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<document ID-DOI="10.11646/phytotaxa.572.1.3" ID-GBIF-Dataset="535a1fdf-518a-4e9d-895f-96a3780511b3" ID-ISSN="1179-3163" ID-Zenodo-Dep="7305618" checkinTime="1667942941898" checkinUser="plazi" docAuthor="Averyanov, Leonid V., Thai, Tran Huy, Nguyen, Van Canh, Truong, Ba Vuong, Maisak, Tatiana V., Doan, Nga Thi &amp; Nguyen, Khang Sinh" docDate="2022" docId="03FA834FDD4A113F5C8CFA8AFE8C9D97" docLanguage="en" docName="phytotaxa.572.1.3.pdf" docOrigin="Phytotaxa 572 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.572.1.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.5:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="5" docTitle="Cleisostoma tatonii Aver. &amp; V. C. Nguyen 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="48" masterDocId="FFC3FB37DD4E113A5C04FFECFFFA981E" masterDocTitle="New orchids in the flora of Vietnam (Orchidaceae, Aeridinae)" masterLastPageNumber="60" masterPageNumber="43" pageNumber="47" updateTime="1668084151424" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New orchids in the flora of Vietnam (Orchidaceae, Aeridinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Averyanov, Leonid V.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8031-2925</mods:nameIdentifier>
<mods:affiliation>Komarov Botanical Institute of the Russian Academy of Sciences, Prof. Popov Str., 2, 197376 St. Petersburg, Russia. &amp; av _ leonid @ mail. ru; https: // orcid. org / 0000 - 0001 - 8031 - 2925</mods:affiliation>
<mods:nameIdentifier type="email">av_leonid@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Thai, Tran Huy</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-1636-9729</mods:nameIdentifier>
<mods:affiliation>Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoang Quoc Viet Road, Cau Giay, Hanoi, Vietnam. &amp; thaiiebr @ yahoo. com. vn; https: // orcid. org / 0000 - 0003 - 1636 - 9729</mods:affiliation>
<mods:nameIdentifier type="email">thaiiebr@yahoo.com.vn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nguyen, Van Canh</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-9578-0342</mods:nameIdentifier>
<mods:affiliation>Institute of Applied Technology, Thu Dau Mot University, No. 6, Tran Van On Street, Phu Hoa Ward, Thu Dau Mot City, Binh Duong Province, Vietnam. &amp; nguyenvancanh @ tdmu. edu. vn; https: // orcid. org / 0000 - 0001 - 9578 - 0342</mods:affiliation>
<mods:nameIdentifier type="email">nguyenvancanh@tdmu.edu.vn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Truong, Ba Vuong</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-3452-8455</mods:nameIdentifier>
<mods:affiliation>Institute of Applied Technology, Thu Dau Mot University, No. 6, Tran Van On Street, Phu Hoa Ward, Thu Dau Mot City, Binh Duong Province, Vietnam. &amp; Institute of Tropical Biology Vietnam Academy of Science and Technology, 85 Tran Quoc Toan, District 3, Ho Chi Minh City, Vietnam. Saint-Petersburg State Forest-Technical University, The Institute Lane 5, 195021, St. Petersburg, Russia. &amp; bavuong 2019 @ yahoo. com; https: // orcid. org / 0000 - 0003 - 3452 - 8455</mods:affiliation>
<mods:nameIdentifier type="email">bavuong2019@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Maisak, Tatiana V.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-5919-6755</mods:nameIdentifier>
<mods:affiliation>Komarov Botanical Institute of the Russian Academy of Sciences, Prof. Popov Str., 2, 197376 St. Petersburg, Russia. &amp; tmaisak @ mail. ru; https: // orcid. org / 0000 - 0001 - 5919 - 6755</mods:affiliation>
<mods:nameIdentifier type="email">tmaisak@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Doan, Nga Thi</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-5685-2953</mods:nameIdentifier>
<mods:affiliation>Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam. &amp; doanthinga 1994 hus @ gmail. com; https: // orcid. org / 0000 - 0002 - 5685 - 2953</mods:affiliation>
<mods:nameIdentifier type="email">doanthinga1994hus@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nguyen, Khang Sinh</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-5171-4140</mods:nameIdentifier>
<mods:affiliation>Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoang Quoc Viet Road, Cau Giay, Hanoi, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoang Quoc Viet Road, Cau Giay, Hanoi, Vietnam. &amp; nskhang @ gmail. com; https: // orcid. org / 0000 - 0001 - 5171 - 4140</mods:affiliation>
<mods:nameIdentifier type="email">nskhang@gmail.com</mods:nameIdentifier>
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<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>572</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>43</mods:start>
<mods:end>60</mods:end>
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<mods:url>http://dx.doi.org/10.11646/phytotaxa.572.1.3</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.572.1.3</mods:identifier>
<mods:identifier type="GBIF-Dataset">535a1fdf-518a-4e9d-895f-96a3780511b3</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7305618</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7305628" ID-Zenodo-Dep="7305628" LSID="urn:lsid:plazi:treatment:03FA834FDD4A113F5C8CFA8AFE8C9D97" httpUri="http://treatment.plazi.org/id/03FA834FDD4A113F5C8CFA8AFE8C9D97" lastPageId="5" lastPageNumber="48" pageId="4" pageNumber="47">
<subSubSection box="[136,696,1382,1409]" pageId="4" pageNumber="47" type="nomenclature">
<paragraph blockId="4.[136,696,1382,1409]" box="[136,696,1382,1409]" pageId="4" pageNumber="47">
<heading box="[136,696,1382,1409]" fontSize="11" level="2" pageId="4" pageNumber="47" reason="3">
<taxonomicName authority="Aver. &amp; V.C.Nguyen" authorityName="Aver. &amp; V. C. Nguyen" authorityYear="2022" box="[136,598,1382,1409]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="47" phylum="Tracheophyta" rank="species" species="tatonii" status="sp. nov.">
<emphasis bold="true" box="[136,358,1382,1409]" italics="true" pageId="4" pageNumber="47">Cleisostoma tatonii</emphasis>
Aver. &amp; V.C.Nguyen
</taxonomicName>
,
<emphasis bold="true" box="[610,696,1383,1409]" italics="true" pageId="4" pageNumber="47">
<taxonomicNameLabel box="[610,696,1383,1409]" pageId="4" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[136,241,1422,1444]" pageId="4" pageNumber="47" type="description">
<paragraph blockId="4.[136,241,1422,1444]" box="[136,241,1422,1444]" pageId="4" pageNumber="47">
(
<figureCitation box="[143,210,1422,1444]" captionStart="FIGURE 3" captionStartId="6.[136,229,1866,1888]" captionTargetBox="[243,1344,190,1841]" captionTargetId="figure-23@6.[243,1344,190,1841]" captionTargetPageId="6" captionText="FIGURE 3. Cleisostoma tatonii Aver. &amp; V.C.Nguyen, sp. nov. A. Flowering plant. B. Inflorescence. C. Portion of inflorescence. D. Intact flower, frontal view. E. Intact and flattened flowers, side views. F. Median sepal, abaxial side. G. Lateral sepals, abaxial and adaxial sides. H. Petals, abaxial and adaxial sides. I. Flattened flower, frontal part of the lip (including spur) removed. J, K. Frontal part of the lip (including spur), view from inside. L. Intact column, frontal, half-side and side views. M. Anther cap, views from different sides. N. Pollinarium. O. Pollinarium stalk (tegula), views from different sides. P. Pollinia. All photos made from the type specimen AL 1591 by Nguyen Van Canh and L. Averyanov, photo correction and design by L.Averyanov and T. Maisak." figureDoi="http://doi.org/10.5281/zenodo.7305626" httpUri="https://zenodo.org/record/7305626/files/figure.png" pageId="4" pageNumber="47">Figs. 3</figureCitation>
,
<figureCitation box="[221,234,1422,1443]" captionStart="FIGURE 4" captionStartId="7.[136,229,1901,1923]" captionTargetBox="[233,1352,193,1874]" captionTargetId="figure-18@7.[232,1356,190,1876]" captionTargetPageId="7" captionText="FIGURE 4. Cleisostoma tatonii Aver. &amp; V.C.Nguyen, sp. nov. A. Flowering plant. B. Leaf, adaxial side. C-E. Intact flower, frontal, halfside, and side views. F. Sagittal lip section. G. Flattened flower, frontal part of the lip (including spur) removed. H, I. Frontal part of the lip (including spur), view from inside and outside. J. Anther cap, views from different sides. K. Pollinarium, frontal view and view from back. L. Pollinarium stalk (tegula), views from different sides. All drawings made from the type specimen AL 1591 by L. Averyanov and T. Maisak." figureDoi="http://doi.org/10.5281/zenodo.7305630" httpUri="https://zenodo.org/record/7305630/files/figure.png" pageId="4" pageNumber="47">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="47" type="materials_examined">
<paragraph blockId="4.[136,1452,1489,2079]" pageId="4" pageNumber="47">
<emphasis bold="true" box="[136,204,1490,1516]" pageId="4" pageNumber="47">Type:</emphasis>
<materialsCitation collectingDate="2021-09-13" collectionCode="S" collectorName="Ta Ton &amp; Nguyen Van Canh" country="Vietnam" location="Ngoc Linh Mountains" municipality="Vietnam" pageId="4" pageNumber="47" specimenCount="1" stateProvince="Kon Tum">
<collectingCountry box="[233,365,1490,1516]" name="Vietnam" pageId="4" pageNumber="47">VIETNAM</collectingCountry>
.
<collectionCode box="[375,391,1490,1516]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="4" pageNumber="47" type="Herbarium">S</collectionCode>
<collectingMunicipality box="[397,495,1489,1515]" pageId="4" pageNumber="47">Vietnam</collectingMunicipality>
,
<collectingRegion box="[506,722,1489,1516]" country="Vietnam" name="Kon Tum" pageId="4" pageNumber="47">Kon Tum Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03FA834FDD4A113F5C8CFA8AFE8C9D97:8E8C6482DD4A113E5EDAFA3EFC229DF5" box="[734,984,1489,1516]" country="Vietnam" municipality="Vietnam" name="Ngoc Linh Mountains" pageId="4" pageNumber="47" stateProvince="Kon Tum">Ngoc Linh Mountains</location>
,
<date box="[995,1216,1489,1516]" pageId="4" pageNumber="47" value="2021-09-13">
<collectingDate box="[995,1216,1489,1516]" pageId="4" pageNumber="47" value="2021-09-13">13 September 2021</collectingDate>
</date>
,
<emphasis italics="true" pageId="4" pageNumber="47">
<collectorName box="[1227,1306,1490,1515]" pageId="4" pageNumber="47">Ta Ton</collectorName>
,
<collectorName pageId="4" pageNumber="47">Nguyen Van Canh</collectorName>
</emphasis>
</materialsCitation>
<emphasis box="[199,203,1525,1551]" italics="true" pageId="4" pageNumber="47">,</emphasis>
<materialsCitation collectionCode="AL, LE" country="Vietnam" pageId="4" pageNumber="47" specimenCode="AL 1591, LE01169337, LE01122992, LE01123534" specimenCount="1" typeStatus="holotype">
<emphasis box="[208,300,1525,1551]" italics="true" pageId="4" pageNumber="47">
<specimenCode box="[208,300,1525,1551]" collectionCode="AL" country="Algeria" lsid="urn:lsid:biocol.org:col:14432" name="Université d'Alger" pageId="4" pageNumber="47" type="Herbarium">AL 1591</specimenCode>
</emphasis>
(
<typeStatus box="[312,412,1525,1551]" pageId="4" pageNumber="47">holotype</typeStatus>
<specimenCode box="[416,561,1524,1550]" collectionCode="LE" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="4" pageNumber="47">LE01169337</specimenCode>
https://herbariumle.ru/?t=occ&amp;id=142648, analytical photos and drawings made from specimen used for preparation of the type specimen
<specimenCode box="[790,934,1560,1586]" collectionCode="LE" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="4" pageNumber="47">LE01122992</specimenCode>
https://en.herbariumle.ru/?t=occ&amp;id=110358,
<specimenCode box="[136,281,1595,1622]" collectionCode="LE" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="4" pageNumber="47">LE01123534</specimenCode>
https://en.herbariumle.ru/?t=occ&amp;id=141837)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[189,1227,1630,1657]" pageId="4" pageNumber="47" type="etymology">
<paragraph blockId="4.[136,1452,1489,2079]" box="[189,1227,1630,1657]" pageId="4" pageNumber="47">
<emphasis bold="true" box="[189,328,1630,1656]" pageId="4" pageNumber="47">Etymology:</emphasis>
—Species name honors its discoverer, an orchid enthusiast, Mr. Nguyen Ta Ton.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="48" pageId="4" pageNumber="47" type="description">
<paragraph blockId="4.[136,1452,1489,2079]" lastBlockId="5.[136,1452,158,1417]" lastPageId="5" lastPageNumber="48" pageId="4" pageNumber="47">
<emphasis bold="true" box="[189,427,1665,1691]" pageId="4" pageNumber="47">Description:—Herb</emphasis>
perennial monopodial epiphytic.
<emphasis bold="true" box="[805,866,1665,1691]" pageId="4" pageNumber="47">Stem</emphasis>
simple or basally few branching, semi-woody, erect or ascending, (4)58(10) cm long, covered throughout by leaf sheaths, with (5)68(10) leaves, in basal half with few thick wiry flexuose white roots; internodes (6)710(11) mm long.
<emphasis bold="true" box="[869,951,1736,1762]" pageId="4" pageNumber="47">Leaves</emphasis>
subdistichous, sessile, dorsiventral, leathery, somewhat succulent, straight or arching, oblong, (14)1518(20) cm long, (1.6)1.82(2.2) cm wide, unequally bilobed at apex, with distinct articulation at the base.
<emphasis bold="true" box="[662,820,1806,1832]" pageId="4" pageNumber="47">Inflorescence</emphasis>
lateral raceme or few branching panicle, (8)1012(14) cm long, scape and rachis dark greenish gray; scape
<quantity box="[723,806,1842,1868]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="47" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">12 cm</quantity>
long, pendulous, panicle branches 13(4), (4)68(10) cm long; rachis (4)58(10) cm long, more or less straight, with many spirally arranged, dense to subdense flowers.
<emphasis bold="true" pageId="4" pageNumber="47">Floral bracts</emphasis>
dark gray greenish, minute, triangular, acute, 11.2(1.4) mm long,
<quantity box="[957,1091,1912,1938]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="4" pageNumber="47" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.60.7 mm</quantity>
wide.
<emphasis bold="true" box="[1164,1376,1912,1938]" pageId="4" pageNumber="47">Pedicel and ovary</emphasis>
terete, longitudinally slightly grooved, (4.5)56(6.5) mm long, (0.7)0.8(0.9) mm in diameter, light green, slightly broadening at the base, glabrous or with few scattered scurfy brownish hairs.
<emphasis bold="true" box="[831,926,1982,2008]" pageId="4" pageNumber="47">Flowers</emphasis>
widely opening, (6.5)78(9) mm across; sepals and petals somewhat fleshy, spreading, dark purple brown, with pale yellowish median vein and yellowish margin; lip yellow with brown purple marks on the lip lobes and spur; column yellow flushed with brown purple at apex; anther cap dark brown purple with two lateral yellowish spots.
<emphasis bold="true" box="[783,859,158,184]" pageId="5" pageNumber="48">Sepals</emphasis>
narrowly ovate to ovate, concave, rounded at apex, (3.4)3.64(4.2) mm long, (1.2)1.41.8(2) mm wide, at the base with few scattered scurfy brownish hairs; median sepal distinctly cucullate; lateral sepals somewhat oblique.
<emphasis bold="true" box="[730,802,229,255]" pageId="5" pageNumber="48">Petals</emphasis>
narrowly obovate to broadly oblanceolate, glabrous, blunt to obtuse at apex, (2)2.22.4(2.6) mm long, (0.5)0.60.8(1) mm wide, slightly oblique.
<emphasis bold="true" box="[1113,1155,264,290]" pageId="5" pageNumber="48">Lip</emphasis>
spurred, (6.2)6.57.5(7.8) mm long (from spur apex to the apex of epichile), 3-lobed; side-lobes triangular, erect,
<quantity box="[1111,1246,299,326]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="5" pageNumber="48" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.50.7 mm</quantity>
long, inside at the base with small round callus and in distal part with erect subulate processus (1)1.21.4(1.6) mm long; median lobe fleshy, arrow shaped, (1)1.2(1.4) mm long, (4.2)4.44.6(4.8) mm wide, with 3 small triangular lobules at apex and 2 large ensiform, lateral, back directed acute lobules (1.4)1.61.8(2) mm long; spur shortly cylindrical, almost parallel to ovary or slightly upcurved, (3)3.23.4(3.6) mm long, (1)1.21.4((1.6) mm in diameter, round at apex, without longitudinal septum inside.
<emphasis bold="true" box="[453,638,475,501]" pageId="5" pageNumber="48">Spur back-wall</emphasis>
inside with Y-shaped callus, (1.6)1.8(2) mm wide, (0.7)0.81(1.1) mm long, shortly hairy or densely glandular at base.
<emphasis bold="true" box="[692,878,510,536]" pageId="5" pageNumber="48">Spur front-wall</emphasis>
inside with two hairy or densely glandular lateral calli, allied to the apex of back-wall callus and almost closing (together with back-wall callus) the entrance to the spur; the base of median lobe with short glabrous rising longitudinal keel.
<emphasis bold="true" box="[930,1026,580,606]" pageId="5" pageNumber="48">Column</emphasis>
short, broad, stout, shortly cylindric, (1.7)1.82(2.1) mm tall and wide, at apex on each side with dense, half-globular bunch of white trichomes
<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="5" pageNumber="48" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 mm</quantity>
in diameter, near apex at front with rostellum supporting large rectangular concave viscidium; stigma at front of column, in form of large prominent disk, (1.8)22.2(2.3) mm in diameter, with small obscurely triangular hollow near proximal margin.
<emphasis bold="true" box="[396,531,721,748]" pageId="5" pageNumber="48">Anther cap</emphasis>
helmet shaped,
<quantity box="[717,854,721,748]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="5" pageNumber="48" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 mm</quantity>
tall, (1.1)1.21.4(1.5) mm wide, with large gibbous, concave, narrowly triangular beak, (1)1.11.2(1.3) mm long.
<emphasis bold="true" box="[832,924,756,782]" pageId="5" pageNumber="48">Pollinia</emphasis>
2, each incompletely split into 2 half-globular bodies.
<emphasis bold="true" box="[222,283,791,817]" pageId="5" pageNumber="48">Stipe</emphasis>
(tegula) in form of narrowly triangular, conduplicate scarious stalk (1.2)1.41.5(1.6) mm long; viscidium large, in form of rectangular concave plate,
<quantity box="[620,755,827,854]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="5" pageNumber="48" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.91.1 mm</quantity>
long,
<quantity box="[823,958,827,854]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="5" pageNumber="48" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.70.8 mm</quantity>
wide (being flattened).
<emphasis bold="true" box="[1222,1284,827,853]" pageId="5" pageNumber="48">Fruit</emphasis>
almost sessile, narrowly ellipsoid, slightly ridged capsule, (9)1012(13) mm long.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="48" type="biology_ecology">
<paragraph blockId="5.[136,1452,158,1417]" pageId="5" pageNumber="48">
<emphasis bold="true" box="[189,468,898,924]" pageId="5" pageNumber="48">Ecology and phenology</emphasis>
:—Epiphyte. Primary evergreen broad-leaved and mixed forests on granite at elevation
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a.s.l. Flowers in September.
</paragraph>
</subSubSection>
<subSubSection box="[189,1241,968,994]" pageId="5" pageNumber="48" type="distribution">
<paragraph blockId="5.[136,1452,158,1417]" box="[189,1241,968,994]" pageId="5" pageNumber="48">
<emphasis bold="true" box="[189,335,968,994]" pageId="5" pageNumber="48">Distribution</emphasis>
:—
<collectingCountry box="[369,504,968,994]" name="Vietnam" pageId="5" pageNumber="48">VIETNAM</collectingCountry>
. S Vietnam,
<collectingRegion box="[645,861,968,994]" country="Vietnam" name="Kon Tum" pageId="5" pageNumber="48">Kon Tum Province</collectingRegion>
, Ngoc Linh Mountains. Endemic.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="48" type="conservation">
<paragraph blockId="5.[136,1452,158,1417]" pageId="5" pageNumber="48">
<emphasis bold="true" box="[189,426,1003,1029]" pageId="5" pageNumber="48">Conservation status</emphasis>
:—Despite many-years, special field searches of professional botanists and numerous orchid enthusiasts only few samples of this extremely rare species were found in one location. Being into consideration vast deforestation in Ngoc Linh Mountains it may be expected that this species stands on the verge of full extinction in the nature. According to available data, having one population with EOO and AOO less than
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and
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<superScript attach="left" box="[1443,1451,1108,1122]" fontSize="6" pageId="5" pageNumber="48">2</superScript>
respectively, expected coming extinction, and very small population with less than 50 mature individuals, it may be preliminarily assessed as globally Critically Endangered CR (B1a,b(ii,iii), B2a,b(ii,iii); C1a(i,ii), C2a(i,ii); D).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="48" type="discussion">
<paragraph blockId="5.[136,1452,158,1417]" pageId="5" pageNumber="48">
<emphasis bold="true" box="[189,256,1214,1240]" pageId="5" pageNumber="48">Notes</emphasis>
:—According to plant and floral morphology, the new species has rather isolated, intermediate taxonomic position between
<emphasis box="[337,363,1250,1276]" italics="true" pageId="5" pageNumber="48">C.</emphasis>
sect.
<taxonomicName authority="(Seidenfaden 1975: 10)" baseAuthorityName="Seidenfaden" baseAuthorityPageNumber="10" baseAuthorityYear="1975" box="[430,844,1249,1276]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="48" phylum="Tracheophyta" rank="genus">
<emphasis box="[430,568,1249,1275]" italics="true" pageId="5" pageNumber="48">Cleisostoma</emphasis>
(
<bibRefCitation author="Seidenfaden, G." box="[585,835,1249,1276]" pageId="5" pageNumber="48" pagination="1 - 80" refId="ref11372" refString="Seidenfaden, G. (1975) Orchid genera in Thailand II. Cleisostoma Bl. Dansk Botanisk Arkiv 29 (3): 1 - 80." type="journal article" year="1975">Seidenfaden 1975: 10</bibRefCitation>
)
</taxonomicName>
and
<emphasis box="[901,927,1250,1276]" italics="true" pageId="5" pageNumber="48">C.</emphasis>
sect.
<taxonomicName authority="Seidenfaden (1975: 34)" authorityName="Seidenfaden" authorityPageNumber="34" authorityYear="1975" box="[995,1396,1249,1276]" class="Insecta" family="Lecithoceridae" genus="Paniculata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis box="[995,1118,1249,1275]" italics="true" pageId="5" pageNumber="48">Paniculata</emphasis>
<bibRefCitation author="Seidenfaden, G." box="[1127,1396,1249,1276]" pageId="5" pageNumber="48" pagination="1 - 80" refId="ref11372" refString="Seidenfaden, G. (1975) Orchid genera in Thailand II. Cleisostoma Bl. Dansk Botanisk Arkiv 29 (3): 1 - 80." type="journal article" year="1975">Seidenfaden (1975: 34)</bibRefCitation>
</taxonomicName>
. The structure of its pollinarium has triangular conduplicate stipe with large rectangular longitudinally concave viscidium, very large discoid stigma broader than the column itself, dense bunches of white hairs at the column apex, and dense inflorescence are features unique in the genus (
<bibRefCitation author="Averyanov, L. V. &amp; Nguyen, T. T. &amp; Nguyen, V. C." box="[677,939,1355,1382]" pageId="5" pageNumber="48" pagination="107 - 116" refId="ref10386" refString="Averyanov, L. V., Nguyen, T. T. &amp; Nguyen, V. C. (2015) New species of the genus Cleisostoma in the flora of Vietnam. Taiwania 60 (3): 107 - 116." type="journal article" year="2015">
Averyanov
<emphasis box="[811,869,1355,1381]" italics="true" pageId="5" pageNumber="48">et al.</emphasis>
2015
</bibRefCitation>
). We see no any close relatives for our plant among its congeners.
</paragraph>
</subSubSection>
</treatment>
</document>