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<document id="7AE6A96069E2F1AEFD28579BB527D401" ID-CLB-Dataset="295801" ID-DOI="10.11646/zootaxa.5447.4.4" ID-GBIF-Dataset="060af3f0-7183-4cbf-8ff8-2d8950649151" ID-ISSN="1175-5326" ID-Zenodo-Dep="11150484" ID-ZooBank="A07DCF7E-7F1A-4FE1-BDE6-AB394522390D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1715181961429" checkinUser="plazi" docAuthor="Shi, Songtao, Mu, Yannan &amp; Zhang, Feng" docDate="2024" docId="03F187FAFFA1FF96FF35ED9CA687FEFD" docLanguage="en" docName="zootaxa.5447.4.4.pdf" docOrigin="Zootaxa 5447 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Otacilia pugnus Shi &amp; Mu &amp; Zhang 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="522" masterDocId="FFC8FF82FFA4FF9FFFA2EF4AA406FF98" masterDocTitle="Description of four new species of Otacilia Thorell, 1897 (Araneae: Phrurolithidae) from southern China" masterLastPageNumber="530" masterPageNumber="513" pageNumber="518" updateTime="1715182649704" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FBD90CB6639100AED7BEC315E75F71BD">Description of four new species of Otacilia Thorell, 1897 (Araneae: Phrurolithidae) from southern China</mods:title>
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<mods:affiliation id="4548DFD59E899B0B367A2BAE4954839A">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China &amp; Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China</mods:affiliation>
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<mods:affiliation id="F3051803FF81E7C7BFE416D86A9A96BC">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China &amp; Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China</mods:affiliation>
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<mods:namePart id="AD8A79C60FDC76A3B44947DF285EC34B">Zhang, Feng</mods:namePart>
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<mods:affiliation id="108C079C90EDE5F2C6F6F2207E4C4ADF">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China &amp; Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China</mods:affiliation>
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<subSubSection id="C3426567FFA1FF9AFF35ED9CA648FD68" box="[151,590,726,753]" pageId="5" pageNumber="518" type="nomenclature">
<paragraph id="8BE736ECFFA1FF9AFF35ED9CA648FD68" blockId="5.[151,590,726,789]" box="[151,590,726,753]" pageId="5" pageNumber="518">
<heading id="D0AF8180FFA1FF9AFF35ED9CA648FD68" bold="true" box="[151,590,726,753]" fontSize="11" level="1" pageId="5" pageNumber="518" reason="1">
<emphasis id="B92CEAFEFFA1FF9AFF35ED9CA648FD68" bold="true" box="[151,590,726,753]" pageId="5" pageNumber="518">
<taxonomicName id="4C584D6FFFA1FF9AFF35ED9CA55FFD69" authority="Shi &amp; Mu &amp; Zhang, 2024" authorityName="Shi &amp; Mu &amp; Zhang" authorityYear="2024" box="[151,345,726,753]" class="Arachnida" family="Phrurolithidae" genus="Otacilia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="518" phylum="Arthropoda" rank="species" species="pugnus" status="sp. nov.">
<emphasis id="B92CEAFEFFA1FF9AFF35ED9CA55FFD69" bold="true" box="[151,345,726,753]" italics="true" pageId="5" pageNumber="518">Otacilia pugnus</emphasis>
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<taxonomicNameLabel id="A21F5785FFA1FF9AFEC2ED9DA5BBFD69" box="[352,445,727,753]" pageId="5" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
(Þff⁂ặ)
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<paragraph id="8BE736ECFFA1FF9AFF35EDB1A524FC8D" blockId="5.[151,590,726,789]" box="[151,290,763,789]" pageId="5" pageNumber="518">
<figureCitation id="13632A69FFA1FF9AFF35EDB1A4FDFC8D" box="[151,251,763,789]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="6.[151,250,1943,1967]" captionStartId-1="7.[151,250,1869,1893]" captionStartId-2="8.[151,250,1943,1967]" captionTargetBox-0="[211,1373,181,1907]" captionTargetBox-1="[173,1416,189,1835]" captionTargetBox-2="[163,1424,189,1910]" captionTargetId-0="figure-23@6.[198,1390,181,1919]" captionTargetId-1="figure-17@7.[170,1416,181,1845]" captionTargetId-2="figure-23@8.[155,1433,181,1919]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURE 4. Habitus of Otacilia pugnus sp. nov.: A. Male holotype, dorsal view; B. Same, ventral view; C. Female paratype, dorsal view; D. Same, ventral view." captionText-1="FIGURE 5. Copulatory organs of Otacilia pugnus sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C— conductor; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT—connecting tube; FD—fertilization duct; MS—median septum; S—spermathecae." captionText-2="FIGURE 6. Copulatory organs of Otacilia pugnus sp. nov.: A. Male left palp, prolateral view; B. same, retrolateral view; C. Same, ventral view; D. same, dorsal view; E. epigyne, ventral view; F. Same, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.11150496" figureDoi-1="http://doi.org/10.5281/zenodo.11150500" figureDoi-2="http://doi.org/10.5281/zenodo.11150506" httpUri-0="https://zenodo.org/record/11150496/files/figure.png" httpUri-1="https://zenodo.org/record/11150500/files/figure.png" httpUri-2="https://zenodo.org/record/11150506/files/figure.png" pageId="5" pageNumber="518">Figs 4–6</figureCitation>
,
<figureCitation id="13632A69FFA1FF9AFEA4EDB1A524FC8D" box="[262,290,763,789]" captionStart="FIGURE 13" captionStartId="16.[151,250,1421,1445]" captionTargetBox="[157,1436,499,1398]" captionTargetId="figure-119@16.[151,1436,489,1398]" captionTargetPageId="16" captionText="FIGURE 13. Distribution of the new species described in this paper." figureDoi="http://doi.org/10.5281/zenodo.11150528" httpUri="https://zenodo.org/record/11150528/files/figure.png" pageId="5" pageNumber="518">13</figureCitation>
</paragraph>
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<paragraph id="8BE736ECFFA1FF9AFF35EC09A112FC19" blockId="5.[151,1437,834,2013]" pageId="5" pageNumber="518">
<emphasis id="B92CEAFEFFA1FF9AFF35EC09A540FCC4" bold="true" box="[151,326,834,861]" pageId="5" pageNumber="518">Type material.</emphasis>
<materialsCitation id="3B303CB1FFA1FF9AFEECEC09A112FC19" collectingDate="2018-10-30" collectionCode="MHBU" collectorName="Z. Li." country="China" county="Shimen Co." elevation="362" latitude="30.004139" location="Hupingshan Town" longLatPrecision="1" longitude="110.47856" municipality="Changde City" pageId="5" pageNumber="518" specimenCount="1" stateProvince="Hunan" typeStatus="holotype">
<emphasis id="B92CEAFEFFA1FF9AFEECEC09A5D1FCC5" bold="true" box="[334,471,835,861]" pageId="5" pageNumber="518">
<typeStatus id="54E3884EFFA1FF9AFEECEC09A5BDFCC5" box="[334,443,835,861]" pageId="5" pageNumber="518" type="holotype">Holotype</typeStatus>
J
</emphasis>
(
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),
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<collectingCountry id="F34F767CFFA1FF9AFDF7EC08A6B1FCC4" box="[597,695,834,860]" name="China" pageId="5" pageNumber="518">CHINA</collectingCountry>
:
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<emphasis id="B92CEAFEFFA1FF9AFD61EC09A77DFCC4" box="[707,891,835,860]" italics="true" pageId="5" pageNumber="518">
<collectingRegion id="499CF80EFFA1FF9AFD61EC09A77DFCC4" box="[707,891,835,860]" country="China" name="Hunan" pageId="5" pageNumber="518">Hunan Province</collectingRegion>
</emphasis>
:
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,
<collectingCounty id="62864E60FFA1FF9AFB91EC09A0BDFCC5" box="[1075,1211,835,861]" pageId="5" pageNumber="518">Shimen Co.</collectingCounty>
,
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, Jiangpinghe Vill.,
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(
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,
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,
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a.s.l.
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),
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<collectingDate id="EFA2E9C4FFA1FF9AFBBAEC2DA09CFC19" box="[1048,1178,871,897]" pageId="5" pageNumber="518" value="2018-10-30">30.10.2018</collectingDate>
</date>
, leg.
<collectorName id="26AD533AFFA1FF9AFB77EC2DA112FC19" box="[1237,1300,871,897]" pageId="5" pageNumber="518">Z. Li.</collectorName>
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</paragraph>
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<materialsCitation id="3B303CB1FFA1FF9AFF65ECC1A74DFC3D" box="[199,843,907,935]" collectingDate="2018-10-30" collectionCode="MHBU" collectorName="Z. Li." country="China" county="Shimen Co." elevation="362" latitude="30.004139" location="Hupingshan Town" longLatPrecision="1" longitude="110.47856" municipality="Changde City" pageId="5" pageNumber="518" specimenCount="7" specimenCount-female="7" stateProvince="Hunan" typeStatus="paratype">
<emphasis id="B92CEAFEFFA1FF9AFF65ECC1A54EFC3D" bold="true" box="[199,328,907,933]" pageId="5" pageNumber="518">
<typeStatus id="54E3884EFFA1FF9AFF65ECC1A545FC3D" box="[199,323,907,933]" pageId="5" pageNumber="518" type="paratype">Paratypes</typeStatus>
:
</emphasis>
2♁
<specimenCount id="9D5EFD65FFA1FF9AFEDBECC1A59BFC3D" box="[377,413,907,933]" count="7" pageId="5" pageNumber="518" type="female">7♀</specimenCount>
(
<collectionCode id="ED49AE29FFA1FF9AFE09ECC1A603FC3D" box="[427,517,907,933]" pageId="5" pageNumber="518">MHBU</collectionCode>
), with same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426567FFA1FF9AFF65ECE5A549FC75" pageId="5" pageNumber="518" type="etymology">
<paragraph id="8BE736ECFFA1FF9AFF65ECE5A549FC75" blockId="5.[151,1437,834,2013]" pageId="5" pageNumber="518">
<emphasis id="B92CEAFEFFA1FF9AFF65ECE5A548FC51" bold="true" box="[199,334,943,969]" pageId="5" pageNumber="518">Etymology.</emphasis>
The specific epithet is derived from the Latin word
<taxonomicName id="4C584D6FFFA1FF9AFC21ECE5A7E6FC50" authorityName="Shi &amp; Mu &amp; Zhang" authorityYear="2024" box="[899,992,943,969]" class="Arachnida" family="Phrurolithidae" genus="Otacilia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="518" phylum="Arthropoda" rank="species" species="pugnus">
“
<emphasis id="B92CEAFEFFA1FF9AFC2DECE5A7E6FC50" box="[911,992,943,968]" italics="true" pageId="5" pageNumber="518">pugnus</emphasis>
</taxonomicName>
”, referring to the fist-shaped retrolateral tibial apophysis.
</paragraph>
</subSubSection>
<subSubSection id="C3426567FFA1FF9AFF65ECBCA537FB39" pageId="5" pageNumber="518" type="diagnosis">
<paragraph id="8BE736ECFFA1FF9AFF65ECBCA537FB39" blockId="5.[151,1437,834,2013]" pageId="5" pageNumber="518">
<emphasis id="B92CEAFEFFA1FF9AFF65ECBCA544FB88" bold="true" box="[199,322,1014,1040]" pageId="5" pageNumber="518">Diagnosis.</emphasis>
The new species resembles
<taxonomicName id="4C584D6FFFA1FF9AFD23ECBDA7ECFB89" authority="Yao, Irfan &amp; Peng, 2019" authorityName="Yao, Irfan &amp; Peng" authorityYear="2019" box="[641,1002,1015,1041]" class="Arachnida" family="Phrurolithidae" genus="Otacilia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="518" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B92CEAFEFFA1FF9AFD23ECBDA6D6FB89" box="[641,720,1015,1041]" italics="true" pageId="5" pageNumber="518">O. lata</emphasis>
<bibRefCitation id="EFC94B1DFFA1FF9AFD74ECBDA7ECFB89" author="Yao, N. &amp; Irfan, M. &amp; Peng, X. J." box="[726,1002,1015,1041]" pageId="5" pageNumber="518" pagination="290 - 304" refId="ref6236" refString="Yao, N., Irfan, M. &amp; Peng, X. J. (2019) Five new species of Otacilia Thorell, 1897 (Araneae: Phrurolithidae) from the Wuling Mountain Range, China. Zootaxa, 4613 (2), 290 - 304. https: // doi. org / 10.11646 / zootaxa. 4613.2.4" type="journal article" year="2019">
Yao, Irfan &amp; Peng,
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</bibRefCitation>
</taxonomicName>
in having a similarly shaped embolus, tegular apophysis and bursa, but can be recognized by: 1) the retrolateral tibial apophysis short, fist-shaped, with digitiform tip (vs long, rectangular, cf.
<figureCitation id="13632A69FFA1FF9AFDECEB75A6A4FBC1" box="[590,674,1086,1113]" captionStart="FIGURE 5" captionStartId="7.[151,250,1869,1893]" captionTargetBox="[173,1416,189,1835]" captionTargetId="figure-17@7.[170,1416,181,1845]" captionTargetPageId="7" captionText="FIGURE 5. Copulatory organs of Otacilia pugnus sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C— conductor; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT—connecting tube; FD—fertilization duct; MS—median septum; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.11150500" httpUri="https://zenodo.org/record/11150500/files/figure.png" pageId="5" pageNumber="518">Fig. 5B</figureCitation>
and fig. 3G in
<bibRefCitation id="EFC94B1DFFA1FF9AFCEFEB75A008FBC1" author="Yao, N. &amp; Irfan, M. &amp; Peng, X. J." box="[845,1038,1087,1113]" pageId="5" pageNumber="518" pagination="290 - 304" refId="ref6236" refString="Yao, N., Irfan, M. &amp; Peng, X. J. (2019) Five new species of Otacilia Thorell, 1897 (Araneae: Phrurolithidae) from the Wuling Mountain Range, China. Zootaxa, 4613 (2), 290 - 304. https: // doi. org / 10.11646 / zootaxa. 4613.2.4" type="journal article" year="2019">
Yao
<emphasis id="B92CEAFEFFA1FF9AFC22EB75A7BFFBC1" box="[896,953,1087,1113]" italics="true" pageId="5" pageNumber="518">et al.</emphasis>
[2019]
</bibRefCitation>
), 2) conductor slender (vs wide, cf.
<figureCitation id="13632A69FFA1FF9AFF35EB29A50AFBE5" box="[151,268,1122,1149]" captionStart="FIGURE 5" captionStartId="7.[151,250,1869,1893]" captionTargetBox="[173,1416,189,1835]" captionTargetId="figure-17@7.[170,1416,181,1845]" captionTargetPageId="7" captionText="FIGURE 5. Copulatory organs of Otacilia pugnus sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C— conductor; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT—connecting tube; FD—fertilization duct; MS—median septum; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.11150500" httpUri="https://zenodo.org/record/11150500/files/figure.png" pageId="5" pageNumber="518">Fig. 5B–C</figureCitation>
and fig. 3F–G in
<bibRefCitation id="EFC94B1DFFA1FF9AFE70EB29A697FBE5" author="Yao, N. &amp; Irfan, M. &amp; Peng, X. J." box="[466,657,1123,1149]" pageId="5" pageNumber="518" pagination="290 - 304" refId="ref6236" refString="Yao, N., Irfan, M. &amp; Peng, X. J. (2019) Five new species of Otacilia Thorell, 1897 (Araneae: Phrurolithidae) from the Wuling Mountain Range, China. Zootaxa, 4613 (2), 290 - 304. https: // doi. org / 10.11646 / zootaxa. 4613.2.4" type="journal article" year="2019">
Yao
<emphasis id="B92CEAFEFFA1FF9AFDA7EB29A63BFBE5" box="[517,573,1123,1149]" italics="true" pageId="5" pageNumber="518">et al.</emphasis>
[2019]
</bibRefCitation>
), 3) copulatory opening round (vs oval, cf.
<figureCitation id="13632A69FFA1FF9AFBD5EB29A0CFFBE5" box="[1143,1225,1122,1149]" captionStart="FIGURE 5" captionStartId="7.[151,250,1869,1893]" captionTargetBox="[173,1416,189,1835]" captionTargetId="figure-17@7.[170,1416,181,1845]" captionTargetPageId="7" captionText="FIGURE 5. Copulatory organs of Otacilia pugnus sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C— conductor; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT—connecting tube; FD—fertilization duct; MS—median septum; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.11150500" httpUri="https://zenodo.org/record/11150500/files/figure.png" pageId="5" pageNumber="518">Fig. 5E</figureCitation>
and fig. 3C in
<bibRefCitation id="EFC94B1DFFA1FF9AFAD2EB29A523FB39" author="Yao, N. &amp; Irfan, M. &amp; Peng, X. J." pageId="5" pageNumber="518" pagination="290 - 304" refId="ref6236" refString="Yao, N., Irfan, M. &amp; Peng, X. J. (2019) Five new species of Otacilia Thorell, 1897 (Araneae: Phrurolithidae) from the Wuling Mountain Range, China. Zootaxa, 4613 (2), 290 - 304. https: // doi. org / 10.11646 / zootaxa. 4613.2.4" type="journal article" year="2019">
Yao
<emphasis id="B92CEAFEFFA1FF9AFF35EBCDA4C9FB39" box="[151,207,1159,1185]" italics="true" pageId="5" pageNumber="518">et al.</emphasis>
[2019]
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3426567FFA1FF96FF65EBE0A67BFED9" lastPageId="9" lastPageNumber="522" pageId="5" pageNumber="518" type="description">
<paragraph id="8BE736ECFFA1FF9AFF65EBE0A0B8FA05" blockId="5.[151,1437,834,2013]" pageId="5" pageNumber="518">
<emphasis id="B92CEAFEFFA1FF9AFF65EBE0A55FFB5C" bold="true" box="[199,345,1194,1220]" pageId="5" pageNumber="518">Description.</emphasis>
Male (
<emphasis id="B92CEAFEFFA1FF9AFE05EBE1A612FB5D" bold="true" box="[423,532,1195,1221]" pageId="5" pageNumber="518">
<typeStatus id="54E3884EFFA1FF9AFE05EBE1A612FB5D" box="[423,532,1195,1221]" pageId="5" pageNumber="518" type="holotype">Holotype</typeStatus>
</emphasis>
): total length 2.69, carapace 1.43 long, 1.23 wide; abdomen 1.26 long, 0.86 wide. Eye sizes and interdistances:AME 0.08, ALE 0.10, PME 0.08, PLE 0.10; AME–AME 0.04, AME–ALE 0.01, ALE– ALE 0.21, PME–PME 0.11, PME–PLE 0.06, PLE–PLE 0.38, ALE–PLE 0.08. EAW 0.50, CRW 0.74, EAW/CRW 0.68, CRW/CW 0.60. MOA 0.25 long, anterior width 0.20, posterior width 0.26.
<collectingCountry id="F34F767CFFA1FF9AFB86EA5DA04DFAA9" box="[1060,1099,1303,1329]" name="Switzerland" pageId="5" pageNumber="518">CH</collectingCountry>
0.14,
<collectingCountry id="F34F767CFFA1FF9AFB30EA5DA0BEFAA9" box="[1170,1208,1303,1329]" name="Switzerland" pageId="5" pageNumber="518">CH</collectingCountry>
/AME 1.75. Labium 0.15 long, 0.21 wide. Sternum 0.84 long, 0.75 wide. Leg measurements: Ⅰ 5.91 (1.52, 0.57, 1.82, 1.46, 0.54), II 4.72 (1.23, 0.49, 1.24, 1.11, 0.65), III 3.98 (1.07, 0.46, 0.84, 1.04, 0.57), IV 6.31 (1.68, 0.58, 1.46, 1.74, 0.85). Spination: femora I–IV d 1, I pl 4, II pl 2, tibiae Ⅰ pv 7 rv 8, II pv 7 rv 6, metatarsi Ⅰ pv 4 rv 4, II pv 4 rv 3.
</paragraph>
<paragraph id="8BE736ECFFA1FF9AFF65EAEDA78AF9B5" blockId="5.[151,1437,834,2013]" pageId="5" pageNumber="518">
Coloration. Carapace yellowish, with broad longitudinal mottled median band, slightly broader than ocular region. Chelicerae with three promarginal and five retromarginal teeth. Sternum longer than wide, shield-shaped. Legs yellow. Abdomen brown, with narrow and wedge-shaped dorsal scutum, three light transverse stripes at posterior of abdomen, venter of abdomen with pattern (
<figureCitation id="13632A69FFA1FF9AFCA1E959A779F9B5" box="[771,895,1555,1581]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[211,1373,181,1907]" captionTargetId="figure-23@6.[198,1390,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Otacilia pugnus sp. nov.: A. Male holotype, dorsal view; B. Same, ventral view; C. Female paratype, dorsal view; D. Same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.11150496" httpUri="https://zenodo.org/record/11150496/files/figure.png" pageId="5" pageNumber="518">Fig. 4A–B</figureCitation>
).
</paragraph>
<paragraph id="8BE736ECFFA1FF9AFF65E97DA7D0F925" blockId="5.[151,1437,834,2013]" pageId="5" pageNumber="518">
Palp as in
<figureCitation id="13632A69FFA1FF9AFE9BE97DA5B0F9C9" box="[313,438,1590,1617]" captionStart="FIGURE 5" captionStartId="7.[151,250,1869,1893]" captionTargetBox="[173,1416,189,1835]" captionTargetId="figure-17@7.[170,1416,181,1845]" captionTargetPageId="7" captionText="FIGURE 5. Copulatory organs of Otacilia pugnus sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C— conductor; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT—connecting tube; FD—fertilization duct; MS—median septum; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.11150500" httpUri="https://zenodo.org/record/11150500/files/figure.png" pageId="5" pageNumber="518">Figs 5A–D</figureCitation>
,
<figureCitation id="13632A69FFA1FF9AFE63E97DA60FF9C9" box="[449,521,1591,1617]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[163,1424,189,1910]" captionTargetId="figure-23@8.[155,1433,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Copulatory organs of Otacilia pugnus sp. nov.: A. Male left palp, prolateral view; B. same, retrolateral view; C. Same, ventral view; D. same, dorsal view; E. epigyne, ventral view; F. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11150506" httpUri="https://zenodo.org/record/11150506/files/figure.png" pageId="5" pageNumber="518">6A–D</figureCitation>
. Femoral apophysis well-developed, nearly at middle of femur. Tibia swollen, half as long as femur, prolateral tibial apophysis nearly half-round; retrolateral tibial apophysis long, fist-shaped, with digitiform tip. Tegulum pyriform, tegular apophysis clavate, slightly curved at end. Sperm duct distinct, U-shaped. Embolus hooked. Conductor membranous, shorter than tegular apophysis.
</paragraph>
<paragraph id="8BE736ECFFA1FF9AFF65E98DA7A1F845" blockId="5.[151,1437,834,2013]" pageId="5" pageNumber="518">
Female (
<typeStatus id="54E3884EFFA1FF9AFE8BE98DA597F979" box="[297,401,1735,1761]" pageId="5" pageNumber="518" type="paratype">paratype</typeStatus>
): total length 3.51, carapace 1.61 long, 1.39 wide; abdomen 1.90 long, 1.30 wide. Eye sizes and interdistances: AME 0.10, ALE 0.10, PME 0.08, PLE 0.09; AME–AME 0.04, AME–ALE 0.01, ALE–ALE 0.24, PME–PME 0.11, PME–PLE 0.05, PLE–PLE 0.38, ALE–PLE 0.08. EAW 0.53, CRW 0.78, EAW/CRW 0.68, CRW/CW 0.56. MOA 0.26 long, anterior width 0.23, posterior width 0.28.
<collectingCountry id="F34F767CFFA1FF9AFC48E879A017F8D5" box="[1002,1041,1843,1869]" name="Switzerland" pageId="5" pageNumber="518">CH</collectingCountry>
0.13,
<collectingCountry id="F34F767CFFA1FF9AFBFAE879A078F8D5" box="[1112,1150,1843,1869]" name="Switzerland" pageId="5" pageNumber="518">CH</collectingCountry>
/AME 1.30. Labium 0.16 long, 0.25 wide. Sternum 0.97 long, 0.84 wide. Leg measurements: Ⅰ 6.33 (1.56, 0.65, 1.97, 1.51, 0.64), II 5.14 (1.34, 0.57, 1.37, 1.20, 0.66), III 4.37 (1.18, 0.52, 0.91, 1.10, 0.66), IV 6.54 (1.74, 0.59, 1.56, 1.76, 0.89). Spination: femora I–IV d 1, I pl 5, II pl 3, tibiae Ⅰ pv 8 rv 9, II pv 8 rv 8, metatarsi Ⅰ pv 4 rv 4, II pv 4 rv 3. Other characters as in male, except darker color and absent of dorsal scutum (
<figureCitation id="13632A69FFA1FF9AFCBDE889A79DF845" box="[799,923,1987,2013]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[211,1373,181,1907]" captionTargetId="figure-23@6.[198,1390,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Otacilia pugnus sp. nov.: A. Male holotype, dorsal view; B. Same, ventral view; C. Female paratype, dorsal view; D. Same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.11150496" httpUri="https://zenodo.org/record/11150496/files/figure.png" pageId="5" pageNumber="518">Fig. 4C, D</figureCitation>
).
</paragraph>
<caption id="DF276664FFA2FF99FF35E8DDA604F84B" ID-DOI="http://doi.org/10.5281/zenodo.11150496" ID-Zenodo-Dep="11150496" httpUri="https://zenodo.org/record/11150496/files/figure.png" pageId="6" pageNumber="519" startId="6.[151,250,1943,1967]" targetBox="[211,1373,181,1907]" targetPageId="6" targetType="figure">
<paragraph id="8BE736ECFFA2FF99FF35E8DDA604F84B" blockId="6.[151,1436,1943,2003]" pageId="6" pageNumber="519">
<emphasis id="B92CEAFEFFA2FF99FF35E8DDA512F837" bold="true" box="[151,276,1943,1967]" pageId="6" pageNumber="519">FIGURE 4.</emphasis>
Habitus of
<taxonomicName id="4C584D6FFFA2FF99FE2CE8DDA637F837" authority="Shi &amp; Mu &amp; Zhang, 2024" authorityName="Shi &amp; Mu &amp; Zhang" authorityYear="2024" box="[398,561,1943,1967]" class="Arachnida" family="Phrurolithidae" genus="Otacilia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="519" phylum="Arthropoda" rank="species" species="pugnus" status="sp. nov.">
<emphasis id="B92CEAFEFFA2FF99FE2CE8DDA637F837" box="[398,561,1943,1967]" italics="true" pageId="6" pageNumber="519">Otacilia pugnus</emphasis>
</taxonomicName>
<emphasis id="B92CEAFEFFA2FF99FD9AE8DDA68CF837" bold="true" box="[568,650,1943,1967]" pageId="6" pageNumber="519">
<taxonomicNameLabel id="A21F5785FFA2FF99FD9AE8DDA68CF837" box="[568,650,1943,1967]" pageId="6" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A. Male holotype, dorsal view; B. Same, ventral view; C. Female paratype, dorsal view; D. Same, ventral view.
</paragraph>
</caption>
<caption id="DF276664FFA3FF98FF35E807A668F86D" ID-DOI="http://doi.org/10.5281/zenodo.11150500" ID-Zenodo-Dep="11150500" httpUri="https://zenodo.org/record/11150500/files/figure.png" pageId="7" pageNumber="520" startId="7.[151,250,1869,1893]" targetBox="[173,1416,189,1835]" targetPageId="7" targetType="figure">
<paragraph id="8BE736ECFFA3FF98FF35E807A668F86D" blockId="7.[151,1437,1869,2037]" pageId="7" pageNumber="520">
<emphasis id="B92CEAFEFFA3FF98FF35E807A512F8FD" bold="true" box="[151,276,1869,1893]" pageId="7" pageNumber="520">FIGURE 5.</emphasis>
Copulatory organs of
<taxonomicName id="4C584D6FFFA3FF98FE59E807A699F8FD" authority="Shi &amp; Mu &amp; Zhang, 2024" authorityName="Shi &amp; Mu &amp; Zhang" authorityYear="2024" box="[507,671,1869,1893]" class="Arachnida" family="Phrurolithidae" genus="Otacilia" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="520" phylum="Arthropoda" rank="species" species="pugnus" status="sp. nov.">
<emphasis id="B92CEAFEFFA3FF98FE59E807A699F8FD" box="[507,671,1869,1893]" italics="true" pageId="7" pageNumber="520">Otacilia pugnus</emphasis>
</taxonomicName>
<emphasis id="B92CEAFEFFA3FF98FD04E807A6FEF8FD" bold="true" box="[678,760,1869,1893]" pageId="7" pageNumber="520">
<taxonomicNameLabel id="A21F5785FFA3FF98FD04E807A6FEF8FD" box="[678,760,1869,1893]" pageId="7" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C— conductor; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT—connecting tube; FD—fertilization duct; MS—median septum; S—spermathecae.
</paragraph>
</caption>
<caption id="DF276664FFACFF97FF35E8DDA021F84B" ID-DOI="http://doi.org/10.5281/zenodo.11150506" ID-Zenodo-Dep="11150506" httpUri="https://zenodo.org/record/11150506/files/figure.png" pageId="8" pageNumber="521" startId="8.[151,250,1943,1967]" targetBox="[163,1424,189,1910]" targetPageId="8" targetType="figure">
<paragraph id="8BE736ECFFACFF97FF35E8DDA021F84B" blockId="8.[151,1437,1943,2004]" pageId="8" pageNumber="521">
<emphasis id="B92CEAFEFFACFF97FF35E8DDA512F837" bold="true" box="[151,276,1943,1967]" pageId="8" pageNumber="521">FIGURE 6.</emphasis>
Copulatory organs of
<taxonomicName id="4C584D6FFFACFF97FE5EE8DDA6A6F837" authority="Shi &amp; Mu &amp; Zhang, 2024" authorityName="Shi &amp; Mu &amp; Zhang" authorityYear="2024" box="[508,672,1943,1967]" class="Arachnida" family="Phrurolithidae" genus="Otacilia" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="521" phylum="Arthropoda" rank="species" species="pugnus" status="sp. nov.">
<emphasis id="B92CEAFEFFACFF97FE5EE8DDA6A6F837" box="[508,672,1943,1967]" italics="true" pageId="8" pageNumber="521">Otacilia pugnus</emphasis>
</taxonomicName>
<emphasis id="B92CEAFEFFACFF97FD0AE8D2A6FFF828" bold="true" box="[680,761,1944,1968]" pageId="8" pageNumber="521">
<taxonomicNameLabel id="A21F5785FFACFF97FD0AE8D2A6FFF828" box="[680,761,1944,1968]" pageId="8" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A. Male left palp, prolateral view; B. same, retrolateral view; C. Same, ventral view; D. same, dorsal view; E. epigyne, ventral view; F. Same, dorsal view.
</paragraph>
</caption>
<paragraph id="8BE736ECFFADFF96FF65EFDDA67BFED9" blockId="9.[151,1436,150,357]" pageId="9" pageNumber="522">
Epigyne as in
<figureCitation id="13632A69FFADFF96FEC1EFDDA5D0FF29" box="[355,470,150,177]" captionStart="FIGURE 5" captionStartId="7.[151,250,1869,1893]" captionTargetBox="[173,1416,189,1835]" captionTargetId="figure-17@7.[170,1416,181,1845]" captionTargetPageId="7" captionText="FIGURE 5. Copulatory organs of Otacilia pugnus sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C— conductor; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT—connecting tube; FD—fertilization duct; MS—median septum; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.11150500" httpUri="https://zenodo.org/record/11150500/files/figure.png" pageId="9" pageNumber="522">Figs 5E–F</figureCitation>
,
<figureCitation id="13632A69FFADFF96FE42EFDDA618FF29" box="[480,542,151,177]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[163,1424,189,1910]" captionTargetId="figure-23@8.[155,1433,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Copulatory organs of Otacilia pugnus sp. nov.: A. Male left palp, prolateral view; B. same, retrolateral view; C. Same, ventral view; D. same, dorsal view; E. epigyne, ventral view; F. Same, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11150506" httpUri="https://zenodo.org/record/11150506/files/figure.png" pageId="9" pageNumber="522">6E–F</figureCitation>
. Median septum distinct, anteriorly wider. Copulatory openings round, separated by more than two times of spermathecae diameters. Copulatory ducts oval, longer than wider. Connecting tubes L-shaped, longer than copulatory ducts. Glandular appendage indistinct. Bursa large, covering more than half of epigynal plate, nearly more egg-shaped. Spermathecae oval, separated by about by one radius. Fertilization ducts located at anterior margin of spermathecae.
</paragraph>
</subSubSection>
<subSubSection id="C3426567FFADFF96FF65EE00A687FEFD" box="[199,641,330,357]" pageId="9" pageNumber="522" type="distribution">
<paragraph id="8BE736ECFFADFF96FF65EE00A687FEFD" blockId="9.[151,1436,150,357]" box="[199,641,330,357]" pageId="9" pageNumber="522">
<emphasis id="B92CEAFEFFADFF96FF65EE00A567FEFC" bold="true" box="[199,353,330,356]" pageId="9" pageNumber="522">Distribution.</emphasis>
<collectingCountry id="F34F767CFFADFF96FEC5EE01A5ACFEFD" box="[359,426,331,357]" name="China" pageId="9" pageNumber="522">China</collectingCountry>
(
<collectingRegion id="499CF80EFFADFF96FE1BEE01A673FEFD" box="[441,629,331,357]" country="China" name="Hunan" pageId="9" pageNumber="522">Hunan Province</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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