239 lines
31 KiB
XML
239 lines
31 KiB
XML
<document id="61599038A82BA8F1E869BCCB9B6D46C6" ID-CLB-Dataset="26032" ID-DOI="10.11646/zootaxa.4656.1.5" ID-GBIF-Dataset="cfb36b3b-46e0-4cfe-83fd-4b1f09d29fb7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3366776" ID-ZooBank="884B53B1-3EC1-4258-A4E6-1928F167632C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565681563246" checkinUser="plazi" docAuthor="Schlüter, Nils, Taherpour-Khalil-Abad, Morteza, Majidifard, Mahmoudreza, Hassanzadeh, Zinat & Taheri, Jafar" docDate="2019" docId="03EF87BA542C3E365595FD14FDAFD925" docLanguage="en" docName="zootaxa.4656.1.5.pdf" docOrigin="Zootaxa 4656 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tetragramma L. Agassiz 1838" docType="treatment" docVersion="10" lastPageNumber="125" masterDocId="FFD6FFC254283E335502FFDBFF91D866" masterDocTitle="Two echinoid species from the early Aptian (Early Cretaceous) of the Kopet-Dagh Basin, NE Iran" masterLastPageNumber="132" masterPageNumber="121" pageNumber="125" updateTime="1698750843938" updateUser="ExternalLinkService">
|
|
<mods:mods id="8783A199EF6A873D4E6EDDC90A06A9E0" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="65E4DA64A22414FD2D5734E0F865F8B6">
|
|
<mods:title id="52B9CA07765C606F86F117281B72FF8D">Two echinoid species from the early Aptian (Early Cretaceous) of the Kopet-Dagh Basin, NE Iran</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="D6A7A5951F9A9BF73BB7A8B8827F57B4" type="personal">
|
|
<mods:role id="BA3C9A3E13EF8CB4EA763896DE53A05F">
|
|
<mods:roleTerm id="D85C6A0BA93AD2559F647EAEE7A2A0C0">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="AE7B752CE7269F929251DED1817C247B">Schlüter, Nils</mods:namePart>
|
|
</mods:name>
|
|
<mods:name id="FC77E5B92436AA4F4955B1946ACE69B9" type="personal">
|
|
<mods:role id="6737DBE427A21CDFEACB0051E933DE65">
|
|
<mods:roleTerm id="9A5768FD54E35C056684D5B1169F5843">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="D68CDFD85910878C6368407C62B3E9F7">Taherpour-Khalil-Abad, Morteza</mods:namePart>
|
|
</mods:name>
|
|
<mods:name id="33918159ED8DB5A58C196A9367C2F52F" type="personal">
|
|
<mods:role id="1DBED45486980B61EA30B2596AD32078">
|
|
<mods:roleTerm id="C09207F02FBBFEF47D2788E7BD1F6585">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="20E898416B3642971F1CF3CEBCEC82D9">Majidifard, Mahmoudreza</mods:namePart>
|
|
</mods:name>
|
|
<mods:name id="A28F0CCDA4E0113916F5F55462B2BA30" type="personal">
|
|
<mods:role id="96771127AC0FD8546C8BCA591EEFEB11">
|
|
<mods:roleTerm id="37CA57E4B38D849230EAE15D4C1FC755">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="2F78B07889B2091671D2AA559FB015A8">Hassanzadeh, Zinat</mods:namePart>
|
|
</mods:name>
|
|
<mods:name id="4F3178E1160E13A57CFFE341D3F5B442" type="personal">
|
|
<mods:role id="B5C21C1AAD5741EF6CDFD1BB3C625459">
|
|
<mods:roleTerm id="F84D9EB7777B96BCB5CCEDE514A4B50E">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="3CE95339F7075FBB8FEADF73224B7AE0">Taheri, Jafar</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource id="6C7FAFECBC443740A72C51BEB02C9ED4">text</mods:typeOfResource>
|
|
<mods:relatedItem id="C30FE85C71C04F277044101801AAD6EC" type="host">
|
|
<mods:titleInfo id="0E9E5CB9BB26BC9DDE6A0FC41B7F4796">
|
|
<mods:title id="DE8329BCD28A0EA6FB34895426D04430">Zootaxa</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="BDDB23DC6BA113820F0D075FC28FE2C4">
|
|
<mods:date id="553950E32781534E9DE65CAD4E73AD9D">2019</mods:date>
|
|
<mods:detail id="F906833DCED870907936C8669A1DCB63" type="pubDate">
|
|
<mods:number id="BB0D12899BCF51888D732AE26A288CBF">2019-08-13</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="8F1ECD4B17E7AF63DE891216F4316CDE" type="volume">
|
|
<mods:number id="73E327BD75A16ABA5BFA90C9CA223839">4656</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="DA365C28D8B2CF03D2B64811AC4D3B78" type="issue">
|
|
<mods:number id="B69AA949B30DC0C347CFC6B16118126C">1</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="C1B31B91381FACBD749266660050ED60" unit="page">
|
|
<mods:start id="6DCD5661EAE6F065F1BA1B441763E942">121</mods:start>
|
|
<mods:end id="00D9A28976D05E57DCBB3D95AB0E656E">132</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:classification id="136E1F96568F7390675232F5624F1813">journal article</mods:classification>
|
|
<mods:identifier id="6EAF55D44745CCB0CEE3FF99159C007D" type="CLB-Dataset">26032</mods:identifier>
|
|
<mods:identifier id="E4328F0429B96C8EAE5F34EC94A83CBA" type="DOI">10.11646/zootaxa.4656.1.5</mods:identifier>
|
|
<mods:identifier id="44354C24BE796FC37A9A66FEB4523670" type="GBIF-Dataset">cfb36b3b-46e0-4cfe-83fd-4b1f09d29fb7</mods:identifier>
|
|
<mods:identifier id="07D2E6D19D19DB1B3E614E759D6D90CA" type="ISSN">1175-5326</mods:identifier>
|
|
<mods:identifier id="9DEFED1DE1F34D0558BC00ECE480BFDA" type="Zenodo-Dep">3366776</mods:identifier>
|
|
<mods:identifier id="8B200EE751137C5A151B35F929C137E0" type="ZooBank">884B53B1-3EC1-4258-A4E6-1928F167632C</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="03EF87BA542C3E365595FD14FDAFD925" ID-GBIF-Taxon="159126035" LSID="urn:lsid:plazi:treatment:03EF87BA542C3E365595FD14FDAFD925" httpUri="http://treatment.plazi.org/id/03EF87BA542C3E365595FD14FDAFD925" lastPageId="5" lastPageNumber="125" pageId="4" pageNumber="125">
|
|
<subSubSection id="C35C6527542C3E375595FD14FEF4DA8F" box="[151,357,719,745]" pageId="4" pageNumber="125" type="nomenclature">
|
|
<paragraph id="8BF936AC542C3E375595FD14FEF4DA8F" blockId="4.[151,357,719,781]" box="[151,357,719,745]" pageId="4" pageNumber="125">
|
|
<heading id="D0B181C0542C3E375595FD14FEF4DA8F" bold="true" box="[151,357,719,745]" fontSize="11" level="1" pageId="4" pageNumber="125" reason="1">
|
|
<emphasis id="B932EABE542C3E375595FD14FEF4DA8F" bold="true" box="[151,357,719,745]" pageId="4" pageNumber="125">
|
|
<taxonomicName id="4C464D2F542C3E375595FD14FEAADA8F" ID-CoL="7Q44S" authorityName="L. Agassiz" authorityYear="1838" box="[151,315,719,745]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="genus">
|
|
<emphasis id="B932EABE542C3E375595FD14FEAADA8F" bold="true" box="[151,315,719,745]" italics="true" pageId="4" pageNumber="125">Tetragramma</emphasis>
|
|
</taxonomicName>
|
|
sp.
|
|
</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C35C6527542C3E375595FD28FF7ADB6B" box="[151,235,754,781]" pageId="4" pageNumber="125" type="description">
|
|
<paragraph id="8BF936AC542C3E375595FD28FF7ADB6B" blockId="4.[151,357,719,781]" box="[151,235,754,781]" pageId="4" pageNumber="125">
|
|
(
|
|
<figureCitation id="137D2A29542C3E3755A2FD28FF73DB6B" box="[160,226,754,781]" captionStart="FIGURE 5" captionStartId="5.[151,250,1278,1303]" captionTargetBox="[151,1436,345,1258]" captionTargetId="figure@5.[151,1436,345,1258]" captionTargetPageId="5" captionText="FIGURE 5. Tetragramma sp., GSINET97FE248, from the vicinity of the village Gelian, NE Iran, (Sarcheshmeh Formation, early Aptian), in apical (A) and oral (B) view, C: detail of the aboral interambulacral plates, D: detail of the perperistomal am- bulacra, E: detail of the adapical ambulacra." httpUri="https://zenodo.org/record/3366784/files/figure.png" pageId="4" pageNumber="125">Fig. 5</figureCitation>
|
|
)
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C35C6527542C3E365595FCE1FDAFD925" lastPageId="5" lastPageNumber="126" pageId="4" pageNumber="125" type="materials_examined">
|
|
<paragraph id="8BF936AC542C3E375595FCE1FC33DB1F" blockId="4.[151,1437,826,2041]" pageId="4" pageNumber="125">
|
|
<emphasis id="B932EABE542C3E375595FCE1FE97DB33" bold="true" box="[151,262,826,853]" pageId="4" pageNumber="125">Material.</emphasis>
|
|
A single worn and incomplete specimen from the vicinity of the village Gelian, NE
|
|
<collectingCountry id="F351763C542C3E3751C0FCE0FB63DB33" box="[1218,1266,827,853]" name="Iran" pageId="4" pageNumber="125">Iran</collectingCountry>
|
|
, (Sarcheshmeh Formation, early Aptian) was available for study (
|
|
<collectionCode id="ED57AE69542C3E3757C7FC84FCBCDB1F" box="[709,813,863,889]" pageId="4" pageNumber="125">GSINET</collectionCode>
|
|
97FE248).
|
|
</paragraph>
|
|
<paragraph id="8BF936AC542C3E3755C5FC59FAECDB83" blockId="4.[151,1437,826,2041]" pageId="4" pageNumber="125">
|
|
<emphasis id="B932EABE542C3E3755C5FC59FEC8DBFB" bold="true" box="[199,345,898,925]" pageId="4" pageNumber="125">Description.</emphasis>
|
|
The slightly distorted individual shows a complete test only above the ambitus without apical disc; most of the oral side is not preserved. The specimen is medium-sized (approximately
|
|
<quantity id="4CBE9B49542C3E37515BFC7DFB39DBA7" box="[1113,1192,934,961]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="4" pageNumber="125" unit="mm" value="52.0">52 mm</quantity>
|
|
in diameter) and flattened (approximately
|
|
<quantity id="4CBE9B49542C3E37548FFC10FE4DDB83" box="[397,476,971,997]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="125" unit="mm" value="22.0">22 mm</quantity>
|
|
in height). The ambitus is at mid-height. The tubercles are perforate and crenulate.
|
|
</paragraph>
|
|
<paragraph id="8BF936AC542C3E3755C5FC34FF6EDC37" blockId="4.[151,1437,826,2041]" pageId="4" pageNumber="125">The peristome is deeply invaginated as it can be seen by the remaining perioral test. The perioral plates are highly abraded; however, distinct buccal notches can be observed. The diameter is approximately 31% of the test diameter.</paragraph>
|
|
<paragraph id="8BF936AC542C3E3755C5FB80FB30DCFF" blockId="4.[151,1437,826,2041]" pageId="4" pageNumber="125">
|
|
The apical opening is large (the longest axis spans
|
|
<emphasis id="B932EABE542C3E3757F4FB87FC98DC13" box="[758,777,1116,1141]" italics="true" pageId="4" pageNumber="125">c.</emphasis>
|
|
38% of the length of the test diameter). Based on the somewhat elongated pentagonal outline, a monocyclic structure of the apical disc can be inferred.
|
|
</paragraph>
|
|
<paragraph id="8BF936AC542C3E3755C5FB78FCA5DD4F" blockId="4.[151,1437,826,2041]" pageId="4" pageNumber="125">The ambulacra taper towards the apical opening and short phyllodes can be observed near the peristome. Adorally, pore pairs run uniserial towards the ambitus, from which they are biserially arranged to the apex. Ambulacral plates carry five pore pairs per plate ambitally and each ambital plate has a single large primary tubercle. Details in plate compounding are not visible due to poor preservation.</paragraph>
|
|
<paragraph id="8BF936AC542C3E3755C5FAE8FD98DDBB" blockId="4.[151,1437,826,2041]" pageId="4" pageNumber="125">The interambulacral plates are much wider than they are tall. Ambitally, four subequal primary tubercles per plate are present. Adapically, a naked V-shaped and slightly depressed zone is present at the interradius, reaching the ambitus. Adapically from the ambitus, the primary tubercles at the adradius are replaced by smaller, vertically aligned secondary tubercles. There are more than 15 interambulacral plates in a single row, estimated by the imprints of the plates on the mould.</paragraph>
|
|
<paragraph id="8BF936AC542C3E3655C5FA3DFDAFD925" blockId="4.[151,1437,826,2041]" lastBlockId="5.[151,1437,153,323]" lastPageId="5" lastPageNumber="126" pageId="4" pageNumber="125">
|
|
<emphasis id="B932EABE542C3E3755C5FA3DFEDDDE67" bold="true" box="[199,332,1510,1537]" pageId="4" pageNumber="125">Discussion.</emphasis>
|
|
Features of this specimen (e.g. four subequal primary tubercles, shape of interambulacral plates, pore pairs arrangement: biserial adapically, short phyllodes) are characteristic of the genus
|
|
<taxonomicName id="4C464D2F542C3E375189F9D7FAB3DE43" authorityName="L. Agassiz" authorityYear="1838" box="[1163,1314,1548,1573]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="genus">
|
|
<emphasis id="B932EABE542C3E375189F9D7FAB3DE43" box="[1163,1314,1548,1573]" italics="true" pageId="4" pageNumber="125">Tetragramma</emphasis>
|
|
</taxonomicName>
|
|
. However, due to the lack of better preserved material, the nomenclature of the here described specimen must remain open. This specimen resembles, in the number and size of primary tubercles in the ambital interambulacra and the invagination of the peristome, the Aptian species
|
|
<taxonomicName id="4C464D2F542C3E37577AF9A3FBEADEF7" authority="Agassiz & Desor, 1846" authorityName="Agassiz & Desor" authorityYear="1846" box="[632,1147,1655,1681]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="species" species="malbosi">
|
|
<emphasis id="B932EABE542C3E37577AF9A3FCFEDEF7" box="[632,879,1655,1681]" italics="true" pageId="4" pageNumber="125">Tetragramma malbosi</emphasis>
|
|
<bibRefCitation id="EFD74B5D542C3E375677F9ACFBEADEF7" author="Agassiz, L. & Desor, E." box="[885,1147,1655,1681]" pageId="4" pageNumber="125" pagination="305 - 374" refId="ref4348" refString="Agassiz, L. & Desor, E. (1846) Catalogue raisonne des especes, des genres, et familles d'echinides. Annales des Sciences Naturelles, Troisieme Serie, Zoologie, 6, 305 - 374." type="journal article" year="1846">Agassiz & Desor, 1846</bibRefCitation>
|
|
</taxonomicName>
|
|
.
|
|
<taxonomicName id="4C464D2F542C3E375187F9A3FE5EDED3" authority="Vaziri & Arab, 2013" authorityName="Vaziri & Arab" authorityYear="2013" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="species" species="tetratuberculatus">
|
|
<emphasis id="B932EABE542C3E375187F9A3FF79DED3" italics="true" pageId="4" pageNumber="125">Tetragramma tetratuberculatus</emphasis>
|
|
<bibRefCitation id="EFD74B5D542C3E3755ECF940FE5EDED3" author="Vaziri, M. R. & Arab, A. L." box="[238,463,1691,1717]" pageId="4" pageNumber="125" pagination="149 - 155" refId="ref6534" refString="Vaziri, M. R. & Arab, A. L. (2013) Echinoids of the Genus Tetragramma Agassiz (Phymosomatoida) from the Aptian Sediments of the Basab Region, Northwest of Kerman, Iran. Journal of Sciences, Islamic Republic of Iran, 24, 149 - 155." type="journal article" year="2013">Vaziri & Arab, 2013</bibRefCitation>
|
|
</taxonomicName>
|
|
has been described from Aptian strata from the
|
|
<collectingRegion id="4982F84E542C3E3756E7F940FB3BDED3" box="[997,1194,1691,1717]" country="Iran" name="Kerman" pageId="4" pageNumber="125">Kerman province</collectingRegion>
|
|
, Southern
|
|
<collectingCountry id="F351763C542C3E375022F940FAC1DED3" box="[1312,1360,1691,1717]" name="Iran" pageId="4" pageNumber="125">Iran</collectingCountry>
|
|
, it was distinguished from
|
|
<taxonomicName id="4C464D2F542C3E37546DF91BFE72DEBF" authorityName="Agassiz & Desor" authorityYear="1846" box="[367,483,1727,1753]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="species" species="malbosi">
|
|
<emphasis id="B932EABE542C3E37546DF91BFE72DEBF" box="[367,483,1727,1753]" italics="true" pageId="4" pageNumber="125">T. malbosi</emphasis>
|
|
</taxonomicName>
|
|
in its more inflated test and the presence of phyllodes and biserial arranged pore pairs adapically. However, the statement of
|
|
<bibRefCitation id="EFD74B5D542C3E37573DF938FCBDDE9B" author="Vaziri, M. R. & Arab, A. L." box="[575,812,1763,1789]" pageId="4" pageNumber="125" pagination="149 - 155" refId="ref6534" refString="Vaziri, M. R. & Arab, A. L. (2013) Echinoids of the Genus Tetragramma Agassiz (Phymosomatoida) from the Aptian Sediments of the Basab Region, Northwest of Kerman, Iran. Journal of Sciences, Islamic Republic of Iran, 24, 149 - 155." type="journal article" year="2013">Vaziri & Arab (2013)</bibRefCitation>
|
|
on pore pairs development in
|
|
<taxonomicName id="4C464D2F542C3E375179F93FFB7EDE9B" authorityName="Agassiz & Desor" authorityYear="1846" box="[1147,1263,1763,1789]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="species" species="malbosi">
|
|
<emphasis id="B932EABE542C3E375179F93FFB7EDE9B" box="[1147,1263,1763,1789]" italics="true" pageId="4" pageNumber="125">T. malbosi</emphasis>
|
|
</taxonomicName>
|
|
was erroneous,
|
|
<taxonomicName id="4C464D2F542C3E375595F8D3FE9DDF47" authorityName="Agassiz & Desor" authorityYear="1846" box="[151,268,1799,1825]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="species" species="malbosi">
|
|
<emphasis id="B932EABE542C3E375595F8D3FE9DDF47" box="[151,268,1799,1825]" italics="true" pageId="4" pageNumber="125">T. malbosi</emphasis>
|
|
</taxonomicName>
|
|
clearly has biserial running pore pairs adapically and phyllodes as it is a diagnostic feature of the genus
|
|
<taxonomicName id="4C464D2F542C3E375595F8F7FDD2DF23" authority="(Smith & Wright, 1993)" baseAuthorityName="Smith & Wright" baseAuthorityYear="1993" box="[151,579,1835,1861]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="genus">
|
|
<emphasis id="B932EABE542C3E375595F8F7FEBFDF23" box="[151,302,1836,1861]" italics="true" pageId="4" pageNumber="125">Tetragramma</emphasis>
|
|
(
|
|
<bibRefCitation id="EFD74B5D542C3E37543FF8F0FDAADF23" author="Smith, A. B. & Wright, C. W." box="[317,571,1835,1861]" pageId="4" pageNumber="125" pagination="199 - 267" refId="ref6146" refString="Smith, A. B. & Wright, C. W. (1993) British Cretaceous Echinoids. Part 3, Stirodonta 2 (Hemicidaroida, Arbacioida and Phymosomatoida, part 1). Monograph of the Palaeontographical Society, 147, 199 - 267." type="journal article" year="1993">Smith & Wright, 1993</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
. Also, the name
|
|
<taxonomicName id="4C464D2F542C3E3757F8F8F7FAAEDF23" authority="Vaziri & Arab, 2013" authorityName="Vaziri & Arab" authorityYear="2013" box="[762,1343,1835,1861]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="species" species="tetratuberculatus">
|
|
<emphasis id="B932EABE542C3E3757F8F8F7FBC9DF23" box="[762,1112,1835,1861]" italics="true" pageId="4" pageNumber="125">Tetragramma tetratuberculatus</emphasis>
|
|
<bibRefCitation id="EFD74B5D542C3E37515FF8F0FAAEDF23" author="Vaziri, M. R. & Arab, A. L." box="[1117,1343,1835,1861]" pageId="4" pageNumber="125" pagination="149 - 155" refId="ref6534" refString="Vaziri, M. R. & Arab, A. L. (2013) Echinoids of the Genus Tetragramma Agassiz (Phymosomatoida) from the Aptian Sediments of the Basab Region, Northwest of Kerman, Iran. Journal of Sciences, Islamic Republic of Iran, 24, 149 - 155." type="journal article" year="2013">Vaziri & Arab, 2013</bibRefCitation>
|
|
</taxonomicName>
|
|
must be treated as a nomen nudum (
|
|
<bibRefCitation id="EFD74B5D542C3E3754C5F894FD21DF0F" author="Kroh, A. & Mooi, R." box="[455,688,1871,1897]" pageId="4" pageNumber="125" refId="ref5178" refString="Kroh, A. & Mooi, R. (2019 a) Tetragramma tetratuberculatus Vaziri & Arab, 2013 †. World Echinoidea Database. World Register of Marine Species. Available from: http: // www. marinespecies. org / aphia. php? p = taxdetails & id = 738296 (accessed 1 February 2019) https: // doi. org / 10.1044 / leader. PPL. 24012019.26" type="url" year="2019">Kroh & Mooi 2019a</bibRefCitation>
|
|
), because no
|
|
<typeStatus id="54FD880E542C3E375641F894FCE2DF0F" box="[835,883,1871,1897]" pageId="4" pageNumber="125">type</typeStatus>
|
|
specimen was designated; which fails to meet the criteria of the
|
|
<collectionCode id="ED57AE69542C3E375435F8A8FEE9DFEB" box="[311,376,1907,1933]" pageId="4" pageNumber="125">ICZN</collectionCode>
|
|
Article 16.4.1.
|
|
<bibRefCitation id="EFD74B5D542C3E375725F8A8FC89DFEB" author="Vaziri, M. R. & Arab, A. L." box="[551,792,1907,1933]" pageId="4" pageNumber="125" pagination="149 - 155" refId="ref6534" refString="Vaziri, M. R. & Arab, A. L. (2013) Echinoids of the Genus Tetragramma Agassiz (Phymosomatoida) from the Aptian Sediments of the Basab Region, Northwest of Kerman, Iran. Journal of Sciences, Islamic Republic of Iran, 24, 149 - 155." type="journal article" year="2013">Vaziri & Arab (2013)</bibRefCitation>
|
|
erected two further Aptian species of
|
|
<taxonomicName id="4C464D2F542C3E3751C5F8AFFACFDFEB" baseAuthorityName="Smith & Wright" baseAuthorityYear="1993" box="[1223,1374,1908,1933]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="genus">
|
|
<emphasis id="B932EABE542C3E3751C5F8AFFACFDFEB" box="[1223,1374,1908,1933]" italics="true" pageId="4" pageNumber="125">Tetragramma</emphasis>
|
|
</taxonomicName>
|
|
from the
|
|
<collectingRegion id="4982F84E542C3E3755C3F84CFE18DFD7" box="[193,393,1943,1969]" country="Iran" name="Kerman" pageId="4" pageNumber="125">Kerman province</collectingRegion>
|
|
:
|
|
<taxonomicName id="4C464D2F542C3E375497F843FC0CDFD7" authority="Vaziri & Arab, 2013" authorityName="Vaziri & Arab" authorityYear="2013" box="[405,925,1943,1969]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="species" species="basabensis">
|
|
<emphasis id="B932EABE542C3E375497F843FD21DFD7" box="[405,688,1943,1969]" italics="true" pageId="4" pageNumber="125">Tetragramma basabensis</emphasis>
|
|
<bibRefCitation id="EFD74B5D542C3E3757B5F84CFC0CDFD7" author="Vaziri, M. R. & Arab, A. L." box="[695,925,1943,1969]" pageId="4" pageNumber="125" pagination="149 - 155" refId="ref6534" refString="Vaziri, M. R. & Arab, A. L. (2013) Echinoids of the Genus Tetragramma Agassiz (Phymosomatoida) from the Aptian Sediments of the Basab Region, Northwest of Kerman, Iran. Journal of Sciences, Islamic Republic of Iran, 24, 149 - 155." type="journal article" year="2013">Vaziri & Arab, 2013</bibRefCitation>
|
|
</taxonomicName>
|
|
has a larger peristome than estimated for the specimen describe here;
|
|
<taxonomicName id="4C464D2F542C3E3754AFF867FC21DFB3" authority="Vaziri & Arab, 2013" authorityName="Vaziri & Arab" authorityYear="2013" box="[429,944,1979,2005]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="species" species="depressum">
|
|
<emphasis id="B932EABE542C3E3754AFF867FD52DFB3" box="[429,707,1979,2005]" italics="true" pageId="4" pageNumber="125">Tetragramma depressum</emphasis>
|
|
<bibRefCitation id="EFD74B5D542C3E3757C9F860FC21DFB3" author="Vaziri, M. R. & Arab, A. L." box="[715,944,1979,2005]" pageId="4" pageNumber="125" pagination="149 - 155" refId="ref6534" refString="Vaziri, M. R. & Arab, A. L. (2013) Echinoids of the Genus Tetragramma Agassiz (Phymosomatoida) from the Aptian Sediments of the Basab Region, Northwest of Kerman, Iran. Journal of Sciences, Islamic Republic of Iran, 24, 149 - 155." type="journal article" year="2013">Vaziri & Arab, 2013</bibRefCitation>
|
|
</taxonomicName>
|
|
was differentiated by
|
|
<bibRefCitation id="EFD74B5D542C3E3751AEF860FA0DDFB3" author="Vaziri, M. R. & Arab, A. L." box="[1196,1436,1979,2005]" pageId="4" pageNumber="125" pagination="149 - 155" refId="ref6534" refString="Vaziri, M. R. & Arab, A. L. (2013) Echinoids of the Genus Tetragramma Agassiz (Phymosomatoida) from the Aptian Sediments of the Basab Region, Northwest of Kerman, Iran. Journal of Sciences, Islamic Republic of Iran, 24, 149 - 155." type="journal article" year="2013">Vaziri & Arab (2013)</bibRefCitation>
|
|
by the increasing numbers of secondary tubercles adorally. However, both species,
|
|
<taxonomicName id="4C464D2F542C3E375135F83BFB5FDF9F" authorityName="Vaziri & Arab" authorityYear="2013" box="[1079,1230,2015,2041]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="species" species="basabensis">
|
|
<emphasis id="B932EABE542C3E375135F83BFB5FDF9F" box="[1079,1230,2015,2041]" italics="true" pageId="4" pageNumber="125">T. basabensis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C464D2F542C3E375001F83BFA04DF9F" authorityName="Vaziri & Arab" authorityYear="2013" box="[1283,1429,2015,2041]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="4" pageNumber="125" phylum="Echinodermata" rank="species" species="depressum">
|
|
<emphasis id="B932EABE542C3E375001F83BFA04DF9F" box="[1283,1429,2015,2041]" italics="true" pageId="4" pageNumber="125">T. depressum</emphasis>
|
|
</taxonomicName>
|
|
, are treated as nomina nuda (
|
|
<bibRefCitation id="EFD74B5D542D3E3654D9FF42FD5DD8D5" author="Kroh, A. & Mooi, R." box="[475,716,153,179]" pageId="5" pageNumber="126" refId="ref5253" refString="Kroh, A. & Mooi, R. (2019 b) Tetragramma basabensis Vaziri & Arab, 2013 †. World Echinoidea Database. World Register of Marine Species. Available from: http: // www. marinespecies. org / aphia. php? p = taxdetails & id = 738294 (accessed 1 February 2019) https: // doi. org / 10.1044 / leader. PPL. 24012019.26" type="url" year="2019">Kroh & Mooi 2019b</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFD74B5D542D3E3657DBFF42FC8ED8D5" author="Kroh, A. & Mooi, R." box="[729,799,153,179]" pageId="5" pageNumber="126" refId="ref5328" refString="Kroh, A. & Mooi, R. (2019 c) Tetragramma depressum Vaziri & Arab, 2013 †. World Echinoidea Database. World Register of Marine Species. Available from: http: // www. marinespecies. org / aphia. php? p = taxdetails & id = 738295 (accessed 1 February 2019) https: // doi. org / 10.1044 / leader. PPL. 24012019.26" type="url" year="2019">2019c</bibRefCitation>
|
|
) as no
|
|
<typeStatus id="54FD880E542D3E365676FF42FC3ED8D5" box="[884,943,153,179]" pageId="5" pageNumber="126">types</typeStatus>
|
|
were designated for each name. Other Ap- tian species,
|
|
<taxonomicName id="4C464D2F542D3E36542BFF65FD4ED8B1" authority="(Lambert, 1902)" baseAuthorityName="Lambert" baseAuthorityYear="1902" box="[297,735,189,215]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="5" pageNumber="126" phylum="Echinodermata" rank="species" species="almerai">
|
|
<emphasis id="B932EABE542D3E36542BFF65FD8ED8B1" box="[297,543,189,215]" italics="true" pageId="5" pageNumber="126">Tetragramma almerai</emphasis>
|
|
(
|
|
<bibRefCitation id="EFD74B5D542D3E36572CFF66FD46D8B1" author="Lambert, J." box="[558,727,189,215]" pageId="5" pageNumber="126" pagination="1 - 54" refId="ref5403" refString="Lambert, J. (1902) Description des Echinides fossiles de la province de Barcelone. Memoires de la Societe geologique de France, Paleontologie, 24, 1 - 54." type="journal article" year="1902">Lambert, 1902</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
from Casa-alta-Castellet (
|
|
<collectingRegion id="4982F84E542D3E36510AFF66FBE9D8B1" box="[1032,1144,189,215]" country="Spain" name="Catalunya" pageId="5" pageNumber="126">Catalonia</collectingRegion>
|
|
,
|
|
<collectingCountry id="F351763C542D3E365186FF66FB54D8BE" box="[1156,1221,189,216]" name="Spain" pageId="5" pageNumber="126">Spain</collectingCountry>
|
|
) and
|
|
<taxonomicName id="4C464D2F542D3E365007FF65FD0CD89D" authority="(Lambert in Lambert & Thiery, 1910)" baseAuthorityName="Lambert in Lambert & Thiery" baseAuthorityYear="1910" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="5" pageNumber="126" phylum="Echinodermata" rank="species" species="dumasi">
|
|
<emphasis id="B932EABE542D3E365007FF65FF79D89D" italics="true" pageId="5" pageNumber="126">Tetragramma dumasi</emphasis>
|
|
(Lambert in
|
|
<bibRefCitation id="EFD74B5D542D3E36547DFF3AFD05D89D" author="Lambert, J. & Thiery, P." box="[383,660,225,251]" pageId="5" pageNumber="126" pagination="81 - 160" refId="ref5486" refString="Lambert, J. & Thiery, P. (1910) Essai de Nomenclature Raisonnee des Echinides. Fasc. 2. L. Ferriere, Chaumont, 80 pp. [pp. 81 - 160, July 1910]" type="book chapter" year="1910">Lambert & Thiéry, 1910</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
from Montaren (
|
|
<collectingRegion id="4982F84E542D3E365662FF3AFBF3D89D" box="[864,1122,225,251]" country="France" name="Languedoc-Roussillon" pageId="5" pageNumber="126">Languedoc-Roussillon</collectingRegion>
|
|
,
|
|
<collectingCountry id="F351763C542D3E36516CFF3AFB2DD89D" box="[1134,1212,225,251]" name="France" pageId="5" pageNumber="126">France</collectingCountry>
|
|
) are very similar to
|
|
<taxonomicName id="4C464D2F542D3E365595FEDDFE9CD979" authorityName="Agassiz & Desor" authorityYear="1846" box="[151,269,261,287]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="5" pageNumber="126" phylum="Echinodermata" rank="species" species="malbosi">
|
|
<emphasis id="B932EABE542D3E365595FEDDFE9CD979" box="[151,269,261,287]" italics="true" pageId="5" pageNumber="126">T. malbosi</emphasis>
|
|
</taxonomicName>
|
|
. However, further investigations are needed to conclude whether
|
|
<taxonomicName id="4C464D2F542D3E3656F9FEDDFBFED979" baseAuthorityName="Lambert" baseAuthorityYear="1902" box="[1019,1135,261,287]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="5" pageNumber="126" phylum="Echinodermata" rank="species" species="almerai">
|
|
<emphasis id="B932EABE542D3E3656F9FEDDFBFED979" box="[1019,1135,261,287]" italics="true" pageId="5" pageNumber="126">T. almerai</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C464D2F542D3E36517DFEDDFB7CD979" baseAuthorityName="Lambert in Lambert & Thiery" baseAuthorityYear="1910" box="[1151,1261,261,287]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="5" pageNumber="126" phylum="Echinodermata" rank="species" species="dumasi">
|
|
<emphasis id="B932EABE542D3E36517DFEDDFB7CD979" box="[1151,1261,261,287]" italics="true" pageId="5" pageNumber="126">T. dumasi</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C464D2F542D3E365025FEDDFA0CD979" authorityName="Agassiz & Desor" authorityYear="1846" box="[1319,1437,261,287]" class="Echinoidea" family="Diplopodiidae" genus="Tetragramma" higherTaxonomySource="CoL" kingdom="Animalia" order="Phymosomatoida" pageId="5" pageNumber="126" phylum="Echinodermata" rank="species" species="malbosi">
|
|
<emphasis id="B932EABE542D3E365025FEDDFA0CD979" box="[1319,1437,261,287]" italics="true" pageId="5" pageNumber="126">T. malbosi</emphasis>
|
|
</taxonomicName>
|
|
should be treated as synonyms or not.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |