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<document id="9C5245F53DF75B5D7BF25585D9714158" ID-CLB-Dataset="45740" ID-DOI="10.11646/zootaxa.3796.3.8" ID-GBIF-Dataset="e6f90dd2-610e-4278-8cd6-b6e4c52d2574" ID-ISSN="1175-5326" ID-Zenodo-Dep="230100" ID-ZooBank="5DC6D761-D857-49B8-A88C-29DEC44E30F7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460421721727" checkinUser="plazi" docAuthor="Boldrini, R. &amp; Pes, A. M. O." docDate="2014" docId="03F55C2CE31D1C78FF2F3F31FA1FFD14" docLanguage="en" docName="zt03796p567.pdf" docOrigin="Zootaxa 3796 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Camelobaetidius carolinae Boldrini &amp; Pes, 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="554" masterDocId="FFCC2454E31B1C71FFB83E7DFF8DFFBC" masterDocTitle="Five new species of Camelobaetidius Demoulin, 1966 (Ephemeroptera: Baetidae), and redescription of Camelobaetidius mexicanus (Traver &amp; Edmunds, 1968)" masterLastPageNumber="567" masterPageNumber="545" pageNumber="551" updateTime="1698330095492" updateUser="ExternalLinkService">
<mods:mods id="20CB5243BFF774AC6CCA2AEE1A73A6FF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9B8533673E416207ADC8BC85F566F68A">
<mods:title id="54179F295E416DE00AD6DEE9CBB8F1B7">Five new species of Camelobaetidius Demoulin, 1966 (Ephemeroptera: Baetidae), and redescription of Camelobaetidius mexicanus (Traver &amp; Edmunds, 1968)</mods:title>
</mods:titleInfo>
<mods:name id="D4266F7F3DA8D93570C980C58A3C62FA" type="personal">
<mods:role id="9364CEAACEAB82FC062B626A5870E013">
<mods:roleTerm id="6B1B0FCEC21A998747CA21423FEA076F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A900F7ED0BD0DB13647D8631F507ADA2">Boldrini, R.</mods:namePart>
</mods:name>
<mods:name id="FA742345710EACE848D931EC4118166B" type="personal">
<mods:role id="B293E3751BBB1A9968D836E28D9B8311">
<mods:roleTerm id="1E9B6AE2E0ACA1585AA4EC47B74D1616">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B5955A4EC264857AC2FE4A489F1E4A80">Pes, A. M. O.</mods:namePart>
</mods:name>
<mods:typeOfResource id="F156CD973CE75289E9056B92E64B0341">text</mods:typeOfResource>
<mods:relatedItem id="8821D4FC12A51091844F649DE2338579" type="host">
<mods:titleInfo id="8FA5E07FA586AB74FD907D711BA38696">
<mods:title id="CB12032A62A10388674AFED59CB6D593">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="076C1C516A2B20C1E9403019E77A0CE2">
<mods:date id="F27EE523689F0C1836A2D6EFAEFF28AF">2014</mods:date>
<mods:detail id="49F399178941E51CEC3587025383D391" type="volume">
<mods:number id="9982700D855AE3AD60E55345B1D5B9E2">3796</mods:number>
</mods:detail>
<mods:detail id="003C692E16EE186E8885FF1D0D2F3A40" type="issue">
<mods:number id="9E5DF5E680C2E86D05B5A1E9BA1757FD">3</mods:number>
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<mods:start id="C3E4CA483E374AAD099D9CFB9718A2BB">545</mods:start>
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</mods:extent>
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<mods:classification id="D54BDCEF2D32AFB7FD71BD0DA0AF2F94">journal article</mods:classification>
<mods:identifier id="A9C7D5D2AA6B2639FD3FBCEA2DFDD0D4" type="CLB-Dataset">45740</mods:identifier>
<mods:identifier id="893D43027B636B0AE648A28569FCC734" type="DOI">10.11646/zootaxa.3796.3.8</mods:identifier>
<mods:identifier id="15AEB47C6165878EBE1E04D80B16FB4C" type="GBIF-Dataset">e6f90dd2-610e-4278-8cd6-b6e4c52d2574</mods:identifier>
<mods:identifier id="A14DE8254D49AEE93307FF03EDE02C92" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="13D8B82165FB6AC240587C2A8BA71274" type="Zenodo-Dep">230100</mods:identifier>
<mods:identifier id="02B7E047823455626C7EE7CE0A291EB1" type="ZooBank">5DC6D761-D857-49B8-A88C-29DEC44E30F7</mods:identifier>
</mods:mods>
<treatment id="03F55C2CE31D1C78FF2F3F31FA1FFD14" ID-DOI="http://doi.org/10.5281/zenodo.6136547" ID-GBIF-Taxon="119427332" ID-Zenodo-Dep="6136547" LSID="urn:lsid:plazi:treatment:03F55C2CE31D1C78FF2F3F31FA1FFD14" httpUri="http://treatment.plazi.org/id/03F55C2CE31D1C78FF2F3F31FA1FFD14" lastPageId="9" lastPageNumber="554" pageId="6" pageNumber="551">
<subSubSection id="C346BEB1E31D1C77FF2F3F31FE8DFD87" pageId="6" pageNumber="551" type="nomenclature">
<paragraph id="8BE3ED3AE31D1C77FF2F3F31FD23FEDA" blockId="6.[151,686,332,391]" box="[151,686,332,358]" pageId="6" pageNumber="551">
<heading id="D0AB5A56E31D1C77FF2F3F31FD23FEDA" bold="true" box="[151,686,332,358]" fontSize="11" level="1" pageId="6" pageNumber="551" reason="1">
<emphasis id="B9283128E31D1C77FF2F3F31FD23FEDA" bold="true" box="[151,686,332,358]" pageId="6" pageNumber="551">
<taxonomicName id="4C5C96B9E31D1C77FF2F3F31FE57FEDA" box="[151,474,332,358]" class="Insecta" family="Baetidae" genus="Camelobaetidius" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B9283128E31D1C77FF2F3F31FE57FEDA" bold="true" box="[151,474,332,358]" italics="true" pageId="6" pageNumber="551">Camelobaetidius carolinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B8C53E31D1C77FE593F31FDB2FEDA" box="[481,575,332,358]" pageId="6" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
Boldrini
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE3ED3AE31D1C77FF2F3F13FEC7FE3B" blockId="6.[151,686,332,391]" box="[151,330,366,391]" pageId="6" pageNumber="551">
(
<figureCitation id="1367F1BFE31D1C77FF273F13FF62FE3B" box="[159,239,366,391]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1968,1991]" captionTargetBox="[182,1401,193,1933]" captionTargetId="figure@2.[175,1402,193,1938]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Camelobaetidius, nymphs (d. v.). 1, C. baumgardneri sp. nov. 2, C. carolinae sp. nov. 3, C. cruzi sp. nov. 4, C. matilei." httpUri="https://zenodo.org/record/230101/files/figure.png" pageId="6" pageNumber="551">Figs. 2</figureCitation>
; 18–27)
</paragraph>
<paragraph id="8BE3ED3AE31D1C77FF2F3FCBFE8DFD87" blockId="6.[151,1437,438,2011]" pageId="6" pageNumber="551">
<emphasis id="B9283128E31D1C77FF2F3FCBFE89FE73" bold="true" box="[151,260,438,463]" pageId="6" pageNumber="551">Diagnose</emphasis>
.
<emphasis id="B9283128E31D1C77FEAC3FCBFEF5FE73" bold="true" box="[276,376,438,463]" pageId="6" pageNumber="551">Nymphs</emphasis>
: 1) Labrum narrowly rounded anteriorly (
<figureCitation id="1367F1BFE31D1C77FCE73FCBFC38FE73" box="[863,949,438,463]" captionStart="FIGURES 18 – 23" captionStartId="7.[151,264,1958,1981]" captionTargetBox="[212,1375,202,1931]" captionTargetId="figure@7.[200,1387,193,1938]" captionTargetPageId="7" captionText="FIGURES 18 – 23. Camelobaetidius carolinae sp. nov., nymph. 18, labrum (left v. v., right d. v.). 19, left mandible. 20, right mandible. 21, hypopharynx. 22, maxilla. 23, Labium (left v. v., right d. v.)." httpUri="https://zenodo.org/record/230105/files/figure.png" pageId="6" pageNumber="551">Fig. 18</figureCitation>
); 2) segment II of labial palp with a short rounded distomedial projection (
<figureCitation id="1367F1BFE31D1C77FDA83FA7FDEAFE4F" box="[528,615,474,499]" captionStart="FIGURES 18 – 23" captionStartId="7.[151,264,1958,1981]" captionTargetBox="[212,1375,202,1931]" captionTargetId="figure@7.[200,1387,193,1938]" captionTargetPageId="7" captionText="FIGURES 18 – 23. Camelobaetidius carolinae sp. nov., nymph. 18, labrum (left v. v., right d. v.). 19, left mandible. 20, right mandible. 21, hypopharynx. 22, maxilla. 23, Labium (left v. v., right d. v.)." httpUri="https://zenodo.org/record/230105/files/figure.png" pageId="6" pageNumber="551">Fig. 23</figureCitation>
); 3) thoracic gill absent at the base of coxae; 4) tarsal claws with six denticles, first denticle distinctly longer than others (
<figureCitation id="1367F1BFE31D1C77FD4D3F83FCC6FDAB" box="[757,843,510,535]" captionStart="FIGURES 24 – 27" captionStartId="8.[151,264,1958,1981]" captionTargetBox="[261,1343,213,1927]" captionTargetId="figure@8.[234,1353,193,1938]" captionTargetPageId="8" captionText="FIGURES 24 – 27. Camelobaetidius carolinae sp. nov., nymph. 24, foreleg. 24 a, detail of tarsal claw. 25, posterior margin of terga IV. 26, gill IV. 27, paraproct." httpUri="https://zenodo.org/record/230106/files/figure.png" pageId="6" pageNumber="551">Fig. 24</figureCitation>
a); 5) posterior margin of terga with rounded spines (
<figureCitation id="1367F1BFE31D1C77FF273C5FFF7EFD87" box="[159,243,546,571]" captionStart="FIGURES 24 – 27" captionStartId="8.[151,264,1958,1981]" captionTargetBox="[261,1343,213,1927]" captionTargetId="figure@8.[234,1353,193,1938]" captionTargetPageId="8" captionText="FIGURES 24 – 27. Camelobaetidius carolinae sp. nov., nymph. 24, foreleg. 24 a, detail of tarsal claw. 25, posterior margin of terga IV. 26, gill IV. 27, paraproct." httpUri="https://zenodo.org/record/230106/files/figure.png" pageId="6" pageNumber="551">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C346BEB1E31D1C78FF7F3C3BFB0CFF44" lastPageId="9" lastPageNumber="554" pageId="6" pageNumber="551" type="description">
<paragraph id="8BE3ED3AE31D1C77FF7F3C3BFC43FD3E" blockId="6.[151,1437,438,2011]" pageId="6" pageNumber="551">
<emphasis id="B9283128E31D1C77FF7F3C3BFE3AFDE3" bold="true" box="[199,439,582,607]" pageId="6" pageNumber="551">Description. Nymph</emphasis>
. Length of body: 5.0 mm; length of antennae:
<quantity id="4CA440DFE31D1C77FC793C3AFB94FDE2" box="[961,1049,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="551" unit="mm" value="1.3">1.3 mm</quantity>
; length of cerci:
<quantity id="4CA440DFE31D1C77FB6C3C3AFAA0FDE2" box="[1236,1325,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="551" unit="mm" value="1.8">1.8 mm</quantity>
; length of terminal filament: 1.6; tibia I:
<quantity id="4CA440DFE31D1C77FE523C16FDCEFD3E" box="[490,579,619,643]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="551" unit="mm" value="0.8">0.8 mm</quantity>
; tibia II:
<quantity id="4CA440DFE31D1C77FD103C17FC8CFD3E" box="[680,769,618,643]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="551" unit="mm" value="0.6">0.6 mm</quantity>
; tibia III:
<quantity id="4CA440DFE31D1C77FCC93C17FC44FD3E" box="[881,969,618,643]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="551" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
<paragraph id="8BE3ED3AE31D1C77FF7F3CF3FB62FD1B" blockId="6.[151,1437,438,2011]" box="[199,1263,654,679]" pageId="6" pageNumber="551">
<emphasis id="B9283128E31D1C77FF7F3CF3FE8AFD1B" bold="true" box="[199,263,654,679]" pageId="6" pageNumber="551">Head</emphasis>
(
<figureCitation id="1367F1BFE31D1C77FEAE3CF3FED6FD1B" box="[278,347,654,679]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1968,1991]" captionTargetBox="[182,1401,193,1933]" captionTargetId="figure@2.[175,1402,193,1938]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Camelobaetidius, nymphs (d. v.). 1, C. baumgardneri sp. nov. 2, C. carolinae sp. nov. 3, C. cruzi sp. nov. 4, C. matilei." httpUri="https://zenodo.org/record/230101/files/figure.png" pageId="6" pageNumber="551">Fig. 2</figureCitation>
). Light brown. Antennae light brown. Lateral branch of epicranial suture straight.
</paragraph>
<paragraph id="8BE3ED3AE31D1C77FF7F3CCFFDCEFC8B" blockId="6.[151,1437,438,2011]" pageId="6" pageNumber="551">
Labrum (
<figureCitation id="1367F1BFE31D1C77FE8D3CCFFE03FD77" box="[309,398,690,715]" captionStart="FIGURES 18 – 23" captionStartId="7.[151,264,1958,1981]" captionTargetBox="[212,1375,202,1931]" captionTargetId="figure@7.[200,1387,193,1938]" captionTargetPageId="7" captionText="FIGURES 18 – 23. Camelobaetidius carolinae sp. nov., nymph. 18, labrum (left v. v., right d. v.). 19, left mandible. 20, right mandible. 21, hypopharynx. 22, maxilla. 23, Labium (left v. v., right d. v.)." httpUri="https://zenodo.org/record/230105/files/figure.png" pageId="6" pageNumber="551">Fig. 18</figureCitation>
). Narrowly rounded anteriorly, broader than long. Length about 0.7× maximum width; anterolateral margin with long, fine, bifid and pectinate setae; medially with fine, bifid and pectinate setae. Dorsally with three short, fine, simple setae scattered over basal area; dorsolateral arc of setae with one long, fine, apically pointed seta on lateral region.
</paragraph>
<paragraph id="8BE3ED3AE31D1C77FF7F3D3FFD85FC1E" blockId="6.[151,1437,438,2011]" pageId="6" pageNumber="551">
Left mandible (
<figureCitation id="1367F1BFE31D1C77FECD3D3FFE44FCE7" box="[373,457,834,859]" captionStart="FIGURES 18 – 23" captionStartId="7.[151,264,1958,1981]" captionTargetBox="[212,1375,202,1931]" captionTargetId="figure@7.[200,1387,193,1938]" captionTargetPageId="7" captionText="FIGURES 18 – 23. Camelobaetidius carolinae sp. nov., nymph. 18, labrum (left v. v., right d. v.). 19, left mandible. 20, right mandible. 21, hypopharynx. 22, maxilla. 23, Labium (left v. v., right d. v.)." httpUri="https://zenodo.org/record/230105/files/figure.png" pageId="6" pageNumber="551">Fig. 19</figureCitation>
). Incisors with seven denticles. Prostheca robust, apically denticulate. Margin between prostheca and mola with three spine-like setae; subtriangular process wide; setae at apex of mola absent. Basal half with six short, fine, simple setae.
</paragraph>
<paragraph id="8BE3ED3AE31D1C77FF7F3DD3FEEFFBB2" blockId="6.[151,1437,438,2011]" pageId="6" pageNumber="551">
Right mandible (
<figureCitation id="1367F1BFE31D1C77FE283DD3FE66FC7B" box="[400,491,942,967]" captionStart="FIGURES 18 – 23" captionStartId="7.[151,264,1958,1981]" captionTargetBox="[212,1375,202,1931]" captionTargetId="figure@7.[200,1387,193,1938]" captionTargetPageId="7" captionText="FIGURES 18 – 23. Camelobaetidius carolinae sp. nov., nymph. 18, labrum (left v. v., right d. v.). 19, left mandible. 20, right mandible. 21, hypopharynx. 22, maxilla. 23, Labium (left v. v., right d. v.)." httpUri="https://zenodo.org/record/230105/files/figure.png" pageId="6" pageNumber="551">Fig. 20</figureCitation>
). Incisors with eleven denticles. Prostheca slender, apically denticulate; margin between prostheca and mola with four spine-like setae; setae at apex of mola present. Basal half with three short, fine, simple setae.
</paragraph>
<paragraph id="8BE3ED3AE31D1C77FF7F3A67FE91FBC7" blockId="6.[151,1437,438,2011]" pageId="6" pageNumber="551">
Hypopharynx (
<figureCitation id="1367F1BFE31D1C77FECA3A67FE4BFB8F" box="[370,454,1050,1075]" captionStart="FIGURES 18 – 23" captionStartId="7.[151,264,1958,1981]" captionTargetBox="[212,1375,202,1931]" captionTargetId="figure@7.[200,1387,193,1938]" captionTargetPageId="7" captionText="FIGURES 18 – 23. Camelobaetidius carolinae sp. nov., nymph. 18, labrum (left v. v., right d. v.). 19, left mandible. 20, right mandible. 21, hypopharynx. 22, maxilla. 23, Labium (left v. v., right d. v.)." httpUri="https://zenodo.org/record/230105/files/figure.png" pageId="6" pageNumber="551">Fig. 21</figureCitation>
). Lingua shorter than superlingua, apex rounded, with tuft of short, fine, simple setae. Superlingua not expanded, with fine, simple setae scattered over lateral and distal margin, external margin cleft subapically.
</paragraph>
<paragraph id="8BE3ED3AE31D1C77FF7F3AFBFD32FAB7" blockId="6.[151,1437,438,2011]" pageId="6" pageNumber="551">
Maxilla (
<figureCitation id="1367F1BFE31D1C77FE953AFBFE0CFB23" box="[301,385,1158,1183]" captionStart="FIGURES 18 – 23" captionStartId="7.[151,264,1958,1981]" captionTargetBox="[212,1375,202,1931]" captionTargetId="figure@7.[200,1387,193,1938]" captionTargetPageId="7" captionText="FIGURES 18 – 23. Camelobaetidius carolinae sp. nov., nymph. 18, labrum (left v. v., right d. v.). 19, left mandible. 20, right mandible. 21, hypopharynx. 22, maxilla. 23, Labium (left v. v., right d. v.)." httpUri="https://zenodo.org/record/230105/files/figure.png" pageId="6" pageNumber="551">Fig. 22</figureCitation>
). Inner dorsal row of setae with two denti-setae, one simple denti-seta and one pectinate dentiseta, apical denti-seta at the same orientation of canines. Medial protuberance of galea with 1 + 3 long, fine and simple setae. Maxillary palp 1.1× length of galea-lacinia. Palp segment II 0.8× length of segment I. Fine and simple setae scattered over segment II of maxillary palp.
</paragraph>
<paragraph id="8BE3ED3AE31D1C77FF7F3B6BFB38FA5E" blockId="6.[151,1437,438,2011]" pageId="6" pageNumber="551">
Labium (
<figureCitation id="1367F1BFE31D1C77FE973B6BFE08FA93" box="[303,389,1302,1327]" captionStart="FIGURES 18 – 23" captionStartId="7.[151,264,1958,1981]" captionTargetBox="[212,1375,202,1931]" captionTargetId="figure@7.[200,1387,193,1938]" captionTargetPageId="7" captionText="FIGURES 18 – 23. Camelobaetidius carolinae sp. nov., nymph. 18, labrum (left v. v., right d. v.). 19, left mandible. 20, right mandible. 21, hypopharynx. 22, maxilla. 23, Labium (left v. v., right d. v.)." httpUri="https://zenodo.org/record/230105/files/figure.png" pageId="6" pageNumber="551">Fig. 23</figureCitation>
). Glossa shorter than paraglossa; inner margin with six spine-like setae increasing in length distally; apex with two long, spine-like setae; outer margin with three long, spine-like setae; ventral and dorsal surface bare. Paraglossa sub-rectangular, curved inward; apex with two rows of long, fine, simple setae; ventral surface with a row of four long, fine, simple setae; dorsal surface with two fine, spine-like setae near to apex. Labial palp segment II with distomedial projection short and rounded; dorsally with a row of four long, spine-like setae; segment III oblong, length 1.2× width, scattered with simple setae, and spine-like setae.
</paragraph>
<paragraph id="8BE3ED3AE31D1C77FF7F3B93FEE2F9CE" blockId="6.[151,1437,438,2011]" pageId="6" pageNumber="551">
<emphasis id="B9283128E31D1C77FF7F3B93FEADF9BB" bold="true" box="[199,288,1518,1543]" pageId="6" pageNumber="551">Thorax</emphasis>
(
<figureCitation id="1367F1BFE31D1C77FE893B93FEF5F9BB" box="[305,376,1518,1543]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1968,1991]" captionTargetBox="[182,1401,193,1933]" captionTargetId="figure@2.[175,1402,193,1938]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Camelobaetidius, nymphs (d. v.). 1, C. baumgardneri sp. nov. 2, C. carolinae sp. nov. 3, C. cruzi sp. nov. 4, C. matilei." httpUri="https://zenodo.org/record/230101/files/figure.png" pageId="6" pageNumber="551">Fig. 2</figureCitation>
). Light brown; metanotum with anterior region brown.
<collectingCountry id="F34BADAAE31D1C77FC4E3B93FBA3F9BB" box="[1014,1070,1518,1543]" name="India" pageId="6" pageNumber="551">Hind</collectingCountry>
wing pad present. Thoracic gill absent at the base of coxae. Femur, tibia and tarsi light brown. Tibia I subequal in length to femur I, tibia II 0.8× length of femur II, tibia III 0.7× length of femur III; tarsi I and II 0.4× length of the respective femur, tarsi III 0.3× length of femur III.
</paragraph>
<paragraph id="8BE3ED3AE31D1C77FF7F3803FEC1F8F6" blockId="6.[151,1437,438,2011]" pageId="6" pageNumber="551">
Forefemur (
<figureCitation id="1367F1BFE31D1C77FEF53803FE2FF92B" box="[333,418,1662,1687]" captionStart="FIGURES 24 – 27" captionStartId="8.[151,264,1958,1981]" captionTargetBox="[261,1343,213,1927]" captionTargetId="figure@8.[234,1353,193,1938]" captionTargetPageId="8" captionText="FIGURES 24 – 27. Camelobaetidius carolinae sp. nov., nymph. 24, foreleg. 24 a, detail of tarsal claw. 25, posterior margin of terga IV. 26, gill IV. 27, paraproct." httpUri="https://zenodo.org/record/230106/files/figure.png" pageId="6" pageNumber="551">Fig. 24</figureCitation>
). Length about 3.7× maximum width; dorsally with a row of c.a. 18 long, spine-like setae; ventrally with five short, fine spine-like setae. Foretibia. Dorsally with few fine, simple setae; anterior surface with four short, spine-like setae; patella-tibial suture present, 0.4× length of tibia. Tarsal claws with six denticles, first denticle distinctly longer than others (
<figureCitation id="1367F1BFE31D1C77FDF83897FD1EF8BF" box="[576,659,1770,1795]" captionStart="FIGURES 24 – 27" captionStartId="8.[151,264,1958,1981]" captionTargetBox="[261,1343,213,1927]" captionTargetId="figure@8.[234,1353,193,1938]" captionTargetPageId="8" captionText="FIGURES 24 – 27. Camelobaetidius carolinae sp. nov., nymph. 24, foreleg. 24 a, detail of tarsal claw. 25, posterior margin of terga IV. 26, gill IV. 27, paraproct." httpUri="https://zenodo.org/record/230106/files/figure.png" pageId="6" pageNumber="551">Fig. 24</figureCitation>
a). Mid femur. Length about 3.4× maximum width; dorsally with row of c.a. 13 long, spine-like setae.
<collectingCountry id="F34BADAAE31D1C77FDBC3973FDB1F89B" box="[516,572,1806,1831]" name="India" pageId="6" pageNumber="551">Hind</collectingCountry>
femur. Length about 2.9× maximum width; dorsally with row of c.a. 8 long, spine-like setae.
</paragraph>
<paragraph id="8BE3ED3AE31D1C78FF7F392BFB0CFF44" blockId="6.[151,1437,438,2011]" lastBlockId="9.[151,1437,151,680]" lastPageId="9" lastPageNumber="554" pageId="6" pageNumber="551">
<emphasis id="B9283128E31D1C77FF7F392BFEB1F8D3" bold="true" box="[199,316,1878,1903]" pageId="6" pageNumber="551">Abdomen</emphasis>
(
<figureCitation id="1367F1BFE31D1C77FEF2392BFE03F8D3" box="[330,398,1878,1903]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1968,1991]" captionTargetBox="[182,1401,193,1933]" captionTargetId="figure@2.[175,1402,193,1938]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Camelobaetidius, nymphs (d. v.). 1, C. baumgardneri sp. nov. 2, C. carolinae sp. nov. 3, C. cruzi sp. nov. 4, C. matilei." httpUri="https://zenodo.org/record/230101/files/figure.png" pageId="6" pageNumber="551">Fig. 2</figureCitation>
). Light brown, posterolateral corners of segments II–III and V–VI brown. Terga with creased surface, posterior margin of tergum IV with rounded spines (
<figureCitation id="1367F1BFE31D1C77FCEA3907FC25F82F" box="[850,936,1914,1939]" captionStart="FIGURES 24 – 27" captionStartId="8.[151,264,1958,1981]" captionTargetBox="[261,1343,213,1927]" captionTargetId="figure@8.[234,1353,193,1938]" captionTargetPageId="8" captionText="FIGURES 24 – 27. Camelobaetidius carolinae sp. nov., nymph. 24, foreleg. 24 a, detail of tarsal claw. 25, posterior margin of terga IV. 26, gill IV. 27, paraproct." httpUri="https://zenodo.org/record/230106/files/figure.png" pageId="6" pageNumber="551">Fig. 25</figureCitation>
). Gills oval. Gills IV (
<figureCitation id="1367F1BFE31D1C77FB143907FB72F82F" box="[1196,1279,1914,1939]" captionStart="FIGURES 24 – 27" captionStartId="8.[151,264,1958,1981]" captionTargetBox="[261,1343,213,1927]" captionTargetId="figure@8.[234,1353,193,1938]" captionTargetPageId="8" captionText="FIGURES 24 – 27. Camelobaetidius carolinae sp. nov., nymph. 24, foreleg. 24 a, detail of tarsal claw. 25, posterior margin of terga IV. 26, gill IV. 27, paraproct." httpUri="https://zenodo.org/record/230106/files/figure.png" pageId="6" pageNumber="551">Fig. 26</figureCitation>
) light brown; margins with narrow spines intercalating short, fine, simple setae; tracheae pigmented. Gill I 1.2× length of segment II; gill IV 1.9× length of segment V; gill VII 1.2× length of segment VIII. Paraproct (
<figureCitation id="1367F1BFE31D1C77FB0C39BFFA84F867" box="[1204,1289,1986,2011]" captionStart="FIGURES 24 – 27" captionStartId="8.[151,264,1958,1981]" captionTargetBox="[261,1343,213,1927]" captionTargetId="figure@8.[234,1353,193,1938]" captionTargetPageId="8" captionText="FIGURES 24 – 27. Camelobaetidius carolinae sp. nov., nymph. 24, foreleg. 24 a, detail of tarsal claw. 25, posterior margin of terga IV. 26, gill IV. 27, paraproct." httpUri="https://zenodo.org/record/230106/files/figure.png" pageId="6" pageNumber="551">Fig. 27</figureCitation>
). With three spines near inner margin; without shagreened area; postero-lateral extension with marginal spines. Cerci light brown, with a brown band at mid length; outer and inner margin of terminal filament and inner margin of cerci with tufts of long, simple setae; outer margin of the cerci with spines at apex on each segment.
</paragraph>
</subSubSection>
<caption id="DF23BDB2E31C1C76FF2F39DBFCF0F860" httpUri="https://zenodo.org/record/230105/files/figure.png" pageId="7" pageNumber="552" targetBox="[212,1375,202,1931]" targetPageId="7">
<paragraph id="8BE3ED3AE31C1C76FF2F39DBFCF0F860" blockId="7.[151,1436,1958,2012]" pageId="7" pageNumber="552">
<emphasis id="B9283128E31C1C76FF2F39DBFEDBF800" bold="true" box="[151,342,1958,1981]" pageId="7" pageNumber="552">FIGURES 18–23.</emphasis>
<taxonomicName id="4C5C96B9E31C1C76FEE639DAFDFFF801" box="[350,626,1959,1981]" class="Insecta" family="Baetidae" genus="Camelobaetidius" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B9283128E31C1C76FEE639DAFDFFF801" box="[350,626,1959,1981]" italics="true" pageId="7" pageNumber="552">Camelobaetidius carolinae</emphasis>
</taxonomicName>
<emphasis id="B9283128E31C1C76FDC339DAFD43F800" bold="true" box="[635,718,1959,1980]" pageId="7" pageNumber="552">
<taxonomicNameLabel id="A21B8C53E31C1C76FDC339DAFD43F800" box="[635,718,1959,1980]" pageId="7" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, nymph. 18, labrum (left v.v., right d.v.). 19, left mandible. 20, right mandible. 21, hypopharynx. 22, maxilla. 23, Labium (left v.v., right d.v.).
</paragraph>
</caption>
<caption id="DF23BDB2E3131C79FF2F39DBFE7FF860" httpUri="https://zenodo.org/record/230106/files/figure.png" pageId="8" pageNumber="553" targetBox="[261,1343,213,1927]" targetPageId="8">
<paragraph id="8BE3ED3AE3131C79FF2F39DBFE7FF860" blockId="8.[151,1436,1958,2012]" pageId="8" pageNumber="553">
<emphasis id="B9283128E3131C79FF2F39DBFED8F800" bold="true" box="[151,341,1958,1981]" pageId="8" pageNumber="553">FIGURES 24–27.</emphasis>
<taxonomicName id="4C5C96B9E3131C79FEE439DAFDE2F801" box="[348,623,1959,1981]" class="Insecta" family="Baetidae" genus="Camelobaetidius" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B9283128E3131C79FEE439DAFDE2F801" box="[348,623,1959,1981]" italics="true" pageId="8" pageNumber="553">Camelobaetidius carolinae</emphasis>
</taxonomicName>
<emphasis id="B9283128E3131C79FDCE39DAFD45F800" bold="true" box="[630,712,1959,1980]" pageId="8" pageNumber="553">
<taxonomicNameLabel id="A21B8C53E3131C79FDCE39DAFD45F800" box="[630,712,1959,1980]" pageId="8" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, nymph. 24, foreleg. 24a, detail of tarsal claw. 25, posterior margin of terga IV. 26, gill IV. 27, paraproct.
</paragraph>
</caption>
<subSubSection id="C346BEB1E3121C78FF7F3F79FD5CFEFC" pageId="9" pageNumber="554" type="etymology">
<paragraph id="8BE3ED3AE3121C78FF7F3F79FD5CFEFC" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="554">
<emphasis id="B9283128E3121C78FF7F3F79FEC5FEA1" bold="true" box="[199,328,260,285]" pageId="9" pageNumber="554">Etymology</emphasis>
: The name of the species is in honor of Dr.
<taxonomicName id="4C5C96B9E3121C78FC823F79FC6EFEA1" authority="Nieto" authorityName="Nieto" box="[826,995,260,285]" class="Insecta" family="Scarabaeidae" genus="Carolina" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="genus">Carolina Nieto</taxonomicName>
, for her significant contributions to the knowledge of
<taxonomicName id="4C5C96B9E3121C78FE8F3F5AFE68FEFC" box="[311,485,295,320]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="order">Ephemeroptera</taxonomicName>
, especially
<taxonomicName id="4C5C96B9E3121C78FDDF3F5AFD40FEFC" box="[615,717,295,320]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C346BEB1E3121C78FF7F3F31FDC2FED9" box="[199,591,332,357]" pageId="9" pageNumber="554" type="distribution">
<paragraph id="8BE3ED3AE3121C78FF7F3F31FDC2FED9" blockId="9.[151,1437,151,680]" box="[199,591,332,357]" pageId="9" pageNumber="554">
<emphasis id="B9283128E3121C78FF7F3F31FED4FED9" bold="true" box="[199,345,332,357]" pageId="9" pageNumber="554">Distribution</emphasis>
.
<collectingCountry id="F34BADAAE3121C78FEDF3F31FE5CFED8" box="[359,465,332,356]" name="Brazil" pageId="9" pageNumber="554">BRAZIL</collectingCountry>
: Rondônia.
</paragraph>
</subSubSection>
<subSubSection id="C346BEB1E3121C78FF7F3F12FCD6FE48" pageId="9" pageNumber="554" type="discussion">
<paragraph id="8BE3ED3AE3121C78FF7F3F12FCD6FE48" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="554">
<emphasis id="B9283128E3121C78FF7F3F12FEC5FE34" bold="true" box="[199,328,367,392]" pageId="9" pageNumber="554">Comments</emphasis>
. The general shape of the labial palp of
<taxonomicName id="4C5C96B9E3121C78FCB13F0DFC19FE34" box="[777,916,368,392]" class="Insecta" family="Baetidae" genus="Camelobaetidius" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B9283128E3121C78FCB13F0DFC19FE34" box="[777,916,368,392]" italics="true" pageId="9" pageNumber="554">C. carolinae</emphasis>
</taxonomicName>
<emphasis id="B9283128E3121C78FC233F12FC79FE34" bold="true" box="[923,1012,367,392]" pageId="9" pageNumber="554">
<taxonomicNameLabel id="A21B8C53E3121C78FC233F12FC79FE34" box="[923,1012,367,392]" pageId="9" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to that found in
<taxonomicName id="4C5C96B9E3121C78FAA33F0DFAF9FE3B" box="[1307,1396,368,391]" class="Insecta" family="Baetidae" genus="Camelobaetidius" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="cruzi" status="sp. nov.">
<emphasis id="B9283128E3121C78FAA33F0DFAF9FE3B" box="[1307,1396,368,391]" italics="true" pageId="9" pageNumber="554">C. cruzi</emphasis>
</taxonomicName>
<emphasis id="B9283128E3121C78FAC23F12FF45FE11" bold="true" pageId="9" pageNumber="554">
<taxonomicNameLabel id="A21B8C53E3121C78FAC23F12FF45FE11" pageId="9" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C5C96B9E3121C78FF453FE9FDE6FE10" authority="Thomas" authorityName="Thomas" authorityYear="2003" box="[253,619,404,429]" class="Insecta" family="Baetidae" genus="Camelobaetidius" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="matilei">
<emphasis id="B9283128E3121C78FF453FE9FEE7FE10" box="[253,362,404,428]" italics="true" pageId="9" pageNumber="554">C. matilei</emphasis>
<bibRefCitation id="EFCD90CBE3121C78FEC93FE9FDE6FE10" author="Thomas" box="[369,619,404,429]" pageId="9" pageNumber="554" refString="Thomas, A., Peru, N. &amp; Horeau, V. (2003) Les Ephemeres de la Guyane Francaise. 8. Description de Camelobaetidius matilei n. sp. et cle de determination de especes guyanensis de ce genre [Ephemeroptera: Baetidae]. Ephemera, 3 (2), 123 - 133. Traver, J. R. &amp; Edmunds, G. F. Jr. (1968) A revision of the Baetidae with spatulate-clawed nymphs (Ephemeroptera). Pacific Insects, (10), 629 - 677." type="journal article" year="2003">
Thomas &amp;
<collectingCountry id="F34BADAAE3121C78FE563FE9FDA9FE10" box="[494,548,404,428]" name="Peru" pageId="9" pageNumber="554">Péru</collectingCountry>
, 2003
</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C5C96B9E3121C78FDCC3FE9FC2EFE10" box="[628,931,404,428]" class="Insecta" family="Baetidae" genus="Camelobaetidius" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B9283128E3121C78FDCC3FE9FC2EFE10" box="[628,931,404,428]" italics="true" pageId="9" pageNumber="554">Camelobaetidius carolinae</emphasis>
</taxonomicName>
<emphasis id="B9283128E3121C78FC113FE9FB8CFE11" bold="true" box="[937,1025,404,429]" pageId="9" pageNumber="554">
<taxonomicNameLabel id="A21B8C53E3121C78FC113FE9FB8CFE11" box="[937,1025,404,429]" pageId="9" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, however, can be distinguished from these two species by the size of the denticles on the tarsal claws [first denticle distinctly longer than others (
<figureCitation id="1367F1BFE3121C78FAD73FCAFF3AFE48" captionStart="FIGURES 24 – 27" captionStartId="8.[151,264,1958,1981]" captionTargetBox="[261,1343,213,1927]" captionTargetId="figure@8.[234,1353,193,1938]" captionTargetPageId="8" captionText="FIGURES 24 – 27. Camelobaetidius carolinae sp. nov., nymph. 24, foreleg. 24 a, detail of tarsal claw. 25, posterior margin of terga IV. 26, gill IV. 27, paraproct." httpUri="https://zenodo.org/record/230106/files/figure.png" pageId="9" pageNumber="554">Fig. 24</figureCitation>
a)], and by a more complex abdominal color pattern (
<figureCitation id="1367F1BFE3121C78FCB13FA1FCC3FE48" box="[777,846,476,501]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1968,1991]" captionTargetBox="[182,1401,193,1933]" captionTargetId="figure@2.[175,1402,193,1938]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Camelobaetidius, nymphs (d. v.). 1, C. baumgardneri sp. nov. 2, C. carolinae sp. nov. 3, C. cruzi sp. nov. 4, C. matilei." httpUri="https://zenodo.org/record/230101/files/figure.png" pageId="9" pageNumber="554">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C346BEB1E3121C78FF7F3F82FA1FFD14" pageId="9" pageNumber="554" type="materials_examined">
<paragraph id="8BE3ED3AE3121C78FF7F3F82FA1FFD14" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="554">
<emphasis id="B9283128E3121C78FF7F3F82FE24FDA4" bold="true" box="[199,425,511,536]" pageId="9" pageNumber="554">Material examined</emphasis>
.
<emphasis id="B9283128E3121C78FE003F82FDA8FDA4" bold="true" box="[440,549,511,536]" pageId="9" pageNumber="554">
<typeStatus id="54E75398E3121C78FE003F82FDA8FDA4" box="[440,549,511,536]" pageId="9" pageNumber="554" type="holotype">Holotype</typeStatus>
</emphasis>
: Nymph (mounted on slides, medium Euparal),
<collectingCountry id="F34BADAAE3121C78FBFD3C7DFB20FDA4" box="[1093,1197,512,536]" name="Brazil" pageId="9" pageNumber="554">BRAZIL</collectingCountry>
, Rondônia, Colorado do Oeste, Eixo Zero, Rio Hermes,
<geoCoordinate id="EE688BFDE3121C78FD893C59FD4EFD81" box="[561,707,548,573]" direction="south" orientation="latitude" pageId="9" pageNumber="554" precision="1" value="-13.202111">13°12'07.6''S</geoCoordinate>
/
<geoCoordinate id="EE688BFDE3121C78FD6F3C59FCF9FD81" box="[727,884,548,573]" direction="west" orientation="longitude" pageId="9" pageNumber="554" precision="1" value="-60.39031">60°23'25.1''W</geoCoordinate>
,
<date id="FFE2CBFAE3121C78FC3B3C59FC73FD81" box="[899,1022,548,573]" pageId="9" pageNumber="554" value="2012-09-06">06.ix.2012</date>
, Boldrini, R., Nascimentos, J.M.C., Fernandes, A.S., Hamada, N. leg (
<collectionCode id="ED4D75FFE3121C78FD823C35FD0DFDDC" box="[570,640,584,608]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="9" pageNumber="554">INPA</collectionCode>
).
<typeStatus id="54E75398E3121C78FD213C3AFC9CFDDC" box="[665,785,583,608]" pageId="9" pageNumber="554" type="paratype">
<emphasis id="B9283128E3121C78FD213C3AFC9CFDDC" bold="true" box="[665,785,583,608]" pageId="9" pageNumber="554">Paratypes</emphasis>
</typeStatus>
: Two immature nymphs (mounted on slides, medium Euparal),
<collectingCountry id="F34BADAAE3121C78FEB13C11FEFCFD38" box="[265,369,620,644]" name="Brazil" pageId="9" pageNumber="554">BRAZIL</collectingCountry>
, Rondônia, Colorado do Oeste, Rio Cabixi, below the bridge,
<geoCoordinate id="EE688BFDE3121C78FBEC3C11FB68FD39" box="[1108,1253,620,645]" direction="south" orientation="latitude" pageId="9" pageNumber="554" precision="1" value="-13.258833">13°15'31.8''S</geoCoordinate>
/
<geoCoordinate id="EE688BFDE3121C78FB423C11FA1AFD39" box="[1274,1431,620,645]" direction="west" orientation="longitude" pageId="9" pageNumber="554" precision="1" value="-60.334667">60°20'04.8''W</geoCoordinate>
,
<date id="FFE2CBFAE3121C78FF2F3CF2FE9CFD14" box="[151,273,655,680]" pageId="9" pageNumber="554" value="2012-09-06">06.ix.2012</date>
, Boldrini, R., Nascimento, J.M.C., Fernandes, A.S., Hamada, N. leg (one in
<collectionCode id="ED4D75FFE3121C78FBD23CEDFB38FD14" box="[1130,1205,656,680]" pageId="9" pageNumber="554">CZNC</collectionCode>
and one in
<collectionCode id="ED4D75FFE3121C78FA8F3CEDFA08FD14" box="[1335,1413,656,680]" pageId="9" pageNumber="554">UFRR</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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