treatments-xml/data/03/EF/87/03EF87A4FFEE7D0AFF97FD28FE22789A.xml
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<mods:title>New look on the geographical variation in Rhinolophus clivosus with description of a new horseshoe bat species from Cyrenaica, Libya</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Benda, Petr</mods:namePart>
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<mods:namePart>Vallo, Peter</mods:namePart>
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<mods:title>Vespertilio</mods:title>
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<mods:date>2012</mods:date>
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<mods:number>16</mods:number>
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<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[117,362,709,733]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="84" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis bold="true" box="[117,362,709,733]" italics="true" pageId="15" pageNumber="84">Rhinolophus horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[369,429,709,733]" pageId="15" pageNumber="84" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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<paragraph blockId="15.[117,1138,761,898]" pageId="15" pageNumber="84">
<treatmentCitationGroup pageId="15" pageNumber="84">
<emphasis box="[117,323,761,782]" italics="true" pageId="15" pageNumber="84">
<taxonomicName authorityName="Lacepede" authorityYear="1799" box="[117,238,761,782]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="84" phylum="Chordata" rank="genus">Rhinolophus</taxonomicName>
clivosus
</emphasis>
Cretzchmar, 1828:
<treatmentCitation author="QUMSIYEH M. B." box="[516,710,761,783]" page="49" pageId="15" pageNumber="84" year="1981">
<bibRefCitation author="QUMSIYEH M. B." box="[516,710,761,783]" journalOrPublisher="N. E. Libya. Bat Research News" pageId="15" pageNumber="84" pagination="49" part="22" refId="ref14788" refString="QUMSIYEH M. B., 1981: The bat fauna of Jebel Al Akhdar, N. E. Libya. Bat Research News, 22 (4): 49." title="The bat fauna of Jebel Al Akhdar" type="journal article" year="1981">Qumsiyeh 1981: 49</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="QUMSIYEH M. B. &amp; SCHLITTER D. A." box="[721,1041,761,783]" page="384" pageId="15" pageNumber="84" year="1982">
<bibRefCitation author="QUMSIYEH M. B. &amp; SCHLITTER D. A." box="[721,1041,761,783]" journalOrPublisher="Annals of Carnegie Museum" pageId="15" pageNumber="84" pagination="377 - 389" part="51" refId="ref14848" refString="QUMSIYEH M. B. &amp; SCHLITTER D. A., 1982: The bat fauna of Jabal Al Akhdar, Northeast Libya. Annals of Carnegie Museum, 51: 377 - 389." title="The bat fauna of Jabal Al Akhdar, Northeast Libya" type="journal article" year="1982">Qumsiyeh &amp; Schlitter 1982: 384</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="LE BERRE M." page="78" pageId="15" pageNumber="84" year="1990">
<bibRefCitation author="LE BERRE M." bookContentInfo="360 pp." journalOrPublisher="Chabaud &amp; Lechevalier, Paris" pageId="15" pageNumber="84" refId="ref14564" refString="LE BERRE M., 1990: Faune du Sahara. 2. Mammiferes. Chabaud &amp; Lechevalier, Paris, 360 pp." title="Faune du Sahara. 2. Mammiferes" type="book" year="1990">Le Berre 1990: 78</bibRefCitation>
</treatmentCitation>
[partim];
<treatmentCitation author="HORACEK I. &amp; HANAK V. &amp; GAISLER J." box="[337,583,787,809]" page="100" pageId="15" pageNumber="84" year="2000">
<bibRefCitation author="HORACEK I. &amp; HANAK V. &amp; GAISLER J." bookContentInfo="280 pp." box="[337,583,787,809]" editor="WOLOSZYN W." journalOrPublisher="Institute of Systematics and Evolution of Animals PAS, Krakow" pageId="15" pageNumber="84" pagination="11 - 157" refId="ref14216" refString="HORACEK I., HANAK V. &amp; GAISLER J., 2000: Bats of the Palearctic Region: a taxonomic and biogegraphic review. Pp.: 11 - 157. In: WOLOSZYN W. (ed.): Proceedings of the VIIIth European Bat Research Conference. Volume I. Approaches to Biogeography and Ecology of Bats. Institute of Systematics and Evolution of Animals PAS, Krakow, 280 pp." title="Bats of the Palearctic Region: a taxonomic and biogegraphic review. Pp" type="book chapter" volumeTitle="Proceedings of the VIIIth European Bat Research Conference. Volume I. Approaches to Biogeography and Ecology of Bats" year="2000">Horáček et al. 2000: 100</bibRefCitation>
</treatmentCitation>
[partim];
<treatmentCitation author="SIMMONS N. B." box="[685,883,787,809]" page="353" pageId="15" pageNumber="84" year="2005">
<bibRefCitation author="SIMMONS N. B." bookContentInfo="743 pp." box="[685,883,787,809]" editor="WILSON D. E. &amp; REEDER D. M." journalOrPublisher="The Johns Hopkins University Press, Baltimore" pageId="15" pageNumber="84" pagination="312 - 529" refId="ref15002" refString="SIMMONS N. B., 2005: Order Chiroptera. Pp.: 312 - 529. In: WILSON D. E. &amp; REEDER D. M. (eds.): Mammal Species of the World. A Taxonomic and Geographic Reference. Third Edition. Volume 1. The Johns Hopkins University Press, Baltimore, xxxv + 743 pp." title="Order Chiroptera. Pp" type="book chapter" volumeTitle="Mammal Species of the World. A Taxonomic and Geographic Reference. Third Edition. Volume 1" year="2005">Simmons 2005: 353</bibRefCitation>
</treatmentCitation>
[partim];
<treatmentCitation author="AULAGNIER S. &amp; HAFFNER P. &amp; MITCHELL- JONES A. J. &amp; MOUTOU F. &amp; ZIMA J." page="72" pageId="15" pageNumber="84" year="2008">
<bibRefCitation author="AULAGNIER S. &amp; HAFFNER P. &amp; MITCHELL- JONES A. J. &amp; MOUTOU F. &amp; ZIMA J." bookContentInfo="271 pp." journalOrPublisher="Delachaux et Niestle SA, Paris" pageId="15" pageNumber="84" refId="ref13190" refString="AULAGNIER S., HAFFNER P., MITCHELL- JONES A. J., MOUTOU F. &amp; ZIMA J., 2008: Guide des mammiferes d ' Europe, d ' Afrique du Nord et du Moyen-Orient. Delachaux et Niestle SA, Paris, 271 pp." title="Guide des mammiferes d ' Europe, d ' Afrique du Nord et du Moyen-Orient" type="book" year="2008">Aulagnier et al. 2008: 72</bibRefCitation>
</treatmentCitation>
[partim].
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="15.[117,1138,761,898]" box="[117,791,845,867]" pageId="15" pageNumber="84">
<treatmentCitationGroup box="[117,791,845,867]" pageId="15" pageNumber="84">
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<taxonomicName authorityName="Lacepede" authorityYear="1799" box="[117,238,845,866]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="84" phylum="Chordata" rank="genus">Rhinolophus</taxonomicName>
clivosus clivosus
</emphasis>
Cretzchmar, 1828:
<treatmentCitation author="QUMSIYEH M. B." box="[595,787,845,867]" page="32" pageId="15" pageNumber="84" year="1985">
<bibRefCitation author="QUMSIYEH M. B." box="[595,787,845,867]" journalOrPublisher="Special Publications, The Texas Tech University" pageId="15" pageNumber="84" pagination="1 - 102" part="23" refId="ref14821" refString="QUMSIYEH M. B., 1985: The bats of Egypt. Special Publications, The Texas Tech University, 23: 1 - 102." title="The bats of Egypt" type="journal article" year="1985">Qumsiyeh 1985: 32</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="15.[117,1138,761,898]" box="[117,1056,876,898]" pageId="15" pageNumber="84">
<treatmentCitationGroup box="[117,1056,876,898]" pageId="15" pageNumber="84">
<emphasis box="[117,471,876,897]" italics="true" pageId="15" pageNumber="84">
<taxonomicName authorityName="Lacepede" authorityYear="1799" box="[117,238,876,897]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="84" phylum="Chordata" rank="genus">Rhinolophus</taxonomicName>
clivosus brachygnathus
</emphasis>
<bibRefCitation author="ANDERSEN K." box="[475,631,876,898]" journalOrPublisher="Annals and Magazine of Natural History" pageId="15" pageNumber="84" pagination="70 - 76" part="15" refId="ref13074" refString="ANDERSEN K., 1905 a: Further descriptions of new Rhinolophi from Africa. Annals and Magazine of Natural History, Series 7, 15: 70 - 76." title="Further descriptions of new Rhinolophi from Africa" type="journal article" year="1905">Andersen, 1905</bibRefCitation>
:
<treatmentCitation author="KOOPMAN K. F." box="[640,828,876,898]" page="54" pageId="15" pageNumber="84" year="1994">
<bibRefCitation author="KOOPMAN K. F." bookContentInfo="217 pp." box="[640,828,876,898]" editor="NIETHAMMER J. &amp; SCHLIEMANN H. &amp; STARCK D." journalOrPublisher="Walter de Gruyter, Berlin &amp; New York" pageId="15" pageNumber="84" refId="ref14513" refString="KOOPMAN K. F., 1994: Chiroptera: Systematics. In: NIETHAMMER J., SCHLIEMANN H. &amp; STARCK D. (eds.): Handbook of Zoology. Volume VIII. Mammalia. Walter de Gruyter, Berlin &amp; New York, 217 pp." title="Chiroptera: Systematics" type="book" volumeTitle="Handbook of Zoology. Volume VIII. Mammalia" year="1994">Koopman 1994: 54</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="CSORBA G. &amp; UJHELYI P. &amp; THOMAS N." box="[838,1052,876,898]" page="35" pageId="15" pageNumber="84" year="2003">
<bibRefCitation author="CSORBA G. &amp; UJHELYI P. &amp; THOMAS N." bookContentInfo="160 pp." box="[838,1052,876,898]" journalOrPublisher="Alana Books, Bishop's Castle" pageId="15" pageNumber="84" refId="ref13634" refString="CSORBA G., UJHELYI P. &amp; THOMAS N., 2003: Horseshoe Bats of the World (Chiroptera: Rhinolophidae). Alana Books, Bishop's Castle, xxxii + 160 pp." title="Horseshoe Bats of the World (Chiroptera: Rhinolophidae)" type="book" year="2003">Csorba et al. 2003: 35</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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TYPE MATERIAL.
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<typeStatus box="[267,360,922,943]" pageId="15" pageNumber="84">Holotype</typeStatus>
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:
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(
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, field No. pb2124 [S+A]), Wadi
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,
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<quantity box="[959,1009,922,944]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="84" unit="km" value="6.0">6 km</quantity>
Sof
</locationDeviation>
<collectingRegion box="[1061,1134,922,944]" country="Libya" name="Darnah" pageId="15" pageNumber="84">Darnah</collectingRegion>
,
<date box="[117,252,948,970]" pageId="15" pageNumber="84" value="2002-05-15">
<collectingDate box="[117,252,948,970]" pageId="15" pageNumber="84" value="2002-05-15">15 May 2002</collectingDate>
</date>
, leg.
<collectorName box="[309,425,948,970]" pageId="15" pageNumber="84">M. Andreas</collectorName>
,
<collectorName box="[438,527,948,970]" pageId="15" pageNumber="84">P. Benda</collectorName>
, V.
<location LSID="urn:lsid:plazi:treatment:03EF87A4FFEE7D0AFF97FD28FE22789A:8E996069FFEE7D10FDD5FC59FCDA7ABB" box="[567,865,948,970]" name="Hanak, A. Reiter &amp; M. Uhrin" pageId="15" pageNumber="84" stateProvince="Darnah">Hanák, A. Reiter &amp; M. Uhrin</location>
.
</materialsCitation>
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<typeStatus box="[897,1000,949,970]" pageId="15" pageNumber="84">Paratypes</typeStatus>
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:
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<specimenCount box="[1041,1071,948,970]" pageId="15" pageNumber="84" type="adult">ad.</specimenCount>
(
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, field No. pb2104 [S+A]),
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,
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<collectingDate box="[539,671,975,997]" pageId="15" pageNumber="84" value="2002-05-12">12 May 2002</collectingDate>
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, leg.
<collectorName box="[724,839,975,997]" pageId="15" pageNumber="84">M. Andreas</collectorName>
,
<collectorName box="[850,938,975,997]" pageId="15" pageNumber="84">P. Benda</collectorName>
, V.
<location LSID="urn:lsid:plazi:treatment:03EF87A4FFEE7D0AFF97FD28FE22789A:8E996069FFEE7D10FC2DFC22FF557D71" municipality="Al Burdi" name="Hanak, A. Reiter &amp; M. Uhrin" pageId="15" pageNumber="84">Hanák, A. Reiter &amp; M. Uhrin</location>
;
</materialsCitation>
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<specimenCount box="[295,325,1002,1024]" pageId="15" pageNumber="84" type="adult">ad.</specimenCount>
(
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<collectingRegion box="[785,858,1002,1024]" country="Libya" name="Darnah" pageId="15" pageNumber="84">Darnah</collectingRegion>
,
<locationDeviation bearing="180.0" box="[870,969,1002,1024]" distance="6000.0" pageId="15" pageNumber="84">
<quantity box="[870,921,1002,1024]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="84" unit="km" value="6.0">6 km</quantity>
Sof
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<collectingRegion box="[977,1050,1002,1024]" country="Libya" name="Darnah" pageId="15" pageNumber="84">Darnah</collectingRegion>
,
<date pageId="15" pageNumber="84" value="2002-05-15">
<collectingDate pageId="15" pageNumber="84" value="2002-05-15">15 May 2002</collectingDate>
</date>
, leg.
<collectorName box="[219,333,1028,1050]" pageId="15" pageNumber="84">M. Andreas</collectorName>
,
<collectorName box="[343,430,1028,1050]" pageId="15" pageNumber="84">P. Benda</collectorName>
, V.
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;
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<specimenCount box="[803,832,1028,1050]" pageId="15" pageNumber="84" type="adult">ad.</specimenCount>
(
<specimenCode box="[844,965,1029,1050]" pageId="15" pageNumber="84">NMP 49882</specimenCode>
, field No. pb2127 [S+A]), Wadi
<collectingRegion box="[256,329,1055,1077]" country="Libya" name="Darnah" pageId="15" pageNumber="84">Darnah</collectingRegion>
,
<locationDeviation bearing="180.0" box="[340,450,1055,1077]" distance="10000.0" pageId="15" pageNumber="84">
<quantity box="[340,402,1055,1077]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="84" unit="km" value="10.0">10 km</quantity>
Sof
</locationDeviation>
<collectingRegion box="[457,532,1055,1077]" country="Libya" name="Darnah" pageId="15" pageNumber="84">Darnah</collectingRegion>
,
<date box="[536,667,1055,1077]" pageId="15" pageNumber="84" value="2002-05-16">
<collectingDate box="[536,667,1055,1077]" pageId="15" pageNumber="84" value="2002-05-16">16 May 2002</collectingDate>
</date>
, leg.
<collectorName box="[721,836,1055,1077]" pageId="15" pageNumber="84">M. Andreas</collectorName>
,
<collectorName box="[848,936,1055,1077]" pageId="15" pageNumber="84">P. Benda</collectorName>
, V.
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;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2960360304" collectingDate="2002-05-20" collectionCode="NMP" collectorName="M. Andreas &amp; P. Benda" location="Hanak, A. Reiter &amp; M. Uhrin" municipality="Wadi Al Kuf" pageId="15" pageNumber="84" specimenCode="NMP 49915" specimenCount="2" specimenCount-adult="1" specimenCount-female="1">
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<specimenCount box="[286,315,1082,1104]" pageId="15" pageNumber="84" type="adult">ad.</specimenCount>
(
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, field No. pb2163 [S+A]),
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,
<date box="[839,969,1082,1103]" pageId="15" pageNumber="84" value="2002-05-20">
<collectingDate box="[839,969,1082,1103]" pageId="15" pageNumber="84" value="2002-05-20">20 May 2002</collectingDate>
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, leg.
<collectorName box="[1020,1134,1082,1104]" pageId="15" pageNumber="84">M. Andreas</collectorName>
,
<collectorName box="[117,204,1108,1130]" pageId="15" pageNumber="84">P. Benda</collectorName>
, V.
<location LSID="urn:lsid:plazi:treatment:03EF87A4FFEE7D0AFF97FD28FE22789A:8E996069FFEE7D10FF0DFBB9FDB07D1B" box="[239,523,1108,1130]" municipality="Wadi Al Kuf" name="Hanak, A. Reiter &amp; M. Uhrin" pageId="15" pageNumber="84">Hanák, A. Reiter &amp; M. Uhrin</location>
.
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</paragraph>
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<materialsCitation ID-GBIF-Occurrence="2960360301" country="Libya" latitude="32.701668" location="Wadi Darnah" longLatPrecision="20" longitude="22.61111" pageId="15" pageNumber="84" specimenCount="1" stateProvince="Wadi Darnah">
TYPE LOCALITY.
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,
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,
<geoCoordinate box="[597,745,1154,1179]" degrees="22" direction="east" minutes="36" orientation="longitude" pageId="15" pageNumber="84" precision="15" seconds="40" value="22.61111">22° 36 40” E</geoCoordinate>
,
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ca.
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Sof Darnah
</locationDeviation>
,
<collectingRegion box="[995,1134,1155,1179]" country="Libya" name="Darnah" pageId="15" pageNumber="84">Darnah Dist.</collectingRegion>
, Cyrenaica,
<collectingCountry box="[239,305,1184,1208]" name="Libya" pageId="15" pageNumber="84">Libya</collectingCountry>
.
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</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="89" pageId="15" pageNumber="84" type="description">
<paragraph blockId="15.[117,1138,1154,1574]" pageId="15" pageNumber="84">
DESCRIPTION.
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<emphasis box="[262,502,1213,1237]" italics="true" pageId="15" pageNumber="84">Rhinolophus horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[512,573,1214,1237]" pageId="15" pageNumber="84" rank="species">sp. n.</taxonomicNameLabel>
is a medium-sized horseshoe bat, in most respects similar to the medium-sized forms of
<emphasis box="[527,543,1243,1266]" italics="true" pageId="15" pageNumber="84">R</emphasis>
.
<emphasis box="[558,644,1243,1267]" italics="true" pageId="15" pageNumber="84">clivosus</emphasis>
Cretzschmar, 1828 from the Middle East and north-eastern Africa, including the structure and relative size of the nose-leaf. Forearm length
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, ear length
<quantity box="[351,507,1301,1325]" metricMagnitude="-2" metricUnit="m" metricValue="2.175" metricValueMax="2.27" metricValueMin="2.08" pageId="15" pageNumber="84" unit="mm" value="21.75" valueMax="22.7" valueMin="20.8">20.822.7 mm</quantity>
, horseshoe width
<quantity box="[691,821,1301,1325]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" metricValueMax="7.5" metricValueMin="6.9" pageId="15" pageNumber="84" unit="mm" value="7.2" valueMax="7.5" valueMin="6.9">6.97.5 mm</quantity>
, condylocanine lengthof skull
<quantity box="[117,274,1330,1354]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" metricValueMax="1.81" metricValueMin="1.75" pageId="15" pageNumber="84" unit="mm" value="17.8" valueMax="18.1" valueMin="17.5">17.518.1 mm</quantity>
, length of the upper tooth-row
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.
</paragraph>
<paragraph blockId="15.[117,1138,1154,1574]" pageId="15" pageNumber="84">
The horseshoe of
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<emphasis box="[336,352,1367,1390]" italics="true" pageId="15" pageNumber="84">R</emphasis>
.
<emphasis box="[364,457,1367,1391]" italics="true" pageId="15" pageNumber="84">horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[464,520,1367,1390]" pageId="15" pageNumber="84" rank="species">sp. n.</taxonomicNameLabel>
is relatively narrow (
<figureCitation box="[746,810,1367,1391]" captionStart="Fig" captionStartId="16.[141,172,766,785]" captionTargetBox="[345,926,173,730]" captionTargetId="figure-121@16.[125,1144,169,736]" captionTargetPageId="16" captionText="Fig. 7. Nose-leaf of Rhinolophus horaceki sp. n. (NMP 49880, holotype), lateral (left) and frontal (right) views. The frontal view is depicted without hairs. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4247732" httpUri="https://zenodo.org/record/4247732/files/figure.png" pageId="15" pageNumber="84">Fig. 7</figureCitation>
), the connecting process of the nose-leaf is high and rounded, the sella is constricted in the middle, tip of the sella is pointed, lancet is hairy and regularly triangular in shape. One medial groove is present in the lower lip.
</paragraph>
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Skull is relatively wide (LaZ
<quantity box="[486,651,1462,1486]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" metricValueMax="1.21" metricValueMin="1.1099999999999999" pageId="15" pageNumber="84" unit="mm" value="11.6" valueMax="12.1" valueMin="11.1">11.112.1 mm</quantity>
; LaZ/LCc
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<geoCoordinate box="[838,916,1461,1485]" degrees="0.619" direction="west" orientation="longitude" pageId="15" pageNumber="84" precision="55" value="-0.619">0.619</geoCoordinate>
), rostral part of the skull inluding the nasal swellings is massive (LaInf
<quantity box="[686,819,1491,1515]" metricMagnitude="-3" metricUnit="m" metricValue="5.699999999999999" metricValueMax="6.1" metricValueMin="5.3" pageId="15" pageNumber="84" unit="mm" value="5.699999999999999" valueMax="6.1" valueMin="5.3">5.36.1 mm</quantity>
; CC
<quantity box="[876,1009,1491,1515]" metricMagnitude="-3" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.3" metricValueMin="5.6" pageId="15" pageNumber="84" unit="mm" value="5.949999999999999" valueMax="6.3" valueMin="5.6">5.66.3 mm</quantity>
; LaInf/CM
<superScript attach="right" box="[1130,1138,1489,1503]" fontSize="6" pageId="15" pageNumber="84">3</superScript>
<geoCoordinate box="[117,179,1520,1544]" degrees="0.762" direction="north" orientation="latitude" pageId="15" pageNumber="84" precision="55" value="0.762">0.762</geoCoordinate>
<geoCoordinate box="[179,254,1520,1544]" degrees="0.786" direction="west" orientation="longitude" pageId="15" pageNumber="84" precision="55" value="-0.786">0.786</geoCoordinate>
; CM
<superScript attach="left" box="[305,313,1519,1533]" fontSize="6" pageId="15" pageNumber="84">3</superScript>
/LCc
<geoCoordinate box="[372,434,1520,1544]" degrees="0.454" direction="north" orientation="latitude" pageId="15" pageNumber="84" precision="55" value="0.454">0.454</geoCoordinate>
<geoCoordinate box="[434,509,1520,1544]" degrees="0.468" direction="west" orientation="longitude" pageId="15" pageNumber="84" precision="55" value="-0.468">0.468</geoCoordinate>
), relatively long and wide (CC/CM
<superScript attach="left" box="[881,889,1519,1533]" fontSize="6" pageId="15" pageNumber="84">3</superScript>
<geoCoordinate box="[894,956,1520,1544]" degrees="0.787" direction="north" orientation="latitude" pageId="15" pageNumber="84" precision="55" value="0.787">0.787</geoCoordinate>
<geoCoordinate box="[956,1030,1520,1544]" degrees="0.821" direction="west" orientation="longitude" pageId="15" pageNumber="84" precision="55" value="-0.821">0.821</geoCoordinate>
; CP
<superScript attach="left" box="[1073,1081,1519,1533]" fontSize="6" pageId="15" pageNumber="84">4</superScript>
/CM
<superScript attach="left" box="[1130,1138,1519,1533]" fontSize="6" pageId="15" pageNumber="84">3</superScript>
<geoCoordinate box="[117,179,1550,1574]" degrees="0.423" direction="north" orientation="latitude" pageId="15" pageNumber="84" precision="55" value="0.423">0.423</geoCoordinate>
<geoCoordinate box="[179,254,1550,1574]" degrees="0.441" direction="west" orientation="longitude" pageId="15" pageNumber="84" precision="55" value="-0.441">0.441</geoCoordinate>
). Sagittal crest ismedium developed, infraorbital foramen islarge and infraorbital bar is long and thin (
<figureCitation box="[303,368,881,905]" captionStart="Fig" captionStartId="16.[149,180,1532,1551]" captionTargetBox="[152,1116,1145,1497]" captionTargetId="figure-119@16.[125,1144,1133,1505]" captionTargetPageId="16" captionText="Fig. 8. Skull in lateral view (left) and rostral part of the skull in dorsal view (right) of Rhinolophus horaceki sp. n. (NMP 49880, holotype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4247734" httpUri="https://zenodo.org/record/4247734/files/figure.png" pageId="16" pageNumber="85">Fig. 8</figureCitation>
). Nasal swellings are rather undeveloped, the posterior median swellings are equally long to the anterior swellings, the anterior lateral swellings are almost equal to the anterior median swellings (
<figureCitation box="[415,480,940,964]" captionStart="Fig" captionStartId="16.[149,180,1532,1551]" captionTargetBox="[152,1116,1145,1497]" captionTargetId="figure-119@16.[125,1144,1133,1505]" captionTargetPageId="16" captionText="Fig. 8. Skull in lateral view (left) and rostral part of the skull in dorsal view (right) of Rhinolophus horaceki sp. n. (NMP 49880, holotype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4247734" httpUri="https://zenodo.org/record/4247734/files/figure.png" pageId="16" pageNumber="85">Fig. 8</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4247732" ID-Zenodo-Dep="4247732" httpUri="https://zenodo.org/record/4247732/files/figure.png" pageId="16" pageNumber="85" startId="16.[141,172,766,785]" targetBox="[345,926,173,730]" targetPageId="16">
<paragraph blockId="16.[141,1129,766,809]" pageId="16" pageNumber="85">
Fig. 7. Nose-leaf of
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[313,501,766,785]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="85" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[313,501,766,785]" italics="true" pageId="16" pageNumber="85">Rhinolophus horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[506,552,766,785]" pageId="16" pageNumber="85" rank="species">sp. n.</taxonomicNameLabel>
(NMP 49880, holotype), lateral (left) and frontal (right) views. The frontal view is depicted without hairs. Scale bar 5 mm.
</paragraph>
</caption>
<paragraph blockId="16.[125,1146,881,1059]" lastBlockId="17.[117,1138,841,986]" lastPageId="17" lastPageNumber="86" pageId="16" pageNumber="85">
The teeth are relatively massive (
<figureCitation box="[518,595,976,1000]" captionStart="Fig" captionStartId="17.[174,205,726,745]" captionTargetBox="[147,1107,174,695]" captionTargetId="figure-155@17.[147,1107,174,695]" captionTargetPageId="17" captionText="Fig. 9. Occlusal view on the mesial part of the left upper tooth-row (I1M1) of Rhinolophus horaceki sp. n. (NMP 49880, holotype). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4247736" httpUri="https://zenodo.org/record/4247736/files/figure.png" pageId="16" pageNumber="85">Figs. 9</figureCitation>
,
<figureCitation box="[609,638,976,1000]" captionStart="Fig" captionStartId="17.[151,182,1534,1553]" captionTargetBox="[147,1107,1053,1506]" captionTargetId="figure-156@17.[147,1107,1053,1506]" captionTargetPageId="17" captionText="Fig. 10. Occlusal view on the right lower unicuspidal teeth (CP4) of Rhinolophus horaceki sp. n. (NMP 49880, holotype). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4247738" httpUri="https://zenodo.org/record/4247738/files/figure.png" pageId="16" pageNumber="85">10</figureCitation>
); upper molars are relatively wide (LaM
<superScript attach="left" box="[1082,1090,974,988]" fontSize="6" pageId="16" pageNumber="85">1</superScript>
/LM
<superScript attach="left" box="[1137,1145,974,988]" fontSize="6" pageId="16" pageNumber="85">1</superScript>
<geoCoordinate box="[125,186,1005,1029]" degrees="1.559" direction="north" orientation="latitude" pageId="16" pageNumber="85" precision="55" value="1.559">1.559</geoCoordinate>
<geoCoordinate box="[186,261,1005,1029]" degrees="1.674" direction="west" orientation="longitude" pageId="16" pageNumber="85" precision="55" value="-1.674">1.674</geoCoordinate>
; LaM
<superScript attach="left" box="[326,334,1004,1018]" fontSize="6" pageId="16" pageNumber="85">3</superScript>
/LM
<superScript attach="left" box="[381,389,1004,1018]" fontSize="6" pageId="16" pageNumber="85">3</superScript>
<geoCoordinate box="[397,457,1005,1029]" degrees="1.691" direction="north" orientation="latitude" pageId="16" pageNumber="85" precision="55" value="1.691">1.691</geoCoordinate>
<geoCoordinate box="[457,534,1005,1029]" degrees="1.735" direction="west" orientation="longitude" pageId="16" pageNumber="85" precision="55" value="-1.735">1.735</geoCoordinate>
), large upper premolars (P
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) are relatively wide and mesio-distally short (LP
<superScript attach="left" box="[355,363,1033,1047]" fontSize="6" pageId="16" pageNumber="85">4</superScript>
/LaP
<superScript attach="left" box="[413,421,1033,1047]" fontSize="6" pageId="16" pageNumber="85">4</superScript>
<geoCoordinate box="[430,492,1034,1058]" degrees="0.569" direction="north" orientation="latitude" pageId="16" pageNumber="85" precision="55" value="0.569">0.569</geoCoordinate>
<geoCoordinate box="[492,568,1034,1058]" degrees="0.610" direction="west" orientation="longitude" pageId="16" pageNumber="85" precision="55" value="-0.61">0.610</geoCoordinate>
), with relatively very shallow concavity in the distal margin of talon (LP
<superScript attach="left" box="[330,349,841,867]" fontSize="6" pageId="17" pageNumber="86">43</superScript>
/LP
<superScript attach="right" box="[390,410,841,867]" fontSize="6" pageId="17" pageNumber="86">41</superScript>
<geoCoordinate box="[418,480,843,867]" degrees="0.524" direction="north" orientation="latitude" pageId="17" pageNumber="86" precision="55" value="0.524">0.524</geoCoordinate>
<geoCoordinate box="[480,557,843,867]" degrees="0.697" direction="west" orientation="longitude" pageId="17" pageNumber="86" precision="55" value="-0.697">0.697</geoCoordinate>
). Large lower premolars (P
<subScript attach="left" box="[849,857,855,869]" fontSize="6" pageId="17" pageNumber="86">4</subScript>
) are absolutely very large (LP
<subScript attach="left" box="[157,165,884,898]" fontSize="6" pageId="17" pageNumber="86">4</subScript>
<quantity box="[172,329,872,896]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.39" metricValueMin="1.31" pageId="17" pageNumber="86" unit="mm" value="1.35" valueMax="1.39" valueMin="1.31">1.311.39 mm</quantity>
) as well as very large in relation to the size of smaller lower premolars (P
<subScript attach="left" box="[1121,1129,884,898]" fontSize="6" pageId="17" pageNumber="86">2</subScript>
) (LP
<subScript attach="right" box="[157,165,913,927]" fontSize="6" pageId="17" pageNumber="86">2</subScript>
×LaP
<subScript attach="left" box="[222,230,913,927]" fontSize="6" pageId="17" pageNumber="86">2</subScript>
/LP
<subScript attach="left" box="[268,276,913,927]" fontSize="6" pageId="17" pageNumber="86">4</subScript>
×LaP
<subScript attach="left" box="[334,342,913,927]" fontSize="6" pageId="17" pageNumber="86">4</subScript>
<geoCoordinate box="[346,407,901,925]" degrees="0.397" direction="north" orientation="latitude" pageId="17" pageNumber="86" precision="55" value="0.397">0.397</geoCoordinate>
<geoCoordinate box="[408,484,901,925]" degrees="0.466" direction="west" orientation="longitude" pageId="17" pageNumber="86" precision="55" value="-0.466">0.466</geoCoordinate>
). The minutesecond lower premolar (P
<subScript attach="left" box="[891,899,913,927]" fontSize="6" pageId="17" pageNumber="86">3</subScript>
) is frequently missing, while the minute first upper premolar (P
<superScript attach="left" box="[548,556,929,943]" fontSize="6" pageId="17" pageNumber="86">2</superScript>
) is frequently present (LP
<superScript attach="left" box="[837,845,929,943]" fontSize="6" pageId="17" pageNumber="86">2</superScript>
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); if present, P
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lies out of the the tooth-row, P
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and P
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are in contact.
</paragraph>
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<paragraph blockId="16.[149,1120,1532,1576]" pageId="16" pageNumber="85">
Fig. 8. Skull in lateral view (left) and rostral part of the skull in dorsal view (right) of
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[880,1069,1532,1551]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="85" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[880,1069,1532,1551]" italics="true" pageId="16" pageNumber="85">Rhinolophus horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1074,1120,1533,1552]" pageId="16" pageNumber="85" rank="species">sp. n.</taxonomicNameLabel>
(NMP 49880, holotype). Scale bar 5 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4247736" ID-Zenodo-Dep="4247736" httpUri="https://zenodo.org/record/4247736/files/figure.png" pageId="17" pageNumber="86" startId="17.[174,205,726,745]" targetBox="[147,1107,174,695]" targetPageId="17">
<paragraph blockId="17.[174,1082,724,769]" pageId="17" pageNumber="86">
Fig. 9. Occlusal view on the mesial part of the left upper tooth-row (I
<superScript attach="left" box="[765,771,724,736]" fontSize="5" pageId="17" pageNumber="86">1</superScript>
M
<superScript attach="left" box="[800,806,724,736]" fontSize="5" pageId="17" pageNumber="86">1</superScript>
) of
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[842,1030,726,745]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="17" pageNumber="86" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[842,1030,726,745]" italics="true" pageId="17" pageNumber="86">Rhinolophus horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1035,1082,726,745]" pageId="17" pageNumber="86" rank="species">sp. n.</taxonomicNameLabel>
(NMP 49880, holotype). Scale bar 1 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4247738" ID-Zenodo-Dep="4247738" httpUri="https://zenodo.org/record/4247738/files/figure.png" pageId="17" pageNumber="86" startId="17.[151,182,1534,1553]" targetBox="[147,1107,1053,1506]" targetPageId="17">
<paragraph blockId="17.[151,1104,1534,1577]" pageId="17" pageNumber="86">
Fig. 10. Occlusal view on the right lower unicuspidal teeth (CP
<subScript attach="left" box="[701,707,1543,1555]" fontSize="5" pageId="17" pageNumber="86">4</subScript>
) of
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[743,931,1534,1553]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="17" pageNumber="86" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[743,931,1534,1553]" italics="true" pageId="17" pageNumber="86">Rhinolophus horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[936,983,1534,1553]" pageId="17" pageNumber="86" rank="species">sp. n.</taxonomicNameLabel>
(NMP 49880, holotype). Scale bar 1 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4247740" ID-Zenodo-Dep="4247740" httpUri="https://zenodo.org/record/4247740/files/figure.png" pageId="18" pageNumber="87" startId="18.[130,161,608,627]" targetBox="[138,1131,174,580]" targetPageId="18">
<paragraph blockId="18.[130,1140,608,748]" pageId="18" pageNumber="87">
Fig. 11. Baculum preparations of
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[417,605,608,627]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="87" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[417,605,608,627]" italics="true" pageId="18" pageNumber="87">Rhinolophus horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[610,657,609,628]" pageId="18" pageNumber="87" rank="species">sp. n.</taxonomicNameLabel>
and two comparative samples of the
<emphasis italics="true" pageId="18" pageNumber="87">ferrumequinum/clivosus</emphasis>
group (for other examined samples of this group see
<bibRefCitation author="HANAK V." box="[642,748,632,651]" journalOrPublisher="Vestnik Ceskoslovenske Spolecnosti Zoologicke" pageId="18" pageNumber="87" pagination="315 - 327" part="33" refId="ref13921" refString="HANAK V., 1969: Zur Kenntnis von Rhinolophus bocharicus Kastchenko et Akimov, 1917 (Mammalia: Chiroptera). Vestnik Ceskoslovenske Spolecnosti Zoologicke, 33: 315 - 327." title="Zur Kenntnis von Rhinolophus bocharicus Kastchenko et Akimov, 1917 (Mammalia: Chiroptera)" type="journal article" year="1969">Hanák 1969</bibRefCitation>
,
<bibRefCitation author="STRELKOV P. P." box="[756,879,632,652]" journalOrPublisher="Zoologiceskij Zurnal" pageId="18" pageNumber="87" pagination="893 - 906" part="50" refId="ref15114" refString="STRELKOV P. P., 1971: Bol'sie (Rhinolophus ferrumequinum) i buharskie (Rh. bocharicus) podkovonosy (Chiroptera) iz pustyni Karakumy [Greater (Rhinolophus ferrumequinum) and Bukhara (Rh. bocharicus) horseshoe bats (Chiroptera) from the Karakum Desert]. Zoologiceskij Zurnal, 50: 893 - 906 (in Russian, with a summary in English)." title="Bol'sie (Rhinolophus ferrumequinum) i buharskie (Rh. bocharicus) podkovonosy (Chiroptera) iz pustyni Karakumy [Greater (Rhinolophus ferrumequinum) and Bukhara (Rh. bocharicus) horseshoe bats (Chiroptera) from the Karakum Desert]" type="journal article" year="1971">Strelkov 1971</bibRefCitation>
,
<bibRefCitation author="THOMAS N. M." bookContentInfo="211 pp." box="[887,1006,632,652]" journalOrPublisher="Harrison Zoological Museum, Seveoaks" pageId="18" pageNumber="87" refId="ref15380" refString="THOMAS N. M., 1997: A Systematic Review of Selected Afro-Asiatic Rhinolophidae (Mammalia: Chiroptera): An Evaluation of Taxonomic Methodologies. Unpubl. PhD Thesis. Harrison Zoological Museum, Seveoaks, v + 211 pp." title="A Systematic Review of Selected Afro-Asiatic Rhinolophidae (Mammalia: Chiroptera): An Evaluation of Taxonomic Methodologies. Unpubl" type="book" year="1997">Thomas 1997</bibRefCitation>
, and
<bibRefCitation author="CSORBA G. &amp; UJHELYI P. &amp; THOMAS N." bookContentInfo="160 pp." journalOrPublisher="Alana Books, Bishop's Castle" pageId="18" pageNumber="87" refId="ref13634" refString="CSORBA G., UJHELYI P. &amp; THOMAS N., 2003: Horseshoe Bats of the World (Chiroptera: Rhinolophidae). Alana Books, Bishop's Castle, xxxii + 160 pp." title="Horseshoe Bats of the World (Chiroptera: Rhinolophidae)" type="book" year="2003">Csorba et al. 2003</bibRefCitation>
). Explanations: a
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[372,471,656,676]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="87" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[372,385,657,675]" italics="true" pageId="18" pageNumber="87">R</emphasis>
.
<emphasis box="[396,471,656,675]" italics="true" pageId="18" pageNumber="87">horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[476,524,657,676]" pageId="18" pageNumber="87" rank="species">sp. n.</taxonomicNameLabel>
, Libya (Cyrenaica), Wadi Darnah, NMP 49879 (left lateral view, right ventral view); b
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[321,420,680,700]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="87" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[321,334,681,699]" italics="true" pageId="18" pageNumber="87">R</emphasis>
.
<emphasis box="[345,420,680,699]" italics="true" pageId="18" pageNumber="87">horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[425,472,681,700]" pageId="18" pageNumber="87" rank="species">sp. n.</taxonomicNameLabel>
, Libya (Cyrenaica), Al Burdi, NMP 49861 (left lateral view, right dorsal view); c
<emphasis box="[244,257,705,723]" italics="true" pageId="18" pageNumber="87">R</emphasis>
.
<emphasis box="[268,337,704,723]" italics="true" pageId="18" pageNumber="87">clivosus</emphasis>
, Egypt, Cairo, Abu Rawash, ZFMK 59.292 (left lateral view, right ventral view); d
<emphasis box="[1099,1112,705,723]" italics="true" pageId="18" pageNumber="87">R</emphasis>
.
<emphasis box="[167,299,729,748]" italics="true" pageId="18" pageNumber="87">ferrumequinum</emphasis>
, Syria, As Salihiyyah, NMP 48974 (left lateral view, right dorsal view). Scale bar 2 mm.
</paragraph>
</caption>
<paragraph blockId="18.[125,1145,803,885]" lastBlockId="20.[125,1146,175,711]" lastPageId="20" lastPageNumber="89" pageId="18" pageNumber="87">
Baculum of
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[286,409,803,827]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="87" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[286,302,803,826]" italics="true" pageId="18" pageNumber="87">R</emphasis>
.
<emphasis box="[316,409,803,827]" italics="true" pageId="18" pageNumber="87">horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[416,473,803,826]" pageId="18" pageNumber="87" rank="species">sp. n.</taxonomicNameLabel>
is a relativelly large bone, dorso-ventrally flattened in its distal two-thirds, creating a lancet-form shape, while its proximal epiphysis is massive and laterally bifurcated (
<figureCitation box="[246,323,861,885]" captionStart="Fig" captionStartId="18.[130,161,608,627]" captionTargetBox="[138,1131,174,580]" captionTargetId="figure-47@18.[125,1144,155,586]" captionTargetPageId="18" captionText="Fig. 11. Baculum preparations of Rhinolophus horaceki sp. n. and two comparative samples of the ferrumequinum/clivosus group (for other examined samples of this group see Hanák 1969, Strelkov 1971, Thomas 1997, and Csorba et al. 2003). Explanations: a R. horaceki sp. n., Libya (Cyrenaica), Wadi Darnah, NMP 49879 (left lateral view, right ventral view); b R. horaceki sp. n., Libya (Cyrenaica), Al Burdi, NMP 49861 (left lateral view, right dorsal view); c R. clivosus, Egypt, Cairo, Abu Rawash, ZFMK 59.292 (left lateral view, right ventral view); d R. ferrumequinum, Syria, As Salihiyyah, NMP 48974 (left lateral view, right dorsal view). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4247740" httpUri="https://zenodo.org/record/4247740/files/figure.png" pageId="18" pageNumber="87">Fig. 11</figureCitation>
). Total length of baculum
<quantity box="[597,726,861,885]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="3.9" metricValueMin="3.7" pageId="18" pageNumber="87" unit="mm" value="3.8" valueMax="3.9" valueMin="3.7">3.73.9 mm</quantity>
, largestwidth of the proximal epiphysis
<quantity box="[125,256,175,199]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="20" pageNumber="89" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.21.5 mm</quantity>
, largest (dorso-ventral) height of the proximal epiphysis
<quantity box="[863,928,175,199]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="89" unit="mm" value="1.0">1 mm</quantity>
, largest width of the lancet
<quantity box="[195,327,204,228]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="20" pageNumber="89" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.80.9 mm</quantity>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4247742" ID-Zenodo-Dep="4247742" box="[343,929,1554,1574]" httpUri="https://zenodo.org/record/4247742/files/figure.png" pageId="18" pageNumber="87" startId="18.[343,374,1554,1573]" targetBox="[316,953,943,1523]" targetPageId="18">
<paragraph blockId="18.[343,929,1554,1574]" box="[343,929,1554,1574]" pageId="18" pageNumber="87">
Fig. 12. Portrait of
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[506,694,1554,1573]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="87" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[506,694,1554,1573]" italics="true" pageId="18" pageNumber="87">Rhinolophus horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[699,746,1555,1574]" pageId="18" pageNumber="87" rank="species">sp. n.</taxonomicNameLabel>
(photo by A. Reiter).
</paragraph>
</caption>
<paragraph blockId="20.[125,1146,175,711]" pageId="20" pageNumber="89">
The dorsal pelage of
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[383,506,240,264]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="89" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[383,399,241,264]" italics="true" pageId="20" pageNumber="89">R</emphasis>
.
<emphasis box="[413,506,240,264]" italics="true" pageId="20" pageNumber="89">horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[513,571,241,264]" pageId="20" pageNumber="89" rank="species">sp. n.</taxonomicNameLabel>
is brown to brownish-grey, ventral pelage is greyish- -beige (
<figureCitation box="[208,289,269,294]" captionStart="Fig" captionStartId="18.[343,374,1554,1573]" captionTargetBox="[316,953,943,1523]" captionTargetId="figure-174@18.[316,953,940,1526]" captionTargetPageId="18" captionText="Fig. 12. Portrait of Rhinolophus horaceki sp. n. (photo by A. Reiter)." figureDoi="http://doi.org/10.5281/zenodo.4247742" httpUri="https://zenodo.org/record/4247742/files/figure.png" pageId="20" pageNumber="89">Fig. 12</figureCitation>
). Nose-leaf and ears are dark brown or dark greyish-brown, distal parts darker than the proximal. Wing membranes are dark brown or greyish-brown.
</paragraph>
<paragraph blockId="20.[125,1146,175,711]" pageId="20" pageNumber="89">
<emphasis bold="true" box="[125,223,335,359]" pageId="20" pageNumber="89">Genetics</emphasis>
. In the group of horseshoe bats of the
<emphasis box="[646,904,335,359]" italics="true" pageId="20" pageNumber="89">ferrumequinum/clivosus</emphasis>
complex (
<emphasis box="[1021,1037,336,359]" italics="true" pageId="20" pageNumber="89">R</emphasis>
.
<emphasis italics="true" pageId="20" pageNumber="89">ferrumequinum</emphasis>
(Schreber, 1774),
<emphasis box="[404,420,365,388]" italics="true" pageId="20" pageNumber="89">R</emphasis>
.
<emphasis box="[435,509,365,388]" italics="true" pageId="20" pageNumber="89">nippon</emphasis>
<bibRefCitation author="TEMMINCK C. J." bookContentInfo="45 pp." box="[516,696,364,388]" journalOrPublisher="C. C. Van Der Hoek, Leiden" pageId="20" pageNumber="89" refId="ref15256" refString="TEMMINCK C. J., 1835: Monographies de mammalogie, ou description de quelques genres de mammiferes, don't les especes ont ete observees dans les differens musees de l'Europe. Tome Second. Huitieme monographie sur le genre rhinolophe; Rhinolophus (Geoff. Cuv. Illig. Desm.), Vespertilio (Linn. Erxleb.), Noctilio (Kuhl.). C. C. Van Der Hoek, Leiden, 45 pp." title="Monographies de mammalogie, ou description de quelques genres de mammiferes, don't les especes ont ete observees dans les differens musees de l'Europe. Tome Second. Huitieme monographie sur le genre rhinolophe; Rhinolophus (Geoff. Cuv. Illig. Desm.), Vespertilio (Linn. Erxleb.), Noctilio (Kuhl.)" type="book" year="1835">Temminck, 1835</bibRefCitation>
and
<emphasis box="[749,765,365,388]" italics="true" pageId="20" pageNumber="89">R</emphasis>
.
<emphasis box="[780,866,364,388]" italics="true" pageId="20" pageNumber="89">clivosus</emphasis>
Cretzschmar, 1828 s.str.),
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[125,247,394,418]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="89" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[125,141,394,417]" italics="true" pageId="20" pageNumber="89">R</emphasis>
.
<emphasis box="[154,247,394,418]" italics="true" pageId="20" pageNumber="89">horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[254,311,394,417]" pageId="20" pageNumber="89" rank="species">sp. n.</taxonomicNameLabel>
shows a unique base position within the mitochondrial gene for cytochrome
<emphasis box="[1132,1145,394,418]" italics="true" pageId="20" pageNumber="89">b</emphasis>
(1140 bp) at 34 sites: 1071 (A+C), 36, 127, 378, 562, 750, 808, 972, 1107, 1134 (A+G), 117 (C+A), 5, 57, 190, 201, 285, 390, 468, 564, 730, 894, 969, 982, 1057 (C+T), 698, 907 (G+ A), 126, 459, 873 (T+C), 864, 1089 (A/C+T), 282, 462 (A/G+C), and 708 (A/C/G+T). With the
<emphasis box="[167,272,511,535]" italics="true" pageId="20" pageNumber="89">fumigatus</emphasis>
group (here,
<emphasis box="[424,440,512,535]" italics="true" pageId="20" pageNumber="89">R</emphasis>
.
<emphasis box="[456,561,511,535]" italics="true" pageId="20" pageNumber="89">fumigatus</emphasis>
<bibRefCitation author="RUPPELL E." box="[571,724,511,535]" journalOrPublisher="Museum Senckenbergianum" pageId="20" pageNumber="89" pagination="129 - 144" part="3" refId="ref14914" refString="RUPPELL E., 1842: Beschreibung mehrerer neuer Saugethiere in der Sammlung der senckenbergischen naturforschenden Gesellschaft befindlich. Museum Senckenbergianum, 3: 129 - 144." title="Beschreibung mehrerer neuer Saugethiere in der Sammlung der senckenbergischen naturforschenden Gesellschaft befindlich" type="journal article" year="1842">Rüppell, 1842</bibRefCitation>
and
<emphasis box="[782,798,512,535]" italics="true" pageId="20" pageNumber="89">R</emphasis>
.
<emphasis box="[814,953,511,535]" italics="true" pageId="20" pageNumber="89">hildebrandtii</emphasis>
<bibRefCitation author="PETERS W." box="[963,1101,511,535]" journalOrPublisher="Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften zu Berlin" pageId="20" pageNumber="89" pagination="194 - 209" part="1878" refId="ref14717" refString="PETERS W., 1878: Uber die von Hrn. J. M. Hildebrandt wahrend seiner letzen ostafrikanischen Reise gesammelten Saugethiere und Amphibien. Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften zu Berlin, 1878: 194 - 209." title="Uber die von Hrn. J. M. Hildebrandt wahrend seiner letzen ostafrikanischen Reise gesammelten Saugethiere und Amphibien" type="journal article" year="1878">Peters, 1878</bibRefCitation>
),
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="89" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[1122,1138,512,535]" italics="true" pageId="20" pageNumber="89">R</emphasis>
.
<emphasis box="[125,218,540,564]" italics="true" pageId="20" pageNumber="89">horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[227,286,541,564]" pageId="20" pageNumber="89" rank="species">sp. n.</taxonomicNameLabel>
shares unique base positions at four sites, which it does not share with bats of the
<emphasis box="[165,422,570,594]" italics="true" pageId="20" pageNumber="89">ferrumequinum/clivosus</emphasis>
complex: 57, 564 (T), 459, and 873 (C); only with
<emphasis box="[983,999,570,593]" italics="true" pageId="20" pageNumber="89">R</emphasis>
.
<emphasis box="[1013,1118,570,594]" italics="true" pageId="20" pageNumber="89">fumigatus</emphasis>
at two sites: 907 (A) and 969 (T); and only with
<emphasis box="[620,636,600,623]" italics="true" pageId="20" pageNumber="89">R</emphasis>
.
<emphasis box="[650,789,599,623]" italics="true" pageId="20" pageNumber="89">hildebrandtii</emphasis>
at three sites: 190, 1089 (T), and 697 (A). With the
<emphasis box="[320,578,628,652]" italics="true" pageId="20" pageNumber="89">ferrumequinum/clivosus</emphasis>
complex,
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[690,813,628,652]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="89" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[690,706,629,652]" italics="true" pageId="20" pageNumber="89">R</emphasis>
.
<emphasis box="[720,813,628,652]" italics="true" pageId="20" pageNumber="89">horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[820,878,629,652]" pageId="20" pageNumber="89" rank="species">sp. n.</taxonomicNameLabel>
shares unique base positions at seven sites, which it does not share with bats of the
<emphasis box="[760,865,658,682]" italics="true" pageId="20" pageNumber="89">fumigatus</emphasis>
group: 141, 591, 681 (A), 105 (C), 640, 835 (G), and 49 (T).
</paragraph>
<paragraph blockId="20.[125,1146,734,1295]" box="[125,1109,734,756]" pageId="20" pageNumber="89">
DIMENSIONS OF THE
<typeStatus box="[305,394,740,754]" pageId="20" pageNumber="89">HOLOTYPE</typeStatus>
(in millimetres).
<emphasis bold="true" box="[562,651,734,755]" pageId="20" pageNumber="89">External</emphasis>
: LC 60; LCd 34; LAt 18.4, LA 21.3; LaFE 6.9.
</paragraph>
<paragraph blockId="20.[125,1146,734,1295]" pageId="20" pageNumber="89">
<emphasis bold="true" box="[125,204,761,783]" pageId="20" pageNumber="89">Cranial</emphasis>
: LCr 20.16; LCc 17.48; LaZ 10.68;
<collectingRegion box="[561,594,761,782]" country="South Sudan" name="Lakes" pageId="20" pageNumber="89">LaI</collectingRegion>
2.42; LaInf 5.69; LaNc 8.43; LaM 9.32; ANc 6.17; ACr 7.54; LBT 3.07; CC 5.84; P
<superScript attach="right" box="[394,401,787,799]" fontSize="5" pageId="20" pageNumber="89">4</superScript>
P
<superScript attach="left" box="[414,473,787,809]" fontSize="5" pageId="20" pageNumber="89">46.48</superScript>
; M
<superScript attach="both" box="[504,511,787,799]" fontSize="5" pageId="20" pageNumber="89">3</superScript>
M
<superScript attach="left" box="[533,591,787,809]" fontSize="5" pageId="20" pageNumber="89">37.71</superScript>
; CM
<superScript attach="left" box="[639,697,787,809]" fontSize="5" pageId="20" pageNumber="89">37.42</superScript>
; M
<superScript attach="both" box="[729,736,787,799]" fontSize="5" pageId="20" pageNumber="89">1</superScript>
M
<superScript attach="left" box="[757,816,787,809]" fontSize="5" pageId="20" pageNumber="89">34.74</superScript>
; CP
<superScript attach="left" box="[855,914,787,809]" fontSize="5" pageId="20" pageNumber="89">43.22</superScript>
; LMd 13.24; ACo 3.28; IM
<subScript attach="left" box="[154,212,814,837]" fontSize="5" pageId="20" pageNumber="89">38.79</subScript>
; CM
<subScript attach="left" box="[259,317,814,837]" fontSize="5" pageId="20" pageNumber="89">38.03</subScript>
; M
<subScript attach="both" box="[348,355,825,837]" fontSize="5" pageId="20" pageNumber="89">1</subScript>
M
<subScript attach="left" box="[376,434,814,837]" fontSize="5" pageId="20" pageNumber="89">35.39</subScript>
; CP
<subScript attach="left" box="[473,531,814,837]" fontSize="5" pageId="20" pageNumber="89">42.94</subScript>
.
</paragraph>
<paragraph blockId="20.[125,1146,734,1295]" pageId="20" pageNumber="89">
<emphasis bold="true" box="[125,193,841,862]" pageId="20" pageNumber="89">Dental</emphasis>
: LCs 1.92; LaCs 1.53; LP
<superScript attach="left" box="[443,500,840,863]" fontSize="5" pageId="20" pageNumber="89">20.35</superScript>
; LP
<superScript attach="left" box="[537,607,840,863]" fontSize="5" pageId="20" pageNumber="89">411.50</superScript>
; LP
<superScript attach="left" box="[644,714,840,862]" fontSize="5" pageId="20" pageNumber="89">420.91</superScript>
; LP
<superScript attach="left" box="[751,820,840,862]" fontSize="5" pageId="20" pageNumber="89">430.79</superScript>
; LaP
<superScript attach="left" box="[868,926,840,863]" fontSize="5" pageId="20" pageNumber="89">42.46</superScript>
; LM
<superScript attach="left" box="[971,1028,840,862]" fontSize="5" pageId="20" pageNumber="89">11.82</superScript>
; LaM
<superScript attach="left" box="[1084,1142,840,863]" fontSize="5" pageId="20" pageNumber="89">13.05</superScript>
; LM
<superScript attach="left" box="[161,219,867,889]" fontSize="5" pageId="20" pageNumber="89">31.20</superScript>
; LaM
<superScript attach="left" box="[275,333,867,889]" fontSize="5" pageId="20" pageNumber="89">32.07</superScript>
; LCi 1.20; LP
<subScript attach="left" box="[469,527,867,891]" fontSize="5" pageId="20" pageNumber="89">20.69</subScript>
; LaP
<subScript attach="left" box="[575,633,868,891]" fontSize="5" pageId="20" pageNumber="89">20.83</subScript>
; LP
<subScript attach="left" box="[671,678,879,891]" fontSize="5" pageId="20" pageNumber="89">3</subScript>
; LP
<subScript attach="left" box="[736,794,868,891]" fontSize="5" pageId="20" pageNumber="89">41.31</subScript>
; LaP
<subScript attach="left" box="[843,901,868,891]" fontSize="5" pageId="20" pageNumber="89">41.10</subScript>
; LM
<subScript attach="left" box="[946,1004,867,891]" fontSize="5" pageId="20" pageNumber="89">12.06</subScript>
.
</paragraph>
<paragraph blockId="20.[125,1146,734,1295]" pageId="20" pageNumber="89">
MITOCHONDRIAL SEQUENCE OF THE
<typeStatus box="[435,524,905,919]" pageId="20" pageNumber="89">HOLOTYPE</typeStatus>
(complete sequence of the mitochondrial gene for cytochrome
<emphasis box="[1127,1139,900,921]" italics="true" pageId="20" pageNumber="89">b</emphasis>
; GenBank Accession Number
<accessionNumber box="[411,520,926,948]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/KC579375" pageId="20" pageNumber="89">KC579375</accessionNumber>
; 5 end). atg atc aac att cgc aag tcc cac cca cta ttc aag att atc aac gac tca ttc gtt gac cta cct gcc cca tca agt atc tct tcc tga tga aacttc gga tcc ctc cta ggg gta tgc cta gcc gtc caa att ctc aca gga ctt ttc cta gca ata cac tac aca tca gat act gcc aca gcc ttc tac tcc gta act cat att tgc cga gac gtc aac tat ggc tga gtc cta cgc tac ctc cac gcc aac gga gcc tct ata ttc ttc atc tgc ctc ttt cta cac gta gga cga gga atc tac tac ggc tcc tat aca ttc tca gaa aca tga aac att gga att atc ctc ctc ttc gcc gtc atg gcc acg gca ttc ata ggt tac gta ctc cca tga ggc caa atg tcc ttc tga ggg gca aca gtc atc aca aac ctt ctc tca gcc atc ccc tac gtt gga aca acc cta gtc gaa tga gtc tga ggc gga ttc tca gtt gat aaa gcc aca ctc acc cga ttc ttc gcc ctg cac ttc cta cta ccc ttt gtt atc gca gcc ata gtt ata gtc cat cta ctt ttc ctc cat gaa aca gga tca aac aac cca acc gga atc cca tca gac gca gac ata atc cca ttc cac ccc tac tac acc att aaa gac atc cta ggc ctc ata cta ata ctt aca gca cta ctg tcc ctg gtc tta ttt gcc ccc gac cta ctg ggc gac cca gac aac tac act cca gcc aac cca cta aat act cca ccc cac att aag cca gaa tga tac ttt cta ttt gcc tac gca atc cta cgc tca atc cca aac aaa ctt ggt gga gtc gta gcc ctg gtc cta tcc att ctc atc cta gcc acc att cca cta ctc cac aca tca aaa caa cgc agc ata gca ttc cga ccc cta agt caa tgt ctg ttc tga ctc tta gta gca gac ctt ctt aca cta acc tga atc gga ggc caa cct gtc gaa cac ccg ttc atc atc atc gga caa tta gcc tcc att ctc tat ttc cta att atc ctt gtc cta ata cca ctt gcg ggc atc gca gaa aac cat cta ttg aag tga aga.
</paragraph>
<paragraph blockId="20.[125,1146,1325,1407]" pageId="20" pageNumber="89">
DERIVATIO NOMINIS. Patronymic; named in honour of Professor Ivan Horáček (
<collectingRegion box="[986,1065,1325,1349]" country="Czech Republic" name="Praha" pageId="20" pageNumber="89">Prague</collectingRegion>
,
<collectingCountry name="Czech Republic" pageId="20" pageNumber="89">Czech Republic</collectingCountry>
) who has significantly contributed to the fauna, taxonomy and ecology of the Mediter- ranean bats.
</paragraph>
<paragraph blockId="20.[125,1129,1459,1575]" box="[125,146,1459,1480]" pageId="20" pageNumber="89">«</paragraph>
<paragraph blockId="20.[125,1129,1459,1575]" pageId="20" pageNumber="89">
Figs. 1316. Sites of occurrence of
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[428,616,1484,1503]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="89" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[428,616,1484,1503]" italics="true" pageId="20" pageNumber="89">Rhinolophus horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[621,668,1484,1503]" pageId="20" pageNumber="89" rank="species">sp. n.</taxonomicNameLabel>
in Cyrenaica,
<collectingCountry box="[791,841,1484,1503]" name="Libya" pageId="20" pageNumber="89">Libya</collectingCountry>
(photos by A. Reiter). 13 dense coniferous forest in the central part of Wadi Al Kuf. 14 Qasr Ash Shahdayn ruins, roost of
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[912,1010,1508,1527]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="89" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[912,925,1509,1527]" italics="true" pageId="20" pageNumber="89">R</emphasis>
.
<emphasis box="[935,1010,1508,1527]" italics="true" pageId="20" pageNumber="89">horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1015,1063,1508,1527]" pageId="20" pageNumber="89" rank="species">sp. n.</taxonomicNameLabel>
, surrounded by dense mountain forests. 15 Wadi
<collectingRegion box="[552,617,1532,1551]" country="Libya" name="Darnah" pageId="20" pageNumber="89">Darnah</collectingRegion>
, mosaic of agricultural areas and Mediterranean
</paragraph>
<paragraph blockId="20.[125,1129,1459,1575]" box="[125,732,1556,1575]" pageId="20" pageNumber="89">woodlands. 16 Al Burdi, shrubland valley in a plateau of dry steppes.</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="90" type="distribution">
<paragraph blockId="21.[117,1138,174,491]" pageId="21" pageNumber="90">
DISTRIBUTION.
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[264,496,174,198]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="21" pageNumber="90" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[264,496,174,198]" italics="true" pageId="21" pageNumber="90">Rhinolophus horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[501,556,175,198]" pageId="21" pageNumber="90" rank="species">sp. n.</taxonomicNameLabel>
isknown from seven sites in northern Cyrenaica (
<bibRefCitation author="QUMSIYEH M. B. &amp; SCHLITTER D. A." journalOrPublisher="Annals of Carnegie Museum" pageId="21" pageNumber="90" pagination="377 - 389" part="51" refId="ref14848" refString="QUMSIYEH M. B. &amp; SCHLITTER D. A., 1982: The bat fauna of Jabal Al Akhdar, Northeast Libya. Annals of Carnegie Museum, 51: 377 - 389." title="The bat fauna of Jabal Al Akhdar, Northeast Libya" type="journal article" year="1982">Qumsiyeh &amp; Schlitter 1982</bibRefCitation>
, original findings), from ca.
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long belt of Mediterranean woodlands and steppes between Wadi Al Kuf in the west and Al Burdi in the east (Figs. 1316). The records are available from altitudes stretching from the sea level up to
<quantity box="[785,853,262,286]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="21" pageNumber="90" unit="m" value="660.0">660 m</quantity>
a. s. l., from the following sites: Al Burdi (
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,
<geoCoordinate box="[402,503,291,316]" degrees="25" direction="east" minutes="05" orientation="longitude" pageId="21" pageNumber="90" precision="925" value="25.083334">25° 05 E</geoCoordinate>
), Qasr Ash Shahdayn (
<geoCoordinate box="[749,855,292,316]" degrees="32" direction="north" minutes="37" orientation="latitude" pageId="21" pageNumber="90" precision="925" value="32.616665">32° 37 N</geoCoordinate>
,
<geoCoordinate box="[865,966,291,316]" degrees="21" direction="east" minutes="35" orientation="longitude" pageId="21" pageNumber="90" precision="925" value="21.583334">21° 35 E</geoCoordinate>
), ruins
<quantity box="[1047,1100,291,315]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="90" unit="km" value="6.0">6 km</quantity>
SE of Qasr [Al] Maqdam (
<geoCoordinate box="[369,478,321,345]" degrees="32" direction="north" minutes="38" orientation="latitude" pageId="21" pageNumber="90" precision="925" value="32.633335">32° 38 N</geoCoordinate>
,
<geoCoordinate box="[490,596,321,345]" degrees="21" direction="east" minutes="36" orientation="longitude" pageId="21" pageNumber="90" precision="925" value="21.6">21° 36 E</geoCoordinate>
;
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), Roman aquaduct at Kufanta (
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,
<geoCoordinate box="[334,437,350,374]" degrees="21" direction="east" minutes="34" orientation="longitude" pageId="21" pageNumber="90" precision="925" value="21.566668">21° 34 E</geoCoordinate>
;
<bibRefCitation author="QUMSIYEH M. B. &amp; SCHLITTER D. A." box="[448,745,350,374]" journalOrPublisher="Annals of Carnegie Museum" pageId="21" pageNumber="90" pagination="377 - 389" part="51" refId="ref14848" refString="QUMSIYEH M. B. &amp; SCHLITTER D. A., 1982: The bat fauna of Jabal Al Akhdar, Northeast Libya. Annals of Carnegie Museum, 51: 377 - 389." title="The bat fauna of Jabal Al Akhdar, Northeast Libya" type="journal article" year="1982">Qumsiyeh &amp; Schlitter 1982</bibRefCitation>
), Wadi
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, gallery ca.
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Sof
<collectingRegion box="[117,195,379,403]" country="Libya" name="Darnah" pageId="21" pageNumber="90">Darnah</collectingRegion>
(
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locality,
<geoCoordinate box="[348,453,380,404]" degrees="32" direction="north" minutes="42" orientation="latitude" pageId="21" pageNumber="90" precision="925" value="32.7">32° 42 N</geoCoordinate>
,
<geoCoordinate box="[462,563,380,404]" degrees="22" direction="east" minutes="37" orientation="longitude" pageId="21" pageNumber="90" precision="925" value="22.616667">22° 37 E</geoCoordinate>
), Wadi
<collectingRegion box="[642,723,379,403]" country="Libya" name="Darnah" pageId="21" pageNumber="90">Darnah</collectingRegion>
, cave ca.
<quantity box="[820,885,379,404]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="90" unit="km" value="10.0">10 km</quantity>
Sof
<collectingRegion box="[937,1015,379,403]" country="Libya" name="Darnah" pageId="21" pageNumber="90">Darnah</collectingRegion>
(
<geoCoordinate box="[1028,1134,380,404]" degrees="32" direction="north" minutes="41" orientation="latitude" pageId="21" pageNumber="90" precision="925" value="32.683334">32° 41 N</geoCoordinate>
,
<geoCoordinate box="[117,218,409,433]" degrees="22" direction="east" minutes="36" orientation="longitude" pageId="21" pageNumber="90" precision="925" value="22.6">22° 36 E</geoCoordinate>
), and Wadi Al Kuf, unnamed cave (
<geoCoordinate box="[598,705,409,433]" degrees="32" direction="north" minutes="41" orientation="latitude" pageId="21" pageNumber="90" precision="925" value="32.683334">32° 41 N</geoCoordinate>
,
<geoCoordinate box="[715,816,409,433]" degrees="21" direction="east" minutes="34" orientation="longitude" pageId="21" pageNumber="90" precision="925" value="21.566668">21° 34 E</geoCoordinate>
). At four sites,
<taxonomicName authority="Benda &amp; Vallo, 2012" authorityName="Benda &amp; Vallo" authorityYear="2012" box="[979,1101,409,433]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="21" pageNumber="90" phylum="Chordata" rank="species" species="horaceki" status="sp. n.">
<emphasis box="[979,995,410,433]" italics="true" pageId="21" pageNumber="90">R</emphasis>
.
<emphasis box="[1008,1101,409,433]" italics="true" pageId="21" pageNumber="90">horaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel pageId="21" pageNumber="90" rank="species">sp. n.</taxonomicNameLabel>
was found roosting; viz., in two natural caves, in an underground part of castle ruins (Fig. 14) and in an abandoned cellar.
</paragraph>
</subSubSection>
</treatment>
</document>