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<document id="2787DA3DD96E9C5ACC2E148A202FB2A3" ID-CLB-Dataset="38962" ID-DOI="10.11646/zootaxa.4117.2.7" ID-GBIF-Dataset="ee367422-f5ce-440f-a72f-4ad984ecff19" ID-ISSN="1175-5326" ID-Zenodo-Dep="265207" ID-ZooBank="A4A656D2-2ABC-4D08-8CC2-A2AD6BE79D85" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464706875399" checkinUser="plazi" docAuthor="Cadena-Castañeda, Oscar J., Garay, Andrea, Castañeda, Didier, Cardona-Granda, Juan Manuel &amp; García, Alexander García" docDate="2016" docId="03F187CAFFCF110A38BEF98AFAA1C1D0" docLanguage="en" docName="zootaxa.4117.2.7.pdf.imf" docOrigin="Zootaxa 4117 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Caenomastax insignis Hebard 1923" docType="treatment" docVersion="8" lastPageNumber="252" masterDocId="FFC8FFB2FFC511013829FFDBFFC6C058" masterDocTitle="Systematics and phylogeny of the genus Caenomastax Hebard, 1923 (Orthoptera: Eumastacidae: Eumastacinae)" masterLastPageNumber="264" masterPageNumber="241" pageNumber="251" updateTime="1698662661435" updateUser="ExternalLinkService">
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<mods:title id="308B1FA1B2C6237A0F61B63C6589EAB6">Systematics and phylogeny of the genus Caenomastax Hebard, 1923 (Orthoptera: Eumastacidae: Eumastacinae)</mods:title>
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<mods:namePart id="7FBDBDFC058890B5C387FEC3DFEDC2E6">Cadena-Castañeda, Oscar J.</mods:namePart>
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<mods:namePart id="E41AA5F07CB2CBF9E6452A4B22C32051">Castañeda, Didier</mods:namePart>
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<mods:namePart id="DEAA5FFE8DE2C1A7535EB58F16D2252F">Cardona-Granda, Juan Manuel</mods:namePart>
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<treatment id="03F187CAFFCF110A38BEF98AFAA1C1D0" ID-DOI="http://doi.org/10.5281/zenodo.6075010" ID-GBIF-Taxon="120733520" ID-Zenodo-Dep="6075010" LSID="urn:lsid:plazi:treatment:03F187CAFFCF110A38BEF98AFAA1C1D0" httpUri="http://treatment.plazi.org/id/03F187CAFFCF110A38BEF98AFAA1C1D0" lastPageId="11" lastPageNumber="252" pageId="10" pageNumber="251">
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<paragraph id="8BE736DCFFCF110B38BEF98AFD94C633" blockId="10.[151,1040,1617,1712]" box="[151,594,1617,1643]" pageId="10" pageNumber="251">
<heading id="D0AF81B0FFCF110B38BEF98AFD94C633" bold="true" box="[151,594,1617,1643]" fontSize="11" level="1" pageId="10" pageNumber="251" reason="1">
<taxonomicName id="4C584D5FFFCF110B38BEF98AFD94C633" authority="Hebard, 1923" authorityName="Hebard" authorityYear="1923" box="[151,594,1617,1643]" class="Insecta" family="Gryllidae" genus="Caenomastax" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B92CEACEFFCF110B38BEF98AFD94C633" bold="true" box="[151,594,1617,1643]" pageId="10" pageNumber="251">
<emphasis id="B92CEACEFFCF110B38BEF98AFE66C633" bold="true" box="[151,416,1617,1643]" italics="true" pageId="10" pageNumber="251">Caenomastax insignis</emphasis>
<bibRefCitation id="EFC94B2DFFCF110B3981F98AFD94C633" author="Hebard" box="[424,594,1617,1643]" pageId="10" pageNumber="251" refString="Hebard, M. (1923) Studies in the Dermaptera and Orthoptera of Colombia. Third paper. Orthopterous family Acrididae. Transactions of the American Entomological Society, 49, 165 - 313, 8 pls." type="journal article" year="1923">Hebard, 1923</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8BE736DCFFCF110B38BEF9AFFD46C6D4" blockId="10.[151,1040,1617,1712]" box="[151,640,1652,1677]" pageId="10" pageNumber="251">
(
<figureCitation id="13632A59FFCF110B38B6F9AFFEDAC6D5" box="[159,284,1652,1677]" captionStart="FIGURES 53 64" captionStartId="12.[151,264,1896,1918]" captionTargetBox="[150,1436,195,1864]" captionTargetId="figure@12.[150,1436,195,1874]" captionTargetPageId="12" captionText="FIGURES 53 64. Males of the species of genus Caenomastax (Head and thorax in dorsal view, terminalia in dorsal and side view respectively). 53 55. C. insignis. 56 58. C. similis n. sp. 59 61. C. bimaculata n. sp. 62 64. C. magdalenensis n. sp." httpUri="https://zenodo.org/record/265213/files/figure.png" pageId="10" pageNumber="251">Fig. 5355</figureCitation>
,
<figureCitation id="13632A59FFCF110B390EF9AFFEB6C6D5" box="[295,368,1652,1677]" captionStart="FIGURES 65 76" captionStartId="13.[151,264,1756,1778]" captionTargetBox="[151,1429,197,1728]" captionTargetId="figure@13.[151,1435,193,1735]" captionTargetPageId="13" captionText="FIGURES 65 76. Phallic complex of the species of Caenomastax. 65 67. C. insignis. 68 70. C. similis n. sp. 71 73. C. bimaculata n. sp. 74 76. C. magdalenensis n. sp." httpUri="https://zenodo.org/record/265214/files/figure.png" pageId="10" pageNumber="251">6567</figureCitation>
,
<figureCitation id="13632A59FFCF110B3955F9AFFE03C6D4" box="[380,453,1652,1676]" captionStart="FIGURES 77 84" captionStartId="14.[151,264,1233,1255]" captionTargetBox="[152,1434,193,1211]" captionTargetId="figure@14.[152,1435,193,1212]" captionTargetPageId="14" captionText="FIGURES 77 84. Head, pronotum and tegmina of Caenomastax females in side and dorsal view, respectively. 77 78. C. insignis. 79 80. C. similis n. sp. 81 82. C. bimaculata n. sp. 83 84. C. magdalenensis n. sp." httpUri="https://zenodo.org/record/265215/files/figure.png" pageId="10" pageNumber="251">7778</figureCitation>
,
<figureCitation id="13632A59FFCF110B39E6F9AFFDB1C6D4" box="[463,631,1652,1677]" captionStart-0="FIGURES 85 96" captionStart-1="FIGURES 19 30" captionStart-10="FIGURES 105 110" captionStart-2="FIGURES 31 40" captionStart-3="FIGURES 41 52" captionStart-4="FIGURES 53 64" captionStart-5="FIGURES 65 76" captionStart-6="FIGURES 77 84" captionStart-7="FIGURES 97 102" captionStart-8="FIGURES 103" captionStart-9="FIGURES 104" captionStartId-0="14.[151,264,1969,1991]" captionStartId-1="6.[151,264,1874,1896]" captionStartId-10="19.[151,264,1883,1905]" captionStartId-2="7.[151,264,1923,1945]" captionStartId-3="8.[151,264,1983,2005]" captionStartId-4="12.[151,264,1896,1918]" captionStartId-5="13.[151,264,1756,1778]" captionStartId-6="14.[151,264,1233,1255]" captionStartId-7="17.[151,264,1968,1990]" captionStartId-8="18.[151,264,821,843]" captionStartId-9="18.[151,264,1622,1644]" captionTargetBox-0="[151,1435,1336,1947]" captionTargetBox-1="[161,1428,201,1839]" captionTargetBox-10="[151,1435,301,1861]" captionTargetBox-2="[168,1423,207,1886]" captionTargetBox-3="[154,1433,471,1960]" captionTargetBox-4="[150,1436,195,1864]" captionTargetBox-5="[151,1429,197,1728]" captionTargetBox-6="[152,1434,193,1211]" captionTargetBox-7="[152,1434,359,1947]" captionTargetBox-8="[153,1436,195,794]" captionTargetBox-9="[151,1435,924,1600]" captionTargetId-0="figure@14.[151,1435,1336,1948]" captionTargetId-1="figure@6.[150,1437,193,1853]" captionTargetId-10="figure@19.[151,1436,301,1861]" captionTargetId-2="figure@7.[151,1435,193,1902]" captionTargetId-3="figure@8.[150,1437,469,1961]" captionTargetId-4="figure@12.[150,1436,195,1874]" captionTargetId-5="figure@13.[151,1435,193,1735]" captionTargetId-6="figure@14.[152,1435,193,1212]" captionTargetId-7="figure@17.[152,1435,353,1947]" captionTargetId-8="figure@18.[153,1438,195,794]" captionTargetId-9="figure@18.[151,1435,924,1601]" captionTargetPageId-0="14" captionTargetPageId-1="6" captionTargetPageId-10="19" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="12" captionTargetPageId-5="13" captionTargetPageId-6="14" captionTargetPageId-7="17" captionTargetPageId-8="18" captionTargetPageId-9="18" captionText-0="FIGURES 85 96. Tegmina, ovipositor and subgenital plate of the females in genus Caenomastax. 85 87. C. insignis 88 90. C. similes n. sp. 91 93. C. immaculate n. sp. 94 96. C. magdalenensis n. sp." captionText-1="FIGURES 19 30. Characters used for the phylogeny (mainly male in dorsal view and phallic complex). 19, 22, 25 and 29. Caenomastax similis n. sp. 20 and 23. Zeromastax selenesii. 21, 24 and 26. Paralethus rowelli. 28. Santanderia lita. 30. Hebardomastax atopa." captionText-10="FIGURES 105 110. Living monkey grasshoppers. 105 106. Santanderia lita (male and female). 107. Homeomastax dereixi (male). 108. Caenomastax similis n. sp. (female) (Photo: Daniel Castro). 109 110. Caenomastax insignis (male and female)." captionText-2="FIGURES 31 40. Head and pronotum of males of the species included in the phylogenetic analysis. 31. Caenomastax insignis. 32. Zeromastax selenesii. 33. Caenomastax similis n. sp. 34. Homeomastax dereixi. 35. Caenomastax bimaculata n. sp. 36. Eumastax restrepoi. 37. Caenomastax magdalenensis n. sp. 38. Santanderia lita. 39. Hebardomastax atopa. 40. Paralethus rowelli." captionText-3="FIGURES 41 52. Dorsal view of terminalia and phallic complex of the species of the out group (except Hebardomastax atopa). 41 43. Homeomastax dereixi. 44 46. Eumastax restrepoi. 47 49. Santanderia lita. 50 52. Zeromastax selenesii." captionText-4="FIGURES 53 64. Males of the species of genus Caenomastax (Head and thorax in dorsal view, terminalia in dorsal and side view respectively). 53 55. C. insignis. 56 58. C. similis n. sp. 59 61. C. bimaculata n. sp. 62 64. C. magdalenensis n. sp." captionText-5="FIGURES 65 76. Phallic complex of the species of Caenomastax. 65 67. C. insignis. 68 70. C. similis n. sp. 71 73. C. bimaculata n. sp. 74 76. C. magdalenensis n. sp." captionText-6="FIGURES 77 84. Head, pronotum and tegmina of Caenomastax females in side and dorsal view, respectively. 77 78. C. insignis. 79 80. C. similis n. sp. 81 82. C. bimaculata n. sp. 83 84. C. magdalenensis n. sp." captionText-7="FIGURES 97 102. Hebardomastax atopa n. comb. 97. Male habit, side view. 98 100. Phallic complex. 101 102. Terminalia in dorsal and lateral view respectively." captionText-8="FIGURES 103. Most parsimonious tree of genus Caenomastax. 145 steps long (CI, 0.93; RI, 0,86). Black circles indicate the synapomorphies and white ones the homoplasies (* C. atopa = Hebardomastax atopa n. comb.)." captionText-9="FIGURES 104. Consensus tree with Bremer (Br), (Br absolute / Br relative), indicated in the upperpart of each branch. Bootstrap value indicated in the lower part of each branch (* C. atopa = Hebardomastax atopa n. comb.)." httpUri-0="https://zenodo.org/record/265216/files/figure.png" httpUri-1="https://zenodo.org/record/265210/files/figure.png" httpUri-10="https://zenodo.org/record/265220/files/figure.png" httpUri-2="https://zenodo.org/record/265211/files/figure.png" httpUri-3="https://zenodo.org/record/265212/files/figure.png" httpUri-4="https://zenodo.org/record/265213/files/figure.png" httpUri-5="https://zenodo.org/record/265214/files/figure.png" httpUri-6="https://zenodo.org/record/265215/files/figure.png" httpUri-7="https://zenodo.org/record/265217/files/figure.png" httpUri-8="https://zenodo.org/record/265218/files/figure.png" httpUri-9="https://zenodo.org/record/265219/files/figure.png" pageId="10" pageNumber="251">8587, 19110</figureCitation>
)
</paragraph>
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<subSubSection id="C3426557FFCF110B38BEF94CFBD6C6E8" box="[151,1040,1687,1712]" pageId="10" pageNumber="251" type="description">
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http://lsid.speciesfile.org/urn:lsid:
<taxonomicName id="4C584D5FFFCF110B3A20F94CFD4FC6E8" box="[521,649,1687,1712]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
.speciesfile.org:TaxonName:40408
</paragraph>
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<subSubSection id="C3426557FFCF110B38BEF904FE33C7AC" pageId="10" pageNumber="251" type="diagnosis">
<paragraph id="8BE736DCFFCF110B38BEF904FD23C73D" blockId="10.[151,1436,1759,2037]" pageId="10" pageNumber="251">
<emphasis id="B92CEACEFFCF110B38BEF904FED7C6A0" bold="true" box="[151,273,1759,1784]" pageId="10" pageNumber="251">Diagnosis.</emphasis>
Posterior projection of the pronotal disc of females reaching half the length of tegmina and blunt-ended (
<figureCitation id="13632A59FFCF110B38B6F8DFFEE5C744" box="[159,291,1796,1821]" captionStart="FIGURES 77 84" captionStartId="14.[151,264,1233,1255]" captionTargetBox="[152,1434,193,1211]" captionTargetId="figure@14.[152,1435,193,1212]" captionTargetPageId="14" captionText="FIGURES 77 84. Head, pronotum and tegmina of Caenomastax females in side and dorsal view, respectively. 77 78. C. insignis. 79 80. C. similis n. sp. 81 82. C. bimaculata n. sp. 83 84. C. magdalenensis n. sp." httpUri="https://zenodo.org/record/265215/files/figure.png" pageId="10" pageNumber="251">Fig. 77, 78</figureCitation>
). Female tegmina 3-lobed (
<figureCitation id="13632A59FFCF110B3A49F8DFFD71C745" box="[608,695,1796,1821]" captionStart="FIGURES 85 96" captionStartId="14.[151,264,1969,1991]" captionTargetBox="[151,1435,1336,1947]" captionTargetId="figure@14.[151,1435,1336,1948]" captionTargetPageId="14" captionText="FIGURES 85 96. Tegmina, ovipositor and subgenital plate of the females in genus Caenomastax. 85 87. C. insignis 88 90. C. similes n. sp. 91 93. C. immaculate n. sp. 94 96. C. magdalenensis n. sp." httpUri="https://zenodo.org/record/265216/files/figure.png" pageId="10" pageNumber="251">Fig. 85</figureCitation>
). Medial projection of subgenital plate tip projecting and longer than lateral projections (
<figureCitation id="13632A59FFCF110B3987F8FCFDC0C718" box="[430,518,1831,1856]" captionStart="FIGURES 85 96" captionStartId="14.[151,264,1969,1991]" captionTargetBox="[151,1435,1336,1947]" captionTargetId="figure@14.[151,1435,1336,1948]" captionTargetPageId="14" captionText="FIGURES 85 96. Tegmina, ovipositor and subgenital plate of the females in genus Caenomastax. 85 87. C. insignis 88 90. C. similes n. sp. 91 93. C. immaculate n. sp. 94 96. C. magdalenensis n. sp." httpUri="https://zenodo.org/record/265216/files/figure.png" pageId="10" pageNumber="251">Fig. 87</figureCitation>
). Abdominal sternites of male black and tomentose, lophi of epiphallus slightly curving towards the latero-external margin (
<figureCitation id="13632A59FFCF110B3AACF897FD1EC73D" box="[645,728,1868,1893]" captionStart="FIGURES 65 76" captionStartId="13.[151,264,1756,1778]" captionTargetBox="[151,1429,197,1728]" captionTargetId="figure@13.[151,1435,193,1735]" captionTargetPageId="13" captionText="FIGURES 65 76. Phallic complex of the species of Caenomastax. 65 67. C. insignis. 68 70. C. similis n. sp. 71 73. C. bimaculata n. sp. 74 76. C. magdalenensis n. sp." httpUri="https://zenodo.org/record/265214/files/figure.png" pageId="10" pageNumber="251">Fig. 65</figureCitation>
).
</paragraph>
<paragraph id="8BE736DCFFCF110B38EEF8B4FE33C7AC" blockId="10.[151,1436,1759,2037]" pageId="10" pageNumber="251">
<emphasis id="B92CEACEFFCF110B38EEF8B4FE0FC7D0" bold="true" box="[199,457,1903,1928]" pageId="10" pageNumber="251">Specimens examined:</emphasis>
<collectingCountry id="F34F764CFFCF110B39F9F8ABFDAFC7D0" box="[464,617,1904,1928]" name="Colombia" pageId="10" pageNumber="251">COLOMBIA</collectingCountry>
. Cundinamarca, La Palma, ♂♀
<date id="FFE6101CFFCF110B3BF8F8ABFB48C7D0" box="[977,1166,1903,1928]" pageId="10" pageNumber="251" value="1995-01-14">14 January 1995</date>
. D. Angel
<emphasis id="B92CEACEFFCF110B3D21F8B4FAF4C7D0" box="[1288,1330,1903,1928]" italics="true" pageId="10" pageNumber="251">leg.</emphasis>
Silvania, ♂ Vereda Azafrán,
<quantity id="4CA09B39FFCF110B3956F84FFE1DC7F4" box="[383,475,1940,1965]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="251" unit="m" value="1500.0">1500 m</quantity>
, in
<taxonomicName id="4C584D5FFFCF110B3A24F84FFD7BC7F5" box="[525,701,1940,1965]" class="Magnoliopsida" family="Rubiaceae" genus="Coffea" kingdom="Plantae" order="Gentianales" pageId="10" pageNumber="251" phylum="Tracheophyta" rank="species" species="arabiga">
<emphasis id="B92CEACEFFCF110B3A24F84FFD7BC7F5" box="[525,701,1940,1965]" italics="true" pageId="10" pageNumber="251">Coffea arabiga</emphasis>
</taxonomicName>
,
<date id="FFE6101CFFCF110B3AF9F84FFC70C7F4" box="[720,950,1940,1965]" pageId="10" pageNumber="251" value="2009-11-28">28 November 2009</date>
. N. Flórez
<emphasis id="B92CEACEFFCF110B3C69F84FFBACC7F5" box="[1088,1130,1940,1965]" italics="true" pageId="10" pageNumber="251">leg.</emphasis>
[UNAB]. Boyacá, Coper,
<specimenCount id="9D5EFD55FFCF110B38BEF863FF0DC788" box="[151,203,1976,2000]" pageId="10" pageNumber="251" type="generic">10♂</specimenCount>
12♀ September of 2009. O. Cadena
<emphasis id="B92CEACEFFCF110B3A41F863FD57C789" box="[616,657,1976,2001]" italics="true" pageId="10" pageNumber="251">leg.</emphasis>
, Maripi ♀
<date id="FFE6101CFFCF110B3B3BF863FC74C788" box="[786,946,1976,2001]" pageId="10" pageNumber="251" value="1974-04-27">27 April 1974</date>
I. Arévalo. Santander, Florián,
<specimenCount id="9D5EFD55FFCF110B3D33F863FA86C788" box="[1306,1344,1976,2000]" pageId="10" pageNumber="251" type="generic">8♂</specimenCount>
7♀
<date id="FFE6101CFFCF110B3D40F862FF15C7AC" pageId="10" pageNumber="251" value="2012-06">June 2012</date>
. J. Cardona
<emphasis id="B92CEACEFFCF110B3975F807FEB8C7AD" box="[348,382,2012,2037]" italics="true" pageId="10" pageNumber="251">leg</emphasis>
. [CAUD].
</paragraph>
</subSubSection>
<subSubSection id="C3426557FFCE110A38EEFF4CFB65C0E8" box="[199,1187,151,176]" pageId="11" pageNumber="252" type="description">
<paragraph id="8BE736DCFFCE110A38EEFF4CFB65C0E8" blockId="11.[151,1436,151,392]" box="[199,1187,151,176]" pageId="11" pageNumber="252">
<emphasis id="B92CEACEFFCE110A38EEFF4CFD83C0E8" bold="true" box="[199,581,151,176]" pageId="11" pageNumber="252">Measurements (mm). (♂/♀). Lt:</emphasis>
14 /15.2;
<emphasis id="B92CEACEFFCE110A3A9CFF4CFD16C0E8" bold="true" box="[693,720,151,176]" pageId="11" pageNumber="252">P:</emphasis>
3/4;
<emphasis id="B92CEACEFFCE110A3B23FF4CFCF4C0E8" bold="true" box="[778,818,151,176]" pageId="11" pageNumber="252">Teg</emphasis>
: 2.8 /3;
<emphasis id="B92CEACEFFCE110A3BB8FF4CFC07C0E8" bold="true" box="[913,961,151,176]" pageId="11" pageNumber="252">HF:</emphasis>
8 /8.3;
<emphasis id="B92CEACEFFCE110A3C3EFF4CFB80C0E8" bold="true" box="[1047,1094,151,176]" pageId="11" pageNumber="252">HT:</emphasis>
8.5/5.2.
</paragraph>
</subSubSection>
<subSubSection id="C3426557FFCE110A38EEFF67FAA1C1D0" pageId="11" pageNumber="252" type="discussion">
<paragraph id="8BE736DCFFCE110A38EEFF67FAA1C1D0" blockId="11.[151,1436,151,392]" pageId="11" pageNumber="252">
<emphasis id="B92CEACEFFCE110A38EEFF67FE89C08D" bold="true" box="[199,335,188,213]" pageId="11" pageNumber="252">Comments.</emphasis>
A complete description of the external morphology of the species was given in its original description. As for phallic structures, they were described by
<bibRefCitation id="EFC94B2DFFCE110A3B66FF04FB80C0A0" author="Rehn" box="[847,1094,223,248]" pageId="11" pageNumber="252" refString="Rehn, J. A. G. &amp; Grant Jr., H. J. (1958) The phallic complex in the subfamilies of New World Eumastacidae and the family Tanaoceridae. Proceedings of the Academy of Natural Sciences, Philadelphia, 110, 301 - 319." type="journal article" year="1958">Rehn &amp; Grant (1958)</bibRefCitation>
. They are commonly found in the understory, among varied vegetation, mostly
<taxonomicName id="4C584D5FFFCE110A3A95FEDFFCB8C145" box="[700,894,260,285]" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="252" phylum="Tracheophyta" rank="family">Melastomataceae</taxonomicName>
and other forbs, in which they can be abundant; they are not found commonly in ferns though in some collection places they inhabited disturbed areas, being found even in grasses in roadsides and overgrown cattle fields. According to
<bibRefCitation id="EFC94B2DFFCE110A3BE2FE97FBA3C13C" author="Varon" box="[971,1125,332,356]" pageId="11" pageNumber="252" refString="Varon, A. (2000) Saltamontes eumastacidos (Insecta: Orthoptera: Caelifera: Eumastacidae) de Colombia. Biota Colombiana, 1 (3), 261 - 266." type="journal article" year="2000">Varón (2000)</bibRefCitation>
, the species is found up to
<quantity id="4CA09B39FFCE110A38BEFEABFF35C1D0" box="[151,243,367,392]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="252" unit="m" value="2000.0">2000 m</quantity>
.a.s.l., but literature data and the material that could be checked points to an upper limit of about 1600.
</paragraph>
</subSubSection>
</treatment>
</document>