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<document id="57238C2EC910376C90712C9F748C6FB1" ID-DOI="10.11646/zootaxa.3990.2.2" ID-GBIF-Dataset="1740ec81-c55e-44ce-9a6d-cd62acd5d898" ID-ISSN="1175-5326" ID-Zenodo-Dep="242947" ID-ZooBank="BDF1BC11-4848-4858-99DC-0336A5FBEE2B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461267666231" checkinUser="plazi" docAuthor="Helwerda, Renate A." docDate="2015" docId="03FA878C951BFE46FF26E05D0135CB5D" docLanguage="en" docName="zt03990p220.pdf" docOrigin="Zootaxa 3990 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bullina virgoides" docType="treatment" docVersion="9" lastPageNumber="208" masterDocId="FFC3FFF49510FE4AFFB1E574036DCA1D" masterDocTitle="Acteonidae, Bullinidae and Ringiculidae (Gastropoda: Heterobranchia) from the Plio-Pleistocene of the Philippines" masterLastPageNumber="220" masterPageNumber="197" pageNumber="208" updateTime="1698610896101" updateUser="plazi">
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<mods:title id="64332A0E40802BD909F80A31B0B912D1">Acteonidae, Bullinidae and Ringiculidae (Gastropoda: Heterobranchia) from the Plio-Pleistocene of the Philippines</mods:title>
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<mods:namePart id="C2DC5C909741CD0E27FE169063C17D83">Helwerda, Renate A.</mods:namePart>
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<mods:date id="358616D45E8DFD572DF31AF42EB8EF2A">2015</mods:date>
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<treatment id="03FA878C951BFE46FF26E05D0135CB5D" ID-DOI="http://doi.org/10.5281/zenodo.6116527" ID-GBIF-Taxon="119638213" ID-Zenodo-Dep="6116527" LSID="urn:lsid:plazi:treatment:03FA878C951BFE46FF26E05D0135CB5D" httpUri="http://treatment.plazi.org/id/03FA878C951BFE46FF26E05D0135CB5D" lastPageId="12" lastPageNumber="208" pageId="11" pageNumber="208">
<subSubSection id="C3496511951BFE41FF26E05D0227CF79" pageId="11" pageNumber="208" type="nomenclature">
<paragraph id="8BEC369A951BFE41FF26E05D0288CF5E" blockId="11.[151,485,1321,1381]" box="[151,485,1321,1347]" pageId="11" pageNumber="208">
<heading id="D0A481F6951BFE41FF26E05D0288CF5E" bold="true" box="[151,485,1321,1347]" fontSize="11" level="1" pageId="11" pageNumber="208" reason="1">
<emphasis id="B927EA88951BFE41FF26E05D0288CF5E" bold="true" box="[151,485,1321,1347]" pageId="11" pageNumber="208">
<taxonomicName id="4C534D19951BFE41FF26E05D020BCF5E" box="[151,358,1321,1347]" class="Gastropoda" family="Bullinidae" genus="Bullina" higherTaxonomySource="CoL" kingdom="Animalia" order="Cephalaspidea" pageId="11" pageNumber="208" phylum="Mollusca" rank="species" species="virgoides">
<emphasis id="B927EA88951BFE41FF26E05D020BCF5E" bold="true" box="[151,358,1321,1347]" italics="true" pageId="11" pageNumber="208">Bullina virgoides</emphasis>
</taxonomicName>
nov. spec.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEC369A951BFE41FF26E0380227CF79" blockId="11.[151,485,1321,1381]" box="[151,330,1356,1381]" pageId="11" pageNumber="208">
(
<figureCitation id="13682A1F951BFE41FF2FE038022FCF79" box="[158,322,1356,1381]" captionStart="FIGURE 19 22. 19" captionStartId="10.[151,250,1886,1908]" captionTargetBox="[214,1373,193,1864]" captionTargetId="figure@10.[214,1373,193,1865]" captionTargetPageId="10" captionText="FIGURE 19 22. 19. Crenilabium pacificus (Kuroda &amp; Habe in Habe, 1961). RGM 961.747. Locality Tiep 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 20. Obrussena bracteata (Iredale, 1925). RGM 961.748. Locality Anda 6. (a) rear view, (b) apertural view, composite, (c) basal view, (d) apical view. 21. Bullina virgoides nov. spec .. Holotype RGM 961.749. Locality Tiep 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 22. Bullina virgoides nov. spec .. Paratype RGM 961.846. Locality Tiep 2. (a) rear view, (b) apertural view." httpUri="https://zenodo.org/record/242951/files/figure.png" pageId="11" pageNumber="208">Figures 2122</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3496511951BFE41FF26E0E002DECFCD" pageId="11" pageNumber="208" type="materials_examined">
<paragraph id="8BEC369A951BFE41FF26E0E0060ACFB0" blockId="11.[151,1437,1428,2029]" box="[151,1383,1428,1453]" pageId="11" pageNumber="208">
<emphasis id="B927EA88951BFE41FF26E0E00228CFB0" bold="true" box="[151,325,1428,1453]" pageId="11" pageNumber="208">
<typeStatus id="54E88838951BFE41FF26E0E003BFCFB0" box="[151,210,1428,1453]" pageId="11" pageNumber="208">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54E88838951BFE41FEFDE0E002D9CFB0" box="[332,436,1428,1453]" pageId="11" pageNumber="208" type="holotype">Holotype</typeStatus>
<collectionCode id="ED42AE5F951BFE41FE0AE0E00296CFB1" box="[443,507,1428,1452]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="208">RGM</collectionCode>
961.749 (
<typeStatus id="54E88838951BFE41FDDAE0E001C9CFB1" box="[619,676,1428,1452]" pageId="11" pageNumber="208">Type</typeStatus>
locality).
<typeStatus id="54E88838951BFE41FCA6E0E10014CFB0" box="[791,889,1429,1453]" pageId="11" pageNumber="208" type="paratype">Paratype</typeStatus>
<collectionCode id="ED42AE5F951BFE41FC31E0E000ADCFB1" box="[896,960,1428,1452]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="208">RGM</collectionCode>
961.846 and 961.738 (
<typeStatus id="54E88838951BFE41FB70E0E00796CFB1" box="[1217,1275,1428,1452]" pageId="11" pageNumber="208">Type</typeStatus>
locality).
</paragraph>
<paragraph id="8BEC369A951BFE41FF76E0C302DECFCD" blockId="11.[151,1437,1428,2029]" box="[199,435,1463,1488]" pageId="11" pageNumber="208">
<emphasis id="B927EA88951BFE41FF76E0C30208CFCD" bold="true" box="[199,357,1463,1488]" pageId="11" pageNumber="208">
<typeStatus id="54E88838951BFE41FF76E0C3026FCFCD" box="[199,258,1463,1488]" pageId="11" pageNumber="208">Type</typeStatus>
locality.
</emphasis>
Tiep2.
</paragraph>
</subSubSection>
<subSubSection id="C3496511951BFE41FF76E0A8064DCFE9" box="[199,1312,1500,1525]" pageId="11" pageNumber="208" type="etymology">
<paragraph id="8BEC369A951BFE41FF76E0A8064DCFE9" blockId="11.[151,1437,1428,2029]" box="[199,1312,1500,1525]" pageId="11" pageNumber="208">
<emphasis id="B927EA88951BFE41FF76E0A802C9CFE8" bold="true" box="[199,420,1500,1525]" pageId="11" pageNumber="208">Derivatio nominis.</emphasis>
Named for its resemblance to
<taxonomicName id="4C534D19951BFE41FD4AE0A80777CFE8" authority="Habe, 1950" authorityName="Habe" authorityYear="1950" box="[763,1050,1500,1525]" class="Gastropoda" family="Bullinidae" genus="Bullina" higherTaxonomySource="CoL" kingdom="Animalia" order="Cephalaspidea" pageId="11" pageNumber="208" phylum="Mollusca" rank="species" species="virgo">
<emphasis id="B927EA88951BFE41FD4AE0A800E2CFE9" box="[763,911,1500,1524]" italics="true" pageId="11" pageNumber="208">Bullina virgo</emphasis>
Habe, 1950
</taxonomicName>
, by adding suffix -
<emphasis id="B927EA88951BFE41FB5DE0A80674CFE9" box="[1260,1305,1500,1524]" italics="true" pageId="11" pageNumber="208">ides</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C3496511951BFE41FF76E08B0231CC98" pageId="11" pageNumber="208" type="diagnosis">
<paragraph id="8BEC369A951BFE41FF76E08B0231CC98" blockId="11.[151,1437,1428,2029]" pageId="11" pageNumber="208">
<emphasis id="B927EA88951BFE41FF76E08B022CCC05" bold="true" box="[199,321,1535,1560]" pageId="11" pageNumber="208">Diagnosis.</emphasis>
Shell elongate; H&gt;
<quantity id="4CAB9B7F951BFE41FD9FE08B01F9CC0A" box="[558,660,1535,1560]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" pageId="11" pageNumber="208" unit="mm" value="3.75">3.75 mm</quantity>
,
<geoCoordinate id="EE67505D951BFE41FD10E3740198CC05" box="[673,757,1536,1560]" direction="west" orientation="longitude" pageId="11" pageNumber="208" precision="555" value="-1.92">
W
<quantity id="4CAB9B7F951BFE41FD75E3740198CC05" box="[708,757,1536,1560]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" pageId="11" pageNumber="208" unit="mm" value="1.92">1.92</quantity>
</geoCoordinate>
mm; protoconch elevated, 1.2 whorl, DP
<quantity id="4CAB9B7F951BFE41FB63E08B0657CC0A" box="[1234,1338,1535,1560]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="11" pageNumber="208" unit="mm" value="0.54">0.54 mm</quantity>
; nucleus partially overhanging first teleoconch whorl, DN
<quantity id="4CAB9B7F951BFE41FD63E3500051CC21" box="[722,828,1572,1596]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="208" unit="mm" value="0.22">0.22 mm</quantity>
; teleoconch sculpted with punctuated spiral grooves, colour whitish with two pink bands, one above centre of whorl, one on base; columella callused, bearing a fold; umbilicus closed.
</paragraph>
</subSubSection>
<subSubSection id="C3496511951BFE46FF76E3FB0135CB5D" lastPageId="12" lastPageNumber="209" pageId="11" pageNumber="208" type="description">
<paragraph id="8BEC369A951BFE41FF76E3FB0199CDB8" blockId="11.[151,1437,1428,2029]" pageId="11" pageNumber="208">
<emphasis id="B927EA88951BFE41FF76E3FB0234CCB5" bold="true" box="[199,345,1679,1704]" pageId="11" pageNumber="208">Description.</emphasis>
Shell tall, pear-shaped, fragile, with convex sides. Largest specimen with intact spire has a total of 3.6 whorls, including a protoconch of 1.2 whorl. Protoconch smooth, globose, inclined, with nucleus partially overhanging first teleoconch whorl, margin impressed. Teleoconch strongly sculpted with punctuated spiral grooves, with oval punctuations on the middle of the whorl and rectangular punctuations on the base and near the suture. Suture channeled. Colour dirty white with two rather broad pinkish spiral bands, one above the middle of the whorl and one on the base. Aperture long, semilunar, narrowing towards the elongated anterior end. Upper part of columella twisted, with a fold, rest of columella straight, lowermost part broken in all specimens. Columella and basal part of shell next to columella bear a thin callus.
</paragraph>
<paragraph id="8BEC369A951BFE46FF76E2C40135CB5D" blockId="11.[151,1437,1428,2029]" lastBlockId="12.[151,1436,151,320]" lastPageId="12" lastPageNumber="209" pageId="11" pageNumber="208">
<emphasis id="B927EA88951BFE41FF76E2C40212CDD4" bold="true" box="[199,383,1968,1993]" pageId="11" pageNumber="208">Differentiation.</emphasis>
This species closely resembles
<taxonomicName id="4C534D19951BFE41FD49E2C40772CDD4" authority="Habe, 1950" authorityName="Habe" authorityYear="1950" box="[760,1055,1968,1993]" class="Gastropoda" family="Bullinidae" genus="Bullina" higherTaxonomySource="CoL" kingdom="Animalia" order="Cephalaspidea" pageId="11" pageNumber="208" phylum="Mollusca" rank="species" species="virgo">
<emphasis id="B927EA88951BFE41FD49E2C400E2CDD5" box="[760,911,1968,1992]" italics="true" pageId="11" pageNumber="208">Bullina virgo</emphasis>
Habe,
<quantity id="4CAB9B7F951BFE41FC56E2C40772CDD4" box="[999,1055,1968,1993]" metricMagnitude="1" metricUnit="m" metricValue="4.953" pageId="11" pageNumber="208" unit="in" value="1950.0">1950</quantity>
</taxonomicName>
in the distribution of the colour bands and the punctuated spiral grooves, but
<taxonomicName id="4C534D19951BFE41FD3FE2A10003CDF0" authority="Habe, 1950" authorityName="Habe" authorityYear="1950" box="[654,878,2004,2029]" class="Gastropoda" family="Bullinidae" genus="Bullina" higherTaxonomySource="CoL" kingdom="Animalia" order="Cephalaspidea" pageId="11" pageNumber="208" phylum="Mollusca" rank="species" species="virgo">
<emphasis id="B927EA88951BFE41FD3FE2A1018ACDF1" box="[654,743,2005,2028]" italics="true" pageId="11" pageNumber="208">B. virgo</emphasis>
Habe, 1950
</taxonomicName>
is much less elongate and does not have a fold on the upper part of the columella.
<taxonomicName id="4C534D19951CFE46FE4FE5ED01E9CAAD" box="[510,644,152,176]" class="Gastropoda" family="Bullinidae" genus="Bullina" higherTaxonomySource="CoL" kingdom="Animalia" order="Cephalaspidea" pageId="12" pageNumber="209" phylum="Mollusca" rank="species" species="virgoides">
<emphasis id="B927EA88951CFE46FE4FE5ED01E9CAAD" box="[510,644,152,176]" italics="true" pageId="12" pageNumber="209">B. virgoides</emphasis>
</taxonomicName>
is unusual in having a fold on the columella, which does not appear to be present in any other
<taxonomicName id="4C534D19951CFE46FE1EE5C8016FCAC9" box="[431,514,188,212]" class="Gastropoda" family="Bullinidae" genus="Bullina" higherTaxonomySource="CoL" kingdom="Animalia" order="Cephalaspidea" pageId="12" pageNumber="209" phylum="Mollusca" rank="genus">
<emphasis id="B927EA88951CFE46FE1EE5C8016FCAC9" box="[431,514,188,212]" italics="true" pageId="12" pageNumber="209">Bullina</emphasis>
</taxonomicName>
species known to date. It is also more elongate than any other
<taxonomicName id="4C534D19951CFE46FB5EE5C8062CCAC9" box="[1263,1345,188,212]" class="Gastropoda" family="Bullinidae" genus="Bullina" higherTaxonomySource="CoL" kingdom="Animalia" order="Cephalaspidea" pageId="12" pageNumber="209" phylum="Mollusca" rank="genus">
<emphasis id="B927EA88951CFE46FB5EE5C8062CCAC9" box="[1263,1345,188,212]" italics="true" pageId="12" pageNumber="209">Bullina</emphasis>
</taxonomicName>
, but its sculpture, colouration and general shape clearly place this species within
<taxonomicName id="4C534D19951CFE46FC58E5940756CAE5" box="[1001,1083,224,248]" class="Gastropoda" family="Bullinidae" genus="Bullina" higherTaxonomySource="CoL" kingdom="Animalia" order="Cephalaspidea" pageId="12" pageNumber="209" phylum="Mollusca" rank="genus">
<emphasis id="B927EA88951CFE46FC58E5940756CAE5" box="[1001,1083,224,248]" italics="true" pageId="12" pageNumber="209">Bullina</emphasis>
</taxonomicName>
.
<taxonomicName id="4C534D19951CFE46FBFDE59403A2CB00" authority="Habe, 1958" authorityName="Habe" authorityYear="1958" class="Gastropoda" family="Bullinidae" genus="Rictaxiella" kingdom="Animalia" pageId="12" pageNumber="209" phylum="Mollusca" rank="species" species="choshiensis">
<emphasis id="B927EA88951CFE46FBFDE594063DCAE5" box="[1100,1360,224,248]" italics="true" pageId="12" pageNumber="209">Rictaxiella choshiensis</emphasis>
<bibRefCitation id="EFC24B6B951CFE46FAEBE5AB03A2CB00" author="Habe" pageId="12" pageNumber="209" refString="Habe, T. (1958) On the Shell-bearing Opisthobranchiate Molluscan Fauna from Off Choshi, Chiba Pref., Japan. Annotationes Zoologicae Japonenses, 31, 117 - 120." type="journal article" year="1958">Habe, 1958</bibRefCitation>
</taxonomicName>
is similar to
<taxonomicName id="4C534D19951CFE46FED3E470014ECB01" box="[354,547,260,284]" class="Gastropoda" family="Bullinidae" genus="Bullina" higherTaxonomySource="CoL" kingdom="Animalia" order="Cephalaspidea" pageId="12" pageNumber="209" phylum="Mollusca" rank="species" species="virgoides">
<emphasis id="B927EA88951CFE46FED3E470014ECB01" box="[354,547,260,284]" italics="true" pageId="12" pageNumber="209">Bullina virgoides</emphasis>
</taxonomicName>
nov. spec. in its elongated shape and its sculpture, but it has no red colouration and no fold and callus on the columella.
</paragraph>
</subSubSection>
</treatment>
</document>