593 lines
73 KiB
XML
593 lines
73 KiB
XML
<document id="4372FEE693C61D0DE1FA98D5BD537320" ID-DOI="10.25221/fee.473.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="7945949" ID-ZooBank="E3519841-F78A-417F-B5F3-ACD59F84A622" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683042918429" checkinUser="felipe" docAuthor="Matalin, A. V." docDate="2023" docId="03F18796FF88DF704FBB3209FD686F48" docLanguage="en" docName="FarEastEntomol.473.1-19.pdf.imf" docOrigin="Far Eastern Entomologist 473" docSource="http://dx.doi.org/10.25221/fee.473.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.12:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="12" docTitle="Cylindera (Ifasina) klimenkoi Matalin 2023, sp. n." docType="treatment" docUuid="E3A67286-64B9-46DF-B6F8-FB53524D2E41" docUuidSource="ZooBank" docVersion="6" lastPageNumber="14" masterDocId="FFC8FFEEFF8DDF7D4F58372AFC506A41" masterDocTitle="An annotated checklist of the species of the subgenus Ifasina Jeannel, 1946, genus Cylindera Westwood, 1831 (Coleoptera, Cicindelidae), from the Philippines, with the description of a new species" masterLastPageNumber="19" masterPageNumber="1" pageNumber="6" updateTime="1712595810384" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="BA698E564845F3C3FF54C833FE848A27">An annotated checklist of the species of the subgenus Ifasina Jeannel, 1946, genus Cylindera Westwood, 1831 (Coleoptera, Cicindelidae), from the Philippines, with the description of a new species</mods:title>
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<mods:namePart id="A2E0C4AB9679FDCFD47EDD1565BF92AB">Matalin, A. V.</mods:namePart>
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<mods:title id="B1F65E67C06FEBB511BA0F06C954AD54">Far Eastern Entomologist</mods:title>
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<mods:date id="A73D06C3DAB0244E786B962ED38D176A">2023</mods:date>
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<mods:number id="44568C1F272646A30B0598AB888FA081">2023-03-31</mods:number>
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<mods:identifier id="91BA883F4090BE26D3F2A2531507FFDF" type="DOI">10.25221/fee.473.1</mods:identifier>
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<treatment id="03F18796FF88DF704FBB3209FD686F48" LSID="urn:lsid:zoobank.org:act:E3A67286-64B9-46DF-B6F8-FB53524D2E41" httpUri="http://treatment.plazi.org/id/03F18796FF88DF704FBB3209FD686F48" lastPageId="13" lastPageNumber="14" pageId="5" pageNumber="6">
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<subSubSection id="C342650BFF88DF784FBB3209F87A6F1D" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="8BE73680FF88DF784FBB3209FE846F7D" blockId="5.[227,1066,1315,1372]" box="[227,724,1315,1340]" pageId="5" pageNumber="6">
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<heading id="D0AF81ECFF88DF784FBB3209FE846F7D" box="[227,724,1315,1340]" fontSize="9" level="2" pageId="5" pageNumber="6" reason="2">
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<emphasis id="B92CEA92FF88DF784FBB3209FE846F7D" bold="true" box="[227,724,1315,1340]" pageId="5" pageNumber="6">
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<taxonomicName id="4C584D03FF88DF784FBB3209FEDD6F7A" authority="Matalin" authorityName="Matalin" authorityYear="2023" box="[227,653,1315,1339]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. nov." subGenus="Ifasina">
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<emphasis id="B92CEA92FF88DF784FBB3209FD1F6F7A" bold="true" box="[227,335,1315,1339]" italics="true" pageId="5" pageNumber="6">Cylindera</emphasis>
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||
(
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<emphasis id="B92CEA92FF88DF784E073209FDFD6F7A" bold="true" box="[351,429,1315,1339]" italics="true" pageId="5" pageNumber="6">Ifasina</emphasis>
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||
)
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<emphasis id="B92CEA92FF88DF784EE53209FE7B6F7A" bold="true" box="[445,555,1315,1339]" italics="true" pageId="5" pageNumber="6">klimenkoi</emphasis>
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Matalin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21F57E9FF88DF784DC1320EFE846F7D" box="[665,724,1316,1340]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BE73680FF88DF784FBB326CF87A6F1D" blockId="5.[227,1066,1315,1372]" box="[227,1066,1350,1372]" pageId="5" pageNumber="6">
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||
https://zoobank.org/NomenclaturalActs/
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<uuid id="FFFE0C55FF88DF784D36326CF87A6F1D" box="[622,1066,1350,1372]" pageId="5" pageNumber="6">E3A67286-64B9-46DF-B6F8-FB53524D2E41</uuid>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C342650BFF88DF784FBB3242FEBA6FC1" box="[227,746,1384,1408]" pageId="5" pageNumber="6" type="description">
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||
<paragraph id="8BE73680FF88DF784FBB3242FEBA6FC1" blockId="5.[227,746,1384,1408]" box="[227,746,1384,1408]" pageId="5" pageNumber="6">Figs 4, 5, 9–11, 13, 16, 17, 20, 21, 30–36, 41–44</paragraph>
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</subSubSection>
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<subSubSection id="C342650BFF88DF754E5132BDFE4F6EF3" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6" type="materials_examined">
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||
<paragraph id="8BE73680FF88DF784E5132BDF8126C07" blockId="5.[227,1127,1430,1606]" pageId="5" pageNumber="6">
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<materialsCitation id="3B303CDDFF88DF784E5132BDFF136F8C" collectingDate="2015-05-20" collectionCode="TYPE, MATERIAL" collectorName="N Luzon & A. Klimenko." country="Philippines" elevation="60" location="Dingalan" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Aurora" typeStatus="holotype">
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<collectionCode id="ED49AE45FF88DF784E5132BDFD1C6FEF" box="[265,332,1431,1454]" pageId="5" pageNumber="6">TYPE</collectionCode>
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<collectionCode id="ED49AE45FF88DF784E0D32BCFDB26FEF" box="[341,482,1430,1454]" pageId="5" pageNumber="6">MATERIAL</collectionCode>
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||
.
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||
<typeStatus id="54E38822FF88DF784EB732BCFE026FEF" box="[495,594,1430,1454]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
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||
–
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<specimenCount id="9D5EFD09FF88DF784D2B32BDFED96FEF" box="[627,649,1431,1454]" count="1" pageId="5" pageNumber="6" type="male">♂</specimenCount>
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||
,
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<emphasis id="B92CEA92FF88DF784DCF32BCFF136FEF" bold="true" box="[663,835,1430,1454]" pageId="5" pageNumber="6">
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||
the
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<collectingCountry id="F34F7610FF88DF784D9C32BCFF136FEF" box="[708,835,1430,1454]" name="Philippines" pageId="5" pageNumber="6">Philippines</collectingCountry>
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||
</emphasis>
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||
:
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<collectorName id="26AD5356FF88DF784C0C32BDFFE86FEF" box="[852,952,1431,1454]" pageId="5" pageNumber="6">N Luzon</collectorName>
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||
,
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<collectingRegion id="499CF862FF88DF784C9D32BCF8436FEF" box="[965,1043,1430,1454]" country="Philippines" name="Aurora" pageId="5" pageNumber="6">Aurora</collectingRegion>
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||
,
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||
<quantity id="4CA09B65FF88DF784B7932BCF8376FEF" box="[1057,1127,1430,1454]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="6" unit="km" value="15.0">15 km</quantity>
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NW
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||
<location id="8E87605BFF88DF784E40329FFD2C6F8C" LSID="urn:lsid:plazi:treatment:03F18796FF88DF704FBB3209FD686F48:8E87605BFF88DF784E40329FFD2C6F8C" box="[280,380,1461,1485]" country="Philippines" name="Dingalan" pageId="5" pageNumber="6" stateProvince="Aurora">Dingalan</location>
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, h =
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||
<quantity id="4CA09B65FF88DF784EEE329EFDA06F8D" box="[438,496,1460,1484]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="6" unit="m" value="60.0">
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||
<elevation id="0075D1B3FF88DF784EEE329EFDA06F8D" box="[438,496,1460,1484]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="6" unit="m" value="60.0">60 m</elevation>
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||
</quantity>
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||
,
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||
<date id="FFE61040FF88DF784EA5329FFE226F8D" box="[509,626,1460,1485]" pageId="5" pageNumber="6" value="2015-05-20">
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||
<collectingDate id="EFA2E9A8FF88DF784EA5329FFE226F8D" box="[509,626,1460,1485]" pageId="5" pageNumber="6" value="2015-05-20">20.V 2015</collectingDate>
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||
</date>
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||
, leg.
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<collectorName id="26AD5356FF88DF784DF6329FFF136F8C" box="[686,835,1461,1485]" pageId="5" pageNumber="6">A. Klimenko.</collectorName>
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||
</materialsCitation>
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||
<materialsCitation id="3B303CDDFF88DF784C13329FF86E6C07" collectionCode="MFNB" collectorName="Coll. Schaum" country="Philippines" pageId="5" pageNumber="6" specimenCode="MFNB 2022" specimenCount="41" specimenCount-female="12" specimenCount-male="29" stateProvince="Aurora" typeStatus="paratype">
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||
<typeStatus id="54E38822FF88DF784C13329FFFE76F8C" box="[843,951,1461,1485]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
|
||
:
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||
<specimenCount id="9D5EFD09FF88DF784C9A329FFFA46F8C" box="[962,1012,1461,1485]" count="29" pageId="5" pageNumber="6" type="male">29♂</specimenCount>
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||
,
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||
<specimenCount id="9D5EFD09FF88DF784B58329FF87F6F8C" box="[1024,1071,1461,1485]" count="10" pageId="5" pageNumber="6" type="female">10♀</specimenCount>
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||
with same labels as
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||
<typeStatus id="54E38822FF88DF784EDE32F9FDB76FAA" box="[390,487,1491,1515]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
;
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||
<specimenCount id="9D5EFD09FF88DF784EAA32F9FE436FAA" box="[498,531,1491,1515]" count="2" pageId="5" pageNumber="6" type="female">2♀</specimenCount>
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||
(digital images from the web-site https://carabidae.org/) with labels “Hist.-Coll. (
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<taxonomicName id="4C584D03FF88DF784EA532DBFE286C48" box="[509,632,1521,1545]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
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||
), Nr 42459,
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||
<taxonomicName id="4C584D03FF88DF784C4932DBF8026C48" authority="Schaum" authorityName="Schaum" authorityYear="1862" box="[785,1106,1521,1545]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="macilenta">
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<emphasis id="B92CEA92FF88DF784C4932DBFFBF6C48" box="[785,1007,1521,1545]" italics="true" pageId="5" pageNumber="6">Cicindela macilenta</emphasis>
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Schaum
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</taxonomicName>
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*, Luzon,
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<collectorName id="26AD5356FF88DF784E64313AFD806C69" box="[316,464,1552,1576]" pageId="5" pageNumber="6">Coll. Schaum</collectorName>
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||
, Zool. Muz. Berlin” [yellow printed label], “
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<typeStatus id="54E38822FF88DF784CBD313AF8326C69" box="[997,1122,1552,1576]" pageId="5" pageNumber="6" type="syntype">SYNTYPE</typeStatus>
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,
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<taxonomicName id="4C584D03FF88DF784FBB3104FE096C07" authority="Schaum, 1862" authorityName="Schaum" authorityYear="1862" box="[227,601,1582,1606]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="macilenta">
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Cicindela macilenta
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<bibRefCitation id="EFC94B71FF88DF784EE63104FE096C07" author="Schaum, H." box="[446,601,1582,1606]" pageId="5" pageNumber="6" pagination="172 - 184" refId="ref8630" refString="Schaum, H. 1862. Die Cicindeliden der philippinischen Inseln. Berliner Entomologische Zeitschrift, 1862: 172 - 184." type="journal article" year="1862">Schaum, 1862</bibRefCitation>
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</taxonomicName>
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, labelled by
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<specimenCode id="DBFE9EFBFF88DF784DB83105FF3B6C07" box="[736,875,1582,1606]" collectionCode="MFNB" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="5" pageNumber="6">MFNB 2022</specimenCode>
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” [red printed label]
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</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8BE73680FF8BDF7B4E513129F85C6C11" blockId="6.[227,1127,1539,1616]" pageId="6" pageNumber="7">
|
||
<taxonomicName id="4C584D03FF8BDF7B4E513129FD9C6C58" authorityName="Jeannel" authorityYear="1946" box="[265,460,1539,1561]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF8BDF7B4E513129FD396C59" box="[265,361,1539,1560]" italics="true" pageId="6" pageNumber="7">Cylindera</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF8BDF7B4EDA3129FD956C59" box="[386,453,1539,1560]" italics="true" pageId="6" pageNumber="7">Ifasina</emphasis>
|
||
)
|
||
</taxonomicName>
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||
spp, habitus, dorsal view (digital images from the web-site https:.carabidae.org\): 1–3 –
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||
<taxonomicName id="4C584D03FF8BDF7B4EA73135FE2C6C75" authorityName="Schaum" authorityYear="1862" box="[511,636,1567,1588]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="macilenta">
|
||
<emphasis id="B92CEA92FF8BDF7B4EA73135FE2C6C75" box="[511,636,1567,1588]" italics="true" pageId="6" pageNumber="7">C. macilenta</emphasis>
|
||
</taxonomicName>
|
||
; 4, 5 –
|
||
<taxonomicName id="4C584D03FF8BDF7B4D963135FF196C75" authority="Matalin, 2023" authorityName="Matalin" authorityYear="2023" box="[718,841,1567,1588]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. n.">
|
||
<emphasis id="B92CEA92FF8BDF7B4D963135FF196C75" box="[718,841,1567,1588]" italics="true" pageId="6" pageNumber="7">C. klimenkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F57E9FF8BDF7B4C0A3135FFDA6C75" box="[850,906,1567,1588]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
|
||
; 1, 3–5 – habitus; 2 – labels; 1 – male; 3–5 – females; 1, 2 –
|
||
<typeStatus id="54E38822FF8BDF7B4D0E3110FEE16C11" box="[598,689,1594,1616]" pageId="6" pageNumber="7" type="lectotype">lectotype</typeStatus>
|
||
; 3 –
|
||
<typeStatus id="54E38822FF8BDF7B4D873110FF336C11" box="[735,867,1594,1616]" pageId="6" pageNumber="7" type="paralectotype">paralectotype</typeStatus>
|
||
; 4, 5 –
|
||
<typeStatus id="54E38822FF8BDF7B4CF13111F8586C11" box="[937,1032,1595,1616]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF276608FF8ADF7A4E51312AFF056C0C" ID-DOI="http://doi.org/10.5281/zenodo.10944998" ID-Zenodo-Dep="10944998" httpUri="https://zenodo.org/record/10944998/files/figure.png" pageId="7" pageNumber="8" startId="7.[265,306,1536,1558]" targetBox="[310,1049,263,1495]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8BE73680FF8ADF7A4E51312AFF056C0C" blockId="7.[227,1127,1536,1613]" pageId="7" pageNumber="8">
|
||
Figs 6–11.
|
||
<taxonomicName id="4C584D03FF8ADF7A4E2E312AFE606C57" authorityName="Jeannel" authorityYear="1946" box="[374,560,1536,1558]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subGenus" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF8ADF7A4E2E312AFD866C54" box="[374,470,1536,1557]" italics="true" pageId="7" pageNumber="8">Cylindera</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF8ADF7A4EBD312AFE786C54" box="[485,552,1536,1557]" italics="true" pageId="7" pageNumber="8">Ifasina</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp, habitus: 6–8 –
|
||
<taxonomicName id="4C584D03FF8ADF7A4DAF312BFF236C54" authorityName="Schaum" authorityYear="1862" box="[759,883,1536,1558]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="macilenta">
|
||
<emphasis id="B92CEA92FF8ADF7A4DAF312BFF236C54" box="[759,883,1536,1558]" italics="true" pageId="7" pageNumber="8">C. macilenta</emphasis>
|
||
</taxonomicName>
|
||
(Luzon, Nueva Vizcaya, Santa Fe); 9–11 –
|
||
<taxonomicName id="4C584D03FF8ADF7A4ECF3136FE416C70" authority="Matalin, 2023" authorityName="Matalin" authorityYear="2023" box="[407,529,1564,1585]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. n.">
|
||
<emphasis id="B92CEA92FF8ADF7A4ECF3136FE416C70" box="[407,529,1564,1585]" italics="true" pageId="7" pageNumber="8">C. klimenkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F57E9FF8ADF7A4D403136FE1F6C70" box="[536,591,1564,1585]" pageId="7" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
|
||
; 6, 9 – males; 7, 8, 10, 11 – females; 9 – holotype; 10, 11 – paratypes; 6, 7, 9, 10 – dorsal view; 8, 11 – left lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE73680FF85DF754E513622F8356BDB" blockId="8.[227,1127,264,411]" pageId="8" pageNumber="9">
|
||
<materialsCitation id="3B303CDDFF85DF754E513622FEEA6B61" box="[265,698,264,288]" collectionCode="TYPE, DEPOSITION" pageId="8" pageNumber="9" specimenCount="1" typeStatus="holotype">
|
||
<collectionCode id="ED49AE45FF85DF754E513622FD1C6B5E" box="[265,332,264,287]" pageId="8" pageNumber="9">TYPE</collectionCode>
|
||
<collectionCode id="ED49AE45FF85DF754E0B3622FDA46B61" box="[339,500,264,288]" pageId="8" pageNumber="9">DEPOSITION</collectionCode>
|
||
. The
|
||
<typeStatus id="54E38822FF85DF754D773622FEDC6B61" box="[559,652,264,288]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
and
|
||
</materialsCitation>
|
||
<materialsCitation id="3B303CDDFF85DF754D9A3622FD836B7F" collectionCode="ZIN" pageId="8" pageNumber="9" specimenCount="22" specimenCount-female="2" specimenCount-male="9" typeStatus="paratype">
|
||
<specimenCount id="9D5EFD09FF85DF754D9A3622FF1A6B61" box="[706,842,264,288]" count="11" pageId="8" pageNumber="9" type="generic" typeStatus="paratype">11 paratypes</specimenCount>
|
||
(
|
||
<specimenCount id="9D5EFD09FF85DF754C033622FF2E6B61" box="[859,894,264,288]" count="9" pageId="8" pageNumber="9" type="male">9♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D5EFD09FF85DF754CD23622FFFB6B61" box="[906,939,264,288]" count="2" pageId="8" pageNumber="9" type="female">2♀</specimenCount>
|
||
) are deposited in the collection of
|
||
<collectionCode id="ED49AE45FF85DF754EFB360DFD836B7F" box="[419,467,295,318]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="9" type="Museum">ZIN</collectionCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B303CDDFF85DF754EB9360DF8696B7E" box="[481,1081,294,319]" collectionCode="MSPU" pageId="8" pageNumber="9" specimenCount="26" specimenCount-female="3" specimenCount-male="10" typeStatus="paratype">
|
||
<specimenCount id="9D5EFD09FF85DF754EB9360DFE3D6B7E" box="[481,621,295,319]" count="13" pageId="8" pageNumber="9" type="generic" typeStatus="paratype">13 paratypes</specimenCount>
|
||
(
|
||
<specimenCount id="9D5EFD09FF85DF754DD8360DFEE26B7E" box="[640,690,295,319]" count="10" pageId="8" pageNumber="9" type="male">10♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D5EFD09FF85DF754D98360DFEB16B7E" box="[704,737,295,319]" count="3" pageId="8" pageNumber="9" type="female">3♀</specimenCount>
|
||
) – in the collection of
|
||
<collectionCode id="ED49AE45FF85DF754CB5360DF8696B7E" box="[1005,1081,295,319]" pageId="8" pageNumber="9">MSPU</collectionCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B303CDDFF85DF754B1F360CFEA76B1C" collectionCode="ZMMU" pageId="8" pageNumber="9" specimenCount="12" specimenCount-female="2" specimenCount-male="4" typeStatus="paratype">
|
||
<specimenCount id="9D5EFD09FF85DF754B1F360CFD196B1F" count="6" pageId="8" pageNumber="9" type="generic" typeStatus="paratype">six paratypes</specimenCount>
|
||
(
|
||
<specimenCount id="9D5EFD09FF85DF754E03366FFDD16B1C" box="[347,385,325,349]" count="4" pageId="8" pageNumber="9" type="male">4♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D5EFD09FF85DF754EDD366FFDF46B1C" box="[389,420,325,349]" count="2" pageId="8" pageNumber="9" type="female">2♀</specimenCount>
|
||
) – in the collection of
|
||
<collectionCode id="ED49AE45FF85DF754DF9366CFEA76B1C" box="[673,759,326,349]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="8" pageNumber="9" type="Museum">ZMMU</collectionCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B303CDDFF85DF754C5C366FFDFE6B3D" collectionCode="SIEE" pageId="8" pageNumber="9" specimenCount="8" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
|
||
<specimenCount id="9D5EFD09FF85DF754C5C366FFFCF6B1F" box="[772,927,325,350]" count="4" pageId="8" pageNumber="9" type="generic" typeStatus="paratype">four paratypes</specimenCount>
|
||
(
|
||
<specimenCount id="9D5EFD09FF85DF754CE8366FFF846B1C" box="[944,980,325,349]" count="2" pageId="8" pageNumber="9" type="male">2♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D5EFD09FF85DF754CB8366FF8516B1C" box="[992,1025,325,349]" count="2" pageId="8" pageNumber="9" type="female">2♀</specimenCount>
|
||
) – in the collection of
|
||
<collectionCode id="ED49AE45FF85DF754E2A364EFDFE6B3D" box="[370,430,356,380]" pageId="8" pageNumber="9">SIEE</collectionCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B303CDDFF85DF754EE2364EFFE76B3D" box="[442,951,356,380]" collectionCode="SZM" pageId="8" pageNumber="9" specimenCount="4" specimenCount-male="2" typeStatus="paratype">
|
||
<specimenCount id="9D5EFD09FF85DF754EE2364EFE006B3D" box="[442,592,356,380]" count="2" pageId="8" pageNumber="9" type="generic" typeStatus="paratype">two paratypes</specimenCount>
|
||
(
|
||
<specimenCount id="9D5EFD09FF85DF754D38364EFED36B3D" box="[608,643,356,380]" count="2" pageId="8" pageNumber="9" type="male">2♂</specimenCount>
|
||
) – in the collection of
|
||
<collectionCode id="ED49AE45FF85DF754C25364EFFE76B3D" box="[893,951,356,380]" country="0" httpUri="http://grbio.org/cool/jbbv-ydyk" name="Saitama Zoogeographical Museum" pageId="8" pageNumber="9">SZM</collectionCode>
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="9D5EFD09FF85DF754C9B364EF8366B3D" box="[963,1126,356,380]" count="3" pageId="8" pageNumber="9" type="generic" typeStatus="paratype">three paratypes</specimenCount>
|
||
(
|
||
<specimenCount id="9D5EFD09FF85DF754FB336A9FD5F6BDA" box="[235,271,387,411]" count="2" pageId="8" pageNumber="9" type="male">2♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D5EFD09FF85DF754E4236A9FD6A6BDA" box="[282,314,387,411]" count="1" pageId="8" pageNumber="9" type="female">1♀</specimenCount>
|
||
) – in the collection of JW;
|
||
<materialsCitation id="3B303CDDFF85DF754D3836A9F8316BDB" box="[608,1121,386,411]" collectionCode="MFNB" pageId="8" pageNumber="9" specimenCount="4" specimenCount-female="2" typeStatus="paratype">
|
||
<specimenCount id="9D5EFD09FF85DF754D3836A9FEA56BDA" box="[608,757,387,411]" count="2" pageId="8" pageNumber="9" type="generic" typeStatus="paratype">two paratypes</specimenCount>
|
||
(
|
||
<specimenCount id="9D5EFD09FF85DF754C5D36A9FF766BDA" box="[773,806,387,411]" count="2" pageId="8" pageNumber="9" type="female">2♀</specimenCount>
|
||
) – in the collection of
|
||
<collectionCode id="ED49AE45FF85DF754B4B36A9F8316BDB" box="[1043,1121,387,410]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="8" pageNumber="9">MFNB</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE73680FF85DF754E513304FE4F6EF3" blockId="8.[227,1127,1070,1203]" pageId="8" pageNumber="9">
|
||
Figs 12–21.
|
||
<taxonomicName id="4C584D03FF85DF754ED83304FE696E05" authorityName="Jeannel" authorityYear="1946" box="[384,569,1070,1092]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subGenus" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF85DF754ED83304FDB06E02" box="[384,480,1070,1091]" italics="true" pageId="8" pageNumber="9">Cylindera</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF85DF754EB63304FE616E02" box="[494,561,1070,1091]" italics="true" pageId="8" pageNumber="9">Ifasina</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp, details: 12, 14, 15, 18, 19 –
|
||
<taxonomicName id="4C584D03FF85DF754C203305FFA36E02" authorityName="Schaum" authorityYear="1862" box="[888,1011,1070,1092]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macilenta">
|
||
<emphasis id="B92CEA92FF85DF754C203305FFA36E02" box="[888,1011,1070,1092]" italics="true" pageId="8" pageNumber="9">C. macilenta</emphasis>
|
||
</taxonomicName>
|
||
(12, 14, 15, 19 – Luzon,
|
||
<collectingRegion id="499CF862FF85DF754E063360FDA46E21" box="[350,500,1098,1120]" country="Philippines" name="Nueva Vizcaya" pageId="8" pageNumber="9">Nueva Vizcaya</collectingRegion>
|
||
, Santa Fe; 18 – Luzon,
|
||
<collectingRegion id="499CF862FF85DF754D833360FF226E21" box="[731,882,1098,1120]" country="Philippines" name="Nueva Vizcaya" pageId="8" pageNumber="9">Nueva Vizcaya</collectingRegion>
|
||
, Imugan); 13, 16, 17, 20, 21 –
|
||
<taxonomicName id="4C584D03FF85DF754E4D334CFDDF6E3B" authority="Matalin, 2023" authorityName="Matalin" authorityYear="2023" box="[277,399,1125,1147]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. n.">
|
||
<emphasis id="B92CEA92FF85DF754E4D334CFDDF6E3B" box="[277,399,1125,1147]" italics="true" pageId="8" pageNumber="9">C. klimenkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F57E9FF85DF754ECF334CFD9D6E3A" box="[407,461,1126,1147]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
|
||
; 12, 13 – head (12 – left half; 13 – right half); 14–17 – labrum; 18– 21 – pronotum, dorsal view; 12–14, 16, 18, 20 – males; 15, 17, 19, 21 – females; 16, 20 –
|
||
<typeStatus id="54E38822FF85DF754FBB33B7FD6A6EF2" box="[227,314,1181,1203]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
; 13, 17, 21 –
|
||
<typeStatus id="54E38822FF85DF754EE433B7FE4B6EF3" box="[444,539,1181,1202]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342650BFF85DF744E5133F2F87B6B1C" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
|
||
<paragraph id="8BE73680FF85DF744E5133F2F87B6B1C" blockId="8.[227,1127,1240,1601]" lastBlockId="9.[227,1127,264,472]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||
DIAGNOSIS. The new species seems to be particularly similar to
|
||
<taxonomicName id="4C584D03FF85DF754C8733F2FD486F4F" authorityName="Schaum" authorityYear="1862" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macilenta" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF85DF754C8733F2FFA86EAE" box="[991,1016,1240,1263]" italics="true" pageId="8" pageNumber="9">C.</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF85DF754B5133F3F8496EB1" box="[1033,1049,1241,1264]" italics="true" pageId="8" pageNumber="9">I.</emphasis>
|
||
)
|
||
<emphasis id="B92CEA92FF85DF754B7233F2FD486F4F" italics="true" pageId="8" pageNumber="9">macilenta</emphasis>
|
||
</taxonomicName>
|
||
, but it can easily be distinguished by the apical part of the elytra being much more strongly, roof-like elevated along the suture, especially so in females in
|
||
<taxonomicName id="4C584D03FF85DF754B7E323FFD1B6F0D" authority="Matalin, 2023" authorityName="Matalin" authorityYear="2023" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. n." subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF85DF754B7E323FF86E6F6D" box="[1062,1086,1301,1324]" italics="true" pageId="8" pageNumber="9">C.</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF85DF754B16323CF80E6F6C" box="[1102,1118,1302,1325]" italics="true" pageId="8" pageNumber="9">I.</emphasis>
|
||
)
|
||
<emphasis id="B92CEA92FF85DF754FBB321EFD1B6F0D" box="[227,331,1332,1356]" italics="true" pageId="8" pageNumber="9">klimenkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F57E9FF85DF754E0A321EFDDC6F0A" box="[338,396,1332,1355]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13632A05FF85DF754EC4321EFDBB6F0D" box="[412,491,1332,1356]" captionStart="Figs 6–11" captionStartId="7.[265,306,1536,1558]" captionTargetBox="[310,1049,263,1495]" captionTargetId="figure-0@7.[286,1068,257,1503]" captionTargetPageId="7" captionText="Figs 6–11. Cylindera (Ifasina) spp, habitus: 6–8 – C. macilenta (Luzon, Nueva Vizcaya, Santa Fe); 9–11 – C. klimenkoi sp. n.; 6, 9 – males; 7, 8, 10, 11 – females; 9 – holotype; 10, 11 – paratypes; 6, 7, 9, 10 – dorsal view; 8, 11 – left lateral view." figureDoi="http://doi.org/10.5281/zenodo.10944998" httpUri="https://zenodo.org/record/10944998/files/figure.png" pageId="8" pageNumber="9">Fig. 11</figureCitation>
|
||
),
|
||
<emphasis id="B92CEA92FF85DF754D59321FFE4E6F0D" box="[513,542,1333,1356]" italics="true" pageId="8" pageNumber="9">vs.</emphasis>
|
||
only slightly elevated in
|
||
<taxonomicName id="4C584D03FF85DF754C6A321EFFB46F0D" authorityName="Schaum" authorityYear="1862" box="[818,996,1332,1356]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macilenta" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF85DF754C6A321EFF1A6F0A" box="[818,842,1332,1355]" italics="true" pageId="8" pageNumber="9">C.</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF85DF754C03321FFF3B6F0D" box="[859,875,1333,1356]" italics="true" pageId="8" pageNumber="9">I.</emphasis>
|
||
)
|
||
<emphasis id="B92CEA92FF85DF754C23321EFFB46F0D" box="[891,996,1332,1356]" italics="true" pageId="8" pageNumber="9">macilenta</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13632A05FF85DF754CAB321EF8656F0D" box="[1011,1077,1332,1356]" captionStart="Figs 6–11" captionStartId="7.[265,306,1536,1558]" captionTargetBox="[310,1049,263,1495]" captionTargetId="figure-0@7.[286,1068,257,1503]" captionTargetPageId="7" captionText="Figs 6–11. Cylindera (Ifasina) spp, habitus: 6–8 – C. macilenta (Luzon, Nueva Vizcaya, Santa Fe); 9–11 – C. klimenkoi sp. n.; 6, 9 – males; 7, 8, 10, 11 – females; 9 – holotype; 10, 11 – paratypes; 6, 7, 9, 10 – dorsal view; 8, 11 – left lateral view." figureDoi="http://doi.org/10.5281/zenodo.10944998" httpUri="https://zenodo.org/record/10944998/files/figure.png" pageId="8" pageNumber="9">Fig. 8</figureCitation>
|
||
); by the colouration and white pattern of the elytra: an evenly delated middle band without lateral portion and the ante-apical portion of the apical lunule connected to the marginal portion at an obtuse angle (100–125
|
||
<superScript id="7C2D9BC8FF85DF754D9432A1FE856FD8" attach="none" box="[716,725,1419,1433]" fontSize="6" pageId="8" pageNumber="9">o</superScript>
|
||
) in
|
||
<taxonomicName id="4C584D03FF85DF754C5932BAFFE16FE9" authority="Matalin, 2023" authorityName="Matalin" authorityYear="2023" box="[769,945,1424,1448]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. n." subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF85DF754C5932BAFF4A6FE6" box="[769,794,1424,1447]" italics="true" pageId="8" pageNumber="9">C.</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF85DF754C7232BBFF6A6FE9" box="[810,826,1425,1448]" italics="true" pageId="8" pageNumber="9">I.</emphasis>
|
||
)
|
||
<emphasis id="B92CEA92FF85DF754C1132BAFFE16FE9" box="[841,945,1424,1448]" italics="true" pageId="8" pageNumber="9">klimenkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F57E9FF85DF754CE032BAFFA26FE6" box="[952,1010,1424,1447]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
|
||
(Figs 4, 5, 9, 10, 30–36),
|
||
<emphasis id="B92CEA92FF85DF754ED13285FDF66F87" box="[393,422,1455,1478]" italics="true" pageId="8" pageNumber="9">vs.</emphasis>
|
||
the middle band usually narrow in the centre and with a short lateral portion, as well as the ante-apical portion of the apical lunule connected to the marginal portion at an acute or virtually right angle (78–95
|
||
<superScript id="7C2D9BC8FF85DF754CDA32CDFFDB6FB4" attach="none" box="[898,907,1511,1525]" fontSize="6" pageId="8" pageNumber="9">o</superScript>
|
||
) in
|
||
<taxonomicName id="4C584D03FF85DF754CEE32C6F8366C45" authorityName="Schaum" authorityYear="1862" box="[950,1126,1516,1540]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macilenta" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF85DF754CEE32C6FF9F6C42" box="[950,975,1516,1539]" italics="true" pageId="8" pageNumber="9">C.</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF85DF754C8632C7FFBE6C45" box="[990,1006,1517,1540]" italics="true" pageId="8" pageNumber="9">I.</emphasis>
|
||
)
|
||
<emphasis id="B92CEA92FF85DF754CA532C6F8366C45" box="[1021,1126,1516,1540]" italics="true" pageId="8" pageNumber="9">macilenta</emphasis>
|
||
</taxonomicName>
|
||
(Figs 1, 3, 6, 7, 22–29); by the large basodiscal metallic mirror in females of
|
||
<taxonomicName id="4C584D03FF85DF754B7D3121FD1D6C00" authority="Matalin, 2023" authorityName="Matalin" authorityYear="2023" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. n." subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF85DF754B7D3121F86E6C63" box="[1061,1086,1547,1570]" italics="true" pageId="8" pageNumber="9">C.</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF85DF754B163121F80E6C63" box="[1102,1118,1547,1570]" italics="true" pageId="8" pageNumber="9">I.</emphasis>
|
||
)
|
||
<emphasis id="B92CEA92FF85DF754FBB3103FD1D6C00" box="[227,333,1577,1601]" italics="true" pageId="8" pageNumber="9">klimenkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F57E9FF85DF754E0C3100FDDE6C00" box="[340,398,1578,1601]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
|
||
(Figs 4, 5, 10, 34, 35, 47),
|
||
<emphasis id="B92CEA92FF85DF754DE03100FE866C00" box="[696,726,1578,1601]" italics="true" pageId="8" pageNumber="9">vs.</emphasis>
|
||
small or extremely to even entirely reduced in females of
|
||
<taxonomicName id="4C584D03FF84DF744E893622FED16B61" authorityName="Schaum" authorityYear="1862" box="[465,641,264,288]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macilenta" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF84DF744E893622FDBA6B5E" box="[465,490,264,287]" italics="true" pageId="9" pageNumber="10">C.</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF84DF744EA13623FE596B61" box="[505,521,265,288]" italics="true" pageId="9" pageNumber="10">I.</emphasis>
|
||
)
|
||
<emphasis id="B92CEA92FF84DF744D403622FED16B61" box="[536,641,264,288]" italics="true" pageId="9" pageNumber="10">macilenta</emphasis>
|
||
</taxonomicName>
|
||
(Figs 3, 7, 27, 28, 47); and by the longer and more strongly curved basal portion of the aedeagus in
|
||
<taxonomicName id="4C584D03FF84DF744C6C360DFFB66B7F" authority="Matalin, 2023" authorityName="Matalin" authorityYear="2023" box="[820,998,294,319]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. n." subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF84DF744C6C360DFF1C6B7F" box="[820,844,295,318]" italics="true" pageId="9" pageNumber="10">C.</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF84DF744C05360DFF3D6B7F" box="[861,877,295,318]" italics="true" pageId="9" pageNumber="10">I.</emphasis>
|
||
)
|
||
<emphasis id="B92CEA92FF84DF744C26360CFFB66B7F" box="[894,998,294,318]" italics="true" pageId="9" pageNumber="10">klimenkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F57E9FF84DF744CB5360DF8776B7F" box="[1005,1063,295,318]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13632A05FF84DF744B6F360CFD796B1C" captionStart="Figs 37–44" captionStartId="11.[265,306,1488,1510]" captionTargetBox="[335,1045,264,1450]" captionTargetId="figure-0@11.[294,1059,257,1455]" captionTargetPageId="11" captionText="Figs 37–44. Cylindera (Ifasina) spp, aedeagus, left lateral view: 37–40 – C. macilenta (specimens from: 37 – Luzon, Nueva Vizcaya, Santa Fe; 38, 39 – Luzon, Nueva Vizcaya, Imugan; 40 – Luzon, Quezon, Mt. Banachao); 41–44 – C. klimenkoi sp. n.; 41 – holotype; 42–44 – paratypes; f – flagellum; mt – median tooth." figureDoi="http://doi.org/10.5281/zenodo.10945004" httpUri="https://zenodo.org/record/10945004/files/figure.png" pageId="9" pageNumber="10">Figs 41–44</figureCitation>
|
||
,
|
||
<figureCitation id="13632A05FF84DF744E6C366FFD026B1C" box="[308,338,325,349]" captionStart="Figs 45–47" captionStartId="12.[265,306,1357,1379]" captionTargetBox="[363,990,257,1335]" captionTargetId="figure-0@12.[340,1013,257,1335]" captionTargetPageId="12" captionText="Figs 45–47. Morphometric features of Cylindera (Ifasina) spp: 45 – relation of the proportions elytra, head and pronotum (circles – C. klimenkoi sp. n.; rhombus – C. macilenta; open symbols – males; solid symbols – females; HT – holotype of C. klimenkoi sp. n.; LT – lectotype of C. macilenta; PLT – paralectotype of C. macilenta; PT – paratype of C. klimenkoi sp. n., former syntype of C. macilenta); 46 – relation of the proportions elytra and aedeagus; 47 – relation of the square of subsutural mirror and basodiscal spot; rectangle – 25–75 percentiles; horizontal line in rectangle – median; whiskers – SD; significantly different samples are marked with different lower case letters, according to Mann–Whitney U-test, p <0.05 (46 – U = 0, z = -3.546, p = 0.000, Monte Carlo p = 0.0001; 47 – U = 0, z = - 2.528, p = 0.012, Monte Carlo p = 0.0045)." figureDoi="http://doi.org/10.5281/zenodo.10945008" httpUri="https://zenodo.org/record/10945008/files/figure.png" pageId="9" pageNumber="10">46</figureCitation>
|
||
),
|
||
<emphasis id="B92CEA92FF84DF744E3C366CFDD16B1C" box="[356,385,326,349]" italics="true" pageId="9" pageNumber="10">vs.</emphasis>
|
||
shorter and poorly curved in
|
||
<taxonomicName id="4C584D03FF84DF744DE4366FFF3C6B1C" authorityName="Schaum" authorityYear="1862" box="[700,876,325,349]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macilenta" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF84DF744DE4366FFE856B1D" box="[700,725,325,348]" italics="true" pageId="9" pageNumber="10">C.</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF84DF744DBC366CFEA46B1C" box="[740,756,326,349]" italics="true" pageId="9" pageNumber="10">I.</emphasis>
|
||
)
|
||
<emphasis id="B92CEA92FF84DF744C5B366FFF3C6B1C" box="[771,876,325,349]" italics="true" pageId="9" pageNumber="10">macilenta</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13632A05FF84DF744C23366FFFA66B1C" box="[891,1014,325,349]" captionStart="Figs 37–44" captionStartId="11.[265,306,1488,1510]" captionTargetBox="[335,1045,264,1450]" captionTargetId="figure-0@11.[294,1059,257,1455]" captionTargetPageId="11" captionText="Figs 37–44. Cylindera (Ifasina) spp, aedeagus, left lateral view: 37–40 – C. macilenta (specimens from: 37 – Luzon, Nueva Vizcaya, Santa Fe; 38, 39 – Luzon, Nueva Vizcaya, Imugan; 40 – Luzon, Quezon, Mt. Banachao); 41–44 – C. klimenkoi sp. n.; 41 – holotype; 42–44 – paratypes; f – flagellum; mt – median tooth." figureDoi="http://doi.org/10.5281/zenodo.10945004" httpUri="https://zenodo.org/record/10945004/files/figure.png" pageId="9" pageNumber="10">Figs 37–40</figureCitation>
|
||
,
|
||
<figureCitation id="13632A05FF84DF744B58366FF84F6B1C" box="[1024,1055,325,349]" captionStart="Figs 45–47" captionStartId="12.[265,306,1357,1379]" captionTargetBox="[363,990,257,1335]" captionTargetId="figure-0@12.[340,1013,257,1335]" captionTargetPageId="12" captionText="Figs 45–47. Morphometric features of Cylindera (Ifasina) spp: 45 – relation of the proportions elytra, head and pronotum (circles – C. klimenkoi sp. n.; rhombus – C. macilenta; open symbols – males; solid symbols – females; HT – holotype of C. klimenkoi sp. n.; LT – lectotype of C. macilenta; PLT – paralectotype of C. macilenta; PT – paratype of C. klimenkoi sp. n., former syntype of C. macilenta); 46 – relation of the proportions elytra and aedeagus; 47 – relation of the square of subsutural mirror and basodiscal spot; rectangle – 25–75 percentiles; horizontal line in rectangle – median; whiskers – SD; significantly different samples are marked with different lower case letters, according to Mann–Whitney U-test, p <0.05 (46 – U = 0, z = -3.546, p = 0.000, Monte Carlo p = 0.0001; 47 – U = 0, z = - 2.528, p = 0.012, Monte Carlo p = 0.0045)." figureDoi="http://doi.org/10.5281/zenodo.10945008" httpUri="https://zenodo.org/record/10945008/files/figure.png" pageId="9" pageNumber="10">46</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342650BFF84DF744E51364EFDB86C0B" pageId="9" pageNumber="10" type="reference_group">
|
||
<paragraph id="8BE73680FF84DF744E51364EFFEB6B99" blockId="9.[227,1127,264,472]" pageId="9" pageNumber="10">
|
||
From
|
||
<taxonomicName id="4C584D03FF84DF744E12364EFECF6B3D" ID-CoL="3327J" authority="Dheurle, 2015" authorityName="Dheurle" authorityYear="2015" box="[330,671,356,380]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mouthiezi" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF84DF744E12364EFD336B3A" box="[330,355,356,379]" italics="true" pageId="9" pageNumber="10">C.</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF84DF744E2B364FFDD46B3D" box="[371,388,357,380]" italics="true" pageId="9" pageNumber="10">I.</emphasis>
|
||
)
|
||
<emphasis id="B92CEA92FF84DF744ECC364EFDAC6B3D" box="[404,508,356,380]" italics="true" pageId="9" pageNumber="10">mouthiezi</emphasis>
|
||
<bibRefCitation id="EFC94B71FF84DF744D5B364EFECF6B3D" author="Dheurle, Ch." box="[515,671,356,380]" pageId="9" pageNumber="10" pagination="123 - 124" refId="ref7702" refString="Dheurle, Ch. 2015. Cylindera (Ifasina) mouthiezi, nouvelle espece des Philippines (Coleoptera, Cicindelidae). L'Entomologiste, 71 (2): 123 - 124." type="journal article" year="2015">Dheurle, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
, the new species clearly differs by a complete apical lunule (Figs 4, 5, 9–11, 30–36),
|
||
<emphasis id="B92CEA92FF84DF744D8E36A9FEA36BDB" box="[726,755,387,410]" italics="true" pageId="9" pageNumber="10">vs.</emphasis>
|
||
incomplete in the former species (
|
||
<bibRefCitation id="EFC94B71FF84DF744FB2368BFDD96BF8" author="Dheurle, Ch." box="[234,393,417,441]" pageId="9" pageNumber="10" pagination="123 - 124" refId="ref7702" refString="Dheurle, Ch. 2015. Cylindera (Ifasina) mouthiezi, nouvelle espece des Philippines (Coleoptera, Cicindelidae). L'Entomologiste, 71 (2): 123 - 124." type="journal article" year="2015">Dheurle, 2015</bibRefCitation>
|
||
: fig. 3; 2017: figs 1–2), as well as by a pale central area of the labrum (Figs 16, 17),
|
||
<emphasis id="B92CEA92FF84DF744E2136EBFDC76B99" box="[377,407,449,472]" italics="true" pageId="9" pageNumber="10">vs.</emphasis>
|
||
black in the former species (
|
||
<bibRefCitation id="EFC94B71FF84DF744D9236EAFF396B99" author="Dheurle, Ch." box="[714,873,448,472]" pageId="9" pageNumber="10" pagination="123 - 124" refId="ref7702" refString="Dheurle, Ch. 2015. Cylindera (Ifasina) mouthiezi, nouvelle espece des Philippines (Coleoptera, Cicindelidae). L'Entomologiste, 71 (2): 123 - 124." type="journal article" year="2015">Dheurle, 2015</bibRefCitation>
|
||
: fig. 1).
|
||
</paragraph>
|
||
<caption id="DF276608FF84DF744E5133FBFE726F14" ID-DOI="http://doi.org/10.5281/zenodo.10945002" ID-Zenodo-Dep="10945002" httpUri="https://zenodo.org/record/10945002/files/figure.png" pageId="9" pageNumber="10" startId="9.[265,306,1233,1255]" targetBox="[233,1118,516,1192]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8BE73680FF84DF744E5133FBFE726F14" blockId="9.[227,1128,1233,1365]" pageId="9" pageNumber="10">
|
||
Figs 22–36.
|
||
<taxonomicName id="4C584D03FF84DF744EDA33FBFE6B6EA7" authorityName="Jeannel" authorityYear="1946" box="[386,571,1233,1254]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subGenus" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF84DF744EDA33FBFDB26EA7" box="[386,482,1233,1254]" italics="true" pageId="9" pageNumber="10">Cylindera</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF84DF744EA833FBFE636EA7" box="[496,563,1233,1254]" italics="true" pageId="9" pageNumber="10">Ifasina</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp, left elytron: 22–29 –
|
||
<taxonomicName id="4C584D03FF84DF744C6533FBFFE96EA7" authorityName="Schaum" authorityYear="1862" box="[829,953,1233,1254]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macilenta">
|
||
<emphasis id="B92CEA92FF84DF744C6533FBFFE96EA7" box="[829,953,1233,1254]" italics="true" pageId="9" pageNumber="10">C. macilenta</emphasis>
|
||
</taxonomicName>
|
||
(specimens from: 22 – Luzon; 23, 28, 29 – Luzon, Nueva Vizcaya, Santa Fe; 24, 25, 27 – Luzon, Nueva Vizcaya, Imugan; 26 – Luzon, Quezon, Mt. Banachao); 30–36 –
|
||
<taxonomicName id="4C584D03FF84DF744C383222FF8A6F5C" authority="Matalin, 2023" authorityName="Matalin" authorityYear="2023" box="[864,986,1288,1309]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. n.">
|
||
<emphasis id="B92CEA92FF84DF744C383222FF8A6F5C" box="[864,986,1288,1309]" italics="true" pageId="9" pageNumber="10">C. klimenkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F57E9FF84DF744CBA3222F8496F5C" box="[994,1049,1288,1309]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
; 22–26, 29, 30–33, 36 – males; 27, 28, 34, 35 – females; 22 – lectotype; 30 – holotype; 31–36 – paratypes; 29, 36 – apical lunule.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE73680FF84DF744E513251FE9C6FAF" blockId="9.[227,1127,1402,1610]" pageId="9" pageNumber="10">
|
||
From
|
||
<taxonomicName id="4C584D03FF84DF744E123251FEC06FD3" ID-CoL="8MMTP" authority="Dheurle, 2016" authorityName="Dheurle" authorityYear="2016" box="[330,656,1402,1426]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="richouxi" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF84DF744E123251FD336FD3" box="[330,355,1403,1426]" italics="true" pageId="9" pageNumber="10">C.</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF84DF744E2B3251FDD46FD3" box="[371,388,1403,1426]" italics="true" pageId="9" pageNumber="10">I.</emphasis>
|
||
)
|
||
<emphasis id="B92CEA92FF84DF744ECC3250FDBD6FD3" box="[404,493,1402,1426]" italics="true" pageId="9" pageNumber="10">richouxi</emphasis>
|
||
<bibRefCitation id="EFC94B71FF84DF744EAC3250FEC06FD3" author="Dheurle, Ch." box="[500,656,1402,1426]" pageId="9" pageNumber="10" pagination="401 - 403" refId="ref7735" refString="Dheurle, Ch. 2016. Cylindera (Ifasina) richouxi, nouvelle espece des Philippines (Coleoptera, Cicindelidae). L'Entomologiste, 72 (6): 401 - 403." type="journal article" year="2016">Dheurle, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
, the new species is reliably different by the humeral lunule separated into a humeral and a basodiscal spot (Figs 4, 5, 9, 10, 30– 35),
|
||
<emphasis id="B92CEA92FF84DF744E4E3292FD646F8E" box="[278,308,1464,1487]" italics="true" pageId="9" pageNumber="10">vs.</emphasis>
|
||
immaculate shoulders in females (
|
||
<bibRefCitation id="EFC94B71FF84DF744DE33292FF306F91" author="Dheurle, Ch." box="[699,864,1464,1488]" pageId="9" pageNumber="10" pagination="401 - 403" refId="ref7735" refString="Dheurle, Ch. 2016. Cylindera (Ifasina) richouxi, nouvelle espece des Philippines (Coleoptera, Cicindelidae). L'Entomologiste, 72 (6): 401 - 403." type="journal article" year="2016">Dheurle, 2016</bibRefCitation>
|
||
: fig. 3) or only a small humeral spot in males (
|
||
<bibRefCitation id="EFC94B71FF84DF744E8432FCFE2A6FAF" author="Dheurle, Ch." box="[476,634,1494,1518]" pageId="9" pageNumber="10" pagination="401 - 403" refId="ref7735" refString="Dheurle, Ch. 2016. Cylindera (Ifasina) richouxi, nouvelle espece des Philippines (Coleoptera, Cicindelidae). L'Entomologiste, 72 (6): 401 - 403." type="journal article" year="2016">Dheurle, 2016</bibRefCitation>
|
||
: fig. 1).
|
||
</paragraph>
|
||
<paragraph id="8BE73680FF84DF744E5132DFFDB86C0B" blockId="9.[227,1127,1402,1610]" pageId="9" pageNumber="10">
|
||
Unlike
|
||
<taxonomicName id="4C584D03FF84DF744E0032DFFEF76C4C" ID-CoL="3326F" authority="(Horn, 1893)" baseAuthorityName="Horn" baseAuthorityYear="1893" box="[344,679,1525,1549]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dilatotarsa" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF84DF744E0032DFFD216C4D" box="[344,369,1525,1548]" italics="true" pageId="9" pageNumber="10">C.</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF84DF744ED932DCFDC16C4C" box="[385,401,1526,1549]" italics="true" pageId="9" pageNumber="10">I.</emphasis>
|
||
)
|
||
<emphasis id="B92CEA92FF84DF744EF832DFFE456C4C" box="[416,533,1525,1549]" italics="true" pageId="9" pageNumber="10">dilatotarsa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC94B71FF84DF744D7B32DFFECE6C4C" author="Horn, W." box="[547,670,1525,1549]" pageId="9" pageNumber="10" pagination="320" refId="ref7904" refString="Horn, W. 1893. Cicindela semperi. Deutsche Entomologische Zeitschrift, 1893 (2): 320." type="journal article" year="1893">Horn, 1893</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the new species has a rounded, basodiscal white elytral spot (Figs 4, 5, 9, 10, 30–35) and a narrow, not widely dilated second fore-tarsomere in males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342650BFF87DF704E513622FD716E4E" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11" type="description">
|
||
<paragraph id="8BE73680FF87DF774E513622FED76B7E" blockId="10.[227,1127,264,1606]" pageId="10" pageNumber="11">
|
||
<collectionCode id="ED49AE45FF87DF774E513622FDEA6B61" box="[265,442,264,288]" pageId="10" pageNumber="11">DESCRIPTION</collectionCode>
|
||
. TL in males 7.0–7.7 mm (mean – 7.39 mm, n = 31), in females 7.4–8.2 mm (mean – 7.89 mm, n = 12).
|
||
</paragraph>
|
||
<paragraph id="8BE73680FF87DF774E51366FF8576983" blockId="10.[227,1127,264,1606]" pageId="10" pageNumber="11">
|
||
Head metallic bluish-green with distinct golden-bronze tinge on finely rugulose vertex and supraorbital plates; frons, antennal plates and clypeus with copper-gold reflection; genae finely striolate, metallic greenish-blue with light golden tinge. Eyes large, protruding, globose, HW/
|
||
<collectionCode id="ED49AE45FF87DF774D66368BFE366BF8" box="[574,614,417,441]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="11">PW</collectionCode>
|
||
(
|
||
<figureCitation id="13632A05FF87DF774D2F368BFE976BF8" box="[631,711,417,441]" captionStart="Figs 45–47" captionStartId="12.[265,306,1357,1379]" captionTargetBox="[363,990,257,1335]" captionTargetId="figure-0@12.[340,1013,257,1335]" captionTargetPageId="12" captionText="Figs 45–47. Morphometric features of Cylindera (Ifasina) spp: 45 – relation of the proportions elytra, head and pronotum (circles – C. klimenkoi sp. n.; rhombus – C. macilenta; open symbols – males; solid symbols – females; HT – holotype of C. klimenkoi sp. n.; LT – lectotype of C. macilenta; PLT – paralectotype of C. macilenta; PT – paratype of C. klimenkoi sp. n., former syntype of C. macilenta); 46 – relation of the proportions elytra and aedeagus; 47 – relation of the square of subsutural mirror and basodiscal spot; rectangle – 25–75 percentiles; horizontal line in rectangle – median; whiskers – SD; significantly different samples are marked with different lower case letters, according to Mann–Whitney U-test, p <0.05 (46 – U = 0, z = -3.546, p = 0.000, Monte Carlo p = 0.0001; 47 – U = 0, z = - 2.528, p = 0.012, Monte Carlo p = 0.0045)." figureDoi="http://doi.org/10.5281/zenodo.10945008" httpUri="https://zenodo.org/record/10945008/files/figure.png" pageId="10" pageNumber="11">Fig. 45</figureCitation>
|
||
) in males 1.5–1.67 (mean – 1.59, n = 31), in females 1.44–1.61 (mean – 1.55 mm, n = 10), supra-orbital plates with 3–4 deep and wide, and 9–10 shallow and narrow furrows, and two setae near each eye: the first seta at anterior edge, the second seta at posterior third (Fig. 13). Labrum pale with a narrow, black-brown, anterior edge and very broad, violet metallic lustered, lateral margins, entirely covered with isodiametric microsculpture, short and transverse, LW/LL in males 2.5–2.86 (mean – 2.56, n = 31), in females 2.2– 2.56 (mean – 2.38, n = 10), with an indistinctly curved anterior edge, a small central tooth and 8–11 (mean – 9, n = 41) submarginal setae placed far away from anterior margin, with neither basal dimples nor a central carina (Figs 16, 17). Mandibles black with greenish-golden-violet tinge at base of molars, pale in basal two-thirds. Labial and maxillary palpi pale (sometimes pre-apical palpomeres darker, yellowish brown) except for metallic, green, apical palpomeres. Antennae very long, extended posteriorly more than two-thirds the length of elytra, black, antennomeres 1–4 metallic greenish blue, 1st antennomere with copper-gold while 2nd–4th ones with purple-violet reflection; scape with a single, long, apical seta, antennomeres 3 and 4 with 4–5 and 1–2 short white spiniform setae, except apical ones, respectively, antennomeres 5–11 black, regularly covered with short yellow pubescence.
|
||
</paragraph>
|
||
<paragraph id="8BE73680FF87DF774E5134E3FF056EB4" blockId="10.[227,1127,264,1606]" pageId="10" pageNumber="11">
|
||
Pronotum longitudinal,
|
||
<collectionCode id="ED49AE45FF87DF774D5734E3FE7E69A0" box="[527,558,969,993]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="10" pageNumber="11" type="Museum">PL</collectionCode>
|
||
/
|
||
<collectionCode id="ED49AE45FF87DF774D6E34E3FE0E69A0" box="[566,606,969,993]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="11">PW</collectionCode>
|
||
in males 1.11–1.19 (mean – 1.14, n = 31), in females 1.0–1.07 (mean – 1.05, n = 12), with two irregular rows of sparse, white, decumbent setae along indistinctly rounded lateral sides and 5–6 setae at each anterior angle; pronotal disc finely and shallowly rugulose, metallic bronze with copper-green reflection; midline shallow and indistinct with a small depression anteriorly; anterior and posterior transverse grooves relatively deep, the latter with small basal pits (Figs 20, 21). Pro-episternum metallic greenish blue basally and copper-bronze with golden luster apically; with sparse white semi-erect hairs. Thoracic sternites metallic bluish green with golden shine, pro- and mesosternum glabrous, metathorax covered with white decumbent hairs.
|
||
</paragraph>
|
||
<paragraph id="8BE73680FF87DF774E5133D6FD966F8A" blockId="10.[227,1127,264,1606]" pageId="10" pageNumber="11">Legs long and slender, entirely violet except for light brown trochanters, with greenish-gold reflection along anterior surface of femora and at the base of tibiae; femora, as well as fore and hind tibiae with sparse, white, erect setae, apical half of mid-tibiae with numerous, very dense, white, erect setae along bottom surface; tarsomeres with short setae on bottom surface, in males fore-tarsomeres 1–3 distinctly thickened, 3rd one clearly dilated, 4th and 5th ones very thin; claws thin and moderately long.</paragraph>
|
||
<paragraph id="8BE73680FF87DF704E5132F8FD0E6839" blockId="10.[227,1127,264,1606]" lastBlockId="13.[227,1127,264,1289]" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11">
|
||
Elytra elongated oval; tapering towards apex; strongly elevated roof-like, especially so in females, along the suture in apical part of elytra (
|
||
<figureCitation id="13632A05FF87DF774CD532DAFF8E6C48" box="[909,990,1520,1545]" captionStart="Figs 6–11" captionStartId="7.[265,306,1536,1558]" captionTargetBox="[310,1049,263,1495]" captionTargetId="figure-0@7.[286,1068,257,1503]" captionTargetPageId="7" captionText="Figs 6–11. Cylindera (Ifasina) spp, habitus: 6–8 – C. macilenta (Luzon, Nueva Vizcaya, Santa Fe); 9–11 – C. klimenkoi sp. n.; 6, 9 – males; 7, 8, 10, 11 – females; 9 – holotype; 10, 11 – paratypes; 6, 7, 9, 10 – dorsal view; 8, 11 – left lateral view." figureDoi="http://doi.org/10.5281/zenodo.10944998" httpUri="https://zenodo.org/record/10944998/files/figure.png" pageId="10" pageNumber="11">Fig. 11</figureCitation>
|
||
); EL/EW in males 1.69–1.88 (mean – 1.79, n = 32), in females 1.64–1.83 (mean – 1.72, n = 12); copper-bronze with golden reflection except for a dull central area and numerous, evenly spaced, bright, blue points; suture with blue tinge, subsutural area green, scutellum bronze with greenish copper edges; a very narrow marginal edge deep blue, apical margin narrowly rounded, finely serrulate, sutural tooth small, epipleura brown with greenish blue tinge in basal quarter; white elytral pattern consisting of a disrupted humeral lunule separated into small a humeral and a basodiscal spot, a relatively narrow, clearly sinuate, median band with a distinct, rounded, apical knob, but without marginal portion, and a complete, narrow, apical lunule with its ante-apical portion connected to marginal portion at an obtuse angle – 100–125
|
||
<superScript id="7C2D9BC8FF80DF704B0636F4F8376BAD" attach="none" box="[1118,1127,478,492]" fontSize="6" pageId="13" pageNumber="14">o</superScript>
|
||
(Figs 4, 5, 9, 10, 30–36); in females additionally with a metallic golden-green patch behind humeral spot and a large, metallic, purple-bronze, subsutural mirror, 3.43– 9.2 (mean – 5.32, n = 12) times large than the white basodiscal spot (Figs 4, 5, 10, 34, 35, 47).
|
||
</paragraph>
|
||
<caption id="DF276608FF86DF764E5132FAFE8C6C78" ID-DOI="http://doi.org/10.5281/zenodo.10945004" ID-Zenodo-Dep="10945004" httpUri="https://zenodo.org/record/10945004/files/figure.png" pageId="11" pageNumber="12" startId="11.[265,306,1488,1510]" targetBox="[335,1045,264,1450]" targetPageId="11" targetType="figure">
|
||
<paragraph id="8BE73680FF86DF764E5132FAFE8C6C78" blockId="11.[227,1127,1488,1593]" pageId="11" pageNumber="12">
|
||
Figs 37–44.
|
||
<taxonomicName id="4C584D03FF86DF764EDE32FAFE126FA7" authorityName="Jeannel" authorityYear="1946" box="[390,578,1488,1510]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subGenus" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF86DF764EDE32FAFDB66FA4" box="[390,486,1488,1509]" italics="true" pageId="11" pageNumber="12">Cylindera</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF86DF764EAF32FBFE6A6FA7" box="[503,570,1489,1510]" italics="true" pageId="11" pageNumber="12">Ifasina</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp, aedeagus, left lateral view: 37–40 –
|
||
<taxonomicName id="4C584D03FF86DF764CB132FBF8376FA4" authorityName="Schaum" authorityYear="1862" box="[1001,1127,1488,1510]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="macilenta">
|
||
<emphasis id="B92CEA92FF86DF764CB132FBF8376FA4" box="[1001,1127,1488,1510]" italics="true" pageId="11" pageNumber="12">C. macilenta</emphasis>
|
||
</taxonomicName>
|
||
(specimens from: 37 – Luzon, Nueva Vizcaya, Santa Fe; 38, 39 – Luzon, Nueva Vizcaya, Imugan; 40 – Luzon, Quezon, Mt. Banachao); 41–44 –
|
||
<taxonomicName id="4C584D03FF86DF764C563122FFD96C5C" authority="Matalin, 2023" authorityName="Matalin" authorityYear="2023" box="[782,905,1544,1565]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. n.">
|
||
<emphasis id="B92CEA92FF86DF764C563122FFD96C5C" box="[782,905,1544,1565]" italics="true" pageId="11" pageNumber="12">C. klimenkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F57E9FF86DF764CCA3122FF9A6C5C" box="[914,970,1544,1565]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
|
||
; 41 – holotype; 42–44 – paratypes;
|
||
<emphasis id="B92CEA92FF86DF764EF83109FDF76C79" box="[416,423,1571,1592]" italics="true" pageId="11" pageNumber="12">f</emphasis>
|
||
– flagellum;
|
||
<emphasis id="B92CEA92FF86DF764D7F310EFE6F6C78" box="[551,575,1572,1593]" italics="true" pageId="11" pageNumber="12">mt</emphasis>
|
||
– median tooth.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF276608FF81DF714E513267FED36C1A" ID-DOI="http://doi.org/10.5281/zenodo.10945008" ID-Zenodo-Dep="10945008" httpUri="https://zenodo.org/record/10945008/files/figure.png" pageId="12" pageNumber="13" startId="12.[265,306,1357,1379]" targetBox="[363,990,257,1335]" targetPageId="12" targetType="figure">
|
||
<paragraph id="8BE73680FF81DF714E513267FED36C1A" blockId="12.[227,1127,1356,1627]" pageId="12" pageNumber="13">
|
||
Figs 45–47. Morphometric features of
|
||
<taxonomicName id="4C584D03FF81DF714DC23267FF096F23" authorityName="Jeannel" authorityYear="1946" box="[666,857,1357,1378]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subGenus" subGenus="Ifasina">
|
||
<emphasis id="B92CEA92FF81DF714DC23267FEAA6F23" box="[666,762,1357,1378]" italics="true" pageId="12" pageNumber="13">Cylindera</emphasis>
|
||
(
|
||
<emphasis id="B92CEA92FF81DF714C563267FF016F23" box="[782,849,1357,1378]" italics="true" pageId="12" pageNumber="13">Ifasina</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp: 45 – relation of the proportions elytra, head and pronotum (circles –
|
||
<taxonomicName id="4C584D03FF81DF714DE13243FF626F3C" authority="Matalin, 2023" authorityName="Matalin" authorityYear="2023" box="[697,818,1384,1406]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. n.">
|
||
<emphasis id="B92CEA92FF81DF714DE13243FF626F3C" box="[697,818,1384,1406]" italics="true" pageId="12" pageNumber="13">C. klimenkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F57E9FF81DF714C613243FF3E6F3F" box="[825,878,1385,1406]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
|
||
; rhombus –
|
||
<taxonomicName id="4C584D03FF81DF714CBD3243F8306F3C" authorityName="Schaum" authorityYear="1862" box="[997,1120,1384,1406]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="macilenta">
|
||
<emphasis id="B92CEA92FF81DF714CBD3243F8306F3C" box="[997,1120,1384,1406]" italics="true" pageId="12" pageNumber="13">C. macilenta</emphasis>
|
||
</taxonomicName>
|
||
; open symbols – males; solid symbols – females; HT – holotype of
|
||
<taxonomicName id="4C584D03FF81DF714C2A32AEFFBC6FD8" authority="Matalin, 2023" authorityName="Matalin" authorityYear="2023" box="[882,1004,1412,1433]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. n.">
|
||
<emphasis id="B92CEA92FF81DF714C2A32AEFFBC6FD8" box="[882,1004,1412,1433]" italics="true" pageId="12" pageNumber="13">C. klimenkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F57E9FF81DF714CAC32AEF87A6FD8" box="[1012,1066,1412,1433]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
|
||
; LT – lectotype of
|
||
<taxonomicName id="4C584D03FF81DF714E30328AFDBA6FF5" authorityName="Schaum" authorityYear="1862" box="[360,490,1439,1461]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="macilenta">
|
||
<emphasis id="B92CEA92FF81DF714E30328AFDBA6FF5" box="[360,490,1439,1461]" italics="true" pageId="12" pageNumber="13">C. macilenta</emphasis>
|
||
</taxonomicName>
|
||
; PLT – paralectotype of
|
||
<taxonomicName id="4C584D03FF81DF714DA3328AFF2D6FF5" authorityName="Schaum" authorityYear="1862" box="[763,893,1439,1461]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="macilenta">
|
||
<emphasis id="B92CEA92FF81DF714DA3328AFF2D6FF5" box="[763,893,1439,1461]" italics="true" pageId="12" pageNumber="13">C. macilenta</emphasis>
|
||
</taxonomicName>
|
||
; PT – paratype of
|
||
<taxonomicName id="4C584D03FF81DF714B09328AFD106F91" authority="Matalin, 2023" authorityName="Matalin" authorityYear="2023" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. n.">
|
||
<emphasis id="B92CEA92FF81DF714B09328AFD106F91" italics="true" pageId="12" pageNumber="13">C. klimenkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F57E9FF81DF714E1F3291FD2D6F91" box="[327,381,1467,1488]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
|
||
, former syntype of
|
||
<taxonomicName id="4C584D03FF81DF714D663291FEEA6F91" authorityName="Schaum" authorityYear="1862" box="[574,698,1467,1488]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="macilenta">
|
||
<emphasis id="B92CEA92FF81DF714D663291FEEA6F91" box="[574,698,1467,1488]" italics="true" pageId="12" pageNumber="13">C. macilenta</emphasis>
|
||
</taxonomicName>
|
||
); 46 – relation of the proportions elytra and aedeagus; 47 – relation of the square of subsutural mirror and basodiscal spot; rectangle – 25–75 percentiles; horizontal line in rectangle – median; whiskers – SD; significantly different samples are marked with different lower case letters, according to Mann–Whitney U-test, p <0.05 (46 – U = 0, z = -3.546, p = 0.000, Monte Carlo p = 0.0001; 47 – U = 0, z = - 2.528, p = 0.012, Monte Carlo p = 0.0045).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE73680FF80DF704E513555FEBF6975" blockId="13.[227,1127,264,1289]" pageId="13" pageNumber="14">
|
||
Abdominal sternites metallic greenish blue, regularly covered with white decumbent hairs. Aedeagus slender, moderately expanded in apical third, with a short, relatively broad apex and a long, thin, clearly curved, basal portion; AL 2.6–2.8 mm (mean – 2.7, n = 31), EL/AL (
|
||
<figureCitation id="13632A05FF80DF704D7D35F7FECF68B4" box="[549,671,733,757]" captionStart="Figs 45–47" captionStartId="12.[265,306,1357,1379]" captionTargetBox="[363,990,257,1335]" captionTargetId="figure-0@12.[340,1013,257,1335]" captionTargetPageId="12" captionText="Figs 45–47. Morphometric features of Cylindera (Ifasina) spp: 45 – relation of the proportions elytra, head and pronotum (circles – C. klimenkoi sp. n.; rhombus – C. macilenta; open symbols – males; solid symbols – females; HT – holotype of C. klimenkoi sp. n.; LT – lectotype of C. macilenta; PLT – paralectotype of C. macilenta; PT – paratype of C. klimenkoi sp. n., former syntype of C. macilenta); 46 – relation of the proportions elytra and aedeagus; 47 – relation of the square of subsutural mirror and basodiscal spot; rectangle – 25–75 percentiles; horizontal line in rectangle – median; whiskers – SD; significantly different samples are marked with different lower case letters, according to Mann–Whitney U-test, p <0.05 (46 – U = 0, z = -3.546, p = 0.000, Monte Carlo p = 0.0001; 47 – U = 0, z = - 2.528, p = 0.012, Monte Carlo p = 0.0045)." figureDoi="http://doi.org/10.5281/zenodo.10945008" httpUri="https://zenodo.org/record/10945008/files/figure.png" pageId="13" pageNumber="14">Figs 45, 46</figureCitation>
|
||
) 1.59–1.74 (mean – 1.67, n = 31); internal sack with ten sclerites, including a long flagellum forming 2.5 turns and a relatively long, slightly curved, median tooth (
|
||
<figureCitation id="13632A05FF80DF704D3E3436FEB36975" box="[614,739,796,820]" captionStart="Figs 37–44" captionStartId="11.[265,306,1488,1510]" captionTargetBox="[335,1045,264,1450]" captionTargetId="figure-0@11.[294,1059,257,1455]" captionTargetPageId="11" captionText="Figs 37–44. Cylindera (Ifasina) spp, aedeagus, left lateral view: 37–40 – C. macilenta (specimens from: 37 – Luzon, Nueva Vizcaya, Santa Fe; 38, 39 – Luzon, Nueva Vizcaya, Imugan; 40 – Luzon, Quezon, Mt. Banachao); 41–44 – C. klimenkoi sp. n.; 41 – holotype; 42–44 – paratypes; f – flagellum; mt – median tooth." figureDoi="http://doi.org/10.5281/zenodo.10945004" httpUri="https://zenodo.org/record/10945004/files/figure.png" pageId="13" pageNumber="14">Figs 41–44</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE73680FF80DF704E513411FD716E4E" blockId="13.[227,1127,264,1289]" pageId="13" pageNumber="14">A V-shaped posterior emargination of sternum VIII of female with 3–4 thick setae on each broadly rounded apex and 5 very small and thin setae along lateral margins. Syntergum 9 and10 weakly sclerotized, 1.25–1.3 times longer than wide, with 25–27 long lateral setae. Second gonapophyses strongly curved; ventral notch of second gonacoxa shallow, with a sparse row of very small and thin setae, the largest four being anterior ones. Bursa copulatrix ovoid, oviduct sclerite mediumsized.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342650BFF80DF704E51333CFDC86E2C" pageId="13" pageNumber="14" type="etymology">
|
||
<paragraph id="8BE73680FF80DF704E51333CFDC86E2C" blockId="13.[227,1127,264,1289]" pageId="13" pageNumber="14">
|
||
ETYMOLOGY. The new species is dedicated to Alexei Klimenko, a Russian entomologist and my late good friend, (1970–2017) who collected the
|
||
<typeStatus id="54E38822FF80DF704C80331CF8556E0F" box="[984,1029,1078,1102]" pageId="13" pageNumber="14">type</typeStatus>
|
||
series of this new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342650BFF80DF704E51335EFD686F48" pageId="13" pageNumber="14" type="distribution">
|
||
<paragraph id="8BE73680FF80DF704E51335EFF576EEA" blockId="13.[227,1127,264,1289]" pageId="13" pageNumber="14">
|
||
<materialsCitation id="3B303CDDFF80DF704E51335EFF536EEA" location="Luzon Island" municipality="Dingalan" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Aurora">
|
||
DISTRIBUTION. Known only from the
|
||
<typeStatus id="54E38822FF80DF704D9D335EFEA36ECD" box="[709,755,1140,1164]" pageId="13" pageNumber="14">type</typeStatus>
|
||
locality – environs of the town of
|
||
<collectingMunicipality id="6B83ACFAFF80DF704FBB33B9FD186EEA" box="[227,328,1171,1195]" pageId="13" pageNumber="14">Dingalan</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="499CF862FF80DF704E0A33B9FE566EEA" box="[338,518,1171,1195]" country="Philippines" name="Aurora" pageId="13" pageNumber="14">Aurora Province</collectingRegion>
|
||
,
|
||
<location id="8E87605BFF80DF704D4833BEFECC6EEA" LSID="urn:lsid:plazi:treatment:03F18796FF88DF704FBB3209FD686F48:8E87605BFF80DF704D4833BEFECC6EEA" box="[528,668,1171,1195]" municipality="Dingalan" name="Luzon Island" pageId="13" pageNumber="14" stateProvince="Aurora">Luzon Island</location>
|
||
(
|
||
<figureCitation id="13632A05FF80DF704DF333B9FEAB6EEA" box="[683,763,1171,1195]" captionStart="Fig" captionStartId="15.[265,300,1427,1449]" captionTargetBox="[370,983,502,1394]" captionTargetId="figure-78@15.[373,980,505,1390]" captionTargetPageId="15" captionText="Fig. 48. Distribution of Cylindera (Ifasina) spp. in the Philippines: red solid circle – C. macilenta; red half-solid circle – C. mouthiezi; red crossed-circle – C. klimenkoi sp. n.; red open circle – C. richouxi; orange solid triangle – C. foveolata; blue solid rhomb – C. semperi; blue half-solid rhomb – C. fabiocassolai; blue crossed-rhomb – C. balabacensis; blue open rhomb – C. rizali; blue asterisk – C. viduata; lilac solid quadrat – C. dilatotarsa; lilac open quadrat – C. discreta elaphroides. Map source: https:.en.wikipedia.org/wiki/Philippines#/ media/File:Relief_Map_Of_The_Philippines.png" figureDoi="http://doi.org/10.5281/zenodo.10945010" httpUri="https://zenodo.org/record/10945010/files/figure.png" pageId="13" pageNumber="14">Fig. 48</figureCitation>
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)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE73680FF80DF704E513399FD686F48" blockId="13.[227,1127,264,1289]" pageId="13" pageNumber="14">
|
||
<collectionCode id="ED49AE45FF80DF704E513399FD046E8A" box="[265,340,1203,1227]" pageId="13" pageNumber="14">NOTE</collectionCode>
|
||
. Due to a new species is similar with
|
||
<taxonomicName id="4C584D03FF80DF704DAC3399FF2E6E8B" authorityName="Schaum" authorityYear="1862" box="[756,894,1202,1226]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="macilenta">
|
||
<emphasis id="B92CEA92FF80DF704DAC3399FF2E6E8B" box="[756,894,1202,1226]" italics="true" pageId="13" pageNumber="14">C. macilenta</emphasis>
|
||
</taxonomicName>
|
||
the specimens of the later one should be re-examined for clarification of the distribution area of these species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |