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<document id="97BB0A403DC2FB703CE68C0D0FDE648E" ID-CLB-Dataset="28429" ID-DOI="10.11646/zootaxa.4564.1.4" ID-GBIF-Dataset="6a6ca226-8343-4208-8cdb-f0bd627c3817" ID-ISSN="1175-5326" ID-Zenodo-Dep="2588770" ID-ZooBank="D0A9801B-8049-4211-A4A7-D7792B9D6936" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552176191024" checkinUser="plazi" docAuthor="Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes &amp; Pitts, James P." docDate="2019" docId="03F187A07C0DCD3827B79975FDC7FA39" docLanguage="en" docName="zootaxa.4564.1.4.pdf" docOrigin="Zootaxa 4564 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lomachaeta eironeia Williams &amp; Cambra &amp; Bartholomay &amp; Luz &amp; Quintero &amp; Pitts 2019, sp. nov." docType="treatment" docVersion="16" lastPageNumber="112" masterDocId="FFC8FFD87C06CD332720985CFFE2FFA6" masterDocTitle="Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations" masterLastPageNumber="136" masterPageNumber="101" pageNumber="112" updateTime="1698700719565" updateUser="ExternalLinkService">
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<mods:title id="3D2781F6DB27C513776DA76AFAA4DAF1">Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations</mods:title>
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<mods:namePart id="C816ACD3CF1BEC0DE22F95A3D3F7AC20">Williams, Kevin A.</mods:namePart>
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<mods:namePart id="A7BCAB82C996A4AE9FBD8018D7D1B04B">Cambra, Roberto A.</mods:namePart>
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<mods:namePart id="1085A502AEA3DB8A007B5F781641EFF3">Bartholomay, Pedro R.</mods:namePart>
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<mods:namePart id="7B7F8A856949006663F28255AD020FC9">Luz, David R.</mods:namePart>
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<mods:namePart id="DC07ACD7923E1F6DDF16D6A7DFD28AFB">Quintero, Diomedes</mods:namePart>
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<mods:namePart id="F270AABAFBA77B65CF911C4D65CB7E86">Pitts, James P.</mods:namePart>
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<treatment id="03F187A07C0DCD3827B79975FDC7FA39" ID-DOI="http://doi.org/10.5281/zenodo.5943919" ID-GBIF-Taxon="154366382" ID-Zenodo-Dep="5943919" LSID="urn:lsid:plazi:treatment:03F187A07C0DCD3827B79975FDC7FA39" httpUri="http://treatment.plazi.org/id/03F187A07C0DCD3827B79975FDC7FA39" lastPageNumber="112" pageId="11" pageNumber="112">
<subSubSection id="C342653D7C0DCD3827B79975FD98FEE5" box="[151,634,297,323]" pageId="11" pageNumber="112" type="nomenclature">
<paragraph id="8BE736B67C0DCD3827B79975FD98FEE5" blockId="11.[151,634,297,357]" box="[151,634,297,323]" pageId="11" pageNumber="112">
<heading id="D0AF81DA7C0DCD3827B79975FD98FEE5" bold="true" box="[151,634,297,323]" fontSize="11" level="1" pageId="11" pageNumber="112" reason="1">
<emphasis id="B92CEAA47C0DCD3827B79975FD98FEE5" bold="true" box="[151,634,297,323]" pageId="11" pageNumber="112">
<taxonomicName id="4C584D357C0DCD3827B79975FDF2FEE5" authority="Williams" authorityName="Williams &amp; Cambra &amp; Bartholomay &amp; Luz &amp; Quintero &amp; Pitts" authorityYear="2019" box="[151,528,297,323]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="eironeia" status="sp. nov.">
<emphasis id="B92CEAA47C0DCD3827B79975FE75FEE5" bold="true" box="[151,407,297,323]" italics="true" pageId="11" pageNumber="112">Lomachaeta eironeia</emphasis>
Williams
</taxonomicName>
,
<taxonomicNameLabel id="A21F57DF7C0DCD38253D9975FD98FEE5" box="[541,634,297,323]" pageId="11" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C342653D7C0DCD3827B79910FEB3FEC3" box="[151,337,332,357]" pageId="11" pageNumber="112" type="description">
<paragraph id="8BE736B67C0DCD3827B79910FEB3FEC3" blockId="11.[151,634,297,357]" box="[151,337,332,357]" pageId="11" pageNumber="112">
(
<figureCitation id="13632A337C0DCD3827BF9910FF17FEC3" box="[159,245,332,357]" captionStart="FIGURES 5760" captionStartId="28.[151,264,1963,1986]" captionTargetBox="[186,1407,798,1949]" captionTargetId="figure@28.[186,1407,798,1949]" captionTargetPageId="28" captionText="FIGURES 5760. Lomachaeta males; lateral habitus. 57) L. theresa; 58) L. snellingella; 59) L. eironeia, sp. nov.; 60) L. polemomechana. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588794/files/figure.png" pageId="11" pageNumber="112">Figs 59</figureCitation>
,
<figureCitation id="13632A337C0DCD3826219910FEABFEC3" box="[257,329,332,357]" captionStart="FIGURES 6572" captionStartId="32.[151,264,1932,1955]" captionTargetBox="[191,1396,193,1898]" captionTargetId="figure@32.[191,1396,193,1898]" captionTargetPageId="32" captionText="FIGURES 6572. Lomachaeta line drawings: female mandible (65, 66), male genitalia, lateral view (67, 69, 71), penis valve, lateral view (68, 70, 72). 65) L. hicksi; 66) L. beadugrimi; 6768) L. calamondin, sp. nov.; 6970) L. eironeia, sp. nov.; 7172) L. juanita, sp. nov." httpUri="https://zenodo.org/record/2588800/files/figure.png" pageId="11" pageNumber="112">69, 70</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C342653D7C0DCD3827B799C8FB0AFD9A" pageId="11" pageNumber="112" type="diagnosis">
<paragraph id="8BE736B67C0DCD3827B799C8FB0AFD9A" blockId="11.[151,1437,404,1440]" pageId="11" pageNumber="112">
<emphasis id="B92CEAA47C0DCD3827B799C8FEF3FE0B" bold="true" box="[151,273,404,429]" pageId="11" pageNumber="112">Diagnosis.</emphasis>
MALE. This species can be immediately recognized by the unique genitalia, wherein the cuspis and lateral paramere margins have numerous elongate (~5 × paramere width) setae. The following characters are also useful for diagnosis: the body is entirely blackish; the head and mesosoma have coarse separated punctures; the mandible is unarmed ventrally; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is sub-disciform; the T2 disc is densely coarsely punctate; the T2 fringe is composed of simple setae.
</paragraph>
</subSubSection>
<subSubSection id="C342653D7C0DCD3827E79A14FE59FBDB" pageId="11" pageNumber="112" type="description">
<paragraph id="8BE736B67C0DCD3827E79A14FA9BFBFF" blockId="11.[151,1437,404,1440]" pageId="11" pageNumber="112">
<emphasis id="B92CEAA47C0DCD3827E79A14FEBBFDC7" bold="true" box="[199,345,584,609]" pageId="11" pageNumber="112">Description.</emphasis>
MALE. Body length
<quantity id="4CA09B537C0DCD38256B9A14FD6DFDC6" box="[587,655,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="112" unit="mm" value="4.0">4 mm</quantity>
.
<emphasis id="B92CEAA47C0DCD3825B99A14FCF6FDC7" box="[665,788,584,609]" italics="true" pageId="11" pageNumber="112">Coloration</emphasis>
(
<figureCitation id="13632A337C0DCD3824029A14FC94FDC6" box="[802,886,584,609]" captionStart="FIGURES 3740" captionStartId="20.[151,264,1148,1171]" captionTargetBox="[167,1420,193,1127]" captionTargetId="figure@20.[167,1420,193,1127]" captionTargetPageId="20" captionText="FIGURES 3740. Lomachaeta females; lateral habitus. 37) L. hedera; 38) L. beadugrimi; 39) L. powelli; 40) L. warneri, sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588786/files/figure.png" pageId="11" pageNumber="112">Fig. 40</figureCitation>
). Entire body black, except mandible, tegula, and legs largely brown. Tibial spurs white. Wings hyaline, veins brown. Body setae whitish, except frons, vertex, and mesoscutum with interspersed erect silvery and brownish brachyplumose setae.
<emphasis id="B92CEAA47C0DCD3823089ACCFB8EFD01" box="[1064,1132,656,681]" italics="true" pageId="11" pageNumber="112">Head.</emphasis>
Rounded posteriorly, with coarse punctures, coarser on face. Mandible tridentate apically, unarmed ventrally. Gena not carinate. Ocelli small; ocellocular distance 6 × length of lateral ocellus, interocellar distance 2 × lateral ocellar diameter. F1 1.0 × pedicel length; F2 1.6 × pedicel length.
<emphasis id="B92CEAA47C0DCD38252C9AA1FD69FCB2" box="[524,651,765,788]" italics="true" pageId="11" pageNumber="112">Mesosoma.</emphasis>
Pronotum and mesoscutum with coarse separated punctures. Tegula smooth, margins setigerously punctate. Mesopleuron with coarse contiguous punctures. Metapleuron glabrous. Scutellum slightly convex, with coarse confluent punctures. Axilla sessile with mesoscutum, with smooth curved sharp posterior tooth. Propodeum areolate dorsally, lateral face mostly smooth with faint areolations posteriorly. Forewing with veins encompassing basal 0.7 × wing.
<emphasis id="B92CEAA47C0DCD3825CC9BD1FC8AFC02" box="[748,872,909,932]" italics="true" pageId="11" pageNumber="112">Metasoma.</emphasis>
T1 shape sub-disciform, with defined anterior and dorsal faces. T2 with coarse dense punctures, intervals smooth; T24 fringe setae slightly thicker than disc setae, but not forming bristles. S2 with coarse separated punctures. T36 punctate. T7 smooth. Hypopygium punctate, sharply bidentate posteriorly.
<emphasis id="B92CEAA47C0DCD3826C39BA4FDAFFBB7" box="[483,589,1016,1041]" italics="true" pageId="11" pageNumber="112">Genitalia</emphasis>
(
<figureCitation id="13632A337C0DCD38257C9BA4FD3CFBB6" box="[604,734,1016,1041]" captionStart="FIGURES 6572" captionStartId="32.[151,264,1932,1955]" captionTargetBox="[191,1396,193,1898]" captionTargetId="figure@32.[191,1396,193,1898]" captionTargetPageId="32" captionText="FIGURES 6572. Lomachaeta line drawings: female mandible (65, 66), male genitalia, lateral view (67, 69, 71), penis valve, lateral view (68, 70, 72). 65) L. hicksi; 66) L. beadugrimi; 6768) L. calamondin, sp. nov.; 6970) L. eironeia, sp. nov.; 7172) L. juanita, sp. nov." httpUri="https://zenodo.org/record/2588800/files/figure.png" pageId="11" pageNumber="112">Figs 69, 70</figureCitation>
). Paramere dorsoventrally flattened, obliquely downcurving to apex, sub-truncate apically, with long (~5 × paramere width) ventral setae around inner and lateral margins, setae longer and denser apically. Cuspis with posteroventral setae ~5 × cuspis length. Penis valve unidentate apically.
</paragraph>
<paragraph id="8BE736B67C0DCD3827E79C39FE59FBDB" blockId="11.[151,1437,404,1440]" box="[199,443,1124,1149]" pageId="11" pageNumber="112">FEMALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C342653D7C0DCD3827E79CD4FEADFB4D" pageId="11" pageNumber="112" type="materials_examined">
<paragraph id="8BE736B67C0DCD3827E79CD4FEADFB4D" blockId="11.[151,1437,404,1440]" pageId="11" pageNumber="112">
<emphasis id="B92CEAA47C0DCD3827E79CD4FE54FB07" bold="true" box="[199,438,1160,1185]" pageId="11" pageNumber="112">Material examined.</emphasis>
<materialsCitation id="3B303CEB7C0DCD3826E39CD4FEADFB4D" ID-GBIF-Occurrence="2012757423" collectingDate="2008-07-23" collectingDateMax="2008-07-26" collectingDateMin="2008-07-23" collectorName="M. E. Irwin &amp; R. L. Wescott" country="Mexico" elevation="40" latitude="27.7285" location="Hornos" longLatPrecision="12" longitude="-109.844" municipality="Hwy" pageId="11" pageNumber="112" specimenCount="1" specimenCount-male="1" stateProvince="Sonora" typeStatus="holotype">
<emphasis id="B92CEAA47C0DCD3826E39CD4FDCBFB07" box="[451,553,1160,1185]" italics="true" pageId="11" pageNumber="112">
<typeStatus id="54E388147C0DCD3826E39CD4FDCBFB07" box="[451,553,1160,1185]" pageId="11" pageNumber="112" type="holotype">Holotype</typeStatus>
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,
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,
<collectingCountry id="F34F76267C0DCD3825459CD4FD34FB06" box="[613,726,1160,1184]" name="Mexico" pageId="11" pageNumber="112">MEXICO</collectingCountry>
,
<emphasis id="B92CEAA47C0DCD3825C89CD5FCDBFB06" box="[744,825,1161,1184]" italics="true" pageId="11" pageNumber="112">
<collectingRegion id="499CF8547C0DCD3825C89CD5FCDBFB06" box="[744,825,1161,1184]" country="Mexico" name="Sonora" pageId="11" pageNumber="112">Sonora</collectingRegion>
</emphasis>
, km 23,
<collectingMunicipality id="6B83ACCC7C0DCD38248D9CD5FC01FB07" box="[941,995,1161,1185]" pageId="11" pageNumber="112">Hwy</collectingMunicipality>
16,
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ENE [Los]
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,
<date id="FFE610767C0DCD3822529CD4FEC1FB61" pageId="11" pageNumber="112" value="2008-07-23" valueMax="2008-07-26" valueMin="2008-07-23">
<collectingDate id="EFA2E99E7C0DCD3822529CD4FEC1FB61" pageId="11" pageNumber="112" value="2008-07-23" valueMax="2008-07-26" valueMin="2008-07-23">23 26.VII.2008</collectingDate>
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27
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43.71N
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109
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50.64W
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,
<collectorName id="26AD53607C0DCD3825B59CECFCF0FB61" box="[661,786,1198,1223]" pageId="11" pageNumber="112">M.E. Irwin</collectorName>
&amp;
<collectorName id="26AD53607C0DCD3824169CECFC2EFB61" box="[822,972,1200,1223]" pageId="11" pageNumber="112">R.L. Wescott</collectorName>
, malaise in dense riparian forest (EMUS,
<figureCitation id="13632A337C0DCD3827B79C8FFF0EFB4A" box="[151,236,1235,1260]" captionStart="FIGURES 5760" captionStartId="28.[151,264,1963,1986]" captionTargetBox="[186,1407,798,1949]" captionTargetId="figure@28.[186,1407,798,1949]" captionTargetPageId="28" captionText="FIGURES 5760. Lomachaeta males; lateral habitus. 57) L. theresa; 58) L. snellingella; 59) L. eironeia, sp. nov.; 60) L. polemomechana. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588794/files/figure.png" pageId="11" pageNumber="112">Figs 59</figureCitation>
,
<figureCitation id="13632A337C0DCD3827D89C8FFEA0FB4D" box="[248,322,1235,1260]" captionStart="FIGURES 6572" captionStartId="32.[151,264,1932,1955]" captionTargetBox="[191,1396,193,1898]" captionTargetId="figure@32.[191,1396,193,1898]" captionTargetPageId="32" captionText="FIGURES 6572. Lomachaeta line drawings: female mandible (65, 66), male genitalia, lateral view (67, 69, 71), penis valve, lateral view (68, 70, 72). 65) L. hicksi; 66) L. beadugrimi; 6768) L. calamondin, sp. nov.; 6970) L. eironeia, sp. nov.; 7172) L. juanita, sp. nov." httpUri="https://zenodo.org/record/2588800/files/figure.png" pageId="11" pageNumber="112">69, 70</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C342653D7C0DCD3827E79CAAFC23FAA9" box="[199,961,1270,1296]" pageId="11" pageNumber="112" type="distribution">
<paragraph id="8BE736B67C0DCD3827E79CAAFC23FAA9" blockId="11.[151,1437,404,1440]" box="[199,961,1270,1296]" pageId="11" pageNumber="112">
<emphasis id="B92CEAA47C0DCD3827E79CAAFE82FAA9" bold="true" box="[199,352,1270,1295]" pageId="11" pageNumber="112">Distribution.</emphasis>
Known only from the
<typeStatus id="54E388147C0DCD3825409CA4FD72FAB6" box="[608,656,1272,1296]" pageId="11" pageNumber="112">type</typeStatus>
locality in
<collectingRegion id="499CF8547C0DCD38242F9CABFC83FAA9" box="[783,865,1271,1295]" country="Mexico" name="Sonora" pageId="11" pageNumber="112">Sonora</collectingRegion>
,
<collectingCountry id="F34F76267C0DCD38244C9CAAFC23FAA9" box="[876,961,1270,1295]" name="Mexico" pageId="11" pageNumber="112">Mexico</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="C342653D7C0DCD3827E79D47FBAAFAFE" pageId="11" pageNumber="112" type="etymology">
<paragraph id="8BE736B67C0DCD3827E79D47FBAAFAFE" blockId="11.[151,1437,404,1440]" pageId="11" pageNumber="112">
<emphasis id="B92CEAA47C0DCD3827E79D47FEACFA92" bold="true" box="[199,334,1307,1332]" pageId="11" pageNumber="112">Etymology.</emphasis>
From the Greek,
<taxonomicName id="4C584D357C0DCD38253C9D40FD9AFA95" authorityName="Williams &amp; Cambra &amp; Bartholomay &amp; Luz &amp; Quintero &amp; Pitts" authorityYear="2019" box="[540,632,1308,1331]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="eironeia">
<emphasis id="B92CEAA47C0DCD38253C9D40FD9AFA95" box="[540,632,1308,1331]" italics="true" pageId="11" pageNumber="112">eironeia</emphasis>
</taxonomicName>
, irony, in reference to the fact that this specimen was recognized as a new species by KAW only a few months after
<bibRefCitation id="EFC94B477C0DCD3825809D63FC78FAF1" author="Williams, K. A. &amp; Pitts, J. P." box="[672,922,1343,1368]" pageId="11" pageNumber="112" pagination="227 - 243" refId="ref25741" refString="Williams, K. A. &amp; Pitts, J. P. (2009) Eight new species of Lomachaeta Mickel and the synonymy of Smicromutilla Mickel (Hymenoptera: Mutillidae). Journal of Hymenoptera Research, 18 (2), 227 - 243." type="journal article" year="2009">Williams &amp; Pitts 2009</bibRefCitation>
was published.
</paragraph>
</subSubSection>
<subSubSection id="C342653D7C0DCD3827E79D3FFDC7FA39" pageId="11" pageNumber="112" type="discussion">
<paragraph id="8BE736B67C0DCD3827E79D3FFDC7FA39" blockId="11.[151,1437,404,1440]" pageId="11" pageNumber="112">
<emphasis id="B92CEAA47C0DCD3827E79D3FFED6FADA" bold="true" box="[199,308,1379,1404]" pageId="11" pageNumber="112">Remarks</emphasis>
. The genitalia of this species are truly bizarre (
<figureCitation id="13632A337C0DCD3824589D3FFC30FADA" box="[888,978,1379,1404]" captionStart="FIGURES 5760" captionStartId="28.[151,264,1963,1986]" captionTargetBox="[186,1407,798,1949]" captionTargetId="figure@28.[186,1407,798,1949]" captionTargetPageId="28" captionText="FIGURES 5760. Lomachaeta males; lateral habitus. 57) L. theresa; 58) L. snellingella; 59) L. eironeia, sp. nov.; 60) L. polemomechana. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2588794/files/figure.png" pageId="11" pageNumber="112">Fig. 59</figureCitation>
), even for a genus with such variable parameres (
<bibRefCitation id="EFC94B477C0DCD3826389DDBFDFBFA39" author="Williams, K. A. &amp; Pitts, J. P." box="[280,537,1415,1440]" pageId="11" pageNumber="112" pagination="227 - 243" refId="ref25741" refString="Williams, K. A. &amp; Pitts, J. P. (2009) Eight new species of Lomachaeta Mickel and the synonymy of Smicromutilla Mickel (Hymenoptera: Mutillidae). Journal of Hymenoptera Research, 18 (2), 227 - 243." type="journal article" year="2009">Williams &amp; Pitts 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>