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<document id="C1C970281609577E4D992F3F7B3A20F3" ID-DOI="10.11646/zootaxa.3990.2.2" ID-GBIF-Dataset="1740ec81-c55e-44ce-9a6d-cd62acd5d898" ID-ISSN="1175-5326" ID-Zenodo-Dep="242947" ID-ZooBank="BDF1BC11-4848-4858-99DC-0336A5FBEE2B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461267666231" checkinUser="plazi" docAuthor="Helwerda, Renate A." docDate="2015" docId="03FA878C951EFE45FF26E0BD0738CBE8" docLanguage="en" docName="zt03990p220.pdf" docOrigin="Zootaxa 3990 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ringicula circumscripta" docType="treatment" docVersion="9" lastPageNumber="212" masterDocId="FFC3FFF49510FE4AFFB1E574036DCA1D" masterDocTitle="Acteonidae, Bullinidae and Ringiculidae (Gastropoda: Heterobranchia) from the Plio-Pleistocene of the Philippines" masterLastPageNumber="220" masterPageNumber="197" pageNumber="211" updateTime="1698610896101" updateUser="plazi">
<mods:mods id="78A1913281B9AFB77194FAA5D8185C8E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="52F51E463037B16F485C27B3ACD2C3B2">
<mods:title id="86BC58D907C55B9238D159CAFABC2ADE">Acteonidae, Bullinidae and Ringiculidae (Gastropoda: Heterobranchia) from the Plio-Pleistocene of the Philippines</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="B3EF7687EB1F352210CF774589F67B54">Author</mods:roleTerm>
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<mods:namePart id="EE299636995C7FA009548F598663835E">Helwerda, Renate A.</mods:namePart>
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<mods:typeOfResource id="0E1B1C391D1E46D89C3A61A07F4DD061">text</mods:typeOfResource>
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<mods:title id="7EC812F0D7D158BFC19BD497C0288345">Zootaxa</mods:title>
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<mods:part id="0B049F06DA236C5E5A51B4244C54232E">
<mods:date id="2AE3981D7511375670C02B25E87112EB">2015</mods:date>
<mods:detail id="36B781F8A6D1790E59C36BFB58701802" type="volume">
<mods:number id="0AE04926CCBB93CF6C1F5DFA9EABC21A">3990</mods:number>
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<mods:classification id="C637379DE815C01DB2DB45BC775853C7">journal article</mods:classification>
<mods:identifier id="51FCBA5A25A4891EA6A49B7C6D750A9B" type="DOI">10.11646/zootaxa.3990.2.2</mods:identifier>
<mods:identifier id="278052A73B50B8EC4C8E36CDD08787BC" type="GBIF-Dataset">1740ec81-c55e-44ce-9a6d-cd62acd5d898</mods:identifier>
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<treatment id="03FA878C951EFE45FF26E0BD0738CBE8" ID-DOI="http://doi.org/10.5281/zenodo.6116537" ID-GBIF-Taxon="119638252" ID-Zenodo-Dep="6116537" LSID="urn:lsid:plazi:treatment:03FA878C951EFE45FF26E0BD0738CBE8" httpUri="http://treatment.plazi.org/id/03FA878C951EFE45FF26E0BD0738CBE8" lastPageId="15" lastPageNumber="212" pageId="14" pageNumber="211">
<subSubSection id="C3496511951EFE44FF26E0BD0278CC19" pageId="14" pageNumber="211" type="nomenclature">
<paragraph id="8BEC369A951EFE44FF26E0BD0154CFFE" blockId="14.[151,569,1481,1541]" box="[151,569,1481,1507]" pageId="14" pageNumber="211">
<heading id="D0A481F6951EFE44FF26E0BD0154CFFE" bold="true" box="[151,569,1481,1507]" fontSize="11" level="1" pageId="14" pageNumber="211" reason="1">
<emphasis id="B927EA88951EFE44FF26E0BD0154CFFE" bold="true" box="[151,569,1481,1507]" pageId="14" pageNumber="211">
<taxonomicName id="4C534D19951EFE44FF26E0BD02D6CFFE" ID-CoL="798R5" box="[151,443,1481,1507]" class="Gastropoda" family="Ringiculidae" genus="Ringicula" higherTaxonomySource="CoL" kingdom="Animalia" order="Ellobiida" pageId="14" pageNumber="211" phylum="Mollusca" rank="species" species="circumscripta">
<emphasis id="B927EA88951EFE44FF26E0BD02D6CFFE" bold="true" box="[151,443,1481,1507]" italics="true" pageId="14" pageNumber="211">Ringicula circumscripta</emphasis>
</taxonomicName>
nov. spec.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEC369A951EFE44FF26E0980278CC19" blockId="14.[151,569,1481,1541]" box="[151,277,1516,1541]" pageId="14" pageNumber="211">
(
<figureCitation id="13682A1F951EFE44FF2FE0980260CC19" box="[158,269,1516,1541]" captionStart="FIGURE 23 – 28. 23" captionStartId="13.[151,250,1781,1803]" captionTargetBox="[239,1346,193,1758]" captionTargetId="figure@13.[239,1347,193,1760]" captionTargetPageId="13" captionText="FIGURE 23 – 28. 23. Ringicula bella nov. spec .. Holotype RGM 961.797. Locality Anda 1. (a) rear view, (b) oblique apertural view, (c) apertural view, (d) basal view, (e) apical view. 24. Ringicula bella nov. spec .. Paratype RGM 961.750. Locality Tiep 2. (a) apical view, (b) apertural view, composite. 25. Ringicula spec 1. RGM 961.817. Locality Tiep 2. (a) rear view, (b) oblique apertural view, (c) apertural view, (d) basal view, (e) apical view. 26. Ringicula cf. spec 1. RGM 961.753. Locality Tiep 2. (a) rear view, (b) oblique apertural view, (c) apertural view, (d) basal view, (e) apical view. 27. Ringicula circumscripta nov. spec .. Holotype RGM 961.754. Locality Roxas. (a) rear view, (b) oblique apertural view, (c) apertural view, (d) basal view, (e) apical view. 28. Ringicula spec 2. RGM 961.751. Locality Tiep 2. (a) rear view, (b) oblique apertural view, (c) apertural view, (d) basal view, (e) apical view." httpUri="https://zenodo.org/record/242952/files/figure.png" pageId="14" pageNumber="211">Figure 27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3496511951EFE44FF26E34002D7CCAA" pageId="14" pageNumber="211" type="materials_examined">
<paragraph id="8BEC369A951EFE44FF26E3400785CC50" blockId="14.[151,1436,1588,2009]" box="[151,1256,1588,1613]" pageId="14" pageNumber="211">
<emphasis id="B927EA88951EFE44FF26E3400228CC50" bold="true" box="[151,325,1588,1613]" pageId="14" pageNumber="211">
<typeStatus id="54E88838951EFE44FF26E34003BFCC50" box="[151,210,1588,1613]" pageId="14" pageNumber="211">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54E88838951EFE44FEFDE34002D9CC50" box="[332,436,1588,1613]" pageId="14" pageNumber="211" type="holotype">Holotype</typeStatus>
<collectionCode id="ED42AE5F951EFE44FE0AE3400296CC51" box="[443,507,1588,1612]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="211">RGM</collectionCode>
961.754 (
<typeStatus id="54E88838951EFE44FDDAE34001C9CC51" box="[619,676,1588,1612]" pageId="14" pageNumber="211">Type</typeStatus>
locality), 3
<typeStatus id="54E88838951EFE44FC9DE34100F5CC50" box="[812,920,1589,1613]" pageId="14" pageNumber="211" type="paratype">paratypes</typeStatus>
<collectionCode id="ED42AE5F951EFE44FC2FE34000B3CC51" box="[926,990,1588,1612]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="211">RGM</collectionCode>
961.854 (AndaDeVos).
</paragraph>
<paragraph id="8BEC369A951EFE44FF76E32302D8CC88" blockId="14.[151,1436,1588,2009]" pageId="14" pageNumber="211">
<emphasis id="B927EA88951EFE44FF76E32302E9CC6D" bold="true" box="[199,388,1623,1648]" pageId="14" pageNumber="211">Other material.</emphasis>
Anda6 (17)
<collectionCode id="ED42AE5F951EFE44FDA9E32C0135CC6D" box="[536,600,1624,1648]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="211">RGM</collectionCode>
961.818, AndaDeVos (25)
<collectionCode id="ED42AE5F951EFE44FC25E32C00B9CC6D" box="[916,980,1624,1648]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="211">RGM</collectionCode>
961.819, AndaClif3 (2)
<collectionCode id="ED42AE5F951EFE44FB40E32C065CCC6D" box="[1265,1329,1624,1648]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="211">RGM</collectionCode>
961.820, Roxas (3)
<collectionCode id="ED42AE5F951EFE44FEBDE3080221CC89" box="[268,332,1660,1684]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="211">RGM</collectionCode>
961.821.
</paragraph>
<paragraph id="8BEC369A951EFE44FF76E3EB02D7CCAA" blockId="14.[151,1436,1588,2009]" box="[199,442,1695,1720]" pageId="14" pageNumber="211">
<emphasis id="B927EA88951EFE44FF76E3EB0208CCA5" bold="true" box="[199,357,1695,1720]" pageId="14" pageNumber="211">
<typeStatus id="54E88838951EFE44FF76E3EB026FCCA5" box="[199,258,1695,1720]" pageId="14" pageNumber="211">Type</typeStatus>
locality.
</emphasis>
Roxas.
</paragraph>
</subSubSection>
<subSubSection id="C3496511951EFE44FF76E3B00249CCE2" pageId="14" pageNumber="211" type="etymology">
<paragraph id="8BEC369A951EFE44FF76E3B00249CCE2" blockId="14.[151,1436,1588,2009]" pageId="14" pageNumber="211">
<emphasis id="B927EA88951EFE44FF76E3B002CBCCC0" bold="true" box="[199,422,1732,1757]" pageId="14" pageNumber="211">Derivatio nominis.</emphasis>
Enclosed, adjective. Named after the prominent ridge that delimits the parietal callus of the aperture.
</paragraph>
</subSubSection>
<subSubSection id="C3496511951EFE44FF76E2780645CD70" pageId="14" pageNumber="211" type="diagnosis">
<paragraph id="8BEC369A951EFE44FF76E2780645CD70" blockId="14.[151,1436,1588,2009]" pageId="14" pageNumber="211">
<emphasis id="B927EA88951EFE44FF76E278022CCD38" bold="true" box="[199,321,1804,1829]" pageId="14" pageNumber="211">Diagnosis.</emphasis>
Shell moderately large; H
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,
<geoCoordinate id="EE67505D951EFE44FD54E2790055CD38" box="[741,824,1804,1829]" direction="west" orientation="longitude" pageId="14" pageNumber="211" precision="555" value="-3.64">
W
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mm; protoconch rather elevated, 1.1 whorl,
<quantity id="4CAB9B7F951EFE44FA81E27806F5CD39" box="[1328,1432,1804,1828]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="14" pageNumber="211" unit="mm" value="0.24">0.24 mm</quantity>
; nucleus small, DN
<quantity id="4CAB9B7F951EFE44FEC9E244028FCD5A" box="[376,482,1840,1864]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="211" unit="mm" value="0.13">0.13 mm</quantity>
; teleoconch with spiral grooves; outer lip thick and crenulated; inner lip callused, delineated by lamella, additional lamella extending into aperture; columella callused and with two folds.
</paragraph>
</subSubSection>
<subSubSection id="C3496511951EFE45FF76E2030738CBE8" lastPageId="15" lastPageNumber="212" pageId="14" pageNumber="211" type="description">
<paragraph id="8BEC369A951EFE45FF76E2030226CB78" blockId="14.[151,1436,1588,2009]" lastBlockId="15.[151,1437,151,501]" lastPageId="15" lastPageNumber="212" pageId="14" pageNumber="211">
<emphasis id="B927EA88951EFE44FF76E2030234CD8D" bold="true" box="[199,345,1911,1936]" pageId="14" pageNumber="211">Description.</emphasis>
Shell rather high-spired, moderately large for its genus. Largest specimen has a total of 6.3 whorls, including a protoconch of 1.1 whorl. Protoconch small, smooth, rather inclined. Teleoconch sculpted with thin spiral grooves and very weak, prosocyrt growth lines. Spiral grooves are regularly spaced, but absent from the upper one-sixth part of the whorl, numbering 14 on the body whorl. Whorls convex, but not globose. Suture obscured by a narrow subsutural ridge, below which is a narrow groove. Aperture covered by thick callus, delineated by a prominent ridge on the inner lip. An additional lamella is present on the parietal callus, extending into the aperture. Columellar lip bears two pronounced, smooth folds. Outer lip thick, crenulated, with a distinct nodule present on the middle of the lip. Exterior of the outer lip bears weak striations. Anterior and posterior sinus well-developed.
</paragraph>
<paragraph id="8BEC369A951FFE45FF76E41B0738CBE8" blockId="15.[151,1437,151,501]" pageId="15" pageNumber="212">
<emphasis id="B927EA88951FFE45FF76E41B0212CB95" bold="true" box="[199,383,367,392]" pageId="15" pageNumber="212">Differentiation.</emphasis>
<taxonomicName id="4C534D19951FFE45FE39E40401F7CB9A" box="[392,666,368,392]" class="Gastropoda" family="Ringiculidae" genus="Ringicula" higherTaxonomySource="CoL" kingdom="Animalia" order="Ellobiida" pageId="15" pageNumber="212" phylum="Mollusca" rank="species" species="circumscripta">
<emphasis id="B927EA88951FFE45FE39E40401F7CB9A" box="[392,666,368,392]" italics="true" pageId="15" pageNumber="212">Ringicula circumscripta</emphasis>
</taxonomicName>
nov. spec. is similar to
<taxonomicName id="4C534D19951FFE45FC03E404070DCB95" box="[946,1120,368,392]" class="Gastropoda" family="Ringiculidae" genus="Ringicula" higherTaxonomySource="CoL" kingdom="Animalia" order="Ellobiida" pageId="15" pageNumber="212" phylum="Mollusca" rank="species" species="bella">
<emphasis id="B927EA88951FFE45FC03E404070DCB95" box="[946,1120,368,392]" italics="true" pageId="15" pageNumber="212">Ringicula bella</emphasis>
</taxonomicName>
nov. spec., but the latter is slightly larger, it has more spiral grooves and it has two or three small nodules on the parietal callus instead of a lamella.
<taxonomicName id="4C534D19951FFE45FF46E4CC01EECBCD" authority="Schepman, 1913" authorityName="Schepman" authorityYear="1913" box="[247,643,439,464]" class="Gastropoda" family="Ringiculidae" genus="Ringicula" higherTaxonomySource="CoL" kingdom="Animalia" order="Ellobiida" pageId="15" pageNumber="212" phylum="Mollusca" rank="species" species="titanica">
<emphasis id="B927EA88951FFE45FF46E4CC02AFCBD2" box="[247,450,440,464]" italics="true" pageId="15" pageNumber="212">Ringicula titanica</emphasis>
<bibRefCitation id="EFC24B6B951FFE45FE79E4C301EECBCD" author="Schepman" box="[456,643,439,464]" pageId="15" pageNumber="212" refString="Schepman, M. M. (1913) The Prosobranchia, Pulmonata and Opisthobranchia Tectibranchiata (Tribe Bullomorpha) of the Siboga Expedition, Part VI, Pulmonata and Opisthobranchia Tectibranchiata (Tribe Bullomorpha). In: Weber, M. (Ed.), Uitkomsten op Zoologisch, Botanisch, Oceanographisch en Geologisch Gebied, verzameld in Nederlands Oost-Indie 1899 - 1900 aan boord H. M. Siboga onder commando van Luitenant ter zee 1 e kl. G. F. Tydeman. Monographie 49. Brill, Leiden, pp. 53 - 494." type="journal article" year="1913">Schepman, 1913</bibRefCitation>
</taxonomicName>
has a similar aperture with a lamella, but it is much larger (H
<quantity id="4CAB9B7F951FFE45FA8EE4C306F5CBD2" box="[1343,1432,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="15" pageNumber="212" unit="mm" value="8.5">8.5 mm</quantity>
, W
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) and it has fewer spiral grooves, present only on the lower half of the whorl.
</paragraph>
</subSubSection>
</treatment>
</document>
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