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<mods:title id="55C618C3A4A945E335AED4E754CDE0FD">Reassessing the phylogeny and divergence times of sloths (Mammalia: Pilosa: Folivora), exploring alternative morphological partitioning and dating models</mods:title>
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<subSubSection id="C35C6525FFF3FF94FC86F8ECC5F99006" box="[809,1256,1839,1863]" pageId="20" pageNumber="1525" type="nomenclature">
<paragraph id="8BF936AEFFF3FF94FC86F8ECC5F99006" blockId="20.[809,1256,1839,1863]" box="[809,1256,1839,1863]" pageId="20" pageNumber="1525">
<taxonomicName id="4C464D2DFFF3FF94FC86F8ECC2E09006" ID-CoL="CH5" authorityName="Ameghino" authorityYear="1889" box="[809,1009,1839,1863]" class="Mammalia" family="Megalonychidae" kingdom="Animalia" order="Pilosa" pageId="20" pageNumber="1525" phylum="Chordata" rank="family">Megalonychidae</taxonomicName>
plus
<taxonomicName id="4C464D2DFFF3FF94FB9BF8ECC5F99006" box="[1076,1256,1839,1863]" class="Mammalia" family="Megatheriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="20" pageNumber="1525" phylum="Chordata" rank="family">Megatheriidae</taxonomicName>
</paragraph>
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<subSubSection id="C35C6525FFF3FF95FC86F89BC21D924D" lastPageId="21" lastPageNumber="1526" pageId="20" pageNumber="1525" type="discussion">
<paragraph id="8BF936AEFFF3FF94FC86F89BC5ED902F" blockId="20.[809,1276,1880,1902]" box="[809,1276,1880,1902]" pageId="20" pageNumber="1525">PP = 100, age = 22.14 Mya (19.0825.29).</paragraph>
<paragraph id="8BF936AEFFF2FF95FF14FF06C0E0952B" blockId="21.[163,780,197,1292]" pageId="21" pageNumber="1526">
The clade, which includes all megatherioids excepting the genera
<taxonomicName id="4C464D2DFFF2FF95FE38FF27C30397B8" authorityName="Ameghino" authorityYear="1891" box="[407,530,228,249]" class="Mammalia" family="Megatheriidae" genus="Pelecyodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="21" pageNumber="1526" phylum="Chordata" rank="genus">
<emphasis id="B932EABCFFF2FF95FE38FF27C30397B8" box="[407,530,228,249]" italics="true" pageId="21" pageNumber="1526">Pelecyodon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C464D2DFFF2FF95FDE5FF27C21597B8" authorityName="Ameghino" authorityYear="1887" box="[586,772,228,249]" class="Mammalia" family="Megatheriidae" genus="Schismotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="21" pageNumber="1526" phylum="Chordata" rank="genus">
<emphasis id="B932EABCFFF2FF95FDE5FF27C21597B8" box="[586,772,228,249]" italics="true" pageId="21" pageNumber="1526">Schismotherium</emphasis>
</taxonomicName>
, was recovered in all Bayesian analyses using H models, and in par_EW and par_IW100, although in these MP analyses,
<taxonomicName id="4C464D2DFFF2FF95FE19FE83C3399614" authorityName="Ameghino" authorityYear="1887" box="[438,552,320,341]" class="Mammalia" family="Megatheriidae" genus="Hapalops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="21" pageNumber="1526" phylum="Chordata" rank="genus">
<emphasis id="B932EABCFFF2FF95FE19FE83C3399614" box="[438,552,320,341]" italics="true" pageId="21" pageNumber="1526">Hapalops</emphasis>
</taxonomicName>
was not recovered closely associated with megalonychids, but as sister to the clade uniting all other megalonychids and megatheriids. In par_IW10 and par_IW5, a distinct arrangement was obtained, with a clade (
<taxonomicName id="4C464D2DFFF2FF95FD3BFE79C215968E" authorityName="Ameghino" authorityYear="1887" box="[660,772,442,463]" class="Mammalia" family="Megatheriidae" genus="Hapalops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="21" pageNumber="1526" phylum="Chordata" rank="genus">
<emphasis id="B932EABCFFF2FF95FD3BFE79C215968E" box="[660,772,442,463]" italics="true" pageId="21" pageNumber="1526">Hapalops</emphasis>
</taxonomicName>
, (
<taxonomicName id="4C464D2DFFF2FF95FF04FE1AC03B96AF" authorityName="Ameghino" authorityYear="1891" box="[171,298,473,494]" class="Mammalia" family="Megatheriidae" genus="Pelecyodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="21" pageNumber="1526" phylum="Chordata" rank="genus">
<emphasis id="B932EABCFFF2FF95FF04FE1AC03B96AF" box="[171,298,473,494]" italics="true" pageId="21" pageNumber="1526">Pelecyodon</emphasis>
</taxonomicName>
,
<taxonomicName id="4C464D2DFFF2FF95FE97FE1AC0E696AF" authorityName="Ameghino" authorityYear="1887" box="[312,503,473,494]" class="Mammalia" family="Megatheriidae" genus="Schismotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="21" pageNumber="1526" phylum="Chordata" rank="genus">
<emphasis id="B932EABCFFF2FF95FE97FE1AC0E696AF" box="[312,503,473,494]" italics="true" pageId="21" pageNumber="1526">Schismotherium</emphasis>
</taxonomicName>
)) recovered as sister to
<taxonomicName id="4C464D2DFFF2FF95FF0CFE3BC040954F" box="[163,337,504,526]" class="Mammalia" family="Megatheriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="21" pageNumber="1526" phylum="Chordata" rank="family">Megatheriidae</taxonomicName>
. In trees from analyses with UN and A models, in which Megatherioidea was recovered as paraphyletic, the clade uniting
<taxonomicName id="4C464D2DFFF2FF95FDB7FDF6C3C6950A" authorityName="Ameghino" authorityYear="1889" box="[536,727,565,587]" class="Mammalia" family="Megalonychidae" kingdom="Animalia" order="Pilosa" pageId="21" pageNumber="1526" phylum="Chordata" rank="family">Megalonychidae</taxonomicName>
and
<taxonomicName id="4C464D2DFFF2FF95FF0CFD97C058952B" box="[163,329,596,618]" class="Mammalia" family="Megatheriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="21" pageNumber="1526" phylum="Chordata" rank="family">Megatheriidae</taxonomicName>
is also absent.
</paragraph>
<paragraph id="8BF936AEFFF2FF95FF14FDB1C21D924D" blockId="21.[163,780,197,1292]" pageId="21" pageNumber="1526">The clade was supported by 16 synapomorphies (11 for both methods and five exclusively for BI): Cf1 and cf1 slightly depressed ventrally relative to molariforms; presence of a well-developed diastema between Cf1/cf1 and molariforms; ovate mediolaterally to rectangular Mf1 cross-section (B1); mandibular symphysis longer than molariform toothrow; horizontal orientation of spout, in lateral view; external opening of mandibular canal opens anterolaterally, on ascending ramus (B1); tip of zygomatic process anterior to frontoparietal suture; nuchal crest splits dorsally into anterior and posterior crests, with a raised triangular area in the dorsal surface of the skull roof (B1); nuchal crest overhangs occiput posteriorly; presence of a distinct neck at the base of occipital condyles (B1); rugose ectotympanic external surface; presence of an ectotympanic medial expansion; presence of pterygoid lateral groove for m. tensor veli palatini; medial extent of trochlear notch of ulna equal to or less than that of the olecranon (B1); straight posterior margin of ulnar diaphysis in lateral view; and lateroproximal process of tuber calcis more distal than medioproximal process.</paragraph>
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