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<document id="3C14331027917C8EAB7FBA2823855A06" ID-DOI="10.5252/cryptogamie-bryologie2020v41a2" ID-ISSN="1776-0992" ID-Zenodo-Dep="12215441" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719006298856" checkinUser="felipe" docAuthor="Reeb, Catherine &amp; Gradstein, Robbert" docDate="2020" docId="03EF87AA6534FFDDC9BEE2FFFC04F987" docLanguage="en" docName="CryptogamieBryologie.41.2.11-34.pdf" docOrigin="Cryptogamie, Bryologie 20 (2)" docSource="http://dx.doi.org/10.5252/cryptogamie-bryologie2020v41a2" docStyle="DocumentStyle:3F4A000F5B047ABA0EAB2943AA9373AE.3:CryptogamieBryologie.2019-.journal_article" docStyleId="3F4A000F5B047ABA0EAB2943AA9373AE" docStyleName="CryptogamieBryologie.2019-.journal_article" docStyleVersion="3" docTitle="Riccardia vohimanensis Reeb &amp; Gradstein 2020, sp. nov." docType="treatment" docVersion="2" lastPageNumber="31" masterDocId="FFD6FFD26522FFCBC97FE62CFFFBFFEA" masterDocTitle="A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key" masterLastPageNumber="34" masterPageNumber="11" pageNumber="31" updateTime="1719020027017" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C9355470C60405D686FF3158606AEC86">A taxonomic revision of Aneuraceae (Marchantiophyta) from eastern Africa with an interactive identification key</mods:title>
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<mods:affiliation id="FDBBA340E4F95EA47458F027955ECEE3">Institut de Systématique Évolution Biodiversité, Sorbonne Université, MNHN, CNRS, EPHE, UMR 7205 Museum national dHistoire naturelle, 57 rue Cuvier 75231 Paris cedex 05 (France)</mods:affiliation>
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<taxonomicName id="4C464D3F6534FFDDC9BEE2FFFD88FB04" ID-CoL="9MFYN" authority="Reeb &amp; Gradst." authorityName="Reeb &amp; Gradstein" authorityYear="2020" box="[193,627,1235,1262]" class="Jungermanniopsida" family="Aneuraceae" genus="Riccardia" kingdom="Plantae" order="Metzgeriales" pageId="22" pageNumber="31" phylum="Marchantiophyta" rank="species" species="vohimanensis" status="sp. nov.">
<emphasis id="B932EAAE6534FFDDC9BEE2FFFE3BFB04" bold="true" box="[193,448,1235,1262]" italics="true" pageId="22" pageNumber="31">Riccardia vohimanensis</emphasis>
Reeb &amp; Gradst.
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<paragraph id="8BF936BC6534FFDDC8F9E2DFFDFEFAE4" blockId="22.[390,517,1267,1294]" box="[390,517,1267,1294]" pageId="22" pageNumber="31">
<heading id="D0B181D06534FFDDC8F9E2DFFDFEFAE4" box="[390,517,1267,1294]" centered="true" fontSize="11" level="3" pageId="22" pageNumber="31" reason="3">
(
<figureCitation id="137D2A396534FFDDC8F2E2DFFE37FAE4" box="[397,460,1267,1294]" captionStart="FIG" captionStartId="18.[132,143,1826,1844]" captionTargetBox="[245,1355,223,1778]" captionTargetId="figure-22@18.[795,1364,782,1353]" captionTargetPageId="18" captionText="FIG. 4. — Riccardia martinii sp. nov.: A, thallus mats; B, dry thallus, showing white margins; C, moistened thallus; D, portion of thallus in dorsal view, showing thickened cell walls; E, apex of ultimate branch, showing papillae; F, main axis in cross section; G, ultimate branches in cross section; H, calyptra; I, portion of thallus with male branches. From Sass-Gyarmati 9613/CW, Vojko 9660/CS, Pócs 9602CU, 6877/K, Bardat JBMAD78 (PC0763890), Riccardia ramosissima (Steph) Grolle: J, thalli; K, main axis in cross section; L, ultimate branches in cross section; M, male branch; N, calyptra; O, upper portion of calyptra, showing umbo. From Reeb CR13Z51 (PC0763864), Pócs 6981/A, Jelinek s.n. (Type of Aneura compacta). Riccardia saccatiflora (Steph.) S.W.Arnell: P, thalli; Q, main axis in cross section; R, ultimate branches in cross section; S, portion of thallus with two female branch (left) and two male branches; T, calyptra. From Wigginton M1758b, Reeb CR11142 (PC0763883), Een M064, Rodriguez 182 (Type of Aneura saccatiflora). Riccardia vohimanensis Reeb &amp; Gradst.: U, thalli; V, main axis in cross section; W, ultimate branch in cross section; X, sporophyte and calyptra with the vegetative expansion at its basis. From Reeb MTM1278 (PC0763879), CR13Z23 (PC0763868), OShea M7327b, Een M028. Scale bars: A-C, H-J, N, P, U, 1 mm; D-G, M, Q, T, V-X, 500 µm; K, L, R, 100 µm; O, S, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.12215449" httpUri="https://zenodo.org/record/12215449/files/figure.png" pageId="22" pageNumber="31">Fig. 4</figureCitation>
U-X)
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. — Monoicous. Thallus rigid, unwinged, with short branches. Axis plano-convex to biconvex, 8-12 cells thick, ventral side with mycorrhiza-like fungi. Immature calyptra strawberry-like.
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. —
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. Vohimana Reserve,
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,
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0771053]).
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. —
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. Alaotra-Mangoro, Zahamena Reserve,
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Reeb
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0763868); Maromizaha Reserve,
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<emphasis id="B932EAAE6534FFDDC8A6E07FFD87F981" box="[473,636,1619,1643]" italics="true" pageId="22" pageNumber="31">Reeb MTM1278*</emphasis>
(
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,
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,
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(
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0763878); Haute Matsiatra, Andringitra Nat. Park, 22°084061”
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21.
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Reeb
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</emphasis>
(
<collectionCode id="ED57AE796534FFDDCBF4E08FFD54F951" box="[651,687,1699,1723]" country="France" lsid="urn:lsid:biocol.org:col:15359" name="Museum National d'Histoire Naturelle, Paris (MNHN) - Non-vascular Plants and Fungi" pageId="22" pageNumber="31" type="Herbarium">PC</collectionCode>
).
</paragraph>
<paragraph id="8BF936BC6534FFDDC9FBE093FDA8F91A" blockId="22.[130,776,1540,1804]" pageId="22" pageNumber="31">
<emphasis id="B932EAAE6534FFDDC9FBE093FF2FF93D" bold="true" box="[132,212,1727,1751]" pageId="22" pageNumber="31">
<collectingCountry id="F351762C6534FFDDC9FBE093FF2FF93D" box="[132,212,1727,1751]" name="Malawi" pageId="22" pageNumber="31">Malawi</collectingCountry>
</emphasis>
.
<collectingRegion id="4982F85E6534FFDDC999E091FEC6F93F" box="[230,317,1725,1749]" country="Malawi" name="Mulanje" pageId="22" pageNumber="31">Mulanje</collectingRegion>
Mt.,
<geoCoordinate id="EE72507B6534FFDDC8FFE091FDEBF93F" box="[384,528,1725,1749]" degrees="15" direction="south" minutes="53" orientation="latitude" pageId="22" pageNumber="31" precision="1" seconds="32.1" value="-15.89225">15°5332.1”S</geoCoordinate>
,
<geoCoordinate id="EE72507B6534FFDDCB60E091FD66F93F" box="[543,669,1725,1749]" degrees="35" direction="east" minutes="29" orientation="longitude" pageId="22" pageNumber="31" precision="15" seconds="24" value="35.49">35°2924”E</geoCoordinate>
,
<quantity id="4CBE9B596534FFDDCBD3E091FCF8F93F" box="[684,771,1725,1749]" metricMagnitude="3" metricUnit="m" metricValue="2.18" pageId="22" pageNumber="31" unit="m" value="2180.0">2180 m</quantity>
,
<date id="FFF8107C6534FFDDC9FBE0F4FF02F91A" box="[132,249,1752,1776]" pageId="22" pageNumber="31" value="1991-06-22">
22.
<collectionCode id="ED57AE796534FFDDC9DCE0F4FF3AF91A" box="[163,193,1752,1776]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="22" pageNumber="31">VI</collectionCode>
.1991
</date>
,
<emphasis id="B932EAAE6534FFDDC87CE0F4FE39F91A" box="[259,450,1752,1776]" italics="true" pageId="22" pageNumber="31">
OShea
<collectionCode id="ED57AE796534FFDDC833E0F4FE98F91A" box="[332,355,1752,1776]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="22" pageNumber="31" type="Herbarium">M</collectionCode>
7327A-2*
</emphasis>
(
<collectionCode id="ED57AE796534FFDDC8AEE0F4FE1BF91A" box="[465,480,1752,1776]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="22" pageNumber="31" type="Herbarium">E</collectionCode>
00430480).
</paragraph>
<paragraph id="8BF936BC6534FFDDC9FBE0D8FD0FF8E0" blockId="22.[130,776,1540,1804]" box="[132,756,1778,1804]" pageId="22" pageNumber="31">
<emphasis id="B932EAAE6534FFDDC9FBE0D8FF1CF8E6" bold="true" box="[132,231,1780,1804]" pageId="22" pageNumber="31">
<collectingCountry id="F351762C6534FFDDC9FBE0D8FF1CF8E6" box="[132,231,1780,1804]" name="Mauritius" pageId="22" pageNumber="31">Mauritius</collectingCountry>
</emphasis>
. Macabé,
<quantity id="4CBE9B596534FFDDC832E0DEFE74F8E0" box="[333,399,1778,1802]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="22" pageNumber="31" unit="m" value="550.0">550 m</quantity>
,
<date id="FFF8107C6534FFDDC8E5E0DEFDFEF8E0" box="[410,517,1778,1802]" pageId="22" pageNumber="31" value="1962-10-03">03.X.1962</date>
,
<emphasis id="B932EAAE6534FFDDCB70E0DFFD84F8E0" box="[527,639,1778,1802]" italics="true" pageId="22" pageNumber="31">
Een
<collectionCode id="ED57AE796534FFDDCB46E0DEFDABF8E0" box="[569,592,1778,1802]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="22" pageNumber="31" type="Herbarium">M</collectionCode>
028*
</emphasis>
(S-B7097).
</paragraph>
<paragraph id="8BF936BC6534FFDDC9FBE10EFD82F8D0" blockId="22.[132,633,1826,1850]" box="[132,633,1826,1850]" pageId="22" pageNumber="31">
<smallCapsWord id="8D1FA0606534FFDDC9FBE10EFEE9F8D2" baselines="1843,1844" box="[132,274,1826,1849]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="22" pageNumber="31">DISTRIBUTION</smallCapsWord>
. —
<collectingCountry id="F351762C6534FFDDC841E10EFE71F8D0" box="[318,394,1826,1850]" name="Malawi" pageId="22" pageNumber="31">Malawi</collectingCountry>
,
<collectingCountry id="F351762C6534FFDDC8ECE10EFDFCF8D0" box="[403,519,1826,1850]" name="Madagascar" pageId="22" pageNumber="31">Madagascar</collectingCountry>
,
<collectingCountry id="F351762C6534FFDDCB6EE10EFD8EF8D0" box="[529,629,1826,1850]" name="Mauritius" pageId="22" pageNumber="31">Mauritius</collectingCountry>
.
</paragraph>
<paragraph id="8BF936BC6534FFDDC9FBE17EFD4EF869" blockId="22.[131,774,1873,1924]" pageId="22" pageNumber="31">
<smallCapsWord id="8D1FA0606534FFDDC9FBE17EFF27F882" baselines="1891,1892" box="[132,220,1874,1897]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Habitat" pageId="22" pageNumber="31">HABITAT</smallCapsWord>
. — On dead logs (bare, smooth and very hard wood) and buttresses, rarely on humid rock, from
<quantity id="4CBE9B596534FFDDCB7FE140FDBBF869" box="[512,576,1900,1924]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="22" pageNumber="31" unit="m" value="550.0">550 m</quantity>
to
<quantity id="4CBE9B596534FFDDCB1EE140FD4AF869" box="[609,689,1900,1924]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="22" pageNumber="31" unit="m" value="2200.0">2200 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C65376534FFDDC9FBE19EFC04F987" pageId="22" pageNumber="31" type="description">
<paragraph id="8BF936BC6534FFDDC9FBE19EFEEEF820" blockId="22.[132,277,1970,1996]" box="[132,277,1970,1996]" pageId="22" pageNumber="31">
<heading id="D0B181D06534FFDDC9FBE19EFEEEF820" box="[132,277,1970,1996]" fontSize="9" level="3" pageId="22" pageNumber="31" reason="5">
<smallCapsWord id="8D1FA0606534FFDDC9FBE19EFEEEF820" baselines="1989,1989" box="[132,277,1970,1996]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="22" pageNumber="31">DESCRIPTION</smallCapsWord>
</heading>
</paragraph>
<paragraph id="8BF936BC6534FFDDC9FBE1FDFDE8F801" blockId="22.[132,531,2001,2028]" box="[132,531,2001,2028]" pageId="22" pageNumber="31">Monoicous (autoicous or paroicous).</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E6FBFC83FF1B" blockId="22.[813,888,215,241]" box="[813,888,215,241]" pageId="22" pageNumber="31">
<heading id="D0B181D06534FFDDCA52E6FBFC83FF1B" box="[813,888,215,241]" fontSize="11" level="3" pageId="22" pageNumber="31" reason="9">
<emphasis id="B932EAAE6534FFDDCA52E6FBFC83FF1B" box="[813,888,215,241]" italics="true" pageId="22" pageNumber="31">Thallus</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E6DBFC29FE7B" blockId="22.[811,1457,247,401]" pageId="22" pageNumber="31">
Leathery and rigid, green to blackish green when fresh and forming rosettes on bare substrates, dull brown to black in herbarium, closely attached to the substrate, to
<quantity id="4CBE9B596534FFDDCC59E71BFA8FFEBB" box="[1318,1396,311,337]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="22" pageNumber="31" unit="mm" value="22.0">22 mm</quantity>
long, 1-2-pinnate, not palmate, branches alternate, not flagelliform, stolons lacking.
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E79BFC6EFE3A" blockId="22.[813,917,439,464]" box="[813,917,439,464]" pageId="22" pageNumber="31">
<heading id="D0B181D06534FFDDCA52E79BFC6EFE3A" box="[813,917,439,464]" fontSize="11" level="3" pageId="22" pageNumber="31" reason="9">
<emphasis id="B932EAAE6534FFDDCA52E79BFC6EFE3A" box="[813,917,439,464]" italics="true" pageId="22" pageNumber="31">Main axis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E7FAFAF1FDBA" blockId="22.[811,1457,470,592]" pageId="22" pageNumber="31">Well-defined, 700-1000(-1200) µm wide, in cross section plano-convex to concave-convex, with mycorrhizal fungi in the ventral part, 8-12 cells thick, margins obtuse, not winged, epidermis cells 2-4× smaller than inner cells.</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E45AFC12FD7A" blockId="22.[813,1001,630,656]" box="[813,1001,630,656]" pageId="22" pageNumber="31">
<heading id="D0B181D06534FFDDCA52E45AFC12FD7A" box="[813,1001,630,656]" fontSize="11" level="3" pageId="22" pageNumber="31" reason="9">
<emphasis id="B932EAAE6534FFDDCA52E45AFC12FD7A" box="[813,1001,630,656]" italics="true" pageId="22" pageNumber="31">Ultimate branches</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E4BAFBFBFCC5" blockId="22.[813,1457,662,815]" pageId="22" pageNumber="31">
Oblong and stocky, to
<quantity id="4CBE9B596534FFDDCD41E4BAFB7FFD5A" box="[1086,1156,662,688]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="31" unit="mm" value="2.0">2 mm</quantity>
long and
<quantity id="4CBE9B596534FFDDCC7FE4BAFABDFD5A" box="[1280,1350,662,688]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="31" unit="mm" value="1.0">1 mm</quantity>
wide, in cross section biconvex to plano-convex, 7-9 cells thick, margins acute, not winged, branch apex rounded and often widened, emarginate to weakly dissected, without overlapping edges.
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E579FC18FC85" blockId="22.[813,995,853,879]" box="[813,995,853,879]" pageId="22" pageNumber="31">
<heading id="D0B181D06534FFDDCA52E579FC18FC85" box="[813,995,853,879]" fontSize="11" level="3" pageId="22" pageNumber="31" reason="9">
<emphasis id="B932EAAE6534FFDDCA52E579FC18FC85" box="[813,995,853,879]" italics="true" pageId="22" pageNumber="31">Mucilage papillae</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E559FC0BFC65" blockId="22.[813,1008,885,911]" box="[813,1008,885,911]" pageId="22" pageNumber="31">Apical and lateral.</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E599FC6EFC25" blockId="22.[813,917,949,975]" box="[813,917,949,975]" pageId="22" pageNumber="31">
<heading id="D0B181D06534FFDDCA52E599FC6EFC25" box="[813,917,949,975]" fontSize="11" level="3" pageId="22" pageNumber="31" reason="9">
<emphasis id="B932EAAE6534FFDDCA52E599FC6EFC25" box="[813,917,949,975]" italics="true" pageId="22" pageNumber="31">Oil bodies</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E5F9FC6EFC05" blockId="22.[813,917,981,1007]" box="[813,917,981,1007]" pageId="22" pageNumber="31">
<collectingRegion id="4982F85E6534FFDDCA52E5F9FCA3FC05" box="[813,856,981,1007]" country="Cameroon" name="North" pageId="22" pageNumber="31">Not</collectingRegion>
seen.
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E238FC34FBC4" blockId="22.[813,975,1044,1070]" box="[813,975,1044,1070]" pageId="22" pageNumber="31">
<heading id="D0B181D06534FFDDCA52E238FC34FBC4" box="[813,975,1044,1070]" fontSize="11" level="3" pageId="22" pageNumber="31" reason="9">
<emphasis id="B932EAAE6534FFDDCA52E238FC34FBC4" box="[813,975,1044,1070]" italics="true" pageId="22" pageNumber="31">Sexual branches</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E219FAB0FBA4" blockId="22.[813,1355,1076,1103]" box="[813,1355,1076,1103]" pageId="22" pageNumber="31">Numerous all along axis, often close to each other.</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E258FC3AFB64" blockId="22.[813,961,1140,1166]" box="[813,961,1140,1166]" pageId="22" pageNumber="31">
<heading id="D0B181D06534FFDDCA52E258FC3AFB64" box="[813,961,1140,1166]" fontSize="11" level="3" pageId="22" pageNumber="31" reason="9">
<emphasis id="B932EAAE6534FFDDCA52E258FC3AFB64" box="[813,961,1140,1166]" italics="true" pageId="22" pageNumber="31">Male branches</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E2B8FBF6FB24" blockId="22.[811,1455,1172,1230]" pageId="22" pageNumber="31">Mainly grouped, sessile, 300-400 µm long, not winged, with 4-8 antheridial pairs.</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E2DFFC2EFAE7" blockId="22.[813,981,1267,1293]" box="[813,981,1267,1293]" pageId="22" pageNumber="31">
<heading id="D0B181D06534FFDDCA52E2DFFC2EFAE7" box="[813,981,1267,1293]" fontSize="11" level="3" pageId="22" pageNumber="31" reason="9">
<emphasis id="B932EAAE6534FFDDCA52E2DFFC2EFAE7" box="[813,981,1267,1293]" italics="true" pageId="22" pageNumber="31">Female branches</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E33FFBD5FA67" blockId="22.[812,1457,1299,1421]" pageId="22" pageNumber="31">Grouped, laterally inserted, 300-500 µm long, base often elongate, female branch bordered by scales or long hair-like, straight cilia, the cilia longer than the width of the axis, archegonia in 4-6 pairs.</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E39FFC7DFA27" blockId="22.[813,902,1459,1485]" box="[813,902,1459,1485]" pageId="22" pageNumber="31">
<heading id="D0B181D06534FFDDCA52E39FFC7DFA27" box="[813,902,1459,1485]" fontSize="11" level="3" pageId="22" pageNumber="31" reason="9">
<emphasis id="B932EAAE6534FFDDCA52E39FFC7DFA27" box="[813,902,1459,1485]" italics="true" pageId="22" pageNumber="31">Calyptra</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E3FFFB34F9E7" blockId="22.[811,1455,1491,1549]" pageId="22" pageNumber="31">
Less than
<quantity id="4CBE9B596534FFDDCAEDE3FFFC2BFA07" box="[914,976,1491,1517]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="31" unit="mm" value="1.0">1 mm</quantity>
long, strawberry-like when immature, densely covered by rounded, thick-walled cells.
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E01FFC72F9A7" blockId="22.[813,905,1587,1613]" box="[813,905,1587,1613]" pageId="22" pageNumber="31">
<heading id="D0B181D06534FFDDCA52E01FFC72F9A7" box="[813,905,1587,1613]" fontSize="11" level="3" pageId="22" pageNumber="31" reason="9">
<emphasis id="B932EAAE6534FFDDCA52E01FFC72F9A7" box="[813,905,1587,1613]" italics="true" pageId="22" pageNumber="31">Gemmae</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936BC6534FFDDCA52E07EFC04F987" blockId="22.[813,1023,1618,1645]" box="[813,1023,1618,1645]" pageId="22" pageNumber="31">Sometimes present.</paragraph>
</subSubSection>
</treatment>
</document>