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<document id="6E4AA703099B109E8C340564739E38D0" ID-CLB-Dataset="30348" ID-DOI="10.11646/zootaxa.4403.2.11" ID-GBIF-Dataset="c4ffa296-a21f-43b9-9771-771e507e2f9e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1212133" ID-ZooBank="15E35735-7BEE-40B9-B8E7-64C69D58C487" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1522830032937" checkinUser="plazi" docAuthor="Takahashi, Naoki" docDate="2018" docId="03F19024FFF5172BFF0D206189E58319" docLanguage="en" docName="zootaxa.4403.2.11.pdf" docOrigin="Zootaxa 4403 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Malthinellus exiguus Takahashi 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="399" masterDocId="FFC8E85CFFF4172FFF9A220B8C79843D" masterDocTitle="A new soldier beetle Malthinellus exiguus sp. nov. (Coleoptera: Cantharidae) from the Ryukyus in southwestern Japan, with a key to Japanese species" masterLastPageNumber="400" masterPageNumber="395" pageNumber="396" updateTime="1698534938659" updateUser="ExternalLinkService">
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<mods:titleInfo id="162033571862072292DBB762F87C637B">
<mods:title id="FF5FB7C546BDD4FDD28B128FB6E81C92">A new soldier beetle Malthinellus exiguus sp. nov. (Coleoptera: Cantharidae) from the Ryukyus in southwestern Japan, with a key to Japanese species</mods:title>
</mods:titleInfo>
<mods:name id="D68DDAB17DEE066BC4BA270A13BAB2AF" type="personal">
<mods:role id="5CAC6A88693BDA9666A0374EE2F1729E">
<mods:roleTerm id="DF7027219C2C4E021D4C7F23F4C1F726">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="42DF564AB7340881FA03C329D9D2B1CC">Takahashi, Naoki</mods:namePart>
</mods:name>
<mods:typeOfResource id="8AD23942E6EF426AC6F785ACF8C813F6">text</mods:typeOfResource>
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<mods:title id="F3CA85F6144FC0A015E7E421A90958F6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="34C0FA30FB72C1748B3525A77928832E">
<mods:date id="210D512AC45D453C2B36F229A41AFD3B">2018</mods:date>
<mods:detail id="E66493E2BF639BD201C08ACCDF939FAC" type="pubDate">
<mods:number id="8B1F690F3A04573A5B10DA48A0976443">2018-04-04</mods:number>
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<mods:number id="D57200D960895A82039583B66A7BBD02">4403</mods:number>
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<mods:number id="266C252EAAA9524B8EC2DBE806AD5E31">2</mods:number>
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<mods:classification id="C23BFE60A873D8E33DB25C0BE2C39BA4">journal article</mods:classification>
<mods:identifier id="F8B22FFB8AE855A84573B48DA7551555" type="CLB-Dataset">30348</mods:identifier>
<mods:identifier id="DC3AB4F46AF1FA9ABB7DB349509932EB" type="DOI">10.11646/zootaxa.4403.2.11</mods:identifier>
<mods:identifier id="3A07EEC860FCCAF11C472054E4D731C9" type="GBIF-Dataset">c4ffa296-a21f-43b9-9771-771e507e2f9e</mods:identifier>
<mods:identifier id="F17AAB5093EA891440103EB889B13EEF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="95617092F8A817C953151A115D68E5A4" type="Zenodo-Dep">1212133</mods:identifier>
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</mods:mods>
<treatment id="03F19024FFF5172BFF0D206189E58319" ID-DOI="http://doi.org/10.5281/zenodo.5965359" ID-GBIF-Taxon="143467282" ID-Zenodo-Dep="5965359" LSID="urn:lsid:plazi:treatment:03F19024FFF5172BFF0D206189E58319" httpUri="http://treatment.plazi.org/id/03F19024FFF5172BFF0D206189E58319" lastPageId="4" lastPageNumber="399" pageId="1" pageNumber="396">
<subSubSection id="C34272B9FFF5172EFF0D20618D73869B" pageId="1" pageNumber="396" type="nomenclature">
<paragraph id="8BE72132FFF5172EFF0D20618EC986B9" blockId="1.[151,688,618,678]" box="[151,688,618,644]" pageId="1" pageNumber="396">
<heading id="D0AF965EFFF5172EFF0D20618EC986B9" bold="true" box="[151,688,618,644]" fontSize="11" level="1" pageId="1" pageNumber="396" reason="1">
<emphasis id="B92CFD20FFF5172EFF0D20618EC986B9" bold="true" box="[151,688,618,644]" pageId="1" pageNumber="396">
<taxonomicName id="4C585AB1FFF5172EFF0D20618E3C86B9" authority="N. Takahashi" authorityName="Takahashi" authorityYear="2018" box="[151,581,618,644]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="species" species="exiguus" status="sp. nov.">
<emphasis id="B92CFD20FFF5172EFF0D20618DEE86B9" bold="true" box="[151,407,618,644]" italics="true" pageId="1" pageNumber="396">Malthinellus exiguus</emphasis>
N. Takahashi
</taxonomicName>
,
<taxonomicNameLabel id="A21F405BFFF5172EFDC820618EC986B9" box="[594,688,618,644]" pageId="1" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE72132FFF5172EFF0D20868D73869B" blockId="1.[151,688,618,678]" box="[151,266,653,678]" pageId="1" pageNumber="396">
(
<figureCitation id="13633DB7FFF5172EFF0520868D78869B" box="[159,257,653,678]" captionStart-0="FIGURES 12" captionStart-1="FIGURES 38" captionStart-2="FIGURE 9" captionStartId-0="2.[151,264,1329,1352]" captionStartId-1="3.[151,264,1621,1644]" captionStartId-2="4.[151,250,1249,1271]" captionTargetBox-0="[154,1424,445,1308]" captionTargetBox-1="[151,1436,661,1601]" captionTargetBox-2="[392,1262,464,1166]" captionTargetId-0="figure@2.[151,1436,445,1308]" captionTargetId-1="figure@3.[151,1436,661,1601]" captionTargetId-2="figure@4.[322,1265,447,1227]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 12. Habitus of Malthinellus exiguus N. Takahashi, sp. nov., dorsal view. 1, male; 2,female.Scales: 1.0 mm." captionText-1="FIGURES 38. Scanning electron microscopy images of the male genitalia of Malthinellus exiguus N. Takahashi, sp. nov. 3, dorsal view; 4, dorso-basal view; 5, dorso-lateral view; 6, dorsal view showing the positions of parts presented in the FIGS 7 8; 7, inner and outer lateral lobes, enlarged dorsal view; 8, outer lateral lobe, enlarged dorso-basal view.The arrow in the right margin explains the meaning of the terms “basal” and “apical”, as used in description: they correspond to the “anterior” and “posterior”, respectively. Abbreviations: bp, basal piece; il, inner lateral lobe; ol, outer lateral lobe; p, penis. Scales: 0.2 mm (FIGS 35); 0.1 mm (FIGS 78)." captionText-2="FIGURE 9. Female terminalia of Malthinellus exiguus N. Takahashi, sp. nov., ventral view (setae omitted on the right side). The arrow representation is the same as that in FIGS 38. Abbreviations: c, coxite; lp, lapel of coxite; pp, paraproct. Scale: 0.125 mm." httpUri-0="https://zenodo.org/record/1212135/files/figure.png" httpUri-1="https://zenodo.org/record/1212137/files/figure.png" httpUri-2="https://zenodo.org/record/1212139/files/figure.png" pageId="1" pageNumber="396">
Figs 1
<emphasis id="B92CFD20FFF5172EFF7F20848C888699" bold="true" box="[229,241,655,676]" pageId="1" pageNumber="396"></emphasis>
9
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34272B9FFF5172EFF0D20DE89238033" pageId="1" pageNumber="396" type="materials_examined">
<paragraph id="8BE72132FFF5172EFF0D20DE8DA6872F" blockId="1.[151,1437,725,2010]" pageId="1" pageNumber="396">
<materialsCitation id="3B302B6FFFF5172EFF0D20DE8DA6872F" ID-GBIF-Occurrence="1837220179" collectingDate="1995-04-16" collectionCode="ELKU" collectorName="Type No. &amp; Kume Is. &amp; N. Takahashi" country="Japan" location="Otake" municipality="Ryukyus" pageId="1" pageNumber="396" specimenCount="3" typeStatus="holotype">
<emphasis id="B92CFD20FFF5172EFF0D20DE8D2B86D3" bold="true" box="[151,338,725,750]" pageId="1" pageNumber="396">
<typeStatus id="54E39F90FFF5172EFF0D20DE8CAB86D3" box="[151,210,725,750]" pageId="1" pageNumber="396">Type</typeStatus>
materials.
</emphasis>
<typeStatus id="54E39F90FFF5172EFEC120DE8DBA86D3" box="[347,451,725,750]" pageId="1" pageNumber="396" type="holotype">Holotype</typeStatus>
(
<collectorName id="26AD44E4FFF5172EFE4E20DD8E3886D0" box="[468,577,726,750]" pageId="1" pageNumber="396">
<typeStatus id="54E39F90FFF5172EFE4E20DD8E7786D3" box="[468,526,726,750]" pageId="1" pageNumber="396">Type</typeStatus>
No.
</collectorName>
3466,
<collectionCode id="ED49B9F7FFF5172EFD0820DD8E9886D3" box="[658,737,726,750]" pageId="1" pageNumber="396">ELKU</collectionCode>
), male, Mt. Daruma—Mt.
<location id="8E8777E9FFF5172EFB8C20DE882586D3" LSID="urn:lsid:plazi:treatment:03F19024FFF5172BFF0D206189E58319:8E8777E9FFF5172EFB8C20DE882586D3" box="[1046,1116,725,750]" country="Japan" municipality="Ryukyus" name="Otake" pageId="1" pageNumber="396">Ôtake</location>
,
<collectorName id="26AD44E4FFF5172EFBF320DD88AD86D0" box="[1129,1236,726,749]" pageId="1" pageNumber="396">Kume Is.</collectorName>
,
<collectingMunicipality id="6B83BB48FFF5172EFB7820DE893E86D3" box="[1250,1351,725,750]" pageId="1" pageNumber="396">Ryukyus</collectingMunicipality>
,
<collectingCountry id="F34F61A2FFF5172EFACF20DD89E186D0" box="[1365,1432,726,749]" name="Japan" pageId="1" pageNumber="396">Japan</collectingCountry>
,
<collectingDate id="EFA2FE1AFFF5172EFF0D20F28D69872F" box="[151,272,761,786]" pageId="1" pageNumber="396" value="1995-04-16">
<date id="FFE607F2FFF5172EFF0D20F28D69872F" box="[151,272,761,786]" pageId="1" pageNumber="396" value="1995-04-16">16.iv.1995</date>
</collectingDate>
,
<collectorName id="26AD44E4FFF5172EFE8120F18DD7872F" box="[283,430,761,786]" pageId="1" pageNumber="396">N. Takahashi</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BE72132FFF5172EFF5D211589238033" blockId="1.[151,1437,725,2010]" pageId="1" pageNumber="396">
<materialsCitation id="3B302B6FFFF5172EFF5D2115884B870B" ID-GBIF-Occurrence="1837220177" box="[199,1074,797,822]" location="Mt. Yonaha" pageId="1" pageNumber="396" specimenCount="68" specimenCount-female="21" specimenCount-male="13" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="9D5EEABBFFF5172EFF5D21158D20870B" box="[199,345,798,822]" pageId="1" pageNumber="396" type="generic" typeStatus="Paratypes">34 Paratypes</specimenCount>
(
<specimenCount id="9D5EEABBFFF5172EFEF021158DAA870B" box="[362,467,797,822]" pageId="1" pageNumber="396" type="male">13 males</specimenCount>
,
<specimenCount id="9D5EEABBFFF5172EFE7B21158E1B870B" box="[481,610,797,822]" pageId="1" pageNumber="396" type="female">21 females</specimenCount>
): RYUKYUS (
<collectingRegion id="499CEFD0FFF5172EFC9721168F0D870B" box="[781,884,797,822]" country="Japan" name="Okinawa" pageId="1" pageNumber="396">Okinawa</collectingRegion>
Is.)
<location id="8E8777E9FFF5172EFC302115884B870B" LSID="urn:lsid:plazi:treatment:03F19024FFF5172BFF0D206189E58319:8E8777E9FFF5172EFC302115884B870B" box="[938,1074,797,822]" name="Mt. Yonaha" pageId="1" pageNumber="396" stateProvince="Okinawa">Mt. Yonaha</location>
</materialsCitation>
,
<materialsCitation id="3B302B6FFFF5172EFBA521168F6E8764" ID-GBIF-Occurrence="1837220175" collectingDate="1990-04-04" collectionCode="KURA" collectorName="Y. Okushima" location="Benoki Dam" pageId="1" pageNumber="396" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<collectingDate id="EFA2FE1AFFF5172EFBA5211688D0870B" box="[1087,1193,797,822]" pageId="1" pageNumber="396" value="1990-04-04">
<date id="FFE607F2FFF5172EFBA5211688D0870B" box="[1087,1193,797,822]" pageId="1" pageNumber="396" value="1990-04-04">4.iv.1990</date>
</collectingDate>
,
<collectorName id="26AD44E4FFF5172EFB2C21158933870B" box="[1206,1354,797,822]" pageId="1" pageNumber="396">Y. Okushima</collectorName>
leg. (
<specimenCount id="9D5EEABBFFF5172EFA0A21158CA98767" pageId="1" pageNumber="396" type="male">1 male</specimenCount>
,
<collectionCode id="ED49B9F7FFF5172EFF4121498D498767" box="[219,304,834,858]" pageId="1" pageNumber="396">KURA</collectionCode>
). Kunigami Vil., Ié-rindô near
<location id="8E8777E9FFF5172EFD1C214A8F6E8764" LSID="urn:lsid:plazi:treatment:03F19024FFF5172BFF0D206189E58319:8E8777E9FFF5172EFD1C214A8F6E8764" box="[646,791,833,858]" name="Benoki Dam" pageId="1" pageNumber="396" stateProvince="Okinawa">Benoki Dam</location>
</materialsCitation>
,
<materialsCitation id="3B302B6FFFF5172EFCB8214A89E18743" ID-GBIF-Occurrence="1837220181" collectingDate="1998-03-21" collectionCode="ELKU" collectorName="N. Takahashi" county="Kume Is." location="Same" municipality="Same" pageId="1" pageNumber="396" specimenCount="13" specimenCount-female="9" specimenCount-male="4" stateProvince="Okinawa" typeStatus="holotype">
<collectingDate id="EFA2FE1AFFF5172EFCB8214A8FE48767" box="[802,925,833,858]" pageId="1" pageNumber="396" value="1998-03-21">
<date id="FFE607F2FFF5172EFCB8214A8FE48767" box="[802,925,833,858]" pageId="1" pageNumber="396" value="1998-03-21">21.iii.1998</date>
</collectingDate>
,
<collectorName id="26AD44E4FFF5172EFC33214988458767" box="[937,1084,833,858]" pageId="1" pageNumber="396">N. Takahashi</collectorName>
leg. (
<specimenCount id="9D5EEABBFFF5172EFBE7214988B08767" box="[1149,1225,833,858]" pageId="1" pageNumber="396" type="male">1 male</specimenCount>
, NTC). (
<collectingCounty id="628659BEFFF5172EFAB1214989EA8767" box="[1323,1427,834,858]" pageId="1" pageNumber="396">Kume Is.</collectingCounty>
)
<collectingMunicipality id="6B83BB48FFF5172EFF0D216D8CAC8743" box="[151,213,870,894]" pageId="1" pageNumber="396">Same</collectingMunicipality>
label as
<typeStatus id="54E39F90FFF5172EFEA1216E8DE48743" box="[315,413,869,894]" pageId="1" pageNumber="396" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D5EEABBFFF5172EFE2A216D8E7F8743" box="[432,518,869,894]" pageId="1" pageNumber="396" type="male">1 males</specimenCount>
and
<specimenCount id="9D5EEABBFFF5172EFDA4216D8ED78743" box="[574,686,869,894]" pageId="1" pageNumber="396" type="female">2 females</specimenCount>
,
<collectionCode id="ED49B9F7FFF5172EFD20216D8F718740" box="[698,776,870,893]" pageId="1" pageNumber="396">ELKU</collectionCode>
;
<specimenCount id="9D5EEABBFFF5172EFC8F216D8F158743" box="[789,876,869,894]" pageId="1" pageNumber="396" type="male">2 males</specimenCount>
and
<specimenCount id="9D5EEABBFFF5172EFC3F216D886C8743" box="[933,1045,869,894]" pageId="1" pageNumber="396" type="female">7 females</specimenCount>
, NTC).
<location id="8E8777E9FFF5172EFBE8216D88C98743" LSID="urn:lsid:plazi:treatment:03F19024FFF5172BFF0D206189E58319:8E8777E9FFF5172EFBE8216D88C98743" box="[1138,1200,870,894]" county="Kume Is." municipality="Same" name="Same" pageId="1" pageNumber="396" stateProvince="Okinawa">Same</location>
locality as
<typeStatus id="54E39F90FFF5172EFAA9216E89E18743" box="[1331,1432,869,894]" pageId="1" pageNumber="396" type="holotype">holotype</typeStatus>
</materialsCitation>
,
<date id="FFE607F2FFF5172EFF0D21828D69879F" box="[151,272,905,930]" pageId="1" pageNumber="396" value="1995-04-14">14.iv.1995</date>
, N. Takahashi leg. (2 females, NTC);
<materialsCitation id="3B302B6FFFF5172EFD2521828CA487FB" ID-GBIF-Occurrence="1837220176" collectingDate="1995-04-15" collectorName="N. Takahashi" location="Mt. Otake" pageId="1" pageNumber="396" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Okinawa" typeStatus="holotype">
<collectingDate id="EFA2FE1AFFF5172EFD2521828F4E879F" box="[703,823,905,930]" pageId="1" pageNumber="396" value="1995-04-15">
<date id="FFE607F2FFF5172EFD2521828F4E879F" box="[703,823,905,930]" pageId="1" pageNumber="396" value="1995-04-15">15.iv.1995</date>
</collectingDate>
,
<collectorName id="26AD44E4FFF5172EFCD921818FAE879F" box="[835,983,905,930]" pageId="1" pageNumber="396">N. Takahashi</collectorName>
leg. (
<specimenCount id="9D5EEABBFFF5172EFB8021818809879F" box="[1050,1136,905,930]" pageId="1" pageNumber="396" type="male">3 males</specimenCount>
and
<specimenCount id="9D5EEABBFFF5172EFB3321818961879F" box="[1193,1304,905,930]" pageId="1" pageNumber="396" type="female">3 females</specimenCount>
, NTC).
<location id="8E8777E9FFF5172EFAEF21818CA487FB" LSID="urn:lsid:plazi:treatment:03F19024FFF5172BFF0D206189E58319:8E8777E9FFF5172EFAEF21818CA487FB" name="Mt. Otake" pageId="1" pageNumber="396" stateProvince="Okinawa">Mt. Ôtake</location>
</materialsCitation>
,
<materialsCitation id="3B302B6FFFF5172EFF7621A68FEF87F8" ID-GBIF-Occurrence="1837220180" box="[236,918,941,966]" collectingDate="1995-04-15" collectorName="N. Takahashi" location="Mt. Daruma" pageId="1" pageNumber="396" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<collectingDate id="EFA2FE1AFFF5172EFF7621A68D1D87FB" box="[236,356,941,966]" pageId="1" pageNumber="396" value="1995-04-15">
<date id="FFE607F2FFF5172EFF7621A68D1D87FB" box="[236,356,941,966]" pageId="1" pageNumber="396" value="1995-04-15">15.iv.1995</date>
</collectingDate>
,
<collectorName id="26AD44E4FFF5172EFEE921A58E7387FB" box="[371,522,941,966]" pageId="1" pageNumber="396">N. Takahashi</collectorName>
leg. (
<specimenCount id="9D5EEABBFFF5172EFDCF21A58EDD87FB" box="[597,676,941,966]" pageId="1" pageNumber="396" type="male">1 male</specimenCount>
, NTC).
<location id="8E8777E9FFF5172EFC9D21A58FEF87F8" LSID="urn:lsid:plazi:treatment:03F19024FFF5172BFF0D206189E58319:8E8777E9FFF5172EFC9D21A58FEF87F8" box="[775,918,942,965]" name="Mt. Daruma" pageId="1" pageNumber="396" stateProvince="Okinawa">Mt. Daruma</location>
</materialsCitation>
,
<date id="FFE607F2FFF5172EFC3F21A6886987FB" box="[933,1040,941,966]" pageId="1" pageNumber="396" value="1990-04-07">7.iv.1990</date>
, T. Ueno leg. (1 female, KURA);
<date id="FFE607F2FFF5172EFF0D21DA8D7887D7" box="[151,257,977,1002]" pageId="1" pageNumber="396" value="1990-04-08">8.iv.1990</date>
, T. Ueno leg. (2 males and 1 female, KURA);
<materialsCitation id="3B302B6FFFF5172EFC9321DA8D5B8030" ID-GBIF-Occurrence="1837220178" collectingDate="1990-04-09" collectionCode="KURA" collectorName="T. Ueno" location="Nr. Mt." pageId="1" pageNumber="396" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<collectingDate id="EFA2FE1AFFF5172EFC9321DA8F0A87D7" box="[777,883,977,1002]" pageId="1" pageNumber="396" value="1990-04-09">
<date id="FFE607F2FFF5172EFC9321DA8F0A87D7" box="[777,883,977,1002]" pageId="1" pageNumber="396" value="1990-04-09">9.iv.1990</date>
</collectingDate>
,
<collectorName id="26AD44E4FFF5172EFCE421D98FA187D4" box="[894,984,978,1001]" pageId="1" pageNumber="396">T. Ueno</collectorName>
leg. (
<specimenCount id="9D5EEABBFFF5172EFB8321D9881A87D7" box="[1049,1123,977,1002]" pageId="1" pageNumber="396" type="male">1 male</specimenCount>
and
<specimenCount id="9D5EEABBFFF5172EFB0321D9897187D7" box="[1177,1288,977,1002]" pageId="1" pageNumber="396" type="female">3 females</specimenCount>
,
<collectionCode id="ED49B9F7FFF5172EFA8921D9891187D7" box="[1299,1384,978,1002]" pageId="1" pageNumber="396">KURA</collectionCode>
).
<location id="8E8777E9FFF5172EFAE321D98CC68030" LSID="urn:lsid:plazi:treatment:03F19024FFF5172BFF0D206189E58319:8E8777E9FFF5172EFAE321D98CC68030" name="Nr. Mt." pageId="1" pageNumber="396" stateProvince="Okinawa">Nr. Mt.</location>
Daruma
</materialsCitation>
,
<date id="FFE607F2FFF5172EFEB421FE8DE18033" box="[302,408,1013,1038]" pageId="1" pageNumber="396" value="1996-04-05">5.iv.1996</date>
, Y. Okushima leg. (2 females, KURA);
<date id="FFE607F2FFF5172EFCC221FE8FBB8033" box="[856,962,1013,1038]" pageId="1" pageNumber="396" value="1996-04-07">7.iv.1996</date>
, Y. Okushima leg. (1 male, KURA).
</paragraph>
</subSubSection>
<subSubSection id="C34272B9FFF5172EFF5D26128D518137" pageId="1" pageNumber="396" type="diagnosis">
<paragraph id="8BE72132FFF5172EFF5D26128D518137" blockId="1.[151,1437,725,2010]" pageId="1" pageNumber="396">
<emphasis id="B92CFD20FFF5172EFF5D26128D38800F" bold="true" box="[199,321,1049,1074]" pageId="1" pageNumber="396">Diagnosis.</emphasis>
Body yellowish orange to orange with brownish black elytra. Male genitalia: basal piece oboval in outline in ventral view, not produced at apex; outer lateral lobes slender in basal part, nearly securiform apically; inner lateral lobes developed, strongly curved medially in basal part, spathulate in apical part; penis composed of a pair of parts in dorsal view, each part somewhat Santoku knife-shaped with carinae along lateral margins in basal part. Female terminalia: paraprocts gently widened basally in outline in ventral view, approximately reaching the basal 2/3 of coxite in profile; coxite with lapel more or less reaching the basal 2/5 of the coxite at the median part in ventral view.
</paragraph>
</subSubSection>
<subSubSection id="C34272B9FFF5172CFF5D271E8EA88378" lastPageId="3" lastPageNumber="398" pageId="1" pageNumber="396" type="description">
<paragraph id="8BE72132FFF5172EFF5D271E8F6681DF" blockId="1.[151,1437,725,2010]" pageId="1" pageNumber="396">
<emphasis id="B92CFD20FFF5172EFF5D271E8D208113" bold="true" box="[199,345,1301,1326]" pageId="1" pageNumber="396">Description.</emphasis>
MALE (
<figureCitation id="13633DB7FFF5172EFE5B271E8E7A8113" box="[449,515,1301,1326]" captionStart="FIGURES 12" captionStartId="2.[151,264,1329,1352]" captionTargetBox="[154,1424,445,1308]" captionTargetId="figure@2.[151,1436,445,1308]" captionTargetPageId="2" captionText="FIGURES 12. Habitus of Malthinellus exiguus N. Takahashi, sp. nov., dorsal view. 1, male; 2,female.Scales: 1.0 mm." httpUri="https://zenodo.org/record/1212135/files/figure.png" pageId="1" pageNumber="396">Fig. 1</figureCitation>
). Head, pronotum, scutellum, ventral side of body, and legs yellowish orange to orange, maxillary and labial palpi darkened, sometimes so on 2nd to 4th tarsal segments, somewhat more yellowish on abdominal sternites except for the two or three apical sternites; antennae dark brown but yellowish on basal halves of 1
<taxonomicName id="4C585AB1FFF5172EFE9527898DF581A7" box="[271,396,1409,1434]" pageId="1" pageNumber="396" rank="subSpecies" subSpecies="antennal">st antennal</taxonomicName>
segments; eyes black; elytra and hind wings dark brownish to black, however darker on the latter; apical halves of mandibles, tibial spurs and tarsal claws reddish brown and sometimes so on hind coxae along posterior margins, near hind angles of metaventrite.
</paragraph>
<paragraph id="8BE72132FFF5172EFF4B27E688D88273" blockId="1.[151,1437,725,2010]" pageId="1" pageNumber="396">Body elongate but somewhat stout, densely covered with pale brownish and recumbent pubescence, fitted with moderately longer pubescence around clypeus; antennae and lateral areas of elytra sparsely intermingled with pale bristles; antennae covered with darker bristles; legs covered with faintly darker bristles.</paragraph>
<paragraph id="8BE72132FFF5172EFF4C24528EF7831B" blockId="1.[151,1437,725,2010]" pageId="1" pageNumber="396">
Head wider than long, 1.17 (in the
<typeStatus id="54E39F90FFF5172EFD0124528F78824F" box="[667,769,1625,1650]" pageId="1" pageNumber="396" type="holotype">holotype</typeStatus>
; range 1.091.21) times wider than pronotum; surface puncticulate, somewhat granulate and rugulose, slightly matt on dorsum though smoother and lustrous before and around vertex, finely granulate and matt on ventral side but almost impunctate and lustrous on stipes of maxillary palpi; eyes medium in size, interocular distance 3.41 (3.263.86) times larger than radius of eye; antennae filiform, slightly extending over elytral apices, relative lengths of antennal segments from base as follows—27.9: 16.1: 19.9: 22.2: 23.6: 24.6: 24.3: 23.9: 22.9: 21.9: 26.3.
</paragraph>
<paragraph id="8BE72132FFF5172DFF5D25398DF284E8" blockId="1.[151,1437,725,2010]" lastBlockId="2.[151,1436,151,392]" lastPageId="2" lastPageNumber="397" pageId="1" pageNumber="396">Pronotum subquadrate, slightly broad, faintly dilated posteriorly, 1.31 (1.231.31) times wider than long; disc puncticulate, somewhat rugulose, lustrous on surface, gently convex medio-dorsally, faintly and transversely depressed at anterior third, feebly depressed in medio-posterial area, marginate on all margins and particularly so on anterior and posterior ones, sub-truncate on anterior margin, obtusely rounded on front angles, weakly arcuate posteriorly on posterior edge, obtuse on hind angles, flattened and gently arcuate laterally and feebly constricted on sides just before hind angles. Scutellum sub-trapezoidal and narrowed posteriorly, finely granulate and covered with fine pubescence.</paragraph>
<paragraph id="8BE72132FFF6172DFF5D22D48E3385B5" blockId="2.[151,1436,151,392]" pageId="2" pageNumber="397">Elytra 2.55 (2.382.55) times longer than wide, reaching last abdominal tergite; each elytron puncticulate, lustrous around humerus, weakly marginate along sutural line and lateral margin except for areas around humerus and apex, longitudinally punctate in rows with several rather undefined costae, faintly rugulose: particularly so in mesal half but smoother around humerus and apex, the punctures becoming indistinct distally in other area though relatively distinct along lateral margin.</paragraph>
<caption id="DF2771BAFFF6172DFF0D273A89198175" box="[151,1376,1329,1352]" httpUri="https://zenodo.org/record/1212135/files/figure.png" pageId="2" pageNumber="397" startId="2.[151,264,1329,1352]" targetBox="[154,1424,445,1308]" targetPageId="2">
<paragraph id="8BE72132FFF6172DFF0D273A89198175" blockId="2.[151,1376,1329,1352]" box="[151,1376,1329,1352]" pageId="2" pageNumber="397">
<emphasis id="B92CFD20FFF6172DFF0D273A8D43817A" bold="true" box="[151,314,1329,1352]" pageId="2" pageNumber="397">FIGURES 12.</emphasis>
Habitus of
<taxonomicName id="4C585AB1FFF6172DFE2B27398F6B8175" authority="N. Takahashi" authorityName="Takahashi" authorityYear="2018" box="[433,786,1330,1352]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="397" phylum="Arthropoda" rank="species" species="exiguus" status="sp. nov.">
<emphasis id="B92CFD20FFF6172DFE2B27398EFD8175" box="[433,644,1330,1352]" italics="true" pageId="2" pageNumber="397">Malthinellus exiguus</emphasis>
N. Takahashi
</taxonomicName>
,
<taxonomicNameLabel id="A21F405BFFF6172DFC8627398F14817A" box="[796,877,1330,1351]" pageId="2" pageNumber="397" rank="species">
<emphasis id="B92CFD20FFF6172DFC8627398F14817A" bold="true" box="[796,877,1330,1351]" pageId="2" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
, dorsal view. 1, male; 2, female. Scales: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BE72132FFF6172DFF4A277F8FE88258" blockId="2.[151,1437,1396,2033]" pageId="2" pageNumber="397">Mesoventrite, mesepimera, metaventrite, and metanepisterna puncticulate and lustrous on their surfaces but sometimes faintly matt on mesoventrite and mesepimera. Abdominal sternites, except for 9th sternite puncticulate, feebly rugulose and lustrous; 8th sternite widely and obtusely emarginate on posterior margin; 9th sternite nearly oblong in outline and gently convex medioventrally in ventral view, gradually narrowed in anterior half which is hidden under the penultimate sternite, more narrowed behind faint angulations on sides in posterior portion, weakly produced posteriorly and bilobate around apex, puncticulate and finely granulate and matt in posterior half on ventral surface, arcuate postero-ventrally in posterior half in profile.</paragraph>
<paragraph id="8BE72132FFF6172DFF5D247B8DF183B9" blockId="2.[151,1437,1396,2033]" pageId="2" pageNumber="397">
Male genitalia (
<figureCitation id="13633DB7FFF6172DFEEC247B8DA682B4" box="[374,479,1648,1673]" captionStart="FIGURES 38" captionStartId="3.[151,264,1621,1644]" captionTargetBox="[151,1436,661,1601]" captionTargetId="figure@3.[151,1436,661,1601]" captionTargetPageId="3" captionText="FIGURES 38. Scanning electron microscopy images of the male genitalia of Malthinellus exiguus N. Takahashi, sp. nov. 3, dorsal view; 4, dorso-basal view; 5, dorso-lateral view; 6, dorsal view showing the positions of parts presented in the FIGS 7 8; 7, inner and outer lateral lobes, enlarged dorsal view; 8, outer lateral lobe, enlarged dorso-basal view.The arrow in the right margin explains the meaning of the terms “basal” and “apical”, as used in description: they correspond to the “anterior” and “posterior”, respectively. Abbreviations: bp, basal piece; il, inner lateral lobe; ol, outer lateral lobe; p, penis. Scales: 0.2 mm (FIGS 35); 0.1 mm (FIGS 78)." httpUri="https://zenodo.org/record/1212137/files/figure.png" pageId="2" pageNumber="397">Figs. 38</figureCitation>
) stout, nearly oboval but apically and basally flattened in outline in dorsal view: basal piece gently convex medio-ventrally, oboval but apically flattened in outline in ventral view, weakly curved dorsally in apical part in profile, medio-longitudinally furrowed on ventral surface with a carina on dorsal surface which reaches just behind the apex; outer lateral lobe elongate and curved medio-dorsally, slender in basal part, and nearly securiform in apical part; inner lateral lobe prolonged medio-dorsally, strongly curved medially in basal part, spathulate but feebly warped dorsally in apical part, and roundly sub-triangulate around apex in profile; penis composed of a pair of parts in dorsal view, each part somewhat Santoku knife-shaped with carinae along lateral margins in basal part.
</paragraph>
<paragraph id="8BE72132FFF6172DFF5D259B8F328395" blockId="2.[151,1437,1396,2033]" box="[199,843,1936,1961]" pageId="2" pageNumber="397">Body length: 3.2 (2.43.9) mm; width: 0.8 (0.71.0) mm.</paragraph>
<paragraph id="8BE72132FFF6172CFF5D25BE8D6885B5" blockId="2.[151,1437,1396,2033]" lastBlockId="3.[151,1437,151,608]" lastPageId="3" lastPageNumber="398" pageId="2" pageNumber="397">
FEMALE (
<figureCitation id="13633DB7FFF6172DFED025BF8DE883F0" box="[330,401,1972,1997]" captionStart="FIGURES 12" captionStartId="2.[151,264,1329,1352]" captionTargetBox="[154,1424,445,1308]" captionTargetId="figure@2.[151,1436,445,1308]" captionTargetPageId="2" captionText="FIGURES 12. Habitus of Malthinellus exiguus N. Takahashi, sp. nov., dorsal view. 1, male; 2,female.Scales: 1.0 mm." httpUri="https://zenodo.org/record/1212135/files/figure.png" pageId="2" pageNumber="397">Fig. 2</figureCitation>
). Similar to male, but body slightly larger in general appearance; last abdominal sternite sometimes faintly more orange colored than the other ones. Head 1.041.16 times wider than pronotum; eyes slightly smaller than in male and interocular distance 3.684.14 times larger than radius of eye; antennae shorter than in male, approximately reaching the basal 2/3 of elytra, and relative lengths of antennal segments, from base to apex, as follows—29.0: 16.2: 19.9: 20.8: 21.4: 22.2: 22.1: 21.7: 20.1: 18.8: 23.1. Pronotum 1.241.31 times wider than long. Elytra 2.342.48 times longer than wide. Eighth abdominal sternite gradually narrowed posteriorly, somewhat more narrowed in about posterior 1/4, obtusely and roundly emarginate in the middle on posterior margin in ventral view, moderately convex medio-ventrally, its surface puncticulate, relatively smooth and bright.
</paragraph>
<paragraph id="8BE72132FFF7172CFF5D239F8E8E8662" blockId="3.[151,1437,151,608]" pageId="3" pageNumber="398">
Female terminalia (
<figureCitation id="13633DB7FFF7172CFE3F239F8D938591" box="[421,490,404,429]" captionStart="FIGURE 9" captionStartId="4.[151,250,1249,1271]" captionTargetBox="[392,1262,464,1166]" captionTargetId="figure@4.[322,1265,447,1227]" captionTargetPageId="4" captionText="FIGURE 9. Female terminalia of Malthinellus exiguus N. Takahashi, sp. nov., ventral view (setae omitted on the right side). The arrow representation is the same as that in FIGS 38. Abbreviations: c, coxite; lp, lapel of coxite; pp, paraproct. Scale: 0.125 mm." httpUri="https://zenodo.org/record/1212139/files/figure.png" pageId="3" pageNumber="398">Fig. 9</figureCitation>
) somewhat stout; paraproct almost divided symmetrically into two parts, each part nearly oval in outline, narrowed dorso-medially in apical part, slightly rounded at apex in dorso-lateral view, approximately reaching the basal 2/3 of coxite in profile; coxite sub-conical but generally flattened in appearance, weakly and obtusely protruded laterally in the middle on outer margin, swollen along outer margin, feebly depressed in medio-longitudinal area along inner margin in ventral view, with a lapel at the base more or less reaching the basal 2/5 of the coxite at the median part.
</paragraph>
<caption id="DF2771BAFFF7172CFF0D245E8D9E8338" httpUri="https://zenodo.org/record/1212137/files/figure.png" pageId="3" pageNumber="398" startId="3.[151,264,1621,1644]" targetBox="[151,1436,661,1601]" targetPageId="3">
<paragraph id="8BE72132FFF7172CFF0D245E8D9E8338" blockId="3.[151,1436,1621,1797]" pageId="3" pageNumber="398">
<emphasis id="B92CFD20FFF7172CFF0D245E8D428256" bold="true" box="[151,315,1621,1644]" pageId="3" pageNumber="398">FIGURES 38.</emphasis>
Scanning electron microscopy images of the male genitalia of
<taxonomicName id="4C585AB1FFF7172CFC5B245D895C8251" authority="N. Takahashi" authorityName="Takahashi" authorityYear="2018" box="[961,1317,1622,1644]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="exiguus" status="sp. nov.">
<emphasis id="B92CFD20FFF7172CFC5B245D88EC8251" box="[961,1173,1622,1644]" italics="true" pageId="3" pageNumber="398">Malthinellus exiguus</emphasis>
N. Takahashi
</taxonomicName>
,
<taxonomicNameLabel id="A21F405BFFF7172CFAAA245D89F88256" box="[1328,1409,1622,1643]" pageId="3" pageNumber="398" rank="species">
<emphasis id="B92CFD20FFF7172CFAAA245D89F88256" bold="true" box="[1328,1409,1622,1643]" pageId="3" pageNumber="398">sp. nov.</emphasis>
</taxonomicNameLabel>
3, dorsal view; 4, dorso-basal view; 5, dorso-lateral view; 6, dorsal view showing the positions of parts presented in the FIGS 7 8; 7, inner and outer lateral lobes, enlarged dorsal view; 8, outer lateral lobe, enlarged dorso-basal view. The arrow in the right margin explains the meaning of the terms “basal” and “apical”, as used in description: they correspond to the “anterior” and “posterior”, respectively. Abbreviations: bp, basal piece; il, inner lateral lobe; ol, outer lateral lobe; p, penis. Scales: 0.2 mm (FIGS 35); 0.1 mm (FIGS 78).
</paragraph>
</caption>
<paragraph id="8BE72132FFF7172CFF5D25278EA88378" blockId="3.[151,1436,1836,2006]" box="[199,721,1836,1861]" pageId="3" pageNumber="398">
Body length:
<quantity id="4CA08CD7FFF7172CFEC525268D908378" box="[351,489,1837,1861]" metricMagnitude="-3" metricUnit="m" metricValue="3.4499999999999997" metricValueMax="4.1" metricValueMin="2.8" pageId="3" pageNumber="398" unit="mm" value="3.4499999999999997" valueMax="4.1" valueMin="2.8">2.84.1 mm</quantity>
; width:
<quantity id="4CA08CD7FFF7172CFDD925268EB48378" box="[579,717,1837,1861]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="3" pageNumber="398" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.81.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34272B9FFF7172CFF5D255B8FC98354" box="[199,944,1872,1897]" pageId="3" pageNumber="398" type="distribution">
<paragraph id="8BE72132FFF7172CFF5D255B8FC98354" blockId="3.[151,1436,1836,2006]" box="[199,944,1872,1897]" pageId="3" pageNumber="398">
<emphasis id="B92CFD20FFF7172CFF5D255B8D198354" bold="true" box="[199,352,1872,1897]" pageId="3" pageNumber="398">Distribution.</emphasis>
<collectingCountry id="F34F61A2FFF7172CFEFD255A8DD28355" box="[359,427,1873,1896]" name="Japan" pageId="3" pageNumber="398">Japan</collectingCountry>
: central Ryukyus (
<collectingRegion id="499CEFD0FFF7172CFDE0255B8E988354" box="[634,737,1872,1897]" country="Japan" name="Okinawa" pageId="3" pageNumber="398">Okinawa</collectingRegion>
Is. and Kume Is.).
</paragraph>
</subSubSection>
<subSubSection id="C34272B9FFF7172BFF5D257E88E18558" lastPageId="4" lastPageNumber="399" pageId="3" pageNumber="398" type="discussion">
<paragraph id="8BE72132FFF7172BFF5D257E8E7584E8" blockId="3.[151,1436,1836,2006]" lastBlockId="4.[151,1437,151,392]" lastPageId="4" lastPageNumber="399" pageId="3" pageNumber="398">
<emphasis id="B92CFD20FFF7172CFF5D257E8D4283B3" bold="true" box="[199,315,1909,1934]" pageId="3" pageNumber="398">Remarks.</emphasis>
This new species is very similar to
<taxonomicName id="4C585AB1FFF7172CFD4D257E892483B0" authority="Takahashi &amp; N. Takahashi, 2007" authorityName="Takahashi &amp; N. Takahashi" authorityYear="2007" box="[727,1373,1909,1934]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="masatakai">
<emphasis id="B92CFD20FFF7172CFD4D257E8F9983B0" box="[727,992,1909,1933]" italics="true" pageId="3" pageNumber="398">Malthinellus masatakai</emphasis>
<bibRefCitation id="EFC95CC3FFF7172CFC73257E892483B0" box="[1001,1373,1909,1934]" pageId="3" pageNumber="398" refId="ref3296" refString="Takahashi, K. &amp; Takahashi, N. (2007) A new species of the genus Malthinellus (Coleoptera, Cantharidae) from Okinawa Island, Southwest Japan, with notes on the genus Malthinellus. Elytra, Tokyo, 35 (1), 142 - 148." type="journal article">Takahashi &amp; N. Takahashi, 2007</bibRefCitation>
</taxonomicName>
from the
<collectingRegion id="499CEFD0FFF7172CFF2525938D5D838C" box="[191,292,1944,1969]" country="Japan" name="Okinawa" pageId="3" pageNumber="398">Okinawa</collectingRegion>
Island in general appearance, but is distinguishable from the latter by the genital structure: in males by the elongate, apically nearly securiform outer lateral lobe, and the long inner lateral lobe and, in females, by the relatively stout coxites and the gently and basally widened paraprocts in outline in ventral view with less warped apical parts in dorso-lateral view.
</paragraph>
<paragraph id="8BE72132FFF0172BFF5D22D48904857D" blockId="4.[151,1437,151,392]" pageId="4" pageNumber="399">
The terminalia of a single female specimen from the Tonaki Island, located between the
<collectingRegion id="499CEFD0FFF0172BFB2322D4896784C5" box="[1209,1310,223,248]" country="Japan" name="Okinawa" pageId="4" pageNumber="399">Okinawa</collectingRegion>
and Kume Islands, shows intermediate morphological features between
<taxonomicName id="4C585AB1FFF0172BFCA5230E8FC08521" authorityName="Takahashi" authorityYear="2018" box="[831,953,261,284]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="B92CFD20FFF0172BFCA5230E8F2F8521" box="[831,854,261,284]" italics="true" pageId="4" pageNumber="399">M</emphasis>
.
<emphasis id="B92CFD20FFF0172BFCFE230E8FC08521" box="[868,953,261,284]" italics="true" pageId="4" pageNumber="399">exiguus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C585AB1FFF0172BFC75230E88FE8521" authorityName="Takahashi &amp; N. Takahashi" authorityYear="2007" box="[1007,1159,260,284]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="masatakai">
<emphasis id="B92CFD20FFF0172BFC75230E887F8521" box="[1007,1030,261,284]" italics="true" pageId="4" pageNumber="399">M</emphasis>
.
<emphasis id="B92CFD20FFF0172BFB8E230F88FE8521" box="[1044,1159,260,284]" italics="true" pageId="4" pageNumber="399">masatakai</emphasis>
</taxonomicName>
. For this reason, it is felt that further investigations are needed to better understand the taxonomic meaning of morphological differences.
</paragraph>
<paragraph id="8BE72132FFF0172BFF5D234688E18558" blockId="4.[151,1437,151,392]" box="[199,1176,332,357]" pageId="4" pageNumber="399">I propose that the Japanese name of this new species is “Kumejima-futairochibi-jôkai”.</paragraph>
</subSubSection>
<subSubSection id="C34272B9FFF0172BFF5D2364896881A2" pageId="4" pageNumber="399" type="etymology">
<paragraph id="8BE72132FFF0172BFF5D236488E185B5" blockId="4.[151,1437,151,392]" box="[199,1176,367,392]" pageId="4" pageNumber="399">
<emphasis id="B92CFD20FFF0172BFF5D23648D3785B5" bold="true" box="[199,334,367,392]" pageId="4" pageNumber="399">Etymology.</emphasis>
Derived from Latin
<taxonomicName id="4C585AB1FFF0172BFDAF237B8EF385BA" authorityName="Takahashi" authorityYear="2018" box="[565,650,368,391]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="B92CFD20FFF0172BFDAF237B8EF385BA" box="[565,650,368,391]" italics="true" pageId="4" pageNumber="399">exiguus</emphasis>
</taxonomicName>
= small; with reference to the small body size.
</paragraph>
<caption id="DF2771BAFFF0172BFF0D26EA8D7D8109" httpUri="https://zenodo.org/record/1212139/files/figure.png" pageId="4" pageNumber="399" startId="4.[151,250,1249,1271]" targetBox="[392,1262,464,1166]" targetPageId="4">
<paragraph id="8BE72132FFF0172BFF0D26EA8D7D8109" blockId="4.[151,1436,1249,1332]" pageId="4" pageNumber="399">
<emphasis id="B92CFD20FFF0172BFF0D26EA8D6D80CB" bold="true" box="[151,276,1249,1271]" pageId="4" pageNumber="399">FIGURE 9.</emphasis>
Female terminalia of
<taxonomicName id="4C585AB1FFF0172BFE6226EA8F2480CA" authority="N. Takahashi" authorityName="Takahashi" authorityYear="2018" box="[504,861,1249,1271]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="exiguus" status="sp. nov.">
<emphasis id="B92CFD20FFF0172BFE6226EA8EB580CA" box="[504,716,1249,1271]" italics="true" pageId="4" pageNumber="399">Malthinellus exiguus</emphasis>
N. Takahashi
</taxonomicName>
,
<taxonomicNameLabel id="A21F405BFFF0172BFCF226E98FC080CA" box="[872,953,1250,1271]" pageId="4" pageNumber="399" rank="species">
<emphasis id="B92CFD20FFF0172BFCF226E98FC080CA" bold="true" box="[872,953,1250,1271]" pageId="4" pageNumber="399">sp. nov.</emphasis>
</taxonomicNameLabel>
, ventral view (setae omitted on the right side). The arrow representation is the same as that in FIGS 38. Abbreviations: c, coxite; lp, lapel of coxite; pp, paraproct. Scale: 0.125 mm.
</paragraph>
</caption>
<paragraph id="8BE72132FFF0172BFF0D278E896881A2" blockId="4.[151,1297,1412,1439]" box="[151,1297,1412,1439]" pageId="4" pageNumber="399">
<emphasis id="B92CFD20FFF0172BFF0D278E8E9381A3" bold="true" box="[151,746,1412,1439]" pageId="4" pageNumber="399">
Determination key to the Japanese
<taxonomicName id="4C585AB1FFF0172BFDD5278F8E9381A3" authorityName="Kiesenwetter" authorityYear="1874" box="[591,746,1412,1438]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="B92CFD20FFF0172BFDD5278F8E9381A3" bold="true" box="[591,746,1412,1438]" italics="true" pageId="4" pageNumber="399">Malthinellus</emphasis>
</taxonomicName>
</emphasis>
(modified from
<bibRefCitation id="EFC95CC3FFF0172BFC36278E897081A2" box="[940,1289,1413,1439]" pageId="4" pageNumber="399" refId="ref3296" refString="Takahashi, K. &amp; Takahashi, N. (2007) A new species of the genus Malthinellus (Coleoptera, Cantharidae) from Okinawa Island, Southwest Japan, with notes on the genus Malthinellus. Elytra, Tokyo, 35 (1), 142 - 148." type="journal article">Takahashi &amp; Takahashi, 2007</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34272B9FFF0172BFF0D27C789E58319" pageId="4" pageNumber="399" type="key">
<key id="FFC97603FFF0172BFF0D27C789E58319" pageId="4" pageNumber="399">
<keyStep id="30AC3997FFF0172BFF0D27C789E281C3" pageId="4" pageNumber="399">
<paragraph id="8BE72132FFF0172BFF0D27C789E281DC" blockId="4.[151,1436,1484,1828]" box="[151,1435,1484,1505]" pageId="4" pageNumber="399">
<keyLead id="30A98207FFF0172BFF0D27C789E281DC" box="[151,1435,1484,1505]" pageId="4" pageNumber="399">1. Pronotum and elytra brownish black or dark brown........................................................... 2</keyLead>
</paragraph>
<paragraph id="8BE72132FFF0172BFF0D27E289E281C3" blockId="4.[151,1436,1484,1828]" box="[151,1435,1513,1534]" pageId="4" pageNumber="399">
<keyLead id="30A98207FFF0172BFF0D27E289E281C3" box="[151,1435,1513,1534]" pageId="4" pageNumber="399">- Pronotum yellowish orange to orange, elytra dark brown...................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AC3997FFF0172BFF0D240D89E58204" pageId="4" pageNumber="399">
<paragraph id="8BE72132FFF0172BFF0D240D89E58226" blockId="4.[151,1436,1484,1828]" box="[151,1436,1542,1563]" pageId="4" pageNumber="399">
<keyLead id="30A98207FFF0172BFF0D240D89E58226" box="[151,1436,1542,1563]" pageId="4" pageNumber="399">
2. Pronotum and elytra brownish black...............................................
<taxonomicName id="4C585AB1FFF0172BFBED240D89E58226" authority="Kiesenwetter, 1874" authorityName="Kiesenwetter" authorityYear="1874" box="[1143,1436,1542,1563]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B92CFD20FFF0172BFBED240D88F28226" box="[1143,1163,1542,1563]" italics="true" pageId="4" pageNumber="399">M</emphasis>
.
<emphasis id="B92CFD20FFF0172BFB0D240D88A58226" box="[1175,1244,1542,1563]" italics="true" pageId="4" pageNumber="399">bicolor</emphasis>
<bibRefCitation id="EFC95CC3FFF0172BFB78240D89E58226" box="[1250,1436,1542,1563]" pageId="4" pageNumber="399" refId="ref3084" refString="Kiesenwetter, H. von (1874) Die Malacodermen Japans nach dem Ergebnisse der Sammlungen des Herrn G. Lewis wahrend der Jahre 1869 - 1871. Berliner Entomologische Zeitschrift, 18, 241 - 288. https: // doi. org / 10.1002 / mmnd. 18740180302" type="journal article">Kiesenwetter, 1874</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE72132FFF0172BFF0D242F89E58204" blockId="4.[151,1436,1484,1828]" box="[151,1436,1572,1593]" pageId="4" pageNumber="399">
<keyLead id="30A98207FFF0172BFF0D242F89E58204" box="[151,1436,1572,1593]" pageId="4" pageNumber="399">
- Pronotum and elytra dark brown...................................................
<taxonomicName id="4C585AB1FFF0172BFB1B242F89E58204" authority="Takahashi, 2016" authorityName="Takahashi" authorityYear="2016" box="[1153,1436,1572,1593]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="tsuchidai">
<emphasis id="B92CFD20FFF0172BFB1B242F88EC8204" box="[1153,1173,1572,1593]" italics="true" pageId="4" pageNumber="399">M</emphasis>
.
<emphasis id="B92CFD20FFF0172BFB3B242F88808204" box="[1185,1273,1572,1593]" italics="true" pageId="4" pageNumber="399">tsuchidai</emphasis>
<bibRefCitation id="EFC95CC3FFF0172BFB65242F89E58204" box="[1279,1436,1572,1593]" pageId="4" pageNumber="399" refId="ref3250" refString="Takahashi, K. (2016) A new species of the genus Malthinellus (Coleoptera, Cantharidae) from Iriomote Island of the Ryukyu Islands, Southwest Japan. Elytra, New Series, Tokyo, 6 (2), 295 - 298." type="journal article">Takahashi, 2016</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AC3997FFF0172BFF0D244A89E2824E" pageId="4" pageNumber="399">
<paragraph id="8BE72132FFF0172BFF0D244A89E5826B" blockId="4.[151,1436,1484,1828]" box="[151,1436,1601,1622]" pageId="4" pageNumber="399">
<keyLead id="30A98207FFF0172BFF0D244A89E5826B" box="[151,1436,1601,1622]" pageId="4" pageNumber="399">
3. Head blackish brown...............................................................
<taxonomicName id="4C585AB1FFF0172BFB3A244A89E5826B" authority="(Wittmer, 1961)" authorityName="Wittmer" authorityYear="1961" baseAuthorityName="Wittmer" baseAuthorityYear="1961" box="[1184,1436,1601,1622]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="chujoi">
<emphasis id="B92CFD20FFF0172BFB3A244A88CD826B" box="[1184,1204,1601,1622]" italics="true" pageId="4" pageNumber="399">M</emphasis>
.
<emphasis id="B92CFD20FFF0172BFB5A244A8885826B" box="[1216,1276,1601,1622]" italics="true" pageId="4" pageNumber="399">chujoi</emphasis>
(Wittmer, 1961)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE72132FFF0172BFF0D245589E2824E" blockId="4.[151,1436,1484,1828]" box="[151,1435,1630,1651]" pageId="4" pageNumber="399">
<keyLead id="30A98207FFF0172BFF0D245589E2824E" box="[151,1435,1630,1651]" pageId="4" pageNumber="399">- Head yellowish orange to orange......................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30AC3997FFF0172BFF0D247789E58319" pageId="4" pageNumber="399">
<paragraph id="8BE72132FFF0172BFF0D247789E582F6" blockId="4.[151,1436,1484,1828]" pageId="4" pageNumber="399">
<keyLead id="30A98207FFF0172BFF0D247789E582F6" pageId="4" pageNumber="399">
4. Male genitalia with poorly developed inner lateral lobes, outer lateral lobes extended vertically; female terminalia with rela- tively elongate coxites in ventral view, paraprocts basally widened in outline in ventral view and faintly warped around apex in dorso-lateral view................................................
<taxonomicName id="4C585AB1FFF0172BFC4324BD89E582F6" authority="Takahashi &amp; N. Takahashi, 2007" authorityName="Takahashi &amp; N. Takahashi" authorityYear="2007" box="[985,1436,1718,1739]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="masatakai">
<emphasis id="B92CFD20FFF0172BFC4324BD8F9482F6" box="[985,1005,1718,1739]" italics="true" pageId="4" pageNumber="399">M</emphasis>
.
<emphasis id="B92CFD20FFF0172BFC6024BD882482F6" box="[1018,1117,1718,1739]" italics="true" pageId="4" pageNumber="399">masatakai</emphasis>
<bibRefCitation id="EFC95CC3FFF0172BFBF824BD89E582F6" box="[1122,1436,1718,1739]" pageId="4" pageNumber="399" refId="ref3296" refString="Takahashi, K. &amp; Takahashi, N. (2007) A new species of the genus Malthinellus (Coleoptera, Cantharidae) from Okinawa Island, Southwest Japan, with notes on the genus Malthinellus. Elytra, Tokyo, 35 (1), 142 - 148." type="journal article">Takahashi &amp; N. Takahashi, 2007</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE72132FFF0172BFF0D24DF89E58319" blockId="4.[151,1436,1484,1828]" pageId="4" pageNumber="399">
<keyLead id="30A98207FFF0172BFF0D24DF89E58319" pageId="4" pageNumber="399">
- Male genitalia with well-developed inner lateral lobes, outer lateral lobes slender in basal part and nearly securiform in apical part; female terminalia with relatively stout coxites in ventral view, paraprocts gently widened basally in outline in ventral view and less warped around apex in dorso-lateral view.............................
<taxonomicName id="4C585AB1FFF0172BFBCB2505893C831E" authority="N. Takahashi" authorityName="Takahashi" authorityYear="2018" box="[1105,1349,1806,1827]" class="Insecta" family="Cantharidae" genus="Malthinellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="exiguus" status="sp. nov.">
<emphasis id="B92CFD20FFF0172BFBCB25058813831E" box="[1105,1130,1806,1827]" italics="true" pageId="4" pageNumber="399">M</emphasis>
.
<emphasis id="B92CFD20FFF0172BFBEC250588C6831E" box="[1142,1215,1806,1827]" italics="true" pageId="4" pageNumber="399">exiguus</emphasis>
N. Takahashi
</taxonomicName>
,
<taxonomicNameLabel id="A21F405BFFF0172BFAD5250589E58319" box="[1359,1436,1806,1828]" pageId="4" pageNumber="399" rank="species">
<emphasis id="B92CFD20FFF0172BFAD5250589E58319" bold="true" box="[1359,1436,1806,1828]" pageId="4" pageNumber="399">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>