude="3" metricUnit="m" metricValue="2.315" metricValueMax="2.5" metricValueMin="2.13" pageId="46" pageNumber="347" unit="m" value="2315.0" valueMax="2500.0" valueMin="2130.0">
|
||
<quantity id="4CC476F0A3437F53FEFBFF128AF0D9E4" box="[267,428,188,213]" metricMagnitude="3" metricUnit="m" metricValue="2.315" metricValueMax="2.5" metricValueMin="2.13" pageId="46" pageNumber="347" unit="m" value="2315.0" valueMax="2500.0" valueMin="2130.0">2130–2500 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFC6043DA3437F53FE4BFF12896CD9E4" box="[443,560,188,212]" pageId="46" pageNumber="347" value="1998-05-19">
|
||
<date id="FF82FDD5A3437F53FE4BFF12896CD9E4" box="[443,560,188,212]" pageId="46" pageNumber="347" value="1998-05-19">19.V.1998</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C9BEC3A3437F53FDB0FF1289CDD9E5" box="[576,657,188,213]" pageId="46" pageNumber="347">Ahrens</collectorName>
|
||
&
|
||
<collectorName id="26C9BEC3A3437F53FD4DFF128850D9E5" box="[701,780,188,213]" pageId="46" pageNumber="347">Fabrizi</collectorName>
|
||
leg. (cJS)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B54D148A3437F53FC67FF128F19D9C8" ID-GBIF-Occurrence="1655986741" collectingDate="98-05-13" collectionCode="ZIN" collectorName="Ahrens & Fabrizi" country="India" elevation="2000" latitude="27.35" location="Kechu Pheri Lake" longLatPrecision="1237" longitude="88.2" pageId="46" pageNumber="347" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="West Bengal">
|
||
<specimenCount id="9D3A109CA3437F53FC67FF12889BD9E4" box="[919,967,188,212]" pageId="46" pageNumber="347" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3A109CA3437F53FC27FF128F59D9E4" box="[983,1029,188,212]" pageId="46" pageNumber="347" type="female">1 ♀</specimenCount>
|
||
, W-Sikkim,
|
||
<location id="8EE38DCEA3437F53FB6DFF128E33D9E5" LSID="urn:lsid:plazi:treatment:03956A03A3407F4CFF67F9D48F0AD8E0:8EE38DCEA3437F53FB6DFF128E33D9E5" box="[1181,1391,188,213]" country="India" latitude="27.35" longLatPrecision="1237" longitude="88.2" name="Kechu Pheri Lake" pageId="46" pageNumber="347" stateProvince="West Bengal">Kechu Pheri Lake</location>
|
||
[ca
|
||
<geoCoordinate id="EE08BDD2A3437F53FF67FF4E8BAAD9C8" box="[151,246,224,248]" direction="north" orientation="latitude" pageId="46" pageNumber="347" precision="925" value="27.35">27°21'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE08BDD2A3437F53FEF2FF4E8A3CD9C8" box="[258,352,224,248]" direction="east" orientation="longitude" pageId="46" pageNumber="347" precision="925" value="88.2">88°12'E</geoCoordinate>
|
||
],
|
||
<elevation id="00113C26A3437F53FE84FF4E894ED9C7" box="[372,530,224,248]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="46" pageNumber="347" unit="m" value="2000.0" valueMax="2100.0" valueMin="1900.0">
|
||
<quantity id="4CC476F0A3437F53FE84FF4E894ED9C7" box="[372,530,224,248]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="46" pageNumber="347" unit="m" value="2000.0" valueMax="2100.0" valueMin="1900.0">1900–2100 m</quantity>
|
||
</elevation>
|
||
, 10–
|
||
<collectingDate id="EFC6043DA3437F53FDB8FF7189C1D9C8" box="[584,669,223,248]" pageId="46" pageNumber="347" value="98-05-13">
|
||
<date id="FF82FDD5A3437F53FDB8FF7189C1D9C8" box="[584,669,223,248]" pageId="46" pageNumber="347" value="98-05-13">13.5.98</date>
|
||
</collectingDate>
|
||
lg.
|
||
<collectorName id="26C9BEC3A3437F53FD3AFF718847D9C8" box="[714,795,223,248]" pageId="46" pageNumber="347">Ahrens</collectorName>
|
||
&
|
||
<collectorName id="26C9BEC3A3437F53FCB1FF7188CCD9C8" box="[833,912,223,248]" pageId="46" pageNumber="347">Fabrizi</collectorName>
|
||
leg. (cJS,
|
||
<collectionCode id="ED2D43D0A3437F53FBF9FF4E8F60D9C8" box="[1033,1084,224,248]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="46" pageNumber="347">ZIN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B54D148A3437F53FBA0FF718F52D82C" ID-GBIF-Occurrence="1655986722" collectingDate="2003-04-03" collectionCode="NME" collectorName="J. Weipert" country="Nepal" county="Taplejung Distr." elevation="2100" location="Amjilosa" municipality="Sekathum" pageId="46" pageNumber="347" specimenCount="1" specimenCount-male="1">
|
||
<collectingCountry id="F32B9B85A3437F53FBA0FF718FCAD9C8" box="[1104,1174,223,248]" name="Nepal" pageId="46" pageNumber="347">
|
||
<emphasis id="B9480707A3437F53FBA0FF718FCAD9C8" bold="true" box="[1104,1174,223,248]" pageId="46" pageNumber="347">Nepal</emphasis>
|
||
</collectingCountry>
|
||
.
|
||
<specimenCount id="9D3A109CA3437F53FB55FF4E8F8FD9C7" box="[1189,1235,224,248]" pageId="46" pageNumber="347" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingCounty id="62E2A399A3437F53FB10FF718EC4D9C8" box="[1248,1432,223,248]" pageId="46" pageNumber="347">Taplejung Distr.</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BE7416FA3437F53FF67FEAA8A5BD82D" box="[151,263,260,285]" pageId="46" pageNumber="347">Sekathum</collectingMunicipality>
|
||
bis
|
||
<location id="8EE38DCEA3437F53FEC9FEAA8AFFD82D" LSID="urn:lsid:plazi:treatment:03956A03A3407F4CFF67F9D48F0AD8E0:8EE38DCEA3437F53FEC9FEAA8AFFD82D" box="[313,419,260,285]" country="Nepal" county="Taplejung Distr." municipality="Sekathum" name="Amjilosa" pageId="46" pageNumber="347">Amjilosa</location>
|
||
,
|
||
<elevation id="00113C26A3437F53FE40FEAA8911D82C" box="[432,589,260,285]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="46" pageNumber="347" unit="m" value="2100.0" valueMax="2500.0" valueMin="1700.0">
|
||
<quantity id="4CC476F0A3437F53FE40FEAA8911D82C" box="[432,589,260,285]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="46" pageNumber="347" unit="m" value="2100.0" valueMax="2500.0" valueMin="1700.0">1700–2500 m</quantity>
|
||
</elevation>
|
||
NN,
|
||
<collectingDate id="EFC6043DA3437F53FD7EFEAA89A3D82C" box="[654,767,260,284]" pageId="46" pageNumber="347" value="2003-04-03">
|
||
<date id="FF82FDD5A3437F53FD7EFEAA89A3D82C" box="[654,767,260,284]" pageId="46" pageNumber="347" value="2003-04-03">3.IV.2003</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C9BEC3A3437F53FCFCFEAB8823D82D" box="[780,895,260,285]" pageId="46" pageNumber="347">J. Weipert</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED2D43D0A3437F53FC32FEAA8F5AD82C" box="[962,1030,260,284]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="46" pageNumber="347">NME</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B54D148A3437F53FBEBFEAA8AADD854" ID-GBIF-Occurrence="1655986744" collectingDate="1988-04-27" collectingDateMax="1988-04-29" collectingDateMin="1988-04-27" collectionCode="SMNS, ZIN" collectorName="J. Martens & W. Schawaller" country="Nepal" county="Taplejung Distr." elevation="1900" location="Kabeli Khola" municipality="Yamputhin" pageId="46" pageNumber="347" specimenCount="5" specimenCount-female="2" specimenCount-male="3">
|
||
<specimenCount id="9D3A109CA3437F53FBEBFEAA8F16D82C" box="[1051,1098,260,284]" pageId="46" pageNumber="347" type="male">3 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3A109CA3437F53FBA8FEAA8FDFD82C" box="[1112,1155,260,284]" pageId="46" pageNumber="347" type="female">2 ♀</specimenCount>
|
||
,
|
||
<collectingCounty id="62E2A399A3437F53FB64FEAA8E11D82D" box="[1172,1357,260,285]" pageId="46" pageNumber="347">Taplejung Distr.</collectingCounty>
|
||
, above
|
||
<collectingMunicipality id="6BE7416FA3437F53FF67FE898A49D870" box="[151,277,295,320]" pageId="46" pageNumber="347">Yamputhin</collectingMunicipality>
|
||
, left bank of
|
||
<location id="8EE38DCEA3437F53FE4FFE898902D870" LSID="urn:lsid:plazi:treatment:03956A03A3407F4CFF67F9D48F0AD8E0:8EE38DCEA3437F53FE4FFE898902D870" box="[447,606,295,320]" country="Nepal" county="Taplejung Distr." municipality="Yamputhin" name="Kabeli Khola" pageId="46" pageNumber="347">Kabeli Khola</location>
|
||
, bushes, open forest,
|
||
<elevation id="00113C26A3437F53FC96FE868F54D80F" box="[870,1032,296,320]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="46" pageNumber="347" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">
|
||
<quantity id="4CC476F0A3437F53FC96FE868F54D80F" box="[870,1032,296,320]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="46" pageNumber="347" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800–2000 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFC6043DA3437F53FBEAFE868F9ED870" box="[1050,1218,296,320]" pageId="46" pageNumber="347" value="1988-04-27" valueMax="1988-04-29" valueMin="1988-04-27">
|
||
<date id="FF82FDD5A3437F53FBEAFE868F9ED870" box="[1050,1218,296,320]" pageId="46" pageNumber="347" value="1988-04-27" valueMax="1988-04-29" valueMin="1988-04-27">27–29.IV.1988</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C9BEC3A3437F53FB23FE868E12D80F" box="[1235,1358,296,319]" pageId="46" pageNumber="347">J. Martens</collectorName>
|
||
&
|
||
<collectorName id="26C9BEC3A3437F53FA8DFE868A48D855" pageId="46" pageNumber="347">W. Schawaller</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED2D43D0A3437F53FEA8FEE28AF5D854" box="[344,425,332,356]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="46" pageNumber="347">SMNS</collectionCode>
|
||
,
|
||
<collectionCode id="ED2D43D0A3437F53FE46FEE28AB5D854" box="[438,489,332,356]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="46" pageNumber="347">ZIN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B54D148A3437F53FE0FFEE28958D8B8" ID-GBIF-Occurrence="1655986739" collectingDate="1999-09-05" collectorName="J. Schmidt" country="Nepal" county="Sindhupalchok Distr." elevation="2200" location="Ting Sang La" municipality="Lapchi Kang" pageId="46" pageNumber="347" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="9D3A109CA3437F53FE0FFEE28970D854" box="[511,556,332,356]" pageId="46" pageNumber="347" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingCounty id="62E2A399A3437F53FDCDFEE28873D855" box="[573,815,332,357]" pageId="46" pageNumber="347">Sindhupalchok Distr.</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BE7416FA3437F53FCCDFEE2888CD855" box="[829,976,332,357]" pageId="46" pageNumber="347">Lapchi Kang</collectingMunicipality>
|
||
range,
|
||
<location id="8EE38DCEA3437F53FBD9FEE28F95D854" LSID="urn:lsid:plazi:treatment:03956A03A3407F4CFF67F9D48F0AD8E0:8EE38DCEA3437F53FBD9FEE28F95D854" box="[1065,1225,332,357]" country="Nepal" county="Sindhupalchok Distr." municipality="Lapchi Kang" name="Ting Sang La" pageId="46" pageNumber="347">Ting Sang La</location>
|
||
, E slope,
|
||
<elevation id="00113C26A3437F53FACEFEE28ECBD854" box="[1342,1431,332,356]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="46" pageNumber="347" unit="m" value="2200.0">
|
||
<quantity id="4CC476F0A3437F53FACEFEE28ECBD854" box="[1342,1431,332,356]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="46" pageNumber="347" unit="m" value="2200.0">2200 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFC6043DA3437F53FF67FEC18A56D8B8" box="[151,266,367,392]" pageId="46" pageNumber="347" value="1999-09-05">
|
||
<date id="FF82FDD5A3437F53FF67FEC18A56D8B8" box="[151,266,367,392]" pageId="46" pageNumber="347" value="1999-09-05">5.IX.1999</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C9BEC3A3437F53FEE6FEDE8AD0D8B8" box="[278,396,367,392]" pageId="46" pageNumber="347">J. Schmidt</collectorName>
|
||
leg. (cJS).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF438B9DA3437F53FF67FB72883EDC21" httpUri="https://zenodo.org/record/919684/files/figure.png" pageId="46" pageNumber="347" targetBox="[301,1277,456,1212]" targetPageId="46">
|
||
<paragraph id="8B83DB15A3437F53FF67FB72883EDC21" blockId="46.[151,1435,1244,1297]" pageId="46" pageNumber="347">
|
||
<emphasis id="B9480707A3437F53FF67FB728A2FDDC1" bold="true" box="[151,371,1244,1267]" pageId="46" pageNumber="347">FIGURES 135–136.</emphasis>
|
||
<taxonomicName id="4C3CA096A3437F53FE8FFB7288F3DDC2" authority="Andrewes, 1930" authorityName="Andrewes" authorityYear="1930" box="[383,943,1244,1266]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="347" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3437F53FE8FFB728956DDC2" box="[383,522,1244,1266]" italics="true" pageId="46" pageNumber="347">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3437F53FDEFFB7289DDDDC2" box="[543,641,1244,1266]" italics="true" pageId="46" pageNumber="347">Bottchrus</emphasis>
|
||
)
|
||
<emphasis id="B9480707A3437F53FD66FB7289A8DDC2" box="[662,756,1244,1266]" italics="true" pageId="46" pageNumber="347">hingstoni</emphasis>
|
||
Andrewes, 1930
|
||
</taxonomicName>
|
||
, median lobe of aedeagus, lateral view (135, lectotype; 136, holotype of
|
||
<taxonomicName id="4C3CA096A3437F53FE5EFB5289D8DC21" authority="Ito, 1996" authorityName="Ito" authorityYear="1996" box="[430,644,1275,1297]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="347" phylum="Arthropoda" rank="species" species="curvatus">
|
||
<emphasis id="B9480707A3437F53FE5EFB528942DC21" box="[430,542,1276,1297]" italics="true" pageId="46" pageNumber="347">T. curvatus</emphasis>
|
||
Ito, 1996
|
||
</taxonomicName>
|
||
). Scale bar = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C326889EA3437F4DFF37FA938ABBDC48" lastPageId="48" lastPageNumber="349" pageId="46" pageNumber="347" type="description">
|
||
<paragraph id="8B83DB15A3437F53FF37FA938F02DC66" blockId="46.[151,1437,1341,2015]" box="[199,1118,1341,1366]" pageId="46" pageNumber="347">
|
||
<emphasis id="B9480707A3437F53FF37FA938A0DDC66" bold="true" box="[199,337,1341,1366]" pageId="46" pageNumber="347">Description</emphasis>
|
||
. Habitus as in
|
||
<figureCitation id="1307C790A3437F53FE08FA938905DC66" box="[504,601,1341,1366]" captionStart="FIGURES 120 – 123" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[229,1356,202,1919]" captionTargetId="figure@41.[227,1360,193,1921]" captionTargetPageId="41" captionText="FIGURES 120 – 123. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 120, T. sikkimensis sp. n., holotype; 121, T. martensi sp. n., holotype; 122, T. hingstoni Andrewes, 1930, male from Kechu Pheri Lake; 123, T. birmanicus Bates, 1892, male from the Kali Gandaki, Tatopani." httpUri="https://zenodo.org/record/919672/files/figure.png" pageId="46" pageNumber="347">Fig. 122</figureCitation>
|
||
. Body length
|
||
<quantity id="4CC476F0A3437F53FD05FA938822DC66" box="[757,894,1341,1366]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.7" metricValueMin="6.8" pageId="46" pageNumber="347" unit="mm" value="7.25" valueMax="7.7" valueMin="6.8">6.8–7.7 mm</quantity>
|
||
, width 3.0–
|
||
<quantity id="4CC476F0A3437F53FBF2FA908F06DC66" box="[1026,1114,1342,1366]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="46" pageNumber="347" unit="mm" value="3.3">3.3 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3437F53FF37FACF8BBADCAE" blockId="46.[151,1437,1341,2015]" pageId="46" pageNumber="347">Colour: Body piceous, dorsum shiny, with light bluish lustre; palpi, antennae and legs brownish yellow or light brown.</paragraph>
|
||
<paragraph id="8B83DB15A3437F53FF37FA078A2ADFD2" blockId="46.[151,1437,1341,2015]" pageId="46" pageNumber="347">
|
||
Head: Large (in males, HWmax/PWmax = 0.70–0.72; HWmin/PWmax = 0.56–0.59; in females, these indices 0.70–0.72 and 0.56–0.59, respectively). Eyes moderately or weakly convex (HWmax/HWmin =
|
||
<quantity id="4CC476F0A3437F53FAFAFA638EC0DCD6" box="[1290,1436,1485,1510]" metricMagnitude="-2" metricUnit="m" metricValue="3.0860999999999996" metricValueMax="3.2004" metricValueMin="2.9718" pageId="46" pageNumber="347" unit="in" value="1.2149999999999999" valueMax="1.26" valueMin="1.17">1.17–1.26 in</quantity>
|
||
males, and
|
||
<quantity id="4CC476F0A3437F53FEE6FA5C8AFFDF3A" box="[278,419,1521,1546]" metricMagnitude="-2" metricUnit="m" metricValue="3.1623" metricValueMax="3.2512" metricValueMin="3.0734" pageId="46" pageNumber="347" unit="in" value="1.245" valueMax="1.28" valueMin="1.21">1.21–1.28 in</quantity>
|
||
female), slightly elongate oval in lateral view. Temporae about one-third or half as long as eye, convex, slopingly or somewhat abruptly fallen to neck. Genae wide, without setae. Supraorbital setigerous pore located just behind level of posterior margin of eye, separated from supraorbital furrow approximately by width of antennomere 3 apically. Labrum moderately concave anteriorly.
|
||
<collectingRegion id="49F815F7A3437F53FC04F9F18F0FDF46" box="[1012,1107,1631,1654]" country="Myanmar" name="Chin" pageId="46" pageNumber="347">Mentum</collectingRegion>
|
||
and submentum completely fused. Left mandible slightly blunted at apex. Antennae extended just to pronotal basal edge, with antennomeres 4– 8 slightly markedly longer than wide. Dorsal microsculpture visible only behind eyes, meshes weakly transverse, nearly isodiametric.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3437F52FF37F9418AA3D9C8" blockId="46.[151,1437,1341,2015]" lastBlockId="47.[151,1436,151,248]" lastPageId="47" lastPageNumber="348" pageId="46" pageNumber="347">Pronotum: Comparatively wide (PWmax/PL = 1.37–1.44), widest approximately at the end of anterior third, moderately narrowed posteriad (PWmax/PWmin = 1.18–1.27), with one lateral seta inserted slightly before widest point. Sides strongly convex apically and more or less straight basally (but never sinuate). Apical margin very weakly arcuately emarginate, bordered almost throughout. Basal margin straight medially, oblique laterally, vaguely bordered, occasionally only with a short gap at middle. Apical angles not prominent, rounded at apex. Basal angles obtuse, somewhat sharp at apex, without any denticles. Pronotal disc convex, not depressed basally, in some specimens slightly depressed at basal angles. Lateral gutter narrow throughout. Basal foveae small and flat, occasionally poorly evident. Base finely punctate mainly in lateral areas; without distinct punctures at apical margin. Microsculpture very fine, visible throughout, consisting of transverse meshes on disc and nearly isodiametric meshes at margins.</paragraph>
|
||
<caption id="DF438B9DA3427F52FF67F9078884DE0A" httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="47" pageNumber="348" targetBox="[194,1406,318,1664]" targetPageId="47">
|
||
<paragraph id="8B83DB15A3427F52FF67F9078884DE0A" blockId="47.[151,1436,1705,1850]" pageId="47" pageNumber="348">
|
||
<emphasis id="B9480707A3427F52FF67F9078A32DF8F" bold="true" box="[151,366,1705,1728]" pageId="47" pageNumber="348">FIGURES 137–145.</emphasis>
|
||
<taxonomicName id="4C3CA096A3427F52FE84F90488D1DFF0" authority="Andrewes, 1930" authorityName="Andrewes" authorityYear="1930" box="[372,909,1706,1728]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="348" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3427F52FE84F9048AA3DFF0" box="[372,511,1706,1728]" italics="true" pageId="47" pageNumber="348">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3427F52FDFFF904892EDFF0" box="[527,626,1706,1728]" italics="true" pageId="47" pageNumber="348">Bottchrus</emphasis>
|
||
)
|
||
<emphasis id="B9480707A3427F52FD71F9048983DFF0" box="[641,735,1706,1728]" italics="true" pageId="47" pageNumber="348">hingstoni</emphasis>
|
||
Andrewes, 1930
|
||
</taxonomicName>
|
||
(137–139, Kechu Pheri Lake, W-Sikkim; 140–143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143–145, middle portion of median lobe. 137–139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143–145, right lateral view. Scale bars = 0.25 mm (A: Figs 137–140, 142–145; B: Fig. 141).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B83DB15A3427F4DFF37F8C98F57D8B7" blockId="47.[151,1436,1895,2028]" lastBlockId="48.[151,1437,151,2013]" lastPageId="48" lastPageNumber="349" pageId="47" pageNumber="348">Elytra: In lateral and caudal view convex, in dorsal view oval, moderately wide (in male, EL/EW = 1.35–1.49; EL/PL = 2.33–2.64; EW/PWmax = 1.23–1.29; in female, these indices 1.45–1.47, 2.42–2.70, and 1.20–27, respectively); widest behind middle, abruptly fallen behind, with subangulate humeri, either without denticle at apex or with a very tiny denticle barely visible from behind. Preapical sinuation shallow, sutural angle sharp at tip, nearly rectangular apically. Basal bead slightly sinuate, meeting lateral margin at distinct obtuse angle. Striae impunctate, not impressed, nearly superficial, at most lateral striae slightly impressed apically. Intervals impunctate, more or less flat. Parascutellar striole about as long as distance from this pore to suture, with apex free or connected with stria 2, with a large parascutellar pore at base. Interval 3 with a small discal pore just behind middle. Marginal umbilicate series widely interrupted medially, consisting of 7–9 pores in anterior group and 8–10 pores in posterior group. Microsculpture strongly obliterate on disc, visible in most specimens only along basal elytral bead, at apex, and on two lateral intervals; meshes very fine, transverse.</paragraph>
|
||
<paragraph id="8B83DB15A35D7F4DFF37FE3A8869D89D" blockId="48.[151,1437,151,2013]" box="[199,821,404,429]" pageId="48" pageNumber="349">Hindwings: Reduced to scales about a quarter of elytra.</paragraph>
|
||
<paragraph id="8B83DB15A35D7F4DFF37FE198AF9DB28" blockId="48.[151,1437,151,2013]" pageId="48" pageNumber="349">
|
||
Ventral side: Metepisterna (
|
||
<figureCitation id="1307C790A35D7F4DFE0DFE198900D8E0" box="[509,604,439,464]" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="48" pageNumber="349">Fig. 137</figureCitation>
|
||
) longer than wide, strongly narrowed posteriad. Apex of last visible (VII) abdominal sternite in male (
|
||
<figureCitation id="1307C790A35D7F4DFE25FE728969D8C4" box="[469,565,476,501]" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="48" pageNumber="349">Fig. 138</figureCitation>
|
||
) subtruncate and more or less clearly concave just at middle, in female (
|
||
<figureCitation id="1307C790A35D7F4DFA9FFE728B9EDB28" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="48" pageNumber="349">Fig. 139</figureCitation>
|
||
) angularly rounded.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A35D7F4DFF37FD8B8AA2DB98" blockId="48.[151,1437,151,2013]" pageId="48" pageNumber="349">Legs: Protibia finely sulcate on dorsal side basally. Profemur with three to five setae on anterio-ventral margin. Metatarsus short, shorter than width of head across neck constriction, with first metatarsomere approximately equal to second and third together. Tarsomere 5 with two pairs of ventro-lateral setae. Pro- and mesotarsi of male comparatively weakly enlarged.</paragraph>
|
||
<paragraph id="8B83DB15A35D7F4DFF37FD1A8F9EDBC0" blockId="48.[151,1437,151,2013]" pageId="48" pageNumber="349">Female genitalia: Laterotergite (hemisternite) with two apical setae. Basal palpomere with one apical seta. Apical stylomere moderately curved, with one peg-like seta in both ventral and dorsal margins.</paragraph>
|
||
<paragraph id="8B83DB15A35D7F4DFF37FD528F7CDA95" blockId="48.[151,1437,151,2013]" pageId="48" pageNumber="349">
|
||
Male genitalia: Median lobe of aedeagus (
|
||
<figureCitation id="1307C790A35D7F4DFD42FD52880FDA25" box="[690,851,764,789]" captionStart="FIGURES 135 – 136" captionStartId="46.[151,264,1244,1267]" captionTargetBox="[301,1277,456,1212]" captionTargetId="figure@46.[271,1316,445,1223]" captionTargetPageId="46" captionText="FIGURES 135 – 136. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930, median lobe of aedeagus, lateral view (135, lectotype; 136, holotype of T. curvatus Ito, 1996). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/919684/files/figure.png" pageId="48" pageNumber="349">Figs 135–136</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A35D7F4DFC92FD528899DA25" box="[866,965,764,789]" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="48" pageNumber="349">140–145</figureCitation>
|
||
) C-shaped, bent behind basal bulb, with terminal lamella very short, slightly narrowed posteriad; apical capitulum in lateral view (
|
||
<figureCitation id="1307C790A35D7F4DFB47FCB18E05DA08" box="[1207,1369,799,824]" captionStart="FIGURES 135 – 136" captionStartId="46.[151,264,1244,1267]" captionTargetBox="[301,1277,456,1212]" captionTargetId="figure@46.[271,1316,445,1223]" captionTargetPageId="46" captionText="FIGURES 135 – 136. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930, median lobe of aedeagus, lateral view (135, lectotype; 136, holotype of T. curvatus Ito, 1996). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/919684/files/figure.png" pageId="48" pageNumber="349">Figs 135–136</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A35D7F4DFA99FC8E8EC8DA08" box="[1385,1428,800,824]" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="48" pageNumber="349">140</figureCitation>
|
||
) clearly oblique, with ventroapical flange located more distally than dorsoapical flange, in caudal view (
|
||
<figureCitation id="1307C790A35D7F4DFAC3FCEA8EC8DA6C" box="[1331,1428,836,861]" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="48" pageNumber="349">Fig. 141</figureCitation>
|
||
) triangular, somewhat horseshoe-shaped, with rather acute apex. Internal sac beside an elongate curved spiny formation on the rigth side (Figs 14 3–145) without additional sclerotic elements.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A35D7F4DFF37FC018ABBDC48" blockId="48.[151,1437,151,2013]" pageId="48" pageNumber="349">
|
||
<emphasis id="B9480707A35D7F4DFF37FC018A06DAF8" bold="true" box="[199,346,943,968]" pageId="48" pageNumber="349">Comparison</emphasis>
|
||
. Among comparatively small brachypterous
|
||
<taxonomicName id="4C3CA096A35D7F4DFCA4FC01889DDAF8" authorityName="Jedlicka" authorityYear="1935" box="[852,961,943,968]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9480707A35D7F4DFCA4FC01889DDAF8" box="[852,961,943,968]" italics="true" pageId="48" pageNumber="349">Bottchrus</emphasis>
|
||
</taxonomicName>
|
||
of the Himalayan region,
|
||
<taxonomicName id="4C3CA096A35D7F4DFB16FC1F8EC0DAF8" authorityName="Andrewes" authorityYear="1930" box="[1254,1436,943,968]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFB16FC1F8FA0DAF8" box="[1254,1276,945,968]" italics="true" pageId="48" pageNumber="349">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFAFBFC1F8E40DAF8" box="[1291,1308,945,968]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFAC4FC018EC0DAF8" box="[1332,1436,943,968]" italics="true" pageId="48" pageNumber="349">hingstoni</emphasis>
|
||
</taxonomicName>
|
||
readily discriminates by the following combination of distinctive characters: larger size (
|
||
<quantity id="4CC476F0A35D7F4DFB41FC7A8E1FDADC" box="[1201,1347,980,1005]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.7" metricValueMin="6.8" pageId="48" pageNumber="349" unit="mm" value="7.25" valueMax="7.7" valueMin="6.8">6.8–7.7 mm</quantity>
|
||
), genae glabrous, metepisterna markedly longer than wide (
|
||
<figureCitation id="1307C790A35D7F4DFD2CFC598862DD20" box="[732,830,1015,1040]" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="48" pageNumber="349">Fig. 137</figureCitation>
|
||
), pronotum usually with rounded sides (never sinuate basally), protibia longitudinally sulcate on dorsal side, apex of last visible (VII) abdominal sternite of male slightly concave at middle (
|
||
<figureCitation id="1307C790A35D7F4DFE8CFB918ABCDD68" box="[380,480,1087,1112]" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="48" pageNumber="349">Fig. 138</figureCitation>
|
||
), and aedeagal median lobe C-shaped, with short terminal lamella, clearly oblique apical capitulum and without any spines in the internal sac (
|
||
<figureCitation id="1307C790A35D7F4DFCBAFBCA88B6DD4D" box="[842,1002,1124,1149]" captionStart="FIGURES 135 – 136" captionStartId="46.[151,264,1244,1267]" captionTargetBox="[301,1277,456,1212]" captionTargetId="figure@46.[271,1316,445,1223]" captionTargetPageId="46" captionText="FIGURES 135 – 136. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930, median lobe of aedeagus, lateral view (135, lectotype; 136, holotype of T. curvatus Ito, 1996). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/919684/files/figure.png" pageId="48" pageNumber="349">Figs 135–136</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A35D7F4DFC07FBCA8F00DD4D" box="[1015,1116,1124,1149]" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="48" pageNumber="349">140–145</figureCitation>
|
||
), only with elongate curved spiny formation on the right side (
|
||
<figureCitation id="1307C790A35D7F4DFDD8FB298996DD90" box="[552,714,1159,1184]" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="48" pageNumber="349">Figs 143–145</figureCitation>
|
||
). In body size and presence of a fine sulcus on dorsal side of protibia,
|
||
<taxonomicName id="4C3CA096A35D7F4DFF0CFB038AECDDF5" authorityName="Andrewes" authorityYear="1930" box="[252,432,1196,1221]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFF0CFB038A4DDDF4" box="[252,273,1197,1220]" italics="true" pageId="48" pageNumber="349">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFED0FB038A6DDDF4" box="[288,305,1197,1220]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFEB8FB028AECDDF5" box="[328,432,1196,1221]" italics="true" pageId="48" pageNumber="349">hingstoni</emphasis>
|
||
</taxonomicName>
|
||
somewhat reminds
|
||
<taxonomicName id="4C3CA096A35D7F4DFD61FB03881BDDF5" box="[657,839,1196,1221]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="cyanellus" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFD61FB0389FADDF4" box="[657,678,1197,1220]" italics="true" pageId="48" pageNumber="349">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFD45FB03899ADDF4" box="[693,710,1197,1220]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFD2DFB02881BDDF5" box="[733,839,1196,1221]" italics="true" pageId="48" pageNumber="349">cyanellus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3CA096A35D7F4DFC8DFB038F56DDF5" authorityName="Ito" authorityYear="1998" box="[893,1034,1196,1221]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="loebli" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFC8DFB0388CEDDF4" box="[893,914,1197,1220]" italics="true" pageId="48" pageNumber="349">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFC52FB0388EFDDF4" box="[930,947,1197,1220]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFC3AFB028F56DDF5" box="[970,1034,1196,1221]" italics="true" pageId="48" pageNumber="349">loebli</emphasis>
|
||
</taxonomicName>
|
||
, but the latter two species distinctly differ in metepisterna as long as wide or shorter, pronotal sides straight or slightly sinuate basally, apex of last visible (VII) abdominal sternite of male rounded or subtruncate, not concave, and aedeagal median lobe somewhat S-shaped. Based on the original description (
|
||
<bibRefCitation id="EFADA6E4A35D7F4DFD64FAB6886FDC00" author="Ito" box="[660,819,1304,1328]" pageId="48" pageNumber="349" refString="Ito, N. (1998) Some species of the genus Trichotichnus from Nepal and the adjascent regions (Coleoptera, Carabidae, Harpalini). Japanese Journal of Systematic Entomology, 4 (2), 277 - 288." type="journal article" year="1998">Ito 1998: 284</bibRefCitation>
|
||
), the sympatric
|
||
<taxonomicName id="4C3CA096A35D7F4DFC19FAB78FD4DC00" authorityName="Ito" authorityYear="1998" box="[1001,1160,1304,1328]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="opacus" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFC19FAB788A2DC00" box="[1001,1022,1305,1328]" italics="true" pageId="48" pageNumber="349">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFBFFFAB78F7CDC00" box="[1039,1056,1305,1328]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFBC9FAB78FD4DC00" box="[1081,1160,1305,1328]" italics="true" pageId="48" pageNumber="349">opacus</emphasis>
|
||
</taxonomicName>
|
||
, known to us only from literature, is distinguished from
|
||
<taxonomicName id="4C3CA096A35D7F4DFDFEFA938997DC65" authorityName="Andrewes" authorityYear="1930" box="[526,715,1340,1365]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFDFEFA93897FDC64" box="[526,547,1341,1364]" italics="true" pageId="48" pageNumber="349">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFDC7FA938914DC64" box="[567,584,1341,1364]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFD93FA928997DC65" box="[611,715,1340,1365]" italics="true" pageId="48" pageNumber="349">hingstoni</emphasis>
|
||
</taxonomicName>
|
||
by smaller size (about
|
||
<quantity id="4CC476F0A35D7F4DFC15FA928F1DDC64" box="[997,1089,1340,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="48" pageNumber="349" unit="mm" value="5.8">5.8 mm</quantity>
|
||
) and almost transverse apical capitulum of the median lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA35D7F4DFF37FA2A8E30DF1D" pageId="48" pageNumber="349" type="distribution">
|
||
<paragraph id="8B83DB15A35D7F4DFF37FA2A8E30DF1D" blockId="48.[151,1437,151,2013]" pageId="48" pageNumber="349">
|
||
<emphasis id="B9480707A35D7F4DFF37FA2A8A05DCAD" bold="true" box="[199,345,1412,1437]" pageId="48" pageNumber="349">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="1307C790A35D7F4DFE9EFA2A8A8FDCAD" box="[366,467,1412,1437]" captionStart="FIGURE 152" captionStartId="53.[151,250,1975,1997]" captionTargetBox="[151,1436,1234,1954]" captionTargetId="figure@53.[151,1436,1234,1954]" captionTargetPageId="53" captionText="FIGURE 152. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. gupchiensis sp. n.; 2, T. brancuccii sp. n.; 3, T. manasluensis sp. n.; 4, T. loebli Ito, 1998; 5, T. sikkimensis sp. n.; 6, T. hingstoni Andrewes, 1930." httpUri="https://zenodo.org/record/919698/files/figure.png" pageId="48" pageNumber="349">Fig. 152</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A35D7F4DFE15FA2A8AAFDCAD" box="[485,499,1412,1437]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="48" pageNumber="349">
|
||
<emphasis id="B9480707A35D7F4DFE15FA2A8AAFDCAD" box="[485,499,1412,1437]" italics="true" pageId="48" pageNumber="349">6</emphasis>
|
||
</figureCitation>
|
||
).
|
||
<collectingRegion id="49F815F7A35D7F4DFDFFFA2A893EDCAD" box="[527,610,1412,1437]" country="Sri Lanka" name="Central" pageId="48" pageNumber="349">Central</collectingRegion>
|
||
part of the Greater Himalaya: up to today known from the Singalila mountain range in East
|
||
<collectingRegion id="49F815F7A35D7F4DFE54FA098AA9DCF0" box="[420,501,1447,1472]" country="India" name="Sikkim" pageId="48" pageNumber="349">Sikkim</collectingRegion>
|
||
and the Darjeeling District of
|
||
<collectingCountry id="F32B9B85A35D7F4DFCBFFA0988D5DCF0" box="[847,905,1447,1472]" name="India" pageId="48" pageNumber="349">India</collectingCountry>
|
||
in the east to the Lapchi Kang mountain range of
|
||
<collectingRegion id="49F815F7A35D7F4DFF45FA628A54DCD5" box="[181,264,1484,1509]" country="Sri Lanka" name="Central" pageId="48" pageNumber="349">Central</collectingRegion>
|
||
<collectingCountry id="F32B9B85A35D7F4DFEFEFA628A0DDCD5" box="[270,337,1484,1509]" name="Nepal" pageId="48" pageNumber="349">Nepal</collectingCountry>
|
||
in the west. The species was found at altitudes of
|
||
<quantity id="4CC476F0A35D7F4DFC72FA628F7CDCD4" box="[898,1056,1484,1509]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="48" pageNumber="349" unit="m" value="2100.0" valueMax="2500.0" valueMin="1700.0">1700–2500 m</quantity>
|
||
. It occurs sympatricly with
|
||
<taxonomicName id="4C3CA096A35D7F4DFAA6FA638BBADF38" authorityName="Ito" authorityYear="1998" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="opacus" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFAA6FA638E37DCD4" box="[1366,1387,1485,1508]" italics="true" pageId="48" pageNumber="349">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFA8AFA638ED7DCD4" box="[1402,1419,1485,1508]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFF67FA5F8BBADF38" box="[151,230,1521,1544]" italics="true" pageId="48" pageNumber="349">opacus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1307C790A35D7F4DFF05FA418A09DF38" box="[245,341,1519,1544]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="48" pageNumber="349">Fig. 151</figureCitation>
|
||
,
|
||
<emphasis id="B9480707A35D7F4DFE91FA5E8A21DF38" box="[353,381,1520,1544]" italics="true" pageId="48" pageNumber="349">14</emphasis>
|
||
) and
|
||
<taxonomicName id="4C3CA096A35D7F4DFE4CFA5F8914DF38" authorityName="Ito" authorityYear="1998" box="[444,584,1519,1544]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="loebli" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFE4CFA5F8A8DDF38" box="[444,465,1521,1544]" italics="true" pageId="48" pageNumber="349">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFE10FA5F8AADDF38" box="[480,497,1521,1544]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFDF8FA418914DF38" box="[520,584,1519,1544]" italics="true" pageId="48" pageNumber="349">loebli</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1307C790A35D7F4DFDA7FA4189EBDF38" box="[599,695,1519,1544]" captionStart="FIGURE 152" captionStartId="53.[151,250,1975,1997]" captionTargetBox="[151,1436,1234,1954]" captionTargetId="figure@53.[151,1436,1234,1954]" captionTargetPageId="53" captionText="FIGURE 152. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. gupchiensis sp. n.; 2, T. brancuccii sp. n.; 3, T. manasluensis sp. n.; 4, T. loebli Ito, 1998; 5, T. sikkimensis sp. n.; 6, T. hingstoni Andrewes, 1930." httpUri="https://zenodo.org/record/919698/files/figure.png" pageId="48" pageNumber="349">Fig. 152</figureCitation>
|
||
,
|
||
<emphasis id="B9480707A35D7F4DFD33FA5E898DDF38" box="[707,721,1520,1544]" italics="true" pageId="48" pageNumber="349">4</emphasis>
|
||
) in the Darjeeling District, and with
|
||
<taxonomicName id="4C3CA096A35D7F4DFB9CFA5F8E46DF38" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1132,1306,1520,1544]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFB9CFA5F8FDDDF38" box="[1132,1153,1521,1544]" italics="true" pageId="48" pageNumber="349">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFB60FA5F8FFDDF38" box="[1168,1185,1521,1544]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFB48FA5F8E46DF38" box="[1208,1306,1521,1544]" italics="true" pageId="48" pageNumber="349">martensi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA35D7F4DFAD1FA418E3CDF38" box="[1313,1376,1519,1544]" pageId="48" pageNumber="349" rank="species">
|
||
<emphasis id="B9480707A35D7F4DFAD1FA418E3CDF38" bold="true" box="[1313,1376,1519,1544]" pageId="48" pageNumber="349">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="1307C790A35D7F4DFA9EFA418B9FDF1D" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="48" pageNumber="349">Fig. 151</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A35D7F4DFF3FF9BA8BB7DF1C" box="[207,235,1556,1580]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="48" pageNumber="349">
|
||
<emphasis id="B9480707A35D7F4DFF3FF9BA8BB7DF1C" box="[207,235,1556,1580]" italics="true" pageId="48" pageNumber="349">13</emphasis>
|
||
</figureCitation>
|
||
),
|
||
<taxonomicName id="4C3CA096A35D7F4DFEF2F9BB8A91DF1C" authorityName="Ito" authorityYear="1998" box="[258,461,1556,1580]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="cyanescens" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFEF2F9BB8A4BDF1C" box="[258,279,1557,1580]" italics="true" pageId="48" pageNumber="349">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFED7F9BB8A64DF1C" box="[295,312,1557,1580]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFEBFF9BB8A91DF1C" box="[335,461,1557,1580]" italics="true" pageId="48" pageNumber="349">cyanescens</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1307C790A35D7F4DFE2DF9BA8961DF1D" box="[477,573,1556,1581]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="48" pageNumber="349">Fig. 151</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A35D7F4DFDB9F9BA8939DF1C" box="[585,613,1556,1580]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="48" pageNumber="349">
|
||
<emphasis id="B9480707A35D7F4DFDB9F9BA8939DF1C" box="[585,613,1556,1580]" italics="true" pageId="48" pageNumber="349">12</emphasis>
|
||
</figureCitation>
|
||
), and probably with
|
||
<taxonomicName id="4C3CA096A35D7F4DFCBFF9BB88B2DF1D" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[847,1006,1556,1581]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="schawalleri" status="sp. nov.">
|
||
<emphasis id="B9480707A35D7F4DFCBFF9BB88B2DF1D" box="[847,1006,1556,1581]" italics="true" pageId="48" pageNumber="349">T. schawalleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA35D7F4DFC04F9BA8F68DF1D" box="[1012,1076,1556,1581]" pageId="48" pageNumber="349" rank="species">
|
||
<emphasis id="B9480707A35D7F4DFC04F9BA8F68DF1D" bold="true" box="[1012,1076,1556,1581]" pageId="48" pageNumber="349">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="1307C790A35D7F4DFBB3F9BA8FFFDF1D" box="[1091,1187,1556,1581]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="48" pageNumber="349">Fig. 151</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A35D7F4DFB5FF9BB8FE1DF1C" box="[1199,1213,1557,1580]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="48" pageNumber="349">
|
||
<emphasis id="B9480707A35D7F4DFB5FF9BB8FE1DF1C" box="[1199,1213,1557,1580]" italics="true" pageId="48" pageNumber="349">5</emphasis>
|
||
</figureCitation>
|
||
) in East
|
||
<collectingCountry id="F32B9B85A35D7F4DFAD1F9BA8E34DF1D" box="[1313,1384,1556,1581]" name="Nepal" pageId="48" pageNumber="349">Nepal</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA35D7F4CFF37F9998F0AD8E0" lastPageId="49" lastPageNumber="350" pageId="48" pageNumber="349" type="discussion">
|
||
<paragraph id="8B83DB15A35D7F4DFF37F9998EC7DEEC" blockId="48.[151,1437,151,2013]" pageId="48" pageNumber="349">
|
||
<emphasis id="B9480707A35D7F4DFF37F9998A67DF60" bold="true" box="[199,315,1591,1616]" pageId="48" pageNumber="349">Remarks.</emphasis>
|
||
The description of
|
||
<taxonomicName id="4C3CA096A35D7F4DFDD5F999883BDF60" authorityName="Andrewes" authorityYear="1930" box="[549,871,1591,1616]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFDD5F99989E3DF60" box="[549,703,1591,1616]" italics="true" pageId="48" pageNumber="349">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFD23F99789B8DF60" box="[723,740,1593,1616]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFD0FF999883BDF60" box="[767,871,1591,1616]" italics="true" pageId="48" pageNumber="349">hingstoni</emphasis>
|
||
</taxonomicName>
|
||
is based on five specimens collected during the Third British Mount Everest Expedition, 1924: four specimens in
|
||
<collectingRegion id="49F815F7A35D7F4DFC87F9F38F5FDF45" box="[887,1027,1628,1653]" country="India" name="West Bengal" pageId="48" pageNumber="349">West Bengal</collectingRegion>
|
||
(Darjeeling) and one in East
|
||
<collectingRegion id="49F815F7A35D7F4DFABBF9F28EC0DF45" box="[1355,1436,1628,1653]" country="India" name="Sikkim" pageId="48" pageNumber="349">Sikkim</collectingRegion>
|
||
(“Gangtok”). The male collected in Darjeeling during
|
||
<date id="FF82FDD5A35D7F4DFD08F92E88CEDFA8" box="[760,914,1664,1688]" pageId="48" pageNumber="349" value="1924-03-01" valueMax="1924-03-10" valueMin="1924-03-01">1–10.III.1924</date>
|
||
is here designated as the
|
||
<typeStatus id="548765B7A35D7F4DFB41F9D18E47DFA8" box="[1201,1307,1663,1688]" pageId="48" pageNumber="349" type="lectotype">lectotype</typeStatus>
|
||
. According to the map published in the paper together with the original description this specimen was most likely collected in the environments of Darjeeling town where the expeditional team rested between
|
||
<date id="FF82FDD5A35D7F4DFBDBF9668FA4DFD0" box="[1067,1272,1735,1760]" pageId="48" pageNumber="349" value="1924-03-07" valueMax="1924-03-25" valueMin="1924-03-07">7. and 25.III.1924</date>
|
||
.
|
||
<taxonomicName id="4C3CA096A35D7F4DFAF2F9698A74DE34" authorityName="N. Ito" authorityYear="1996" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="curvatus" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFAF2F9698EC0DFD0" box="[1282,1436,1735,1760]" italics="true" pageId="48" pageNumber="349">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFF50F9438BEDDE34" box="[160,177,1773,1796]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFF38F9438A74DE34" box="[200,296,1773,1796]" italics="true" pageId="48" pageNumber="349">curvatus</emphasis>
|
||
</taxonomicName>
|
||
was described from two males, including the
|
||
<typeStatus id="548765B7A35D7F4DFCD8F94288D1DE35" box="[808,909,1772,1797]" pageId="48" pageNumber="349" type="holotype">holotype</typeStatus>
|
||
, from east Thieme (Darjeeling) and one female from forest S. Manisingma [= Mansingma], Khandbari, located in the middle Arun Valley, east of Num and Mure, East
|
||
<collectingCountry id="F32B9B85A35D7F4DFF24F89A8A46DE7D" box="[212,282,1844,1869]" name="Nepal" pageId="48" pageNumber="349">Nepal</collectingCountry>
|
||
, without comparison with
|
||
<taxonomicName id="4C3CA096A35D7F4DFDA9F89B8845DE7D" authorityName="Andrewes" authorityYear="1930" box="[601,793,1844,1869]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFDA9F89B8931DE7C" box="[601,621,1845,1868]" italics="true" pageId="48" pageNumber="349">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFD73F89B89C8DE7C" box="[643,660,1845,1868]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFD41F89A8845DE7D" box="[689,793,1844,1869]" italics="true" pageId="48" pageNumber="349">hingstoni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFADA6E4A35D7F4DFCDDF89B88FFDE7D" author="Ito" box="[813,931,1844,1869]" pageId="48" pageNumber="349" refString="Ito, N. (1996) Some species of the genus Trichotichnus (Coleoptera, Carabidae, Harpalini) from Asia. Elytra, 24 (2), 203 - 211." type="journal article" year="1996">Ito (1996)</bibRefCitation>
|
||
considered the label data “O. Thieme” as information about the locality within Darjeeling, but in fact the
|
||
<typeStatus id="548765B7A35D7F4DFC70F8F688ECDE40" box="[896,944,1880,1904]" pageId="48" pageNumber="349">type</typeStatus>
|
||
locality should be Darjeeling because “O. Thieme” means the name of the German entomologist Otto Thieme (
|
||
<date id="FF82FDD5A35D7F4DFC6FF8D28F7EDEA5" box="[927,1058,1916,1941]" pageId="48" pageNumber="349" value="1836" valueMax="1907">1836–1907</date>
|
||
), the collection of which is partly deposited in MFNB. The comparison of the
|
||
<typeStatus id="548765B7A35D7F4DFD45F80F89ACDE89" box="[693,752,1953,1977]" pageId="48" pageNumber="349">types</typeStatus>
|
||
of these two species and the additional specimens from Darjeeling,
|
||
<collectingRegion id="49F815F7A35D7F4DFEECF86A8A31DEED" box="[284,365,1988,2013]" country="India" name="Sikkim" pageId="48" pageNumber="349">Sikkim</collectingRegion>
|
||
and East
|
||
<collectingCountry id="F32B9B85A35D7F4DFE2FF86A897EDEED" box="[479,546,1988,2013]" name="Nepal" pageId="48" pageNumber="349">Nepal</collectingCountry>
|
||
revealed that
|
||
<taxonomicName id="4C3CA096A35D7F4DFD33F86B882FDEEC" authorityName="N. Ito" authorityYear="1996" box="[707,883,1988,2012]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="349" phylum="Arthropoda" rank="species" species="curvatus" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A35D7F4DFD33F86B898BDEEC" box="[707,727,1989,2012]" italics="true" pageId="48" pageNumber="349">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFD19F86B89A6DEEC" box="[745,762,1989,2012]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A35D7F4DFCE3F86B882FDEEC" box="[787,883,1989,2012]" italics="true" pageId="48" pageNumber="349">curvatus</emphasis>
|
||
</taxonomicName>
|
||
should be treated as a junior synonym of
|
||
<emphasis id="B9480707A35D7F4DFAA4F86B8E35DEEC" box="[1364,1385,1989,2012]" italics="true" pageId="48" pageNumber="349">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A35D7F4DFA8AF86B8ED7DEEC" box="[1402,1419,1989,2012]" italics="true" pageId="48" pageNumber="349">B</emphasis>
|
||
.)
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A35C7F4CFF67FF398F0AD8E0" blockId="49.[151,1436,151,464]" pageId="49" pageNumber="350">
|
||
<emphasis id="B9480707A35C7F4CFF67FF398A5BD980" box="[151,263,151,176]" italics="true" pageId="49" pageNumber="350">
|
||
<taxonomicName id="4C3CA096A35C7F4CFF67FF398A5FD980" authorityName="Andrewes" authorityYear="1930" box="[151,259,151,176]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="350" phylum="Arthropoda" rank="species" species="hingstoni">hingstoni</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The specimens from various localities are very similar to each other in their morphology, including the configuration of the median lobe of the aedeagus in the males, but demonstrate geographical variation in the elytral length: the elytra in the examined specimens from East and
|
||
<collectingRegion id="49F815F7A35C7F4CFCC8FF7188D7D9C8" box="[824,907,223,248]" country="Sri Lanka" name="Central" pageId="49" pageNumber="350">Central</collectingRegion>
|
||
<collectingCountry id="F32B9B85A35C7F4CFC61FF718888D9C8" box="[913,980,223,248]" name="Nepal" pageId="49" pageNumber="350">Nepal</collectingCountry>
|
||
are relatively longer (in male, EL/EW = 1.44–1.49; EL/PL = 2.60–2.69; EW/PWmax = 1.25–1.29; in female, these indices 1.46–1.47, 2.62–2.70, and 1.25– 27, respectively) than in the specimens from Darjeeling and
|
||
<collectingRegion id="49F815F7A35C7F4CFCCEFE8988D3D870" box="[830,911,295,320]" country="India" name="Sikkim" pageId="49" pageNumber="350">Sikkim</collectingRegion>
|
||
(in male, EL/EW = 1.35–1.42; EL/PL = 2.33– 2.55; EW/PWmax = 1.23–1.28; in female, these indices 1.45, 2.42, and 1.20, respectively). In addition, the pronotum in the specimens from
|
||
<collectingCountry id="F32B9B85A35C7F4CFDFFFEC1890ED8B8" box="[527,594,367,392]" name="Nepal" pageId="49" pageNumber="350">Nepal</collectingCountry>
|
||
is usually with sides straighter in the basal half and with basal angles less obtuse, especially so in the examined male from Ting Sang La. Because so few specimens are available from few localities, the status of the populations from
|
||
<collectingCountry id="F32B9B85A35C7F4CFD78FE198997D8E0" box="[648,715,439,464]" name="Nepal" pageId="49" pageNumber="350">Nepal</collectingCountry>
|
||
is obscure and needs further study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |