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<mods:title id="134014118195AA4926C05A927A220A8D">The Living Columbariinae (Gastropoda: Neogastropoda: Turbinellidae) of New Zealand</mods:title>
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<paragraph id="8BE736EBFFA3FF92FF4211FEFD9CFF76" blockId="23.[151,583,151,177]" box="[151,583,151,177]" pageId="23" pageNumber="24">
<heading id="D0AF8187FFA3FF92FF4211FEFD9CFF76" bold="true" box="[151,583,151,177]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="B92CEAF9FFA3FF92FF4211FEFD9CFF76" bold="true" box="[151,583,151,177]" pageId="23" pageNumber="24">
Genus
<taxonomicName id="4C584D68FFA3FF92FF3B11FEFD9CFF76" ID-CoL="7P445" authority="Grabau, 1904" authorityName="Grabau" authorityYear="1904" box="[238,583,151,177]" class="Gastropoda" family="Columbariidae" genus="Fulgurofusus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
<emphasis id="B92CEAF9FFA3FF92FF3B11FEFE4FFF76" bold="true" box="[238,404,151,177]" italics="true" pageId="23" pageNumber="24">Fulgurofusus</emphasis>
<bibRefCitation id="EFC94B1AFFA3FF92FE4E11FEFD9CFF76" author="Grabau, A. W." box="[411,583,151,177]" pageId="23" pageNumber="24" pagination="1 - 157" refId="ref26173" refString="Grabau, A. W. (1904) Phylogeny of Fusus and its allies. Smithsonian Miscellaneous Collections, 44 (1417), 1 - 157, pls. 1 - 18." type="journal article" year="1904">Grabau, 1904</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE736EBFFA3FF92FF4211B7FED6FF33" blockId="23.[151,1409,222,306]" box="[151,269,222,244]" pageId="23" pageNumber="24">Synonymy:</paragraph>
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<paragraph id="8BE736EBFFA3FF92FF421194FDA7FEF6" blockId="23.[151,1409,222,306]" pageId="23" pageNumber="24">
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<treatmentCitation id="0AF910FAFFA3FF92FF421194FE0BFED4" author="Grabau, A. W." box="[151,464,253,275]" page="86" pageId="23" pageNumber="24" year="1904">
<taxonomicName id="4C584D68FFA3FF92FF421194FE0BFED4" ID-CoL="7P445" authority="Grabau, 1904: 86" authorityName="Grabau" authorityPageNumber="86" authorityYear="1904" box="[151,464,253,275]" class="Gastropoda" family="Columbariidae" genus="Fulgurofusus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
<emphasis id="B92CEAF9FFA3FF92FF421194FEC6FED4" box="[151,285,253,275]" italics="true" pageId="23" pageNumber="24">Fulgurofusus</emphasis>
<bibRefCitation id="EFC94B1AFFA3FF92FEF61194FE0BFED4" author="Grabau, A. W." box="[291,464,253,275]" pageId="23" pageNumber="24" pagination="1 - 157" refId="ref26173" refString="Grabau, A. W. (1904) Phylogeny of Fusus and its allies. Smithsonian Miscellaneous Collections, 44 (1417), 1 - 157, pls. 1 - 18." type="journal article" year="1904">Grabau, 1904:86</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AF910FAFFA3FF92FE0E1197FD4EFED4" author="Wenz, W." box="[475,661,253,275]" page="1086" pageId="23" pageNumber="24" year="1941">
<bibRefCitation id="EFC94B1AFFA3FF92FE0E1197FD4EFED4" author="Wenz, W." box="[475,661,253,275]" pageId="23" pageNumber="24" pagination="961 - 1200" refId="ref28335" refString="Wenz, W. (1941) Gastropoda. Teil I: Allgemeiner Teil und Prosobranchia In: Schindewolf, O. H. (Ed.), Handbuch der Palaozoologie, Gebruder Borntraeger, Berlin, Band 6, Tiel 5, 961 - 1200." type="journal article" year="1941">Wenz, 1941: 1086</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF910FAFFA3FF92FD751194FC1CFED4" author="Shimer, H. W. &amp; Shrock, R. R." box="[672,967,253,275]" page="507" pageId="23" pageNumber="24" year="1944">
<bibRefCitation id="EFC94B1AFFA3FF92FD751194FC1CFED4" author="Shimer, H. W. &amp; Shrock, R. R." box="[672,967,253,275]" pageId="23" pageNumber="24" refId="ref27716" refString="Shimer, H. W. &amp; Shrock, R. R. (1944) Index fossils of North America. Wiley and Sons, New York, ix + 837 pp." type="book" year="1944">Shimer &amp; Shrock, 1944: 507</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF910FAFFA3FF92FC041194FB56FED4" author="Darragh, T. A." box="[977,1165,253,275]" page="99" pageId="23" pageNumber="24" year="1969">
<bibRefCitation id="EFC94B1AFFA3FF92FC041194FB56FED4" author="Darragh, T. A." box="[977,1165,253,275]" pageId="23" pageNumber="24" pagination="63 - 119" refId="ref25856" refString="Darragh, T. A. (1969) A review of the family Columbariidae (Mollusca: Gastropoda). Proceedings of the Royal Society of Victoria, 83, 63 - 119." type="journal article" year="1969">Darragh, 1969: 99</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF910FAFFA3FF92FB4D1194FAA6FED4" author="Harasewych, M. G." box="[1176,1405,253,275]" page="5" pageId="23" pageNumber="24" year="1983">
<bibRefCitation id="EFC94B1AFFA3FF92FB4D1194FAA6FED4" author="Harasewych, M. G." box="[1176,1405,253,275]" pageId="23" pageNumber="24" pagination="1 - 42" refId="ref26374" refString="Harasewych, M. G. (1983 b) A Review of the Columbariinae (Gastropoda: Turbinellidae) of the western Atlantic with notes on the anatomy and systematic relationships of the subfamily. Nemouria, 27, 1 - 42." type="journal article" year="1983">Harasewych, 1983b: 5</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AF910FAFFA3FF92FF421075FDA3FEF6" author="Bayer, F. M." box="[151,632,283,306]" page="170" pageId="23" pageNumber="24" year="1971">
<taxonomicName id="4C584D68FFA3FF92FF421075FDA3FEF6" authority="Bayer, 1971: 170" authorityName="Bayer" authorityPageNumber="170" authorityYear="1971" box="[151,632,283,306]" class="Gastropoda" family="Columbariidae" genus="Columbarium" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="subGenus" subGenus="Fulgurofusus">
<emphasis id="B92CEAF9FFA3FF92FF421075FEFFFEF5" box="[151,292,284,306]" italics="true" pageId="23" pageNumber="24">Columbarium</emphasis>
(
<emphasis id="B92CEAF9FFA3FF92FEE71075FE63FEF5" box="[306,440,284,306]" italics="true" pageId="23" pageNumber="24">Fulgurofusus</emphasis>
)
<bibRefCitation id="EFC94B1AFFA3FF92FE121075FDA3FEF6" author="Bayer, F. M." box="[455,632,283,305]" pageId="23" pageNumber="24" pagination="111 - 236" refId="ref25305" refString="Bayer, F. M. (1971) New and unusual mollusks collected by R / V John Elliott Pillsbury and R / V Gerda in the tropical western Atlantic. Bulletin of Marine Science, 21, 111 - 236." type="journal article" year="1971">Bayer, 1971: 170</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
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<paragraph id="8BE736EBFFA3FF92FF421036FC47FEB1" blockId="23.[151,1437,350,1166]" box="[151,924,350,375]" pageId="23" pageNumber="24">
<emphasis id="B92CEAF9FFA3FF92FF421036FEE9FEB0" bold="true" box="[151,306,350,375]" inLineHeading="true" pageId="23" pageNumber="24" reason="1">
<typeStatus id="54E38849FFA3FF92FF421036FF0AFEB0" box="[151,209,351,375]" pageId="23" pageNumber="24">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C584D68FFA3FF92FEEC1036FD53FEB1" authority="Harris, 1896" authorityName="Harris" authorityYear="1896" box="[313,648,350,374]" class="Gastropoda" family="Fasciolariidae" genus="Fusus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="quercollis">
<emphasis id="B92CEAF9FFA3FF92FEEC1036FE29FEB1" box="[313,498,350,374]" italics="true" pageId="23" pageNumber="24">Fusus quercollis</emphasis>
<bibRefCitation id="EFC94B1AFFA3FF92FE2C1037FD53FEB1" author="Harris, G. D." box="[505,648,350,374]" pageId="23" pageNumber="24" pagination="117 - 270" refId="ref26645" refString="Harris, G. D. (1896) The Midway Stage. Bulletins of American Paleontology, No. 4. 1, 117 - 270, pls. 11 - 25." type="journal article" year="1896">Harris, 1896</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C3426560FFA3FF92FF1210EBFF39FCC5" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8BE736EBFFA3FF92FF1210EBFF39FCC5" blockId="23.[151,1437,350,1166]" pageId="23" pageNumber="24">
<emphasis id="B92CEAF9FFA3FF92FF1210EBFE9AFE5C" bold="true" box="[199,321,386,411]" inLineHeading="true" pageId="23" pageNumber="24" reason="1">Diagnosis.</emphasis>
Shell of moderate size (to
<quantity id="4CA09B0EFFA3FF92FD5310EBFD06FE5D" box="[646,733,386,410]" metricMagnitude="-2" metricUnit="m" metricValue="8.6" pageId="23" pageNumber="24" unit="mm" value="86.0">86 mm</quantity>
), fusiform, with short or tall, conical spire, rounded whorls, peripheral keel that may be prominent and flange-like or a weakly nodulose cord, weak anterior carina, and stout, axial siphonal canal of moderate length. Protoconch cylindrical to weakly conical, of 1½3 whorls. First whorl bulbous, deviated. Transition to teleoconch generally indistinct. Suture adpressed onto or slightly above anterior carina. Spiral sculpture of numerous fine threads and weak cords extending from suture to middle of siphonal canal. Early whorls with two closely adjacent cords along periphery, forming an incised furrow between them. Axial sculpture of small nodules along periphery, or low ribs extending from suture to siphonal canal. Outer lip porcellaneous, furrowed beneath periphery. Inner lip smooth, with outermost layers of previous whorls resorbed prior to deposition of a thin glaze that does not produce a raised lamina. Shell color uniformly white. Rachidian teeth of radula with 3 cusps on a U-shaped basal plate with prominent lateral expansions (e.g.
<bibRefCitation id="EFC94B1AFFA3FF92FB6913AFFA43FD19" author="Harasewych, M. G." box="[1212,1432,710,734]" pageId="23" pageNumber="24" pagination="1 - 42" refId="ref26374" refString="Harasewych, M. G. (1983 b) A Review of the Columbariinae (Gastropoda: Turbinellidae) of the western Atlantic with notes on the anatomy and systematic relationships of the subfamily. Nemouria, 27, 1 - 42." type="journal article" year="1983">Harasewych 1983b</bibRefCitation>
: fig. 9).
</paragraph>
</subSubSection>
<subSubSection id="C3426560FFA3FF92FF121266FE5AFB49" pageId="23" pageNumber="24" type="discussion">
<paragraph id="8BE736EBFFA3FF92FF121266FDA2FB81" blockId="23.[151,1437,350,1166]" pageId="23" pageNumber="24">
<emphasis id="B92CEAF9FFA3FF92FF121266FEE0FCE0" bold="true" box="[199,315,783,807]" inLineHeading="true" pageId="23" pageNumber="24" reason="1">Remarks.</emphasis>
The genus
<taxonomicName id="4C584D68FFA3FF92FE111267FD83FCE1" authorityName="Grabau" authorityYear="1904" box="[452,600,782,806]" class="Gastropoda" family="Columbariidae" genus="Fulgurofusus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
<emphasis id="B92CEAF9FFA3FF92FE111267FD83FCE1" box="[452,600,782,806]" italics="true" pageId="23" pageNumber="24">Fulgurofusus</emphasis>
</taxonomicName>
is based on a Paleocene
<typeStatus id="54E38849FFA3FF92FCA81266FC76FCE0" box="[893,941,783,807]" pageId="23" pageNumber="24">type</typeStatus>
species from the Gulf Coast of the
<collectingCountry id="F34F767BFFA3FF92FA851267FF06FC8D" name="United States of America" pageId="23" pageNumber="24">United States</collectingCountry>
. Several Eocene species have been reported from western North America and
<collectingCountry id="F34F767BFFA3FF92FBAD125BFB2BFC8D" box="[1144,1264,818,842]" name="Antarctica" pageId="23" pageNumber="24">Antarctica</collectingCountry>
. In the Recent fauna,
<taxonomicName id="4C584D68FFA3FF92FF31123FFEA3FCA9" authorityName="Grabau" authorityYear="1904" box="[228,376,854,878]" class="Gastropoda" family="Columbariidae" genus="Fulgurofusus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
<emphasis id="B92CEAF9FFA3FF92FF31123FFEA3FCA9" box="[228,376,854,878]" italics="true" pageId="23" pageNumber="24">Fulgurofusus</emphasis>
</taxonomicName>
is the most wide ranging genus within
<taxonomicName id="4C584D68FFA3FF92FCEB123FFC3CFCA9" authorityName="Tomlin" authorityYear="1928" box="[830,999,854,878]" class="Gastropoda" family="Columbariidae" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="subFamily" subFamily="Columbariinae">Columbariinae</taxonomicName>
, extending from the Bering Sea to the Bounty Plateau along the western Pacific, and from the Blake Plateau to the Scotia Sea in the western Atlantic.
<taxonomicName id="4C584D68FFA3FF92FF4212F7FD11FC71" authority="McLean &amp; Andrade, 1982" authorityName="McLean &amp; Andrade" authorityYear="1982" box="[151,714,926,950]" class="Gastropoda" family="Columbariidae" genus="Columbarium" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="tomicici">
<emphasis id="B92CEAF9FFA3FF92FF4212F7FE4EFC71" box="[151,405,926,950]" italics="true" pageId="23" pageNumber="24">Columbarium tomicici</emphasis>
<bibRefCitation id="EFC94B1AFFA3FF92FE4912F6FD11FC71" author="McLean, J. H. &amp; Andrade V, H." box="[412,714,926,950]" pageId="23" pageNumber="24" pagination="1 - 20" refId="ref27302" refString="McLean, J. H. &amp; Andrade V, H. (1982) Large archibenthal gastropods of central Chile: collections from an expedition of the R / V Anton Bruun and the Chilean shrimp fishery. Natural History Museum of Los Angeles County, Contributions in Science, 342, 1 - 20." type="journal article" year="1982">McLean &amp; Andrade, 1982</bibRefCitation>
</taxonomicName>
, from lower bathyal depths off
<collectingCountry id="F34F767BFFA3FF92FBE412F7FBB5FC71" box="[1073,1134,926,950]" name="Chile" pageId="23" pageNumber="24">Chile</collectingCountry>
is likely referrable to
<taxonomicName id="4C584D68FFA3FF92FAB912F7FEDFFC1C" authorityName="Grabau" authorityYear="1904" class="Gastropoda" family="Columbariidae" genus="Fulgurofusus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
<emphasis id="B92CEAF9FFA3FF92FAB912F7FEDFFC1C" italics="true" pageId="23" pageNumber="24">Fulgurofusus</emphasis>
</taxonomicName>
[based on the lack of a raised columellar lamina and the presence of rachidian teeth with a laterally expanded basal plate (
<bibRefCitation id="EFC94B1AFFA3FF92FE46128FFD65FC39" author="McLean, J. H. &amp; Andrade V, H." box="[403,702,998,1022]" pageId="23" pageNumber="24" pagination="1 - 20" refId="ref27302" refString="McLean, J. H. &amp; Andrade V, H. (1982) Large archibenthal gastropods of central Chile: collections from an expedition of the R / V Anton Bruun and the Chilean shrimp fishery. Natural History Museum of Los Angeles County, Contributions in Science, 342, 1 - 20." type="journal article" year="1982">McLean &amp; Andrade 1982</bibRefCitation>
: figs. 2630)], further expanding the range of this genus. Unlike other columbariine genera, the bathymetric range of
<taxonomicName id="4C584D68FFA3FF92FD3F1562FCA5FBE4" authorityName="Grabau" authorityYear="1904" box="[746,894,1035,1059]" class="Gastropoda" family="Columbariidae" genus="Fulgurofusus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
<emphasis id="B92CEAF9FFA3FF92FD3F1562FCA5FBE4" box="[746,894,1035,1059]" italics="true" pageId="23" pageNumber="24">Fulgurofusus</emphasis>
</taxonomicName>
extends to the lower continental slope and onto the abyssal plain, especially near the poles.
</paragraph>
<paragraph id="8BE736EBFFA3FF92FF12153BFE5AFB49" blockId="23.[151,1437,350,1166]" pageId="23" pageNumber="24">
<bibRefCitation id="EFC94B1AFFA3FF92FF12153BFE3FFBAD" author="Finlay, H. J." box="[199,484,1106,1130]" pageId="23" pageNumber="24" pagination="259 - 270" refId="ref26098" refString="Finlay, H. J. (1930 b) Revision of the New Zealand shells referred to Fusinus. Transactions of the New Zealand Institute, 61, 259 - 270, pls. 47 - 49." type="journal article" year="1930">Finlay (1930b: 267268)</bibRefCitation>
considered
<taxonomicName id="4C584D68FFA3FF92FDA4153AFCDEFBAC" authorityName="Grabau" authorityYear="1904" box="[625,773,1107,1131]" class="Gastropoda" family="Columbariidae" genus="Fulgurofusus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
<emphasis id="B92CEAF9FFA3FF92FDA4153AFCDEFBAC" box="[625,773,1107,1131]" italics="true" pageId="23" pageNumber="24">Fulgurofusus</emphasis>
</taxonomicName>
to be less advanced than
<taxonomicName id="4C584D68FFA3FF92FBE1153AFB4BFBAC" authorityName=", Allan" authorityYear="1926" box="[1076,1168,1107,1131]" class="Gastropoda" family="Columbariidae" genus="Coluzea" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
<emphasis id="B92CEAF9FFA3FF92FBE1153AFB4BFBAC" box="[1076,1168,1107,1131]" italics="true" pageId="23" pageNumber="24">Coluzea</emphasis>
</taxonomicName>
, and presumed it to be ancestral to
<taxonomicName id="4C584D68FFA3FF92FECB151FFEA1FB49" authorityName=", Allan" authorityYear="1926" box="[286,378,1142,1166]" class="Gastropoda" family="Columbariidae" genus="Coluzea" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
<emphasis id="B92CEAF9FFA3FF92FECB151FFEA1FB49" box="[286,378,1142,1166]" italics="true" pageId="23" pageNumber="24">Coluzea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>