treatments-xml/data/03/F1/87/03F187C42248FF9EFE38175EFA0CFAC6.xml
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<document ID-DOI="10.1590/S0101-81752003000300023" ID-GBIF-Dataset="f8ccd15f-6063-45e2-82ff-6fd9cc63f07f" ID-GBIF-Taxon="160762815" ID-Zenodo-Dep="3583048" approvedBy="poa" checkinTime="1576682755828" checkinUser="carolina" docAuthor="Grazia, Jocélia &amp; Frey-da-Silva, Angélica" docDate="2003" docId="03F187C42248FF9EFE38175EFA0CFAC6" docLanguage="en" docName="RevBrasZool.20.3.507-510.pdf" docOrigin="Revista Brasileira de Zoologia 20 (3)" docStyle="DocumentStyle{}" docTitle="Rideriana Grazia &amp; Frey-da-Silva 2003, gen. nov" docType="treatment" docVersion="6" lastPageId="1" lastPageNumber="508" masterDocId="FFC8FFBC2248FF9FFFB9114CFFA8FFF8" masterDocTitle="Rideriana amazonica gen. nov. and sp. nov. of Pentatomini (Heteroptera, Pentatomidae)" masterLastPageNumber="510" masterPageNumber="507" pageId="0" pageNumber="507" updateTime="1643592369552" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Rideriana amazonica gen. nov. and sp. nov. of Pentatomini (Heteroptera, Pentatomidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grazia, Jocélia</mods:namePart>
<mods:affiliation>Universidade Federal do Rio Grande do Sul, Instituto de Biociências, Departamento de Zoologia, Prédio 43435. Av. Bento Gonçalves 9500, 91501 - 970. Porto Alegre, RS, Brasil.</mods:affiliation>
<mods:nameIdentifier type="email">jocelia@ufrgs.br</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Frey-da-Silva, Angélica</mods:namePart>
<mods:affiliation>Universidade Federal do Rio Grande do Sul, Instituto de Biociências, Departamento de Zoologia, Prédio 43435. Av. Bento Gonçalves 9500, 91501 - 970. Porto Alegre, RS, Brasil.</mods:affiliation>
<mods:nameIdentifier type="email">angelzi@zipmail.com.br</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Revista Brasileira de Zoologia</mods:title>
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<mods:part>
<mods:date>2003</mods:date>
<mods:detail type="pubDate">
<mods:number>2003-09-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>20</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
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<mods:start>507</mods:start>
<mods:end>510</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/S0101-81752003000300023</mods:identifier>
<mods:identifier type="GBIF-Dataset">f8ccd15f-6063-45e2-82ff-6fd9cc63f07f</mods:identifier>
<mods:identifier type="Zenodo-Dep">3583048</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="160762815" LSID="urn:lsid:plazi:treatment:03F187C42248FF9EFE38175EFA0CFAC6" httpUri="http://treatment.plazi.org/id/03F187C42248FF9EFE38175EFA0CFAC6" lastPageId="1" lastPageNumber="508" pageId="0" pageNumber="507">
<subSubSection box="[385,618,1554,1582]" pageId="0" pageNumber="507" type="nomenclature">
<paragraph blockId="0.[385,618,1554,1582]" box="[385,618,1554,1582]" pageId="0" pageNumber="507">
<heading bold="true" box="[385,618,1554,1582]" centered="true" fontSize="11" level="1" pageId="0" pageNumber="507" reason="0">
<emphasis bold="true" box="[385,618,1554,1582]" pageId="0" pageNumber="507">
<taxonomicName authority="Grazia &amp; Frey-da-Silva, 2003" authorityName="Grazia &amp; Frey-da-Silva" authorityYear="2003" box="[385,487,1554,1582]" class="Insecta" family="Pentatomidae" genus="Rideriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="507" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[385,487,1554,1582]" italics="true" pageId="0" pageNumber="507">Rideriana</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[495,613,1554,1582]" pageId="0" pageNumber="507" rank="genus">gen. nov</taxonomicNameLabel>
nov.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="507" type="type_taxon">
<paragraph blockId="0.[189,813,1597,1825]" pageId="0" pageNumber="507">
<typeStatus box="[250,298,1598,1617]" pageId="0" pageNumber="507">Type</typeStatus>
species:
<taxonomicName authority="Grazia &amp; Frey-da-Silva" authorityName="Grazia &amp; Frey-da-Silva" authorityYear="2003" box="[383,809,1597,1617]" class="Insecta" family="Pentatomidae" genus="Rideriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="507" phylum="Arthropoda" rank="species" species="amazonica">
<emphasis box="[383,581,1597,1616]" italics="true" pageId="0" pageNumber="507">Rideriana amazonica</emphasis>
Grazia &amp; Frey-da-Silva
</taxonomicName>
, here described.
</paragraph>
</subSubSection>
<subSubSection lastPageId="1" lastPageNumber="508" pageId="0" pageNumber="507" type="description">
<paragraph blockId="0.[189,813,1597,1825]" lastBlockId="0.[850,1475,924,1825]" pageId="0" pageNumber="507">
Dorsal surface of body intense and uniformly punctured; ventral surface of the thorax with more sparse and bigger punctures. Head wider than long. Juga longer than and confluent before tylus; lateral margins of juga slightly emarginate; dorsal surface with transverse furrows. Antennal tubercles with rhomboid lateral spine. Antennal segment I surpassing apex of head, densely punctured; segment II extremely reduced. Relative length of antennal segments: V&gt; III
<emphasis bold="true" box="[920,932,949,969]" pageId="0" pageNumber="507"></emphasis>
IV&gt; I&gt; II. Bucculaesubrectilinear, evanescentatbase of head with well developed anterior tooth. Rostrum short, attaining mesosternum. Rostral segment Ishorter than bucculae; segments II and III subequal in length, longer than I and IV.
</paragraph>
<paragraph blockId="0.[850,1475,924,1825]" pageId="0" pageNumber="507">Pronotum subtrapezoidal, anterior 2/3 strongly declivent. Anterolateral margins subrectilinear, scarcely deflected and with angles distinctly toothed; posterolateral margins sinuous. Humeral angles not developed, rounded. Posterior margin subrectilinear. Dorsal surface with scattered yellowish maculae; 1+1 maculae at lateral angles of cicatrices. Prosternum flat between crests limiting this area; mesosternum carinate, anteriorly acute and projected between procoxae; metasternum elevate, anteriorly acute and projected between mesocoxae, posteriorly bifurcate. Ostiolar rugae short, clavate, length 1/4 of evaporatory areas width. Evaporatory areas infuscate, slightly rugose near ostiolum, covering 3/4 of metapleura and almost apical half of mesopleura.</paragraph>
<paragraph blockId="0.[850,1475,924,1825]" pageId="0" pageNumber="507">Scutellum longer than wide at base; basal angles foveate, apex slightly acute reaching connexival segment VI at middle. Posterior angle of corium subacute, reaching posterior margin of connexival segment VI; posterior margin of corium subrectilinear. Apex of vein Radius with small pale yellow callus followed by dark-castaneous macula. Hemelytra surface with scattered yellowish maculae; membrane infuscate. Femur toothed at limits of apical fourth; tibia dorsally sulcate. Femur and tibia with dark castaneous punctures; tarsi immaculate.</paragraph>
<paragraph blockId="0.[850,1475,924,1825]" pageId="0" pageNumber="507">Connexivum scarcely exposed. Postero-lateral angles in black tine spines. Abdominal spine extending between metacoxae, apposed to posterior margin of metasternum. Spiracles sided entad by yellowish callus. Trichobothria placed laterad to the imaginary longitudinal line tangential to the spiracles. Abdominal median third, sutures and pseudo-sutures with black maculae; disc with scattered punctures.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3583050" ID-Zenodo-Dep="3583050" box="[159,848,1015,1035]" httpUri="https://zenodo.org/record/3583050/files/figure.png" pageId="1" pageNumber="508" targetBox="[325,1271,242,988]" targetPageId="1">
<paragraph blockId="1.[159,853,1015,1035]" box="[159,848,1015,1035]" pageId="1" pageNumber="508">
Figure 1. Habitus.
<taxonomicName authority="Grazia &amp; Frey-da-Silva, 2003" authorityName="Grazia &amp; Frey-da-Silva" authorityYear="2003" box="[336,529,1015,1035]" class="Insecta" family="Pentatomidae" genus="Rideriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="species" species="amazonica" status="sp. nov.">
<emphasis box="[336,529,1015,1035]" italics="true" pageId="1" pageNumber="508">Rideriana amazonica</emphasis>
</taxonomicName>
<emphasis bold="true" box="[536,615,1015,1035]" pageId="1" pageNumber="508">
<taxonomicNameLabel box="[536,615,1015,1035]" pageId="1" pageNumber="508" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype. Scale: 1 mm
</paragraph>
</caption>
<paragraph blockId="1.[159,853,1015,1035]" box="[849,853,1015,1035]" pageId="1" pageNumber="508">.</paragraph>
<paragraph blockId="1.[159,784,1113,1832]" pageId="1" pageNumber="508">Male. Pygophore quadrangular, posterolateral angles produced posteriorly; dorsal rim (dr) shallowly escavate; ventral rim (vr) escavate with strongly developed hypandrium. Segment X (proctiger) pyriform in dorsal view, apex ventrally directed. Parameres (pa) well developed.</paragraph>
<paragraph blockId="1.[159,784,1113,1832]" pageId="1" pageNumber="508">
<emphasis box="[220,293,1245,1264]" italics="true" pageId="1" pageNumber="508">Phallus</emphasis>
. Basal plate (bp) well developed longer than phallotheca (ph). Phallotheca opened postero-ventrally; postero-lateral angles produced.
<emphasis box="[500,670,1297,1315]" italics="true" pageId="1" pageNumber="508">Processus capitati</emphasis>
(pc) wide, almost twice the length of dorsal connective (dc). Vesica (v) well developed, almost twice the length of phallotheca, with processes ventrally projected. Conjunctiva absent.
<emphasis italics="true" pageId="1" pageNumber="508">Ductus seminis distalis</emphasis>
(dsd) long, twice the length of phallotheca, slightly wider at basal half.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="508" type="discussion">
<paragraph blockId="1.[159,784,1113,1832]" lastBlockId="1.[820,1444,1113,1342]" pageId="1" pageNumber="508">
Comments.
<taxonomicName authority="Grazia &amp; Frey-da-Silva, 2003" authorityName="Grazia &amp; Frey-da-Silva" authorityYear="2003" box="[343,433,1454,1473]" class="Insecta" family="Pentatomidae" genus="Rideriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis box="[343,433,1454,1473]" italics="true" pageId="1" pageNumber="508">Rideriana</emphasis>
</taxonomicName>
<emphasis bold="true" box="[440,537,1454,1473]" pageId="1" pageNumber="508">
<taxonomicNameLabel box="[440,537,1454,1473]" pageId="1" pageNumber="508" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
has the “facies” of three known genera belonging to Section 3 of
<bibRefCitation author="ROLSTON, L. H. &amp; F. J. D. MCDONALD &amp; D. B. THOMAS JR." box="[573,779,1479,1499]" journalOrPublisher="Journal of the New York Entomological Society, New York" pageId="1" pageNumber="508" pagination="120 - 132" part="88" refId="ref1996" refString="ROLSTON, L. H.; F. J. D. MCDONALD &amp; D. B. THOMAS JR. 1980. A conspectus of Pentatomini genera of the Western Hemisphere. Part I (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, New York, 88 (2): 120 - 132." title="A conspectus of Pentatomini genera of the Western Hemisphere. Part I (Hemiptera: Pentatomidae)" type="journal article" year="1980">
ROLSTON
<emphasis box="[656,697,1480,1499]" italics="true" pageId="1" pageNumber="508">et al</emphasis>
. (1980)
</bibRefCitation>
:
<taxonomicName authorityName="Stal" authorityYear="1858" box="[159,218,1505,1524]" class="Insecta" family="Pentatomidae" genus="Serdia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[159,218,1505,1524]" italics="true" pageId="1" pageNumber="508">Serdia</emphasis>
</taxonomicName>
,
<taxonomicName authority="Froeschner, 1981" authorityName="Froeschner" authorityYear="1981" box="[234,494,1505,1525]" class="Insecta" family="Pentatomidae" genus="Elsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[234,304,1505,1524]" italics="true" pageId="1" pageNumber="508">Elsiella</emphasis>
Froeschner, 1981
</taxonomicName>
, and
<taxonomicName authority="Fortes &amp; Grazia, 1998" authorityName="Fortes &amp; Grazia" authorityYear="1998" box="[557,670,1505,1524]" class="Insecta" family="Pentatomidae" genus="Semiliserdia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[557,670,1505,1524]" italics="true" pageId="1" pageNumber="508">Similiserdia</emphasis>
</taxonomicName>
. It can be separated from these genera by the extremely reduced second antennal segment, and, at least from
<taxonomicName authority="Fortes &amp; Grazia, 1998" authorityName="Fortes &amp; Grazia" authorityYear="1998" box="[530,640,1556,1575]" class="Insecta" family="Pentatomidae" genus="Semiliserdia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[530,640,1556,1575]" italics="true" pageId="1" pageNumber="508">Similiserdia</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Stal" authorityYear="1858" box="[689,746,1556,1575]" class="Insecta" family="Pentatomidae" genus="Serdia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[689,746,1556,1575]" italics="true" pageId="1" pageNumber="508">Serdia</emphasis>
</taxonomicName>
, by the presence of well-developed
<emphasis box="[483,598,1582,1601]" italics="true" pageId="1" pageNumber="508">hypandrium</emphasis>
on ventral rim of pygophore (
<taxonomicName authorityName="Froeschner" authorityYear="1981" box="[281,349,1608,1627]" class="Insecta" family="Pentatomidae" genus="Elsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[281,349,1608,1627]" italics="true" pageId="1" pageNumber="508">Elsiella</emphasis>
</taxonomicName>
is known only from females). In
<taxonomicName authorityName="Stal" authorityYear="1858" box="[681,738,1608,1627]" class="Insecta" family="Pentatomidae" genus="Serdia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[681,738,1608,1627]" italics="true" pageId="1" pageNumber="508">Serdia</emphasis>
</taxonomicName>
and one of the species of
<taxonomicName authority="Fortes &amp; Grazia, 1998" authorityName="Fortes &amp; Grazia" authorityYear="1998" box="[390,501,1633,1652]" class="Insecta" family="Pentatomidae" genus="Semiliserdia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[390,501,1633,1652]" italics="true" pageId="1" pageNumber="508">Similiserdia</emphasis>
</taxonomicName>
(
<emphasis box="[518,631,1633,1653]" italics="true" pageId="1" pageNumber="508">S. aciculata</emphasis>
) the antennal segment II is almost equal in length with segment I, and never surpassing 1/3 of III; in
<taxonomicName authority="Grazia &amp; Frey-da-Silva, 2003" authorityName="Grazia &amp; Frey-da-Silva" authorityYear="2003" box="[427,521,1684,1703]" class="Insecta" family="Pentatomidae" genus="Rideriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis box="[427,521,1684,1703]" italics="true" pageId="1" pageNumber="508">Rideriana</emphasis>
</taxonomicName>
<emphasis bold="true" box="[531,635,1685,1704]" pageId="1" pageNumber="508">
<taxonomicNameLabel box="[531,635,1685,1704]" pageId="1" pageNumber="508" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
the antennal segment II corresponds to 1/3 of antennal segment Iand almost 1/9 of segment III. Also
<taxonomicName authority="Grazia &amp; Frey-da-Silva, 2003" authorityName="Grazia &amp; Frey-da-Silva" authorityYear="2003" box="[389,478,1736,1755]" class="Insecta" family="Pentatomidae" genus="Rideriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis box="[389,478,1736,1755]" italics="true" pageId="1" pageNumber="508">Rideriana</emphasis>
</taxonomicName>
<emphasis bold="true" box="[483,576,1736,1755]" pageId="1" pageNumber="508">
<taxonomicNameLabel box="[483,576,1736,1755]" pageId="1" pageNumber="508" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName authority="Fortes &amp; Grazia, 1998" authorityName="Fortes &amp; Grazia" authorityYear="1998" box="[217,327,1761,1780]" class="Insecta" family="Pentatomidae" genus="Semiliserdia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[217,327,1761,1780]" italics="true" pageId="1" pageNumber="508">Similiserdia</emphasis>
</taxonomicName>
by the presence of mesosternal carina, and jugae with lateral margins slightly deflected. In
<taxonomicName authority="Fortes &amp; Grazia, 1998" authorityName="Fortes &amp; Grazia" authorityYear="1998" box="[634,744,1787,1806]" class="Insecta" family="Pentatomidae" genus="Semiliserdia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[634,744,1787,1806]" italics="true" pageId="1" pageNumber="508">Similiserdia</emphasis>
</taxonomicName>
the mesosternum is sulcate and the lateral margins of jugae are not deflected. From
<taxonomicName authorityName="Stal" authorityYear="1858" box="[1028,1085,1114,1133]" class="Insecta" family="Pentatomidae" genus="Serdia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[1028,1085,1114,1133]" italics="true" pageId="1" pageNumber="508">Serdia</emphasis>
</taxonomicName>
and
<emphasis box="[1139,1212,1113,1133]" italics="true" pageId="1" pageNumber="508">
<taxonomicName authorityName="Froeschner" authorityYear="1981" box="[1139,1208,1113,1133]" class="Insecta" family="Pentatomidae" genus="Elsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">Elsiella</taxonomicName>
,
</emphasis>
the new genus can be separated by the antennal segment Isurpassing apex of head; both genera have antennal segment I not, or at least, reaching apex of head. Finally,
<taxonomicName authority="Grazia &amp; Frey-da-Silva, 2003" authorityName="Grazia &amp; Frey-da-Silva" authorityYear="2003" box="[1049,1139,1191,1210]" class="Insecta" family="Pentatomidae" genus="Rideriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis box="[1049,1139,1191,1210]" italics="true" pageId="1" pageNumber="508">Rideriana</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1147,1247,1191,1210]" pageId="1" pageNumber="508">
<taxonomicNameLabel box="[1147,1247,1191,1210]" pageId="1" pageNumber="508" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
has short rostrum, attaining mesosternum, a character shared with
<taxonomicName authority="Fortes &amp; Grazia, 1998" authorityName="Fortes &amp; Grazia" authorityYear="1998" box="[1292,1401,1216,1235]" class="Insecta" family="Pentatomidae" genus="Semiliserdia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[1292,1401,1216,1235]" italics="true" pageId="1" pageNumber="508">Similiserdia</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Stal" authorityYear="1858" box="[820,877,1242,1261]" class="Insecta" family="Pentatomidae" genus="Serdia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[820,877,1242,1261]" italics="true" pageId="1" pageNumber="508">Serdia</emphasis>
</taxonomicName>
; in
<taxonomicName authorityName="Froeschner" authorityYear="1981" box="[914,982,1242,1261]" class="Insecta" family="Pentatomidae" genus="Elsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis box="[914,982,1242,1261]" italics="true" pageId="1" pageNumber="508">Elsiella</emphasis>
</taxonomicName>
the rostrum just surpasses the anterior margin of third abdominal sternite.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="508" type="etymology">
<paragraph blockId="1.[820,1444,1113,1342]" pageId="1" pageNumber="508">Etymology. The genus is named in honour to Dr. David Rider for his continued effort in cataloguing the pentatomids.</paragraph>
</subSubSection>
</treatment>
</document>