341 lines
42 KiB
XML
341 lines
42 KiB
XML
<document id="48E3A76B117E6CF1A9B9F6A3FE9E3A47" ID-CLB-Dataset="20167" ID-DOI="10.5852/ejt.2022.803.1691" ID-GBIF-Dataset="af09c8e1-b481-4de0-b9d0-83ba26bf9876" ID-ISSN="2118-9773" ID-Zenodo-Dep="6359066" ID-ZooBank="8B66C8AE-F00A-42F6-9641-26B0ECC49F78" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1647350252755" checkinUser="felipe" docAuthor="Enghoff, Henrik" docDate="2022" docId="03F15C39D672066DFD8CFE55C05BFBD2" docLanguage="en" docName="ejt-803.1-136.pdf" docOrigin="European Journal of Taxonomy 803" docSource="http://dx.doi.org/10.5852/ejt.2022.803.1691" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Spinotarsus axeli Enghoff 2022, sp. nov." docType="treatment" docUuid="9B6978B5-2771-4984-AE66-A93A2C07E764" docUuidSource="ZooBank" docVersion="8" lastPageNumber="91" masterDocId="FFC82441D6250637FFF3FFE3C52CFFD8" masterDocTitle="Mountains of millipedes. The family Odontopygidae in the Eastern Arc Mountains of Tanzania (Diplopoda, Spirostreptida)" masterLastPageNumber="136" masterPageNumber="1" pageNumber="88" updateTime="1699318071475" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="70082CA83A69D9FEEFC1F9A4848A7B48">Mountains of millipedes. The family Odontopygidae in the Eastern Arc Mountains of Tanzania (Diplopoda, Spirostreptida)</mods:title>
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<mods:name id="F2537AF76E7474C813E29E32C0922E14" type="personal">
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<mods:roleTerm id="252ED8222AFFF5E5D692D45C57D5D8C0">Author</mods:roleTerm>
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<mods:namePart id="45998447C2C8DB1B48188B166D029BAD">Enghoff, Henrik</mods:namePart>
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<mods:affiliation id="B84657708DC55D38A569D2E71F308AB4">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen Ø, Denmark.</mods:affiliation>
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<mods:nameIdentifier id="753A7486439AF3240EE6D281044695FA" type="email">henghoff@snm.ku.dk</mods:nameIdentifier>
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<mods:typeOfResource id="F9BA5CDD254600B5F15543BCF1EF7B8F">text</mods:typeOfResource>
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<mods:title id="8A4EF2364DC5F57D2AA4903DE32D613F">European Journal of Taxonomy</mods:title>
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<mods:part id="252DC8DD691878E98CA2997E5FBD2EFA">
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<mods:date id="8DB64976559A06DD898D520499AC00F5">2022</mods:date>
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<mods:number id="D32C7ECAEEB15339E31463CEE3619A5A">2022-03-14</mods:number>
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<mods:number id="4AA9332CE53A9EE5CE0AA8A3C92F2F5B">803</mods:number>
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<mods:url id="FE42761EA733E7E697FA7684C89633A9">http://dx.doi.org/10.5852/ejt.2022.803.1691</mods:url>
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<mods:classification id="C50FE19C198BFC26C0B520CE8FE6B334">journal article</mods:classification>
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<mods:identifier id="50C444B7ECA0C70B08B8C5A5326656AF" type="CLB-Dataset">20167</mods:identifier>
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<mods:identifier id="5F2E48364E737BD5D28DFE2060EF4451" type="DOI">10.5852/ejt.2022.803.1691</mods:identifier>
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<treatment id="03F15C39D672066DFD8CFE55C05BFBD2" ID-DOI="http://doi.org/10.5281/zenodo.6373717" ID-GBIF-Taxon="194181618" ID-Zenodo-Dep="6373717" LSID="urn:lsid:zoobank.org:act:9B6978B5-2771-4984-AE66-A93A2C07E764" httpUri="http://treatment.plazi.org/id/03F15C39D672066DFD8CFE55C05BFBD2" lastPageId="90" lastPageNumber="91" pageId="87" pageNumber="88">
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<subSubSection id="C342BEA4D6720660FD8CFE55C699FE08" box="[639,949,437,464]" pageId="87" pageNumber="88" type="nomenclature">
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<paragraph id="8BE7ED2FD6720660FD8CFE55C699FE08" blockId="87.[372,1215,437,537]" box="[639,949,437,464]" pageId="87" pageNumber="88">
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<heading id="D0AF5A43D6720660FD8CFE55C699FE08" box="[639,949,437,464]" centered="true" fontSize="11" level="2" pageId="87" pageNumber="88" reason="2">
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<taxonomicName id="4C5896ACD6720660FD8CFE55C67FFE08" ID-CoL="98ZHS" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[639,851,437,464]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="axeli" status="sp. nov.">
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<emphasis id="B92C313DD6720660FD8CFE55C67FFE08" bold="true" box="[639,851,437,464]" italics="true" pageId="87" pageNumber="88">Spinotarsus axeli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21F8C46D6720660FCA9FE55C699FE08" box="[858,949,438,464]" pageId="87" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C342BEA4D6720660FE87FE39C6A2FDC1" pageId="87" pageNumber="88" type="description">
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<paragraph id="8BE7ED2FD6720660FE87FE39C193FE2D" blockId="87.[372,1215,437,537]" box="[372,1215,474,501]" pageId="87" pageNumber="88">
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<uri id="FFC9E12DD6720660FE87FE39C193FE2D" box="[372,1215,474,501]" pageId="87" pageNumber="88">
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urn:lsid:zoobank.org:act:
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<uuid id="FFFED7FAD6720660FD66FE39C193FE2D" box="[661,1215,474,501]" pageId="87" pageNumber="88">9B6978B5-2771-4984-AE66-A93A2C07E764</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BE7ED2FD6720660FD55FE1CC6A2FDC1" blockId="87.[372,1215,437,537]" box="[678,910,510,537]" pageId="87" pageNumber="88">
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<figureCitation id="1363F1AAD6720660FD55FE1CC629FDC1" box="[678,773,511,537]" captionStart="Fig" captionStartId="7.[189,232,1891,1917]" captionTargetBox="[189,1398,1308,1830]" captionTargetId="figure-396@7.[189,1398,1308,1830]" captionTargetPageId="7" captionText="Fig. 3. Heads of odontopygid males, arrows point to distal margin of mandibular stipes. A. Spinotarsus axelisp. nov.,holotype(NHMD621761),distalmarginof stipesconcave/bilobed,as inmostodontopygids. B. Callistodontopyge latifolia (Attems, 1914), specimen from Kiverenge FR (NHMD 621713), distal margin of stipes trilobed as characteristic of this genus. Scale bars=1 mm. Photo S.G. Selvantharan." figureDoi="http://doi.org/10.5281/zenodo.6359077" httpUri="https://zenodo.org/record/6359077/files/figure.png" pageId="87" pageNumber="88">Figs 3A</figureCitation>
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,
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<figureCitation id="1363F1AAD6720660FCE1FE1CC614FDC1" box="[786,824,511,537]" captionStart="Fig" captionStartId="9.[191,234,1714,1740]" captionTargetBox="[189,1398,504,1672]" captionTargetId="figure-242@9.[189,1398,504,1672]" captionTargetPageId="9" captionText="Fig. 4. Telsons of odontopygids. A. Spinotarsus axeli sp. nov., holotype, ♂ (NHMD 621761), telson and last six body rings, lateral view. B. Xystopyge bentemarieae sp. nov., paratype, ♂ (NHMD 621767), telson, lateral view, showing two setae per anal valve as characteristic of the genus Xystopyge. C–D. Aquattuor udzungwensis Enghoff, 2015, telson and last body ring (from Enghoff & Frederiksen 2015), showing three setae per anal valve as characteristic of most odontopygids; setae in this species placed on ravelins on raised valve margins. C. Lateral view, also showing the rectangular limbus flaps characteristic of the genus Aquattuor. D. Posterior view. Stars indicate the dorsal anal valve spines characteristic of family Odontopygidae; white arrows indicate anal valve setae. Scale bars: A=1 mm; B–D=0.1 mm. A–B: Photo S.G. Selvantharan." figureDoi="http://doi.org/10.5281/zenodo.6359081" httpUri="https://zenodo.org/record/6359081/files/figure.png" pageId="87" pageNumber="88">4A</figureCitation>
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,
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<figureCitation id="1363F1AAD6720660FCB7FE1DC6A2FDC1" box="[836,910,510,537]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="88.[189,232,1857,1883]" captionStartId-1="89.[189,232,1816,1842]" captionTargetBox-0="[189,1398,265,1833]" captionTargetBox-1="[189,1398,265,1763]" captionTargetId-0="figure-97@88.[189,1398,265,1833]" captionTargetId-1="figure-81@89.[189,1398,265,1763]" captionTargetPageId-0="88" captionTargetPageId-1="89" captionText-0="Fig. 54. Spinotarsus axeli sp. nov., holotype, ♂ (NHMD 621761). A–C. Left gonopod coxa. A. Anterior (-mesal) view. B. Posterior view. C. Mesal view. D. Midbody dorsal limbus. E–G. First pair of legs. E. Anterior view. F. Ventral view. G. Sublateral view. H. Sternum 9. Abbreviations: CXS=coxosternal seta; LCS= lateral coxal spine;MF =metaplical flange; ml =metaplical lobe;PPL =proplical lobe. Scale bars: A–C, E–F = 0.2 mm; G–H= 0.1 mm; D =0.01 mm." captionText-1="Fig. 55. Spinotarsus axeli sp. nov., holotype, ♂ (NHMD 621761), left gonopod telopodite. A. Anterior view, inset: tip of telomere. B.Posterior view. C. Venral view. D. Dorsal view. E. Anterior-dorso-apical view. F. Posterior-dorsal view. Abbreviations: bla= basal telomeral lamella; BSS =basal solenomeral spine; PN= post-torsal narrowing; PTS =post-torsal spine; SLM =solenomere; TM =telomere. Scale bars =0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6359235" figureDoi-1="http://doi.org/10.5281/zenodo.6359237" httpUri-0="https://zenodo.org/record/6359235/files/figure.png" httpUri-1="https://zenodo.org/record/6359237/files/figure.png" pageId="87" pageNumber="88">54–55</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C342BEA4D6720660FF4EFDA2C478FD77" pageId="87" pageNumber="88" type="diagnosis">
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||
<paragraph id="8BE7ED2FD6720660FF4EFDA2C41AFD83" blockId="87.[189,310,577,603]" box="[189,310,577,603]" pageId="87" pageNumber="88">
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||
<heading id="D0AF5A43D6720660FF4EFDA2C41AFD83" bold="true" box="[189,310,577,603]" fontSize="11" level="3" pageId="87" pageNumber="88" reason="3">
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||
<emphasis id="B92C313DD6720660FF4EFDA2C41AFD83" bold="true" box="[189,310,577,603]" pageId="87" pageNumber="88">Diagnosis</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BE7ED2FD6720660FF4EFD92C478FD77" blockId="87.[189,1398,625,687]" pageId="87" pageNumber="88">
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Differs from other species of
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<taxonomicName id="4C5896ACD6720660FDD0FD92C782FD53" authorityName="Attems" authorityYear="1909" box="[547,686,625,651]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
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<emphasis id="B92C313DD6720660FDD0FD92C782FD53" box="[547,686,625,651]" italics="true" pageId="87" pageNumber="88">Spinotarsus</emphasis>
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</taxonomicName>
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by the huge, slightly curved spine (
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<emphasis id="B92C313DD6720660FB90FD92C1BAFD53" box="[1123,1174,625,651]" italics="true" pageId="87" pageNumber="88">LCS</emphasis>
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) which is directed latero-basad.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C342BEA4D6720660FF4EFD3BC07DFCFA" box="[189,1361,728,802]" pageId="87" pageNumber="88" type="etymology">
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<paragraph id="8BE7ED2FD6720660FF4EFD3BC468FD2A" blockId="87.[189,324,728,754]" box="[189,324,728,754]" pageId="87" pageNumber="88">
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||
<heading id="D0AF5A43D6720660FF4EFD3BC468FD2A" bold="true" box="[189,324,728,754]" fontSize="11" level="3" pageId="87" pageNumber="88" reason="3">
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<emphasis id="B92C313DD6720660FF4EFD3BC468FD2A" bold="true" box="[189,324,728,754]" pageId="87" pageNumber="88">Etymology</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BE7ED2FD6720660FF4EFCEBC07DFCFA" blockId="87.[189,1361,776,802]" box="[189,1361,776,802]" pageId="87" pageNumber="88">After Axel Kristen Enghoff (1924–2014), the author’s father and ‘landlord’ of ‘Hotel Dolichoiulus’.</paragraph>
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</subSubSection>
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<subSubSection id="C342BEA4D6720660FF4EFCA9C11EFB9C" pageId="87" pageNumber="88" type="materials_examined">
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<paragraph id="8BE7ED2FD6720660FF4EFCA9C76FFCBD" blockId="87.[189,579,842,869]" box="[189,579,842,869]" pageId="87" pageNumber="88">
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||
<emphasis id="B92C313DD6720660FF4EFCA9C486FCBC" bold="true" box="[189,426,842,868]" pageId="87" pageNumber="88">Material examined</emphasis>
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(total
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<specimenCount id="9D5E26A6D6720660FE04FCA8C716FCBD" box="[503,570,843,869]" pageId="87" pageNumber="88" type="male">3 ♂♂</specimenCount>
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)
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</paragraph>
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<paragraph id="8BE7ED2FD6720660FF19FC99C161FC05" blockId="87.[189,1398,890,989]" pageId="87" pageNumber="88">
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||
<materialsCitation id="3B30E772D6720660FF19FC99C161FC05" ID-GBIF-Occurrence="3712026347" collectedFrom="submontane forest" collectingDate="2000-09" collectionCode="NHMD" collectorName="Uluguru Mountains Biodiversity Conservation Project & Nike Doggart" country="Tanzania" location="Shikurufumi FR" longitude="7.15" municipality="Mountains Biodiversity Conservation Project" pageId="87" pageNumber="88" specimenCode="NHMD 621761" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||
<emphasis id="B92C313DD6720660FF19FC99C470FC4C" bold="true" box="[234,348,890,916]" pageId="87" pageNumber="88">
|
||
<typeStatus id="54E3538DD6720660FF19FC99C470FC4C" box="[234,348,890,916]" pageId="87" pageNumber="88" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F34FADBFD6720660FF4EFC7CC47EFC61" box="[189,338,927,953]" name="Tanzania" pageId="87" pageNumber="88">TANZANIA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D5E26A6D6720660FE81FC7CC4A7FC61" box="[370,395,927,953]" pageId="87" pageNumber="88" type="male">♂</specimenCount>
|
||
;
|
||
<location id="8E87BBF4D6720660FE6FFC7DC74FFC61" LSID="urn:lsid:plazi:treatment:03F15C39D672066DFD8CFE55C05BFBD2:8E87BBF4D6720660FE6FFC7DC74FFC61" box="[412,611,926,953]" country="Tanzania" longitude="7.15" municipality="Mountains Biodiversity Conservation Project" name="Shikurufumi FR" pageId="87" pageNumber="88">Shikurufumi FR</location>
|
||
;
|
||
<geoCoordinate id="EE6C8BE8D6720660FD87FC7EC7CCFC61" box="[628,736,925,953]" degrees="37" direction="east" minutes="31" orientation="longitude" pageId="87" pageNumber="88" precision="925" value="37.516666">37°31′ E</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE6C8BE8D6720660FD02FC7EC67CFC61" box="[753,848,925,953]" degrees="7" direction="east" minutes="09" orientation="longitude" pageId="87" pageNumber="88" precision="925" value="7.15">7°09′ E</geoCoordinate>
|
||
;
|
||
<date id="FFE6CBEFD6720660FC92FC7CC6F2FC61" box="[865,990,927,953]" pageId="87" pageNumber="88" value="2000-09">
|
||
<collectingDate id="EFA23207D6720660FC92FC7CC6F2FC61" box="[865,990,927,953]" pageId="87" pageNumber="88" value="2000-09">Sep. 2000</collectingDate>
|
||
</date>
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||
;
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||
<collectorName id="26AD88F9D6720660FC1CFC7DC49BFC05" pageId="87" pageNumber="88">Uluguru Mountains Biodiversity Conservation Project</collectorName>
|
||
,
|
||
<collectorName id="26AD88F9D6720660FE30FC20C74FFC05" box="[451,611,963,989]" pageId="87" pageNumber="88">Nike Doggart</collectorName>
|
||
leg.;
|
||
<collectedFrom id="3844AED2D6720660FD56FC20C6ADFC05" box="[677,897,963,989]" pageId="87" pageNumber="88">submontane forest</collectedFrom>
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||
;
|
||
<specimenCode id="DBFE4554D6720660FC7FFC20C165FC05" box="[908,1097,963,989]" collectionCode="NHMD" pageId="87" pageNumber="88">NHMD 621761</specimenCode>
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||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BE7ED2FD6720660FF19FBE5C11EFB9C" blockId="87.[189,1074,1030,1092]" pageId="87" pageNumber="88">
|
||
<materialsCitation id="3B30E772D6720660FF19FBE5C102FB9C" ID-GBIF-Occurrence="3712026374" collectedFrom="submontane forest" collectingDate="2000-09" collectionCode="NHMD" collectorName="Shikurufumi FR & Nike Doggart" country="Tanzania" location="TANZANIA" longitude="7.15" municipality="Mountains Biodiversity Conservation Project" pageId="87" pageNumber="88" specimenCode="NHMD 621762" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
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||
<emphasis id="B92C313DD6720660FF19FBE5C44BFBF8" bold="true" box="[234,359,1030,1056]" pageId="87" pageNumber="88">
|
||
<typeStatus id="54E3538DD6720660FF19FBE5C44BFBF8" box="[234,359,1030,1056]" pageId="87" pageNumber="88" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F34FADBFD6720660FF4EFBC9C47EFB9C" box="[189,338,1066,1092]" name="Tanzania" pageId="87" pageNumber="88">TANZANIA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D5E26A6D6720660FE9AFBC9C482FB9C" box="[361,430,1066,1092]" pageId="87" pageNumber="88" type="male">2 ♂♂</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="DBFE4554D6720660FC82FBC9C102FB9C" box="[881,1070,1066,1092]" collectionCode="NHMD" pageId="87" pageNumber="88">NHMD 621762</specimenCode>
|
||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342BEA4D672066DFF4EFB8FC6BBFE0B" lastPageId="90" lastPageNumber="91" pageId="87" pageNumber="88" type="description">
|
||
<paragraph id="8BE7ED2FD6720660FF4EFB8FC48EFB5F" blockId="87.[189,418,1132,1159]" box="[189,418,1132,1159]" pageId="87" pageNumber="88">
|
||
<emphasis id="B92C313DD6720660FF4EFB8FC462FB5E" bold="true" box="[189,334,1132,1158]" pageId="87" pageNumber="88">Description</emphasis>
|
||
(male)
|
||
</paragraph>
|
||
<paragraph id="8BE7ED2FD6720660FF4EFB7FC05BFB6E" blockId="87.[189,1399,1180,1207]" box="[189,1399,1180,1207]" pageId="87" pageNumber="88">
|
||
SIZE. Length
|
||
<quantity id="4CA040CAD6720660FEA5FB7FC4FBFB6F" box="[342,471,1180,1207]" metricMagnitude="-2" metricUnit="m" metricValue="6.9" metricValueMax="7.0" metricValueMin="6.8" pageId="87" pageNumber="88" unit="mm" value="69.0" valueMax="70.0" valueMin="68.0">68–70 mm</quantity>
|
||
, diameter
|
||
<quantity id="4CA040CAD6720660FDBCFB7FC7F3FB6F" box="[591,735,1180,1207]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.1" metricValueMin="4.6" pageId="87" pageNumber="88" unit="mm" value="4.85" valueMax="5.1" valueMin="4.6">4.6–5.1 mm</quantity>
|
||
, 57–59 podous rings, no apodous rings in front of telson.
|
||
</paragraph>
|
||
<paragraph id="8BE7ED2FD6720660FF4EFB3CC7FFFB21" blockId="87.[189,723,1247,1273]" box="[189,723,1247,1273]" pageId="87" pageNumber="88">COLOUR. After 18 years in alcohol all whitish.</paragraph>
|
||
<paragraph id="8BE7ED2FD6720660FF4EFAC1C4CDFAE3" blockId="87.[189,481,1313,1340]" box="[189,481,1313,1340]" pageId="87" pageNumber="88">SUPRALABRAL SETAE. 6–7.</paragraph>
|
||
<paragraph id="8BE7ED2FD6720660FF4EFA86C7BDFAA7" blockId="87.[189,657,1380,1407]" box="[189,657,1380,1407]" pageId="87" pageNumber="88">MANDIBULAR STIPES. Shallowly concave.</paragraph>
|
||
<paragraph id="8BE7ED2FD6720660FF4EFA44C44DFA3D" blockId="87.[189,1399,1447,1510]" pageId="87" pageNumber="88">ANAL VALVES. With large dorsal spine and distinct ventral ‘corner’; margins barely raised, three sessile setae on each.</paragraph>
|
||
<paragraph id="8BE7ED2FD6720660FF4EF9EDC1C3F9F0" blockId="87.[189,1263,1550,1577]" box="[189,1263,1550,1577]" pageId="87" pageNumber="88">
|
||
LIMBUS (
|
||
<figureCitation id="1363F1AAD6720660FED1F9EDC4A7F9F0" box="[290,395,1550,1577]" captionStart="Fig" captionStartId="88.[189,232,1857,1883]" captionTargetBox="[189,1398,265,1833]" captionTargetId="figure-97@88.[189,1398,265,1833]" captionTargetPageId="88" captionText="Fig. 54. Spinotarsus axeli sp. nov., holotype, ♂ (NHMD 621761). A–C. Left gonopod coxa. A. Anterior (-mesal) view. B. Posterior view. C. Mesal view. D. Midbody dorsal limbus. E–G. First pair of legs. E. Anterior view. F. Ventral view. G. Sublateral view. H. Sternum 9. Abbreviations: CXS=coxosternal seta; LCS= lateral coxal spine;MF =metaplical flange; ml =metaplical lobe;PPL =proplical lobe. Scale bars: A–C, E–F = 0.2 mm; G–H= 0.1 mm; D =0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.6359235" httpUri="https://zenodo.org/record/6359235/files/figure.png" pageId="87" pageNumber="88">Fig. 54D</figureCitation>
|
||
). Margin with slender pointed-triangular lobes, ca twice as long as broad.
|
||
</paragraph>
|
||
<paragraph id="8BE7ED2FD6720660FF4EF9B2C477F957" blockId="87.[189,1398,1616,1679]" pageId="87" pageNumber="88">LEGS. Postfemoral and tibial pads present from leg-pair 6 backward, diminishing and eventually disappearing.</paragraph>
|
||
<paragraph id="8BE7ED2FD6720660FF4EF95BC74CF8C2" blockId="87.[189,1399,1719,1818]" pageId="87" pageNumber="88">
|
||
FIRST PAIR OF LEGS (
|
||
<figureCitation id="1363F1AAD6720660FE53F954C703F90A" box="[416,559,1719,1746]" captionStart="Fig" captionStartId="88.[189,232,1857,1883]" captionTargetBox="[189,1398,265,1833]" captionTargetId="figure-97@88.[189,1398,265,1833]" captionTargetPageId="88" captionText="Fig. 54. Spinotarsus axeli sp. nov., holotype, ♂ (NHMD 621761). A–C. Left gonopod coxa. A. Anterior (-mesal) view. B. Posterior view. C. Mesal view. D. Midbody dorsal limbus. E–G. First pair of legs. E. Anterior view. F. Ventral view. G. Sublateral view. H. Sternum 9. Abbreviations: CXS=coxosternal seta; LCS= lateral coxal spine;MF =metaplical flange; ml =metaplical lobe;PPL =proplical lobe. Scale bars: A–C, E–F = 0.2 mm; G–H= 0.1 mm; D =0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.6359235" httpUri="https://zenodo.org/record/6359235/files/figure.png" pageId="87" pageNumber="88">Fig. 54E–G</figureCitation>
|
||
). Prefemoral lobes broadly rounded-triangular in ventral view. A single coxosternal seta (
|
||
<emphasis id="B92C313DD6720660FE7DF93FC4EEF92E" box="[398,450,1756,1782]" italics="true" pageId="87" pageNumber="88">CXS</emphasis>
|
||
) close to lateral margin of coxosternum, well separated from prefemoral lobes. Prefemoral setae/sensilla indistinct.
|
||
</paragraph>
|
||
<paragraph id="8BE7ED2FD6720660FF4EF8A0C763F885" blockId="87.[189,591,1859,1886]" box="[189,591,1859,1886]" pageId="87" pageNumber="88">
|
||
STERNUM 9 (
|
||
<figureCitation id="1363F1AAD6720660FEBEF8A0C496F886" box="[333,442,1859,1886]" captionStart="Fig" captionStartId="88.[189,232,1857,1883]" captionTargetBox="[189,1398,265,1833]" captionTargetId="figure-97@88.[189,1398,265,1833]" captionTargetPageId="88" captionText="Fig. 54. Spinotarsus axeli sp. nov., holotype, ♂ (NHMD 621761). A–C. Left gonopod coxa. A. Anterior (-mesal) view. B. Posterior view. C. Mesal view. D. Midbody dorsal limbus. E–G. First pair of legs. E. Anterior view. F. Ventral view. G. Sublateral view. H. Sternum 9. Abbreviations: CXS=coxosternal seta; LCS= lateral coxal spine;MF =metaplical flange; ml =metaplical lobe;PPL =proplical lobe. Scale bars: A–C, E–F = 0.2 mm; G–H= 0.1 mm; D =0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.6359235" httpUri="https://zenodo.org/record/6359235/files/figure.png" pageId="87" pageNumber="88">Fig. 54H</figureCitation>
|
||
). Triangular.
|
||
</paragraph>
|
||
<paragraph id="8BE7ED2FD6720660FF4EF865C403F831" blockId="87.[189,1399,1925,2025]" pageId="87" pageNumber="88">
|
||
GONOPOD COXA (
|
||
<figureCitation id="1363F1AAD6720660FE77F866C734F878" box="[388,536,1925,1952]" captionStart="Fig" captionStartId="88.[189,232,1857,1883]" captionTargetBox="[189,1398,265,1833]" captionTargetId="figure-97@88.[189,1398,265,1833]" captionTargetPageId="88" captionText="Fig. 54. Spinotarsus axeli sp. nov., holotype, ♂ (NHMD 621761). A–C. Left gonopod coxa. A. Anterior (-mesal) view. B. Posterior view. C. Mesal view. D. Midbody dorsal limbus. E–G. First pair of legs. E. Anterior view. F. Ventral view. G. Sublateral view. H. Sternum 9. Abbreviations: CXS=coxosternal seta; LCS= lateral coxal spine;MF =metaplical flange; ml =metaplical lobe;PPL =proplical lobe. Scale bars: A–C, E–F = 0.2 mm; G–H= 0.1 mm; D =0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.6359235" httpUri="https://zenodo.org/record/6359235/files/figure.png" pageId="87" pageNumber="88">Fig. 54A–C</figureCitation>
|
||
). With a huge latero-basad, slightly curved spine (
|
||
<emphasis id="B92C313DD6720660FB73F865C19FF878" box="[1152,1203,1926,1952]" italics="true" pageId="87" pageNumber="88">LCS</emphasis>
|
||
). Proplical lobe (
|
||
<emphasis id="B92C313DD6720660FF34F848C5D7F81C" box="[199,251,1963,1988]" italics="true" pageId="87" pageNumber="88">PPL</emphasis>
|
||
) not covered. Metaplica with a large metaplical flange (
|
||
<emphasis id="B92C313DD6720660FC55F848C6FDF81C" box="[934,977,1963,1988]" italics="true" pageId="87" pageNumber="88">MF</emphasis>
|
||
) and more distally, a semi-circular lobe (
|
||
<emphasis id="B92C313DD6720660FEF2F82DC432F830" box="[257,286,1998,2024]" italics="true" pageId="87" pageNumber="88">ml</emphasis>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF27BDA7D67D066FFF4EF8A2C667F831" ID-DOI="http://doi.org/10.5281/zenodo.6359235" ID-Zenodo-Dep="6359235" httpUri="https://zenodo.org/record/6359235/files/figure.png" pageId="88" pageNumber="89" startId="88.[189,232,1857,1883]" targetBox="[189,1398,265,1833]" targetPageId="88">
|
||
<paragraph id="8BE7ED2FD67D066FFF4EF8A2C667F831" blockId="88.[185,1399,1857,2025]" pageId="88" pageNumber="89">
|
||
<emphasis id="B92C313DD67D066FFF4EF8A2C436F884" bold="true" box="[189,282,1857,1884]" pageId="88" pageNumber="89">Fig. 54.</emphasis>
|
||
<taxonomicName id="4C5896ACD67D066FFED3F8A1C4C0F884" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[288,492,1858,1884]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="axeli" status="sp. nov.">
|
||
<emphasis id="B92C313DD67D066FFED3F8A1C4C0F884" box="[288,492,1858,1884]" italics="true" pageId="88" pageNumber="89">Spinotarsus axeli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F8C46D67D066FFE01F8A1C762F884" box="[498,590,1858,1884]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (NHMD 621761).
|
||
<emphasis id="B92C313DD67D066FFC30F8A2C6D0F883" bold="true" box="[963,1020,1857,1883]" pageId="88" pageNumber="89">A–C</emphasis>
|
||
. Left gonopod coxa. A. Anterior (-mesal) view. B. Posterior view. C. Mesal view.
|
||
<emphasis id="B92C313DD67D066FFCE2F886C608F8A7" bold="true" box="[785,804,1893,1919]" pageId="88" pageNumber="89">D</emphasis>
|
||
. Midbody dorsal limbus.
|
||
<emphasis id="B92C313DD67D066FFBACF886C1BAF8A7" bold="true" box="[1119,1174,1893,1919]" pageId="88" pageNumber="89">E–G</emphasis>
|
||
. First pair of legs. E. Anterior view. F. Ventral view. G. Sublateral view.
|
||
<emphasis id="B92C313DD67D066FFCCEF86BC67FF87A" bold="true" box="[829,851,1928,1954]" pageId="88" pageNumber="89">H</emphasis>
|
||
. Sternum 9. Abbreviations:
|
||
<emphasis id="B92C313DD67D066FFB57F86AC1F4F87B" box="[1188,1240,1929,1955]" italics="true" pageId="88" pageNumber="89">CXS</emphasis>
|
||
=coxosternal seta;
|
||
<emphasis id="B92C313DD67D066FFF09F84FC401F81E" box="[250,301,1964,1990]" italics="true" pageId="88" pageNumber="89">LCS</emphasis>
|
||
= lateral coxal spine;
|
||
<emphasis id="B92C313DD67D066FFDD6F84FC77CF81D" box="[549,592,1964,1989]" italics="true" pageId="88" pageNumber="89">MF</emphasis>
|
||
=metaplical flange;
|
||
<emphasis id="B92C313DD67D066FFCB0F848C64CF81D" box="[835,864,1963,1989]" italics="true" pageId="88" pageNumber="89">ml</emphasis>
|
||
=metaplical lobe;
|
||
<emphasis id="B92C313DD67D066FFBC4F84FC147F81D" box="[1079,1131,1964,1989]" italics="true" pageId="88" pageNumber="89">PPL</emphasis>
|
||
=proplical lobe. Scale bars: A–C, E–F = 0.2 mm; G–H= 0.1 mm; D =0.01 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF27BDA7D67C066EFF4EF8FBC44BF867" ID-DOI="http://doi.org/10.5281/zenodo.6359237" ID-Zenodo-Dep="6359237" httpUri="https://zenodo.org/record/6359237/files/figure.png" pageId="89" pageNumber="90" startId="89.[189,232,1816,1842]" targetBox="[189,1398,265,1763]" targetPageId="89">
|
||
<paragraph id="8BE7ED2FD67C066EFF4EF8FBC44BF867" blockId="89.[185,1399,1816,1983]" pageId="89" pageNumber="90">
|
||
<emphasis id="B92C313DD67C066EFF4EF8FBC437F8EA" bold="true" box="[189,283,1816,1842]" pageId="89" pageNumber="90">Fig. 55.</emphasis>
|
||
<taxonomicName id="4C5896ACD67C066EFED1F8FBC4DCF8EA" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[290,496,1816,1842]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="axeli" status="sp. nov.">
|
||
<emphasis id="B92C313DD67C066EFED1F8FBC4DCF8EA" box="[290,496,1816,1842]" italics="true" pageId="89" pageNumber="90">Spinotarsus axeli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F8C46D67C066EFE04F8FBC779F8EA" box="[503,597,1816,1842]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (NHMD 621761), left gonopod telopodite.
|
||
<emphasis id="B92C313DD67C066EFB03F8FBC029F8EA" bold="true" box="[1264,1285,1816,1842]" pageId="89" pageNumber="90">A</emphasis>
|
||
. Anterior view, inset: tip of telomere.
|
||
<emphasis id="B92C313DD67C066EFDF4F8D8C736F88D" bold="true" box="[519,538,1851,1877]" pageId="89" pageNumber="90">B</emphasis>
|
||
.Posterior view.
|
||
<emphasis id="B92C313DD67C066EFD29F8D8C7C3F88D" bold="true" box="[730,751,1851,1877]" pageId="89" pageNumber="90">C</emphasis>
|
||
. Venral view.
|
||
<emphasis id="B92C313DD67C066EFC64F8D8C680F88D" bold="true" box="[919,940,1851,1877]" pageId="89" pageNumber="90">D</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="B92C313DD67C066EFBABF8D8C147F88D" bold="true" box="[1112,1131,1851,1877]" pageId="89" pageNumber="90">E</emphasis>
|
||
. Anterior-dorso-apical view.
|
||
<emphasis id="B92C313DD67C066EFEF5F8BDC434F8A0" bold="true" box="[262,280,1886,1912]" pageId="89" pageNumber="90">F</emphasis>
|
||
. Posterior-dorsal view. Abbreviations:
|
||
<emphasis id="B92C313DD67C066EFD02F8BDC63BF8A0" box="[753,791,1886,1912]" italics="true" pageId="89" pageNumber="90">bla</emphasis>
|
||
= basal telomeral lamella;
|
||
<emphasis id="B92C313DD67C066EFBA6F8BCC1A9F8A1" box="[1109,1157,1887,1913]" italics="true" pageId="89" pageNumber="90">BSS</emphasis>
|
||
=basal solenomeral spine;
|
||
<emphasis id="B92C313DD67C066EFEE2F861C41BF843" box="[273,311,1922,1947]" italics="true" pageId="89" pageNumber="90">PN</emphasis>
|
||
= post-torsal narrowing;
|
||
<emphasis id="B92C313DD67C066EFD90F861C7B8F844" box="[611,660,1922,1948]" italics="true" pageId="89" pageNumber="90">PTS</emphasis>
|
||
=post-torsal spine;
|
||
<emphasis id="B92C313DD67C066EFC79F861C6EEF844" box="[906,962,1922,1948]" italics="true" pageId="89" pageNumber="90">SLM</emphasis>
|
||
=solenomere;
|
||
<emphasis id="B92C313DD67C066EFB84F861C18CF843" box="[1143,1184,1922,1947]" italics="true" pageId="89" pageNumber="90">TM</emphasis>
|
||
=telomere. Scale bars =0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE7ED2FD67F066DFF4EFEEAC6BBFE0B" blockId="90.[189,1399,263,467]" pageId="90" pageNumber="91">
|
||
GONOPOD TELOPODITE (
|
||
<figureCitation id="1363F1AAD67F066DFE30FEEAC737FEFC" box="[451,539,265,292]" captionStart="Fig" captionStartId="89.[189,232,1816,1842]" captionTargetBox="[189,1398,265,1763]" captionTargetId="figure-81@89.[189,1398,265,1763]" captionTargetPageId="89" captionText="Fig. 55. Spinotarsus axeli sp. nov., holotype, ♂ (NHMD 621761), left gonopod telopodite. A. Anterior view, inset: tip of telomere. B.Posterior view. C. Venral view. D. Dorsal view. E. Anterior-dorso-apical view. F. Posterior-dorsal view. Abbreviations: bla= basal telomeral lamella; BSS =basal solenomeral spine; PN= post-torsal narrowing; PTS =post-torsal spine; SLM =solenomere; TM =telomere. Scale bars =0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6359237" httpUri="https://zenodo.org/record/6359237/files/figure.png" pageId="90" pageNumber="91">Fig. 55</figureCitation>
|
||
). Arculus 90°. Torsotope compact, almost completely hidden by 1½ coil of very long post-torsal spine (
|
||
<emphasis id="B92C313DD67F066DFDDFFECEC771FE9F" box="[556,605,301,327]" italics="true" pageId="90" pageNumber="91">PTS</emphasis>
|
||
). Post-torsal narrowing (
|
||
<emphasis id="B92C313DD67F066DFC74FECEC681FE9E" box="[903,941,301,326]" italics="true" pageId="90" pageNumber="91">PN</emphasis>
|
||
) very pronounced. Solenomere (
|
||
<emphasis id="B92C313DD67F066DFAC6FECEC041FE9F" box="[1333,1389,301,327]" italics="true" pageId="90" pageNumber="91">SLM</emphasis>
|
||
) long, ca same length as telomere, simple, whiplike, coiling irregularly, without outgrowths, with a long straight spine (
|
||
<emphasis id="B92C313DD67F066DFE9FFE90C4B0FE55" box="[364,412,371,397]" italics="true" pageId="90" pageNumber="91">BSS</emphasis>
|
||
) at base. Telomere (
|
||
<emphasis id="B92C313DD67F066DFD7EFE90C79AFE54" box="[653,694,371,396]" italics="true" pageId="90" pageNumber="91">TM</emphasis>
|
||
) with a rounded-rectangular, inconspicuously spinose basal lamella (
|
||
<emphasis id="B92C313DD67F066DFED5FE76C460FE77" box="[294,332,405,431]" italics="true" pageId="90" pageNumber="91">bla</emphasis>
|
||
), its distal part a broad, longitudinally folded sheet bent into U-shape, narrowing towards tip, one margin near tip with coarse denticles (
|
||
<figureCitation id="1363F1AAD67F066DFD2DFE5BC666FE0A" box="[734,842,439,466]" captionStart="Fig" captionStartId="89.[189,232,1816,1842]" captionTargetBox="[189,1398,265,1763]" captionTargetId="figure-81@89.[189,1398,265,1763]" captionTargetPageId="89" captionText="Fig. 55. Spinotarsus axeli sp. nov., holotype, ♂ (NHMD 621761), left gonopod telopodite. A. Anterior view, inset: tip of telomere. B.Posterior view. C. Venral view. D. Dorsal view. E. Anterior-dorso-apical view. F. Posterior-dorsal view. Abbreviations: bla= basal telomeral lamella; BSS =basal solenomeral spine; PN= post-torsal narrowing; PTS =post-torsal spine; SLM =solenomere; TM =telomere. Scale bars =0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6359237" httpUri="https://zenodo.org/record/6359237/files/figure.png" pageId="90" pageNumber="91">Fig. 55A</figureCitation>
|
||
, inset)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342BEA4D67F066DFF4EFE1BC1B0FD99" pageId="90" pageNumber="91" type="distribution">
|
||
<paragraph id="8BE7ED2FD67F066DFF4EFE1BC4C2FDCA" blockId="90.[189,494,504,531]" box="[189,494,504,531]" pageId="90" pageNumber="91">
|
||
<heading id="D0AF5A43D67F066DFF4EFE1BC4C2FDCA" bold="true" box="[189,494,504,531]" fontSize="11" level="3" pageId="90" pageNumber="91" reason="3">
|
||
<emphasis id="B92C313DD67F066DFF4EFE1BC4C2FDCA" bold="true" box="[189,494,504,531]" pageId="90" pageNumber="91">Distribution and habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE7ED2FD67F066DFF4EFDC4C1B0FD99" blockId="90.[189,1180,550,577]" box="[189,1180,550,577]" pageId="90" pageNumber="91">
|
||
Known only from the Shikurufumi FR in the Uluguru Mts, cf.
|
||
<bibRefCitation id="EFC990DED67F066DFC6EFDC4C1B4FD99" author="Doggart N. & Lovett J. & Mhoro B. & Kiure J. & Burgess N." box="[925,1176,550,577]" pageId="90" pageNumber="91" refId="ref47101" refString="Doggart N., Lovett J., Mhoro B., Kiure J. & Burgess N. 2004. Biodiversity Surveys in the Forest Reserves of the Uluguru Mountains. Part I: An Overview of the Biodiversity of the Uluguru Mountains. The Wildlife Conservation Society of Tanzania." type="book" year="2004">
|
||
Doggart
|
||
<emphasis id="B92C313DD67F066DFBF7FDC4C113FD98" box="[1028,1087,550,576]" italics="true" pageId="90" pageNumber="91">et al.</emphasis>
|
||
(2004)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C342BEA4D67F066DFF4EFD84C05BFBD2" pageId="90" pageNumber="91" type="discussion">
|
||
<paragraph id="8BE7ED2FD67F066DFF4EFD84C403FD59" blockId="90.[189,303,615,641]" box="[189,303,615,641]" pageId="90" pageNumber="91">
|
||
<heading id="D0AF5A43D67F066DFF4EFD84C403FD59" bold="true" box="[189,303,615,641]" fontSize="11" level="3" pageId="90" pageNumber="91" reason="3">
|
||
<emphasis id="B92C313DD67F066DFF4EFD84C403FD59" bold="true" box="[189,303,615,641]" pageId="90" pageNumber="91">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE7ED2FD67F066DFF4EFD76C05BFBD2" blockId="90.[187,1400,661,1034]" pageId="90" pageNumber="91">
|
||
The basal lamella (
|
||
<emphasis id="B92C313DD67F066DFE64FD76C490FD77" box="[407,444,661,687]" italics="true" pageId="90" pageNumber="91">bla</emphasis>
|
||
) which is very inconspicuously spinose (spinules not visible on
|
||
<figureCitation id="1363F1AAD67F066DFB6CFD76C028FD68" box="[1183,1284,661,688]" captionStart="Fig" captionStartId="89.[189,232,1816,1842]" captionTargetBox="[189,1398,265,1763]" captionTargetId="figure-81@89.[189,1398,265,1763]" captionTargetPageId="89" captionText="Fig. 55. Spinotarsus axeli sp. nov., holotype, ♂ (NHMD 621761), left gonopod telopodite. A. Anterior view, inset: tip of telomere. B.Posterior view. C. Venral view. D. Dorsal view. E. Anterior-dorso-apical view. F. Posterior-dorsal view. Abbreviations: bla= basal telomeral lamella; BSS =basal solenomeral spine; PN= post-torsal narrowing; PTS =post-torsal spine; SLM =solenomere; TM =telomere. Scale bars =0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6359237" httpUri="https://zenodo.org/record/6359237/files/figure.png" pageId="90" pageNumber="91">Fig. 55B</figureCitation>
|
||
) suggests placement in the large genus
|
||
<taxonomicName id="4C5896ACD67F066DFDFAFD5BC7B8FD0A" authorityName="Attems" authorityYear="1909" box="[521,660,696,722]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92C313DD67F066DFDFAFD5BC7B8FD0A" box="[521,660,696,722]" italics="true" pageId="90" pageNumber="91">Spinotarsus</emphasis>
|
||
</taxonomicName>
|
||
, and no other characters are in conflict with this allocation. Like the only other species of
|
||
<taxonomicName id="4C5896ACD67F066DFE17FD38C743FD2D" authorityName="Attems" authorityYear="1909" box="[484,623,731,757]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92C313DD67F066DFE17FD38C743FD2D" box="[484,623,731,757]" italics="true" pageId="90" pageNumber="91">Spinotarsus</emphasis>
|
||
</taxonomicName>
|
||
known from the Eastern Arc Mts,
|
||
<taxonomicName id="4C5896ACD67F066DFBF5FD38C05EFD2D" authority="Enghoff, 2018" authorityName="Enghoff" authorityYear="2018" box="[1030,1394,730,757]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="fortehamatus">
|
||
<emphasis id="B92C313DD67F066DFBF5FD38C193FD2C" box="[1030,1215,730,757]" italics="true" pageId="90" pageNumber="91">S. fortehamatus</emphasis>
|
||
Enghoff, 2018
|
||
</taxonomicName>
|
||
, the new species lacks a darkly sclerotized ridge on the posterior surface of the telomere, a character found in very many congeners. In the key to
|
||
<taxonomicName id="4C5896ACD67F066DFD3AFCC3C69CFCE2" box="[713,944,800,826]" pageId="90" pageNumber="91">
|
||
<emphasis id="B92C313DD67F066DFD3AFCC3C678FCE2" box="[713,852,800,826]" italics="true" pageId="90" pageNumber="91">Spinotarsus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFC990DED67F066DFC2FFCC3C156FCE3" author="Kraus O." box="[988,1146,800,827]" pageId="90" pageNumber="91" pagination="1 - 143" refId="ref48892" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." type="journal article" year="1966">Kraus (1966)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C5896ACD67F066DFB76FCC3C1F7FCE2" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[1157,1243,800,826]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="axeli" status="sp. nov.">
|
||
<emphasis id="B92C313DD67F066DFB76FCC3C1F7FCE2" box="[1157,1243,800,826]" italics="true" pageId="90" pageNumber="91">S. axeli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F8C46D67F066DFB10FCC3C011FCE2" box="[1251,1341,800,826]" pageId="90" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
runs without problems to couplet 60, as follows. Couplet 1: Gonopodal metaplica with
|
||
<emphasis id="B92C313DD67F066DFB61FCA1C192FC84" bold="true" box="[1170,1214,834,860]" pageId="90" pageNumber="91">one</emphasis>
|
||
or several dark sclerotized lateral spines which stand clearly out in strict oral view → couplet 38:
|
||
<emphasis id="B92C313DD67F066DFB8FFC86C19CFCA7" bold="true" box="[1148,1200,869,895]" pageId="90" pageNumber="91">One</emphasis>
|
||
or more spine(s) (“Postfemoraldorn”) at base of solenomere present → couplet 59:
|
||
<emphasis id="B92C313DD67F066DFC3DFC64C12EFC79" bold="true" box="[974,1026,903,929]" pageId="90" pageNumber="91">One</emphasis>
|
||
spine at base of solenomere → couplet 60. Couplet 60 (cf. Enghoff 2018) leads to couplet 65 where the choice is between the lateral metaplical spine being directed more or less apicad, or being horizontal (strictly laterad); however, in
|
||
<taxonomicName id="4C5896ACD67F066DFF4CFC13C438FBD1" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[191,276,1007,1034]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="axeli" status="sp. nov.">
|
||
<emphasis id="B92C313DD67F066DFF4CFC13C438FBD1" box="[191,276,1007,1034]" italics="true" pageId="90" pageNumber="91">S. axeli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F8C46D67F066DFEEBFC13C45EFBD2" box="[280,370,1008,1034]" pageId="90" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
the lateral spine is directed obliquely laterobasad which seems to be unique in the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |