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<document id="6188C2601C40A49DB6F51FA9512C878D" ID-DOI="10.5281/zenodo.282224" ID-GBIF-Dataset="3c9db890-c1df-4340-8be1-8e5cd631aeed" ID-ISSN="1175-5326" ID-Zenodo-Dep="282224" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460519716872" checkinUser="plazi" docAuthor="Plisko, Jadwiga Danuta" docDate="2012" docId="03F54361FF97FFBFE68BD0CBBCB96C9B" docLanguage="en" docName="zt03458p058.pdf" docOrigin="Zootaxa 3458" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Acanthodrilus sclateri Beddard 1897" docType="treatment" docVersion="8" lastPageNumber="52" masterDocId="FFCC3B19FFB8FF8FE61CD62FBD796E71" masterDocTitle="An annotated checklist of the South African Acanthodrilidae (Oligochaeta: Acanthodrilidae: Acanthodrilinae, Benhamiinae)" masterLastPageNumber="58" masterPageNumber="4" pageNumber="51" updateTime="1698314471110" updateUser="plazi">
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<mods:title id="E9B3285B89F988BBBFDD238BE10B179E">An annotated checklist of the South African Acanthodrilidae (Oligochaeta: Acanthodrilidae: Acanthodrilinae, Benhamiinae)</mods:title>
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<subSubSection id="C346A1FCFF97FFA0E68BD0CBBE5A688C" box="[151,803,1763,1790]" pageId="47" pageNumber="51" type="nomenclature">
<paragraph id="8BE3F277FF97FFA0E68BD0CBBE5A688C" blockId="47.[151,803,1763,1790]" box="[151,803,1763,1790]" pageId="47" pageNumber="51">
<heading id="D0AB451BFF97FFA0E68BD0CBBE5A688C" bold="true" box="[151,803,1763,1790]" fontSize="11" level="1" pageId="47" pageNumber="51" reason="1">
<emphasis id="B9282E65FF97FFA0E68BD0CBBE5A688C" bold="true" box="[151,803,1763,1790]" pageId="47" pageNumber="51">
<taxonomicName id="4C5C89F4FF97FFA0E68BD0CBBF26688C" authority="Beddard, 1897" authorityName="Beddard" authorityYear="1897" box="[151,607,1763,1790]" class="Clitellata" family="Acanthodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="47" pageNumber="51" phylum="Annelida" rank="species" species="sclateri">
<emphasis id="B9282E65FF97FFA0E68BD0CBBCDA688F" bold="true" box="[151,419,1764,1790]" italics="true" pageId="47" pageNumber="51">Acanthodrilus sclateri</emphasis>
<bibRefCitation id="EFCD8F86FF97FFA0E7B6D0CCBF26688C" author="Beddard" box="[426,607,1763,1789]" pageId="47" pageNumber="51" refString="Beddard, F. E. (1897) On a collection of earthworms from South Africa belonging to the genus Acanthodrilus. With an Appendix: on a new genus of earthworms belonging to the family Eudrilidae. Proceedings of the Zoological Society of London, 1897, 336 - 349." type="proceedings paper" year="1897">Beddard, 1897</bibRefCitation>
</taxonomicName>
—species dubia
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C346A1FCFF97FFA0E68BD105BE7269CA" pageId="47" pageNumber="51" type="reference_group">
<paragraph id="8BE3F277FF97FFA0E68BD105BF376931" blockId="47.[151,977,1834,1979]" box="[151,590,1834,1856]" pageId="47" pageNumber="51">
<treatmentCitationGroup id="AB4CD559FF97FFA0E68BD105BF376931" box="[151,590,1834,1856]" pageId="47" pageNumber="51">
<taxonomicName id="4C5C89F4FF97FFA0E68BD105BF336931" authority="Beddard, 1897: 324" authorityName="Beddard" authorityPageNumber="324" authorityYear="1897" box="[151,586,1834,1856]" class="Clitellata" family="Acanthodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="47" pageNumber="51" phylum="Annelida" rank="species" species="sclateri">
<emphasis id="B9282E65FF97FFA0E68BD105BC016931" box="[151,376,1834,1856]" italics="true" pageId="47" pageNumber="51">Acanthodrilus sclateri</emphasis>
<treatmentCitation id="0AFDD466FF97FFA0E763D105BF336931" author="Beddard" box="[383,586,1834,1856]" page="324" pageId="47" pageNumber="51" year="1897">
<bibRefCitation id="EFCD8F86FF97FFA0E763D105BF6E6931" author="Beddard" box="[383,535,1834,1856]" pageId="47" pageNumber="51" refString="Beddard, F. E. (1897) On a collection of earthworms from South Africa belonging to the genus Acanthodrilus. With an Appendix: on a new genus of earthworms belonging to the family Eudrilidae. Proceedings of the Zoological Society of London, 1897, 336 - 349." type="proceedings paper" year="1897">Beddard, 1897</bibRefCitation>
: 324
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE3F277FF97FFA0E68BD166BF53692E" blockId="47.[151,977,1834,1979]" box="[151,554,1865,1887]" pageId="47" pageNumber="51">
<treatmentCitationGroup id="AB4CD559FF97FFA0E68BD166BF53692E" box="[151,554,1865,1887]" pageId="47" pageNumber="51">
<taxonomicName id="4C5C89F4FF97FFA0E68BD166BF5F692E" authority="Michaelsen 1900: 148" authorityName="Michaelsen" authorityPageNumber="148" authorityYear="1900" box="[151,550,1865,1887]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="47" pageNumber="51" phylum="Annelida" rank="species" species="sclateri">
<emphasis id="B9282E65FF97FFA0E68BD166BC4C692E" box="[151,309,1865,1887]" italics="true" pageId="47" pageNumber="51">Chilota sclateri</emphasis>
:
<treatmentCitation id="0AFDD466FF97FFA0E75ED166BF5F692E" author="Michaelsen" box="[322,550,1865,1887]" page="148" pageId="47" pageNumber="51" year="1900">
<bibRefCitation id="EFCD8F86FF97FFA0E75ED166BC8A692E" author="Michaelsen" box="[322,499,1865,1887]" pageId="47" pageNumber="51" refString="Michaelsen, W. (1900) Oligochaeta. In: Das Tierreich. Lief. 10. Berlin: R. Friedlander &amp; Sohn, 575 pp." type="book" year="1900">Michaelsen 1900</bibRefCitation>
: 148
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE3F277FF97FFA0E68BD147BFC969ED" blockId="47.[151,977,1834,1979]" pageId="47" pageNumber="51">
<treatmentCitationGroup id="AB4CD559FF97FFA0E68BD147BFC969ED" pageId="47" pageNumber="51">
<taxonomicName id="4C5C89F4FF97FFA0E68BD147BCF0690F" box="[151,393,1896,1918]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="47" pageNumber="51" phylum="Annelida" rank="species" species="sclateri" subGenus="Acanthodrilus">
(
<emphasis id="B9282E65FF97FFA0E683D147BC57690F" box="[159,302,1896,1918]" italics="true" pageId="47" pageNumber="51">Acanthodrilus</emphasis>
)
<emphasis id="B9282E65FF97FFA0E721D147BCF0690F" box="[317,393,1896,1918]" italics="true" pageId="47" pageNumber="51">sclateri</emphasis>
</taxonomicName>
sp. inquir. aut spuria:
<treatmentCitation id="0AFDD466FF97FFA0E477D147BE20690F" author="Michaelsen" box="[619,857,1896,1918]" page="533" pageId="47" pageNumber="51" year="1913">
<bibRefCitation id="EFCD8F86FF97FFA0E477D147BE51690C" author="Michaelsen" box="[619,808,1896,1918]" pageId="47" pageNumber="51" refString="Michaelsen, W. (1913 a) Die Oligochaeten des Kaplandes. Zoologische Jahrbucher, Abteilung fur Systematik, 34, 473 - 556." type="journal article" year="1913" yearSuffix="a">Michaelsen 1913a</bibRefCitation>
: 533
</treatmentCitation>
, 1913b: 59.
<taxonomicName id="4C5C89F4FF97FFA0E68BD1A8BFD569ED" authority="Reynolds &amp; Cook 1976: 169" authorityName="Reynolds &amp; Cook" authorityPageNumber="169" authorityYear="1976" box="[151,684,1926,1949]" class="Clitellata" family="Acanthodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="47" pageNumber="51" phylum="Annelida" rank="species" species="sclateri">
<emphasis id="B9282E65FF97FFA0E68BD1A8BC0169EC" box="[151,376,1927,1949]" italics="true" pageId="47" pageNumber="51">Acanthodrilus sclateri</emphasis>
:
<treatmentCitation id="0AFDD466FF97FFA0E79AD1A9BFD569ED" author="Reynolds" box="[390,684,1926,1948]" page="169" pageId="47" pageNumber="51" year="1976">
<bibRefCitation id="EFCD8F86FF97FFA0E79AD1A9BF0369ED" author="Reynolds" box="[390,634,1926,1948]" pageId="47" pageNumber="51" refString="Reynolds J. W. &amp; Cook, D. G. (1976) Nomenclatura Oligochaetologica. A catalogue of names, descriptions and type specimens of the Oligochaeta. Ottawa: Runge Press. 217 pp." type="book" year="1976">Reynolds &amp; Cook 1976</bibRefCitation>
: 169
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE3F277FF97FFA0E68BD18ABE7269CA" blockId="47.[151,977,1834,1979]" box="[151,779,1957,1979]" pageId="47" pageNumber="51">
<treatmentCitationGroup id="AB4CD559FF97FFA0E68BD18ABE7269CA" box="[151,779,1957,1979]" pageId="47" pageNumber="51">
<emphasis id="B9282E65FF97FFA0E68BD18ABDD869CA" box="[151,161,1957,1979]" italics="true" pageId="47" pageNumber="51">?</emphasis>
<taxonomicName id="4C5C89F4FF97FFA0E6BCD18ABC0669CA" authority="(Parachilota)" baseAuthorityName="Parachilota" box="[160,383,1957,1979]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="47" pageNumber="51" phylum="Annelida" rank="genus">
<emphasis id="B9282E65FF97FFA0E6BCD18ABD9669CA" box="[160,239,1957,1979]" italics="true" pageId="47" pageNumber="51">Chilota</emphasis>
(
<emphasis id="B9282E65FF97FFA0E6E2D18ABC0E69CA" box="[254,375,1957,1979]" italics="true" pageId="47" pageNumber="51">Parachilota</emphasis>
)
</taxonomicName>
<emphasis id="B9282E65FF97FFA0E79AD18ABCAB69CA" box="[390,466,1957,1979]" italics="true" pageId="47" pageNumber="51">sclateri</emphasis>
sp. dubia:
<treatmentCitation id="0AFDD466FF97FFA0E45DD18ABE7E69CA" author="Pickford" box="[577,775,1957,1979]" page="575" pageId="47" pageNumber="51" year="1937">
<bibRefCitation id="EFCD8F86FF97FFA0E45DD18ABFAD69CA" author="Pickford" box="[577,724,1957,1979]" pageId="47" pageNumber="51" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer &amp; Sons Ltd. 612 pp." type="book" year="1937">Pickford 1937</bibRefCitation>
: 575
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C346A1FCFF88FFBFE68BD6B7BFC26EDE" box="[151,699,151,176]" pageId="48" pageNumber="52" type="materials_examined">
<paragraph id="8BE3F277FF88FFBFE68BD6B7BFC26EDE" blockId="48.[151,1436,151,392]" box="[151,699,151,176]" pageId="48" pageNumber="52">
<emphasis id="B9282E65FF88FFBFE68BD6B7BC4C6EC1" bold="true" box="[151,309,151,176]" pageId="48" pageNumber="52">
<typeStatus id="54E74CD5FF88FFBFE68BD6B7BDAB6EC1" box="[151,210,152,176]" pageId="48" pageNumber="52">Type</typeStatus>
locality.
</emphasis>
Knysna (after
<bibRefCitation id="EFCD8F86FF88FFBFE7C0D6B8BFD76EDE" author="Michaelsen" box="[476,686,151,175]" pageId="48" pageNumber="52" refString="Michaelsen, W. (1913 b) Report upon the Oligochaeta in the South African Museum at Cape Town. Annals of the South African Museum, 13, 43 - 62." type="journal article" year="1913" yearSuffix="b">Michaelsen 1913b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C346A1FCFF88FFBFE6D5D693BCB96C9B" pageId="48" pageNumber="52" type="discussion">
<paragraph id="8BE3F277FF88FFBFE6D5D693B8426FF9" blockId="48.[151,1436,151,392]" pageId="48" pageNumber="52">
<emphasis id="B9282E65FF88FFBFE6D5D693BC4F6EA5" bold="true" box="[201,310,188,212]" pageId="48" pageNumber="52">Remarks</emphasis>
. The confused original species description (
<bibRefCitation id="EFCD8F86FF88FFBFE53ED693BE856EA5" author="Beddard" box="[802,1020,188,212]" pageId="48" pageNumber="52" refString="Beddard, F. E. (1897) On a collection of earthworms from South Africa belonging to the genus Acanthodrilus. With an Appendix: on a new genus of earthworms belonging to the family Eudrilidae. Proceedings of the Zoological Society of London, 1897, 336 - 349." type="proceedings paper" year="1897">Beddard 1897: 342</bibRefCitation>
) and the heterogeneity of the 19
<typeStatus id="54E74CD5FF88FFBFE377D692B8E56EA4" box="[1387,1436,189,213]" pageId="48" pageNumber="52">type</typeStatus>
specimens led
<bibRefCitation id="EFCD8F86FF88FFBFE727D6F0BF6A6E86" author="Michaelsen" box="[315,531,223,247]" pageId="48" pageNumber="52" refString="Michaelsen, W. (1913 a) Die Oligochaeten des Kaplandes. Zoologische Jahrbucher, Abteilung fur Systematik, 34, 473 - 556." type="journal article" year="1913" yearSuffix="a">Michaelsen (1913a</bibRefCitation>
,
<bibRefCitation id="EFCD8F86FF88FFBFE404D6F0BF5E6E86" author="Michaelsen" box="[536,551,223,247]" pageId="48" pageNumber="52" refString="Michaelsen, W. (1913 b) Report upon the Oligochaeta in the South African Museum at Cape Town. Annals of the South African Museum, 13, 43 - 62." type="journal article" year="1913" yearSuffix="b">b</bibRefCitation>
) to consider this species as species inquirenda, if not species spuria. The
<typeStatus id="54E74CD5FF88FFBFE370D6CFB8E56E89" box="[1388,1436,224,248]" pageId="48" pageNumber="52">type</typeStatus>
material consists of few decomposed, fractioned specimens, labelled as
<typeStatus id="54E74CD5FF88FFBFE5ECD72BB92E6F6D" box="[1008,1111,260,284]" pageId="48" pageNumber="52" type="syntype">syntypes</typeStatus>
BMNH 1904.10.5.891893
<taxonomicName id="4C5C89F4FF88FFBFE68BD707BCEA6F31" box="[151,403,296,320]" class="Clitellata" family="Acanthodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="48" pageNumber="52" phylum="Annelida" rank="species" species="sclateri">
<emphasis id="B9282E65FF88FFBFE68BD707BCEA6F31" box="[151,403,296,320]" italics="true" pageId="48" pageNumber="52">Acanthodrilus sclateri</emphasis>
</taxonomicName>
… this is a part of number 891-3. Occasionally this species name was associated with a variety of other species as a synonym.
<bibRefCitation id="EFCD8F86FF88FFBFE477D763BE196F15" author="Pickford" box="[619,864,332,356]" pageId="48" pageNumber="52" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer &amp; Sons Ltd. 612 pp." type="book" year="1937">Pickford (1937: 575)</bibRefCitation>
after comprehensive examination of the
<taxonomicName id="4C5C89F4FF88FFBFE354D763B8E56F15" box="[1352,1436,332,356]" class="Clitellata" family="Acanthodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="48" pageNumber="52" phylum="Annelida" rank="species" species="sclateri">
<emphasis id="B9282E65FF88FFBFE354D763B8E56F15" box="[1352,1436,332,356]" italics="true" pageId="48" pageNumber="52">sclateri</emphasis>
</taxonomicName>
description and fractions of material declared as syntypes, confirmed
<taxonomicName id="4C5C89F4FF88FFBFE5A8D75FB9716FF9" box="[948,1032,368,392]" class="Clitellata" family="Acanthodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="48" pageNumber="52" phylum="Annelida" rank="species" species="sclateri">
<emphasis id="B9282E65FF88FFBFE5A8D75FB9716FF9" box="[948,1032,368,392]" italics="true" pageId="48" pageNumber="52">sclateri</emphasis>
</taxonomicName>
as a species
<emphasis id="B9282E65FF88FFBFE285D75EB84D6FF9" box="[1177,1332,368,392]" italics="true" pageId="48" pageNumber="52">incertae sedis</emphasis>
.
</paragraph>
<paragraph id="8BE3F277FF88FFBFE6DBD798B8756C69" blockId="48.[151,1436,439,536]" pageId="48" pageNumber="52">
Furthermore,
<bibRefCitation id="EFCD8F86FF88FFBFE77FD798BFF36FBE" author="Pickford" box="[355,650,439,463]" pageId="48" pageNumber="52" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer &amp; Sons Ltd. 612 pp." type="book" year="1937">Pickford (1937: 580583)</bibRefCitation>
indicated several, possible new Acanthodrilinae species, but did not name them properly, suggesting only possible relationships to known species. Specimens were immature and poorly preserved, and their short descriptions are not satisfactory for species identification. These are:
</paragraph>
<paragraph id="8BE3F277FF88FFBFE68BD46CBCA76C2B" blockId="48.[151,1436,578,746]" box="[151,478,578,603]" pageId="48" pageNumber="52">
<taxonomicName id="4C5C89F4FF88FFBFE68BD46CBC646C2A" box="[151,285,579,603]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="48" pageNumber="52" phylum="Annelida" rank="genus">
<emphasis id="B9282E65FF88FFBFE68BD46CBC646C2A" box="[151,285,579,603]" italics="true" pageId="48" pageNumber="52">Parachilota</emphasis>
</taxonomicName>
sp. aff.
<emphasis id="B9282E65FF88FFBFE76BD46BBCAA6C2A" box="[375,467,579,603]" italics="true" pageId="48" pageNumber="52">P. p a i s h i</emphasis>
,
</paragraph>
<paragraph id="8BE3F277FF88FFBFE68BD449BF556C0F" blockId="48.[151,1436,578,746]" box="[151,556,614,639]" pageId="48" pageNumber="52">
<taxonomicName id="4C5C89F4FF88FFBFE68BD449BC646C0F" box="[151,285,614,638]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="48" pageNumber="52" phylum="Annelida" rank="genus">
<emphasis id="B9282E65FF88FFBFE68BD449BC646C0F" box="[151,285,614,638]" italics="true" pageId="48" pageNumber="52">Parachilota</emphasis>
</taxonomicName>
sp. aff.
<taxonomicName id="4C5C89F4FF88FFBFE76BD447BF5C6C0F" box="[375,549,614,639]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="48" pageNumber="52" phylum="Annelida" rank="species" species="eodriliformis">
<emphasis id="B9282E65FF88FFBFE76BD447BF5C6C0F" box="[375,549,614,639]" italics="true" pageId="48" pageNumber="52">P. eodriliformis</emphasis>
</taxonomicName>
,
</paragraph>
<paragraph id="8BE3F277FF88FFBFE68BD4A4BC906CD3" blockId="48.[151,1436,578,746]" box="[151,489,650,675]" pageId="48" pageNumber="52">
<taxonomicName id="4C5C89F4FF88FFBFE68BD4A4BC646CD2" box="[151,285,651,675]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="48" pageNumber="52" phylum="Annelida" rank="genus">
<emphasis id="B9282E65FF88FFBFE68BD4A4BC646CD2" box="[151,285,651,675]" italics="true" pageId="48" pageNumber="52">Parachilota</emphasis>
</taxonomicName>
sp. aff.
<emphasis id="B9282E65FF88FFBFE76BD4A3BCA66CD2" box="[375,479,651,675]" italics="true" pageId="48" pageNumber="52">P. t h e ro n i</emphasis>
.
</paragraph>
<paragraph id="8BE3F277FF88FFBFE6DBD481BCB96C9B" blockId="48.[151,1436,578,746]" pageId="48" pageNumber="52">Descriptions of these species are too insufficient to consider them as valid species, and they are recorded here for historical reasons only.</paragraph>
</subSubSection>
</treatment>
</document>