283 lines
31 KiB
XML
283 lines
31 KiB
XML
<document id="BC9B2010FFF5E1F4BA6721CC47425DC5" ID="10.11646/zootaxa.4139.2.12" ID-DOI="10.11646/zootaxa.4139.2.12" ID-GBIF-Dataset="19d9d549-a90a-4812-a420-658b019a7c36" ID-ISSN="1175-5326" ID-Zenodo-Dep="264822" ID-ZooBank="D4B4C1E3-F897-411D-B03B-0C0C054B4EA6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468999433724" checkinUser="plazi" docAuthor="Vaz-De-Mello, Fernando Z. & Nunes, Rafael V." docDate="2016" docId="03FA1339FFD5FFB3FF70FD914040FDCD" docLanguage="en" docName="zootaxa.4139.2.12.pdf" docOrigin="Zootaxa 4139 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dichotomius (Luederwaldtinia) benesi Vaz-De-Mello & Nunes, 2016, n. sp." docType="treatment" docVersion="7" lastPageNumber="297" masterDocId="FFC36B41FFD4FFB0FFE7FFB5455AFFF0" masterDocTitle="New species, redescription and taxonomic notes in the Dichotomius (Luederwaldtinia) batesi (Harold) species-group (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="300" masterPageNumber="294" pageNumber="295" updateTime="1698365383601" updateUser="plazi">
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<mods:titleInfo id="9670BAAC4E9E338EA71A1CFB9A385399">
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<mods:title id="C1F0FC6534370BC863EEFA84B567AC8E">New species, redescription and taxonomic notes in the Dichotomius (Luederwaldtinia) batesi (Harold) species-group (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="12500D56566591C19B2419D5219EF709">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:namePart id="EA64DAF8D01B2E1C63D42E767B774228">Nunes, Rafael V.</mods:namePart>
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<mods:title id="BF6F875A3BE0CF80275CF702FB25DBCB">Zootaxa</mods:title>
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<mods:date id="7C26ED15AC4A3AD19091533B3DC6E2E4">2016</mods:date>
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<mods:number id="B9AA8089CA2AF41E993AEA230B32968A">4139</mods:number>
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<treatment id="03FA1339FFD5FFB3FF70FD914040FDCD" ID-DOI="http://doi.org/10.5281/zenodo.6063810" ID-GBIF-Taxon="123198501" ID-Zenodo-Dep="6063810" LSID="urn:lsid:plazi:treatment:03FA1339FFD5FFB3FF70FD914040FDCD" httpUri="http://treatment.plazi.org/id/03FA1339FFD5FFB3FF70FD914040FDCD" lastPageId="3" lastPageNumber="297" pageId="1" pageNumber="295">
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<paragraph id="8BECA22FFFD5FFB1FF70FD9147EAFDCE" blockId="1.[151,688,548,607]" box="[151,688,548,574]" pageId="1" pageNumber="295">
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<emphasis id="B9277E3DFFD5FFB1FF70FD9147EAFDCE" bold="true" box="[151,688,548,574]" pageId="1" pageNumber="295">
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<taxonomicName id="4C53D9ACFFD5FFB1FF70FD91473DFDCE" ID-CoL="35NB9" box="[151,615,548,574]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="species" species="benesi" status="sp. nov." subGenus="Luederwaldtinia">
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<emphasis id="B9277E3DFFD5FFB1FF70FD91473DFDCE" bold="true" box="[151,615,548,574]" italics="true" pageId="1" pageNumber="295">Dichotomius (Luederwaldtinia) benesi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A214C346FFD5FFB1FD8AFD9147EAFDCE" box="[621,688,548,574]" pageId="1" pageNumber="295" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8BECA22FFFD5FFB1FF70FDF34583FDAF" blockId="1.[151,688,548,607]" box="[151,217,582,607]" pageId="1" pageNumber="295">
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<figureCitation id="1368BEAAFFD5FFB1FF70FDF34583FDAF" box="[151,217,582,607]" captionStart="FIGURE 1" captionStartId="2.[151,250,1930,1952]" captionTargetBox="[221,1350,220,1862]" captionTargetId="figure@2.[196,1368,195,1878]" captionTargetPageId="2" captionText="FIGURE 1. (A) Dorsal habitus of the males of Dichotomius (L.) benesi n. sp. (B) Ventral habitus of the male of D. benesi. (C) Dorsal habitus of the female of D. benesi. (D) Dorsal habitus of the male genitalia of D. benesi. (E) lateral habitus of the male genitalia of D. benesi." httpUri="https://zenodo.org/record/264823/files/figure.png" pageId="1" pageNumber="295">Fig. 1</figureCitation>
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</paragraph>
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<subSubSection id="C349F1A4FFD5FFB1FF70FD3B4620FD1F" pageId="1" pageNumber="295" type="materials_examined">
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<paragraph id="8BECA22FFFD5FFB1FF70FD3B4620FD1F" blockId="1.[151,1437,654,2011]" pageId="1" pageNumber="295">
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Material studied:
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:
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</emphasis>
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Ƌ
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:
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<collectingRegion id="49976CCDFFD5FFB1FD46FD3B464EFD57" box="[673,788,654,679]" country="Panama" name="Veraguas" pageId="1" pageNumber="295">Veraguas</collectingRegion>
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:
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</emphasis>
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<collectingRegion id="49976CCDFFD5FFB1FCC2FD3A46D7FD57" box="[805,909,655,679]" country="Argentina" name="Santa Fe" pageId="1" pageNumber="295">Santa Fé</collectingRegion>
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,
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<quantity id="4CAB0FCAFFD5FFB1FC7AFD3A46B3FD56" box="[925,1001,655,679]" metricMagnitude="2" metricUnit="m" metricValue="8.11" pageId="1" pageNumber="295" unit="m" value="811.0">811 m</quantity>
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,
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<geoCoordinate id="EE67C4E8FFD5FFB1FC1EFD3A4113FD57" box="[1017,1097,655,679]" direction="north" orientation="latitude" pageId="1" pageNumber="295" precision="925" value="8.483334">8°29'N</geoCoordinate>
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,
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<geoCoordinate id="EE67C4E8FFD5FFB1FBBDFD3B41E7FD57" box="[1114,1213,654,679]" direction="west" orientation="longitude" pageId="1" pageNumber="295" precision="925" value="-81.1">81°06'W</geoCoordinate>
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,
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<date id="FFED84EFFFD5FFB1FB2BFD3B4023FD57" box="[1228,1401,654,679]" pageId="1" pageNumber="295" value="2013-06-16" valueMax="2013-06-18" valueMin="2013-06-16">16-18-VI-2013</date>
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. V Benes lgt [
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<collectionCode id="ED423AEAFFD5FFB1FEEBFD064438FD3B" box="[268,354,691,715]" pageId="1" pageNumber="295">CEMT</collectionCode>
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].
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<emphasis id="B9277E3DFFD5FFB1FE93FD0744AEFD3B" bold="true" box="[372,500,690,715]" pageId="1" pageNumber="295">
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<typeStatus id="54E81C8DFFD5FFB1FE93FD0744AAFD3B" box="[372,496,690,715]" pageId="1" pageNumber="295" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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same data as
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<typeStatus id="54E81C8DFFD5FFB1FD69FD0747AAFD3B" box="[654,752,690,715]" pageId="1" pageNumber="295" type="holotype">holotype</typeStatus>
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||
[10 Ƌ,
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<specimenCount id="9D5569A6FFD5FFB1FCA2FD064635FD3A" box="[837,879,691,714]" pageId="1" pageNumber="295" type="female">7 ♀</specimenCount>
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<collectionCode id="ED423AEAFFD5FFB1FC92FD064693FD3B" box="[885,969,691,715]" pageId="1" pageNumber="295">CEMT</collectionCode>
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]; same but Safanda lgt [1Ƌ,
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<specimenCount id="9D5569A6FFD5FFB1FAE4FD064077FD3A" box="[1283,1325,691,715]" pageId="1" pageNumber="295" type="female">3 ♀</specimenCount>
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<collectionCode id="ED423AEAFFD5FFB1FAD4FD0640D9FD3B" box="[1331,1411,691,715]" pageId="1" pageNumber="295">CEMT</collectionCode>
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; 2 Ƌ
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<specimenCount id="9D5569A6FFD5FFB1FF54FD624587FD1E" box="[179,221,727,751]" pageId="1" pageNumber="295" type="female">9 ♀</specimenCount>
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<collectionCode id="ED423AEAFFD5FFB1FF03FD624476FD1F" box="[228,300,727,751]" pageId="1" pageNumber="295">SPCP</collectionCode>
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]; same but iv-v-2014, V Benes lgt [1 Ƌ
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<specimenCount id="9D5569A6FFD5FFB1FD0CFD62464EFD1E" box="[747,788,727,751]" pageId="1" pageNumber="295" type="female">1 ♀</specimenCount>
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<collectionCode id="ED423AEAFFD5FFB1FCFBFD62462AFD1F" box="[796,880,727,751]" pageId="1" pageNumber="295">CEMT</collectionCode>
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].
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C349F1A4FFD5FFB1FF20FD4F413EFBA7" pageId="1" pageNumber="295" type="diagnosis">
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<paragraph id="8BECA22FFFD5FFB1FF20FD4F413EFBA7" blockId="1.[151,1437,654,2011]" pageId="1" pageNumber="295">
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<emphasis id="B9277E3DFFD5FFB1FF20FD4F4419FCE3" bold="true" box="[199,323,762,787]" pageId="1" pageNumber="295">Diagnosis:</emphasis>
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<emphasis id="B9277E3DFFD5FFB1FEB5FD4E4765FCE3" box="[338,575,763,787]" italics="true" pageId="1" pageNumber="295">
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||
<taxonomicName id="4C53D9ACFFD5FFB1FEB5FD4E44BFFCE3" box="[338,485,763,787]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="genus">Dichotomius</taxonomicName>
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.
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<taxonomicName id="4C53D9ACFFD5FFB1FE10FD4E4765FCE3" box="[503,575,763,787]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="species" species="benesi" status="sp. nov.">benesi</taxonomicName>
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</emphasis>
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||
<emphasis id="B9277E3DFFD5FFB1FDA9FD4F47CFFCE3" bold="true" box="[590,661,762,787]" pageId="1" pageNumber="295">
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<taxonomicNameLabel id="A214C346FFD5FFB1FDA9FD4F47CFFCE3" box="[590,661,762,787]" pageId="1" pageNumber="295" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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belongs to the subgenus
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<taxonomicName id="4C53D9ACFFD5FFB1FC3EFD4E41C9FCE3" box="[985,1171,763,787]" class="Insecta" family="Scarabaeidae" genus="Luederwaldtinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="genus">
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<emphasis id="B9277E3DFFD5FFB1FC3EFD4E41C9FCE3" box="[985,1171,763,787]" italics="true" pageId="1" pageNumber="295">Luederwaldtinia</emphasis>
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</taxonomicName>
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due to the following combined characters: two clypeal teeth; ventral clypeal process bifurcated; clypeo-genal junction rounded, lacking angulation; abdominal ventrites having setae; last abdominal sternite of females simply widened, lacking tubercles or projections (Vaz-de-Mello
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<emphasis id="B9277E3DFFD5FFB1FE04FCD2474EFC8E" box="[483,532,870,894]" italics="true" pageId="1" pageNumber="295">et al</emphasis>
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. 2011;
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<bibRefCitation id="EFC2DFDEFFD5FFB1FD8FFCD246E8FC8F" author="Nunes" box="[616,946,870,895]" pageId="1" pageNumber="295" refString="Nunes, R. V. & Vaz-de-Mello, F. Z. (2013) New brachypterous species of Dichotomius Hope, with taxonomic notes in the subgenus Luederwaldtinia Martinez (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa, 3609 (4), 411 - 420. http: // dx. doi. org / 10.11646 / zootaxa. 3609.4.3" type="journal article" year="2013">Nunes & Vaz-de-Mello 2013</bibRefCitation>
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). Among this subgenus, it is assigned to the
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<taxonomicName id="4C53D9ACFFD5FFB1FF70FC3E4580FC53" box="[151,218,907,931]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="species" species="batesi">
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<emphasis id="B9277E3DFFD5FFB1FF70FC3E4580FC53" box="[151,218,907,931]" italics="true" pageId="1" pageNumber="295">batesi</emphasis>
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</taxonomicName>
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species-group since its pronotal disc bears ocellate punctures all over its extension and males calcar is not sickle-shaped.(
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<bibRefCitation id="EFC2DFDEFFD5FFB1FEDAFC1B474AFC37" author="Luederwaldt" box="[317,528,942,967]" pageId="1" pageNumber="295" refString="Luederwaldt, H. (1929) As especies Brasileiras do genero Pinotus. Revista do Museu Paulista, 16, 1 - 173." type="journal article" year="1929">Luederwaldt 1929</bibRefCitation>
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,
|
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<bibRefCitation id="EFC2DFDEFFD5FFB1FDFAFC1B47EAFC37" author="Pereira" box="[541,688,942,967]" pageId="1" pageNumber="295" refString="Pereira, F. S. (1947) Pinotus da secao Batesi. Arquivos do Museu Paranaense, 6, 317 - 328." type="journal article" year="1947">Pereira 1947</bibRefCitation>
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,
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||
<bibRefCitation id="EFC2DFDEFFD5FFB1FD5AFC1A4151FC37" author="Nunes" box="[701,1035,942,967]" pageId="1" pageNumber="295" refString="Nunes, R. V. & Vaz-de-Mello, F. Z. (2013) New brachypterous species of Dichotomius Hope, with taxonomic notes in the subgenus Luederwaldtinia Martinez (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa, 3609 (4), 411 - 420. http: // dx. doi. org / 10.11646 / zootaxa. 3609.4.3" type="journal article" year="2013">Nunes & Vaz-de-Mello 2013</bibRefCitation>
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||
). This new species is differentiated from the others of the group by the combination of the following characters: atrophied hind wing lacking the anal vein and never having more than the elytra length; pronotal disc punctures equally spaced; cephalic process consisting of a unique central tubercle; longitudinal groove present along all the posterior margin of the meso and meta femura; elytral striae lacking punctures and apexes of the paramera membranous.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349F1A4FFD5FFB3FF20FBD7462AFF08" lastPageId="3" lastPageNumber="297" pageId="1" pageNumber="295" type="materials_examined">
|
||
<paragraph id="8BECA22FFFD5FFB1FF20FBD74034F82B" blockId="1.[151,1437,654,2011]" pageId="1" pageNumber="295">
|
||
<emphasis id="B9277E3DFFD5FFB1FF20FBD74467FB8B" bold="true" box="[199,317,1122,1147]" pageId="1" pageNumber="295">
|
||
<typeStatus id="54E81C8DFFD5FFB1FF20FBD74462FB8B" box="[199,312,1122,1147]" pageId="1" pageNumber="295" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
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||
male, length:
|
||
<quantity id="4CAB0FCAFFD5FFB1FE0EFBD64764FB8A" box="[489,574,1123,1147]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="295" unit="mm" value="14.0">14 mm</quantity>
|
||
. Pronotum width:
|
||
<quantity id="4CAB0FCAFFD5FFB1FCFEFBD6463BFB8A" box="[793,865,1123,1147]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="295" unit="mm" value="8.0">8 mm</quantity>
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||
. Dorsal and ventral surface barely shinny, black.
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||
<emphasis id="B9277E3DFFD5FFB1FF70FB3345BAFB6F" bold="true" box="[151,224,1158,1183]" pageId="1" pageNumber="295">Head:</emphasis>
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||
anterior margin of the clypeous with a deep “u” shaped emargination forming two rounded teeth. Clypeogenal junction rounded, not angulating. Clypeal surface smooth at the front and with ocellate punctures anteriorly and laterally. Gena and intraocular space also bearing ocellate punctures equally spaced. Clypeo-genal suture present. Cephalic process consisting of a small central tubercle. Antenal club with brown light tumescence.
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<emphasis id="B9277E3DFFD5FFB1FF70FAA34443FADF" bold="true" box="[151,281,1302,1327]" pageId="1" pageNumber="295">Pronotum:</emphasis>
|
||
wider than long and wider than elytra. Lacking knobs, horns or excavations. Pronotal disc strongly convex with ocellate punctures equally spaced by their diameter. Posterior margin bearing ocellate elliptical punctures larger than those found on the rest of the pronotum. Anterior angles acute.
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||
<emphasis id="B9277E3DFFD5FFB1FBB7FAEB41BDFA87" bold="true" box="[1104,1255,1374,1399]" pageId="1" pageNumber="295">Hypomeron:</emphasis>
|
||
whole structure covered by ocellate setose punctures—setae short, scattered and not present in all punctures.
|
||
<emphasis id="B9277E3DFFD5FFB1FB4EFA37401BFA6B" bold="true" box="[1193,1345,1410,1435]" pageId="1" pageNumber="295">Prosternum:</emphasis>
|
||
bearing ocellate punctures all over its extension.
|
||
<emphasis id="B9277E3DFFD5FFB1FD6FFA13466FFA4F" bold="true" box="[648,821,1446,1471]" pageId="1" pageNumber="295">Mesosternum:</emphasis>
|
||
also bearing ocellate punctures almost all over its extension, except at the middle, where is narrower, smooth and shinny.
|
||
<emphasis id="B9277E3DFFD5FFB1FC29FA7F4123FA13" bold="true" box="[974,1145,1482,1507]" pageId="1" pageNumber="295">Metasternum:</emphasis>
|
||
with few scattered setae, almost glabrous. Mesometasternal suture distinct. Anterior portion of the metasternal lobe bearing ocellate punctures. Anterior lobe width half the length of metafemur. Longitudinal sulcus feebly marked. Fine punctures present near the region between metacoxae. Metasternal sides bearing ocellate punctures similar to those found at the prosternum, mesosternum and sides of the abdominal segments.
|
||
<emphasis id="B9277E3DFFD5FFB1FC4EF9EF46A5F983" bold="true" box="[937,1023,1626,1651]" pageId="1" pageNumber="295">Elytra:</emphasis>
|
||
having humeral callus and strongly convex at lateral view. Striae ill defined and lacking distinct punctures. Interstriae with chagrinated microsculpture (
|
||
<date id="FFED84EFFFD5FFB1FF78F9164594F94B" box="[159,206,1699,1723]" pageId="1" pageNumber="295" value="1920-10">20x</date>
|
||
).
|
||
<emphasis id="B9277E3DFFD5FFB1FF04F9174431F94B" bold="true" box="[227,363,1698,1723]" pageId="1" pageNumber="295">Hind wing:</emphasis>
|
||
length
|
||
<quantity id="4CAB0FCAFFD5FFB1FE24F9164753F94A" box="[451,521,1699,1723]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="295" unit="mm" value="8.0">8 mm</quantity>
|
||
. Cubital venation interrupted and anal vein absent.
|
||
<emphasis id="B9277E3DFFD5FFB1FBB0F91741C2F94B" bold="true" box="[1111,1176,1698,1723]" pageId="1" pageNumber="295">Legs:</emphasis>
|
||
anterior tibiae bearing four teeth. Anterior calcar curved at the apical third, posterior calcar emarginated. Ventral surface of the anterior femur bearing ocellate setose punctures all over its extension—setae short, sparse and not present in all punctures. On the medium and posterior femur these punctures are restricted to their apical portion. Meso and meta femur bearing a longitudinal groove almost all over its extension—this sulcus gets wider as it goes to the distal part.
|
||
<emphasis id="B9277E3DFFD5FFB1FF70F8E3444FF89F" bold="true" box="[151,277,1878,1903]" pageId="1" pageNumber="295">Abdomen:</emphasis>
|
||
abdominal sternites with short-scattered setae mainly concentrated on sides; ocellate punctures present close to the anterior margin.
|
||
<emphasis id="B9277E3DFFD5FFB1FE18F8CF4721F863" bold="true" box="[511,635,1914,1939]" pageId="1" pageNumber="295">Pygidium:</emphasis>
|
||
twice wider then long and covered by ocellate coarse punctures.
|
||
<emphasis id="B9277E3DFFD5FFB1FF70F82B447AF847" bold="true" box="[151,288,1950,1975]" pageId="1" pageNumber="295">Phalobasis:</emphasis>
|
||
apex broadly arcuate, anterior angles acute.
|
||
<emphasis id="B9277E3DFFD5FFB1FCCBF82B46F7F847" bold="true" box="[812,941,1950,1975]" pageId="1" pageNumber="295">Paramera:</emphasis>
|
||
on dorsal view, each paramere is strongly narrowed medially. Apexes truncated and membranous. On lateral view, apex of the paramera bent downward.
|
||
</paragraph>
|
||
<caption id="DF2CF2A7FFD6FFB2FF70F83F442FF82D" httpUri="https://zenodo.org/record/264823/files/figure.png" pageId="2" pageNumber="296" targetBox="[221,1350,220,1862]" targetPageId="2">
|
||
<paragraph id="8BECA22FFFD6FFB2FF70F83F442FF82D" blockId="2.[151,1436,1929,2013]" pageId="2" pageNumber="296">
|
||
<emphasis id="B9277E3DFFD6FFB2FF70F83F4466F850" bold="true" box="[151,316,1929,1952]" pageId="2" pageNumber="296">FIGURE 1. (A)</emphasis>
|
||
Dorsal habitus of the males of
|
||
<taxonomicName id="4C53D9ACFFD6FFB2FD9EF83F4637F86F" box="[633,877,1930,1951]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="benesi" status="sp. nov." subGenus="Luederwaldtinia">
|
||
<emphasis id="B9277E3DFFD6FFB2FD9EF83F4637F86F" box="[633,877,1930,1951]" italics="true" pageId="2" pageNumber="296">Dichotomius (L.) benesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9277E3DFFD6FFB2FC94F83F468EF850" bold="true" box="[883,980,1929,1952]" pageId="2" pageNumber="296">
|
||
<taxonomicNameLabel id="A214C346FFD6FFB2FC94F83F46F6F86F" box="[883,940,1930,1951]" pageId="2" pageNumber="296" rank="species">n. sp.</taxonomicNameLabel>
|
||
(B)
|
||
</emphasis>
|
||
Ventral habitus of the male of
|
||
<taxonomicName id="4C53D9ACFFD6FFB2FAEBF83E4036F86F" box="[1292,1388,1930,1952]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="benesi">
|
||
<emphasis id="B9277E3DFFD6FFB2FAEBF83E4036F86F" box="[1292,1388,1930,1952]" italics="true" pageId="2" pageNumber="296">D. benesi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B9277E3DFFD6FFB2FA9EF83C40C6F850" bold="true" box="[1401,1436,1929,1952]" pageId="2" pageNumber="296">(C)</emphasis>
|
||
Dorsal habitus of the female of
|
||
<taxonomicName id="4C53D9ACFFD6FFB2FE3DF81C4760F84E" box="[474,570,1961,1982]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="benesi">
|
||
<emphasis id="B9277E3DFFD6FFB2FE3DF81C4760F84E" box="[474,570,1961,1982]" italics="true" pageId="2" pageNumber="296">D. benesi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B9277E3DFFD6FFB2FDA0F81D4730F84F" bold="true" box="[583,618,1960,1983]" pageId="2" pageNumber="296">(D)</emphasis>
|
||
Dorsal habitus of the male genitalia of
|
||
<taxonomicName id="4C53D9ACFFD6FFB2FC19F81C4104F84E" box="[1022,1118,1961,1982]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="benesi">
|
||
<emphasis id="B9277E3DFFD6FFB2FC19F81C4104F84E" box="[1022,1118,1961,1982]" italics="true" pageId="2" pageNumber="296">D. benesi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B9277E3DFFD6FFB2FB8CF81D41D7F84F" bold="true" box="[1131,1165,1960,1983]" pageId="2" pageNumber="296">(E)</emphasis>
|
||
lateral habitus of the male genitalia of
|
||
<taxonomicName id="4C53D9ACFFD6FFB2FEF7F87D442AF82C" box="[272,368,1991,2013]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="benesi">
|
||
<emphasis id="B9277E3DFFD6FFB2FEF7F87D442AF82C" box="[272,368,1991,2013]" italics="true" pageId="2" pageNumber="296">D. benesi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BECA22FFFD7FFB3FF20FF22462AFF08" blockId="3.[151,1436,151,573]" pageId="3" pageNumber="297">
|
||
<emphasis id="B9277E3DFFD7FFB3FF20FF2244AEFF40" bold="true" box="[199,500,151,176]" pageId="3" pageNumber="297">Morphological variation.</emphasis>
|
||
This species has few variations, those are mainly related to the body length (
|
||
<quantity id="4CAB0FCAFFD7FFB3FA91FF2D45BAFF24" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="3" pageNumber="297" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">12- 14mm</quantity>
|
||
) and pronotum width (
|
||
<quantity id="4CAB0FCAFFD7FFB3FE38FF09471AFF24" box="[479,576,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="3" pageNumber="297" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
|
||
). Females are distinguished from males by the sixth abdominal sternite—that has more than twice the width than any other abdominal ventrite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349F1A4FFD7FFB3FF20FEB1461AFEEC" box="[199,832,260,285]" pageId="3" pageNumber="297" type="distribution">
|
||
<paragraph id="8BECA22FFFD7FFB3FF20FEB1461AFEEC" blockId="3.[151,1436,151,573]" box="[199,832,260,285]" pageId="3" pageNumber="297">
|
||
<emphasis id="B9277E3DFFD7FFB3FF20FEB1443AFEED" bold="true" box="[199,352,260,285]" pageId="3" pageNumber="297">Distribution.</emphasis>
|
||
<collectingRegion id="49976CCDFFD7FFB3FE80FEB14492FEEC" box="[359,456,260,284]" country="Argentina" name="Santa Fe" pageId="3" pageNumber="297">Santa Fé</collectingRegion>
|
||
in
|
||
<collectingRegion id="49976CCDFFD7FFB3FE0BFEB047E6FEED" box="[492,700,260,285]" country="Panama" name="Veraguas" pageId="3" pageNumber="297">Veraguas province</collectingRegion>
|
||
in
|
||
<collectingCountry id="F344E2BFFFD7FFB3FD38FEB04661FEEC" box="[735,827,261,284]" name="Panama" pageId="3" pageNumber="297">
|
||
<collectingRegion id="49976CCDFFD7FFB3FD38FEB04661FEEC" box="[735,827,261,284]" country="Panama" name="Panama" pageId="3" pageNumber="297">Panama</collectingRegion>
|
||
</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349F1A4FFD7FFB3FF20FE924407FE95" pageId="3" pageNumber="297" type="etymology">
|
||
<paragraph id="8BECA22FFFD7FFB3FF20FE924407FE95" blockId="3.[151,1436,151,573]" pageId="3" pageNumber="297">
|
||
<emphasis id="B9277E3DFFD7FFB3FF20FE924414FEB0" bold="true" box="[199,334,295,320]" pageId="3" pageNumber="297">Etymology.</emphasis>
|
||
Named after Vladimir Beneš, Czech scarab and carabid collector and friend, who collected most of the
|
||
<typeStatus id="54E81C8DFFD7FFB3FF39FEF84454FE95" box="[222,270,333,357]" pageId="3" pageNumber="297">type</typeStatus>
|
||
series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349F1A4FFD7FFB3FF20FEDA4040FDCD" pageId="3" pageNumber="297" type="discussion">
|
||
<paragraph id="8BECA22FFFD7FFB3FF20FEDA4040FDCD" blockId="3.[151,1436,151,573]" pageId="3" pageNumber="297">
|
||
<emphasis id="B9277E3DFFD7FFB3FF20FEDA4461FE78" bold="true" box="[199,315,367,392]" pageId="3" pageNumber="297">Remarks.</emphasis>
|
||
<taxonomicName id="4C53D9ACFFD7FFB3FEA5FEC444F7FE78" box="[322,429,368,392]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="297" phylum="Arthropoda" rank="species" species="benesi" status="sp. nov.">
|
||
<emphasis id="B9277E3DFFD7FFB3FEA5FEC444F7FE78" box="[322,429,368,392]" italics="true" pageId="3" pageNumber="297">D. benesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9277E3DFFD7FFB3FE53FEDA44AEFE78" bold="true" box="[436,500,367,392]" pageId="3" pageNumber="297">
|
||
<taxonomicNameLabel id="A214C346FFD7FFB3FE53FEDA44AEFE78" box="[436,500,367,392]" pageId="3" pageNumber="297" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is close to
|
||
<taxonomicName id="4C53D9ACFFD7FFB3FD92FEC44646FE78" box="[629,796,368,392]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="297" phylum="Arthropoda" rank="species" species="gamboensis">
|
||
<emphasis id="B9277E3DFFD7FFB3FD92FEC44646FE78" box="[629,796,368,392]" italics="true" pageId="3" pageNumber="297">D. gamboensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B9277E3DFFD7FFB3FCB3FEC446B4FE78" box="[852,1006,368,392]" italics="true" pageId="3" pageNumber="297">
|
||
<taxonomicName id="4C53D9ACFFD7FFB3FCB3FEC446B0FE78" box="[852,1002,368,392]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="297" phylum="Arthropoda" rank="species" species="femoratus">D. femoratus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
both described from
|
||
<collectingCountry id="F344E2BFFFD7FFB3FB07FEC54063FE77" box="[1248,1337,368,391]" name="Panama" pageId="3" pageNumber="297">
|
||
<collectingRegion id="49976CCDFFD7FFB3FB07FEC54063FE77" box="[1248,1337,368,391]" country="Panama" name="Panama" pageId="3" pageNumber="297">Panama</collectingRegion>
|
||
</collectingCountry>
|
||
at lower altitudes (
|
||
<bibRefCitation id="EFC2DFDEFFD7FFB3FEE2FE21474CFE5C" author="Howden" box="[261,534,404,429]" pageId="3" pageNumber="297" refString="Howden, H. & Young, O. P. (1981) Panamanian Scarabaeinae: Taxonomy, distribution and habits. Contributions American Entomological Institute, 18 (1), 1 - 204." type="journal article" year="1981">Howden & Young 1981</bibRefCitation>
|
||
). These three species share the following characters: central unique tubercle on the head; pronotal disc punctures equally spaced; chagrinated elytral interstriae; longitudinal sulcus along the posterior margin of meso and metafemura; apexes of the paramera membranous on dorsal view and apexes of the paramera and bent downward on lateral view (
|
||
<figureCitation id="1368BEAAFFD7FFB3FD54FE4A47AFFDE8" box="[691,757,511,536]" captionStart="FIGURE 1" captionStartId="2.[151,250,1930,1952]" captionTargetBox="[221,1350,220,1862]" captionTargetId="figure@2.[196,1368,195,1878]" captionTargetPageId="2" captionText="FIGURE 1. (A) Dorsal habitus of the males of Dichotomius (L.) benesi n. sp. (B) Ventral habitus of the male of D. benesi. (C) Dorsal habitus of the female of D. benesi. (D) Dorsal habitus of the male genitalia of D. benesi. (E) lateral habitus of the male genitalia of D. benesi." httpUri="https://zenodo.org/record/264823/files/figure.png" pageId="3" pageNumber="297">Fig. 1</figureCitation>
|
||
D, 1E). However, the pronotum of
|
||
<taxonomicName id="4C53D9ACFFD7FFB3FB77FDB441A1FDE8" box="[1168,1275,512,536]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="297" phylum="Arthropoda" rank="species" species="benesi">
|
||
<emphasis id="B9277E3DFFD7FFB3FB77FDB441A1FDE8" box="[1168,1275,512,536]" italics="true" pageId="3" pageNumber="297">D. benesi</emphasis>
|
||
</taxonomicName>
|
||
is wider than elytra and the hind wing lack the anal vein; both characteristics are related to the absence of hind wing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |