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<document id="E330B5824A8EFB452DD8FCDF4CC8B0C8" ID-DOI="10.11646/zootaxa.3681.4.1" ID-GBIF-Dataset="17afa4e8-b105-41f8-bf86-e856fc41a3a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="222798" ID-ZooBank="ABBE4175-C2E8-4BB5-9B61-599BD3D0F632" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460640219132" checkinUser="plazi" docAuthor="Colloff, Matthew J. &amp; Cameron, Stephen L." docDate="2013" docId="03EF6064E467FFEF6AEA063CFBB97183" docLanguage="en" docName="zt03681p346.pdf" docOrigin="Zootaxa 3681 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrphonothrus maritimus Colloff &amp; Cameron, 2013, sp. nov." docType="treatment" docVersion="10" lastPageNumber="314" masterDocId="FFD6181CE46BFFE26A7D0410FF8B764E" masterDocTitle="A phylogenetic analysis and taxonomic revision of the oribatid mite family Malaconothridae (Acari: Oribatida), with new species of Tyrphonothrus and Malaconothrus from Australia" masterLastPageNumber="346" masterPageNumber="301" pageNumber="313" updateTime="1698335724857" updateUser="plazi">
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<mods:title id="89B7486FE65F7369F5573110E6C97B09">A phylogenetic analysis and taxonomic revision of the oribatid mite family Malaconothridae (Acari: Oribatida), with new species of Tyrphonothrus and Malaconothrus from Australia</mods:title>
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<mods:namePart id="10E78598DF2F801C0110243E6D3DCBB2">Colloff, Matthew J.</mods:namePart>
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<mods:namePart id="603A5FE7585337865A9C4FD447E54BB7">Cameron, Stephen L.</mods:namePart>
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<mods:title id="CA48F6BF0B5B63A0F807F10D3C415831">Zootaxa</mods:title>
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<mods:date id="883FC86F9EE5D6771FC5451643602250">2013</mods:date>
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<treatment id="03EF6064E467FFEF6AEA063CFBB97183" ID-DOI="http://doi.org/10.5281/zenodo.6163956" ID-GBIF-Taxon="119571179" ID-Zenodo-Dep="6163956" LSID="urn:lsid:plazi:treatment:03EF6064E467FFEF6AEA063CFBB97183" httpUri="http://treatment.plazi.org/id/03EF6064E467FFEF6AEA063CFBB97183" lastPageId="13" lastPageNumber="314" pageId="12" pageNumber="313">
<subSubSection id="C35C82F9E467FFEE6AEA063CFAAC74E1" pageId="12" pageNumber="313" type="nomenclature">
<paragraph id="8BF9D172E467FFEE6AEA063CFDBF7408" blockId="12.[151,564,556,615]" box="[151,564,556,582]" pageId="12" pageNumber="313">
<heading id="D0B1661EE467FFEE6AEA063CFDBF7408" bold="true" box="[151,564,556,582]" fontSize="11" level="1" pageId="12" pageNumber="313" reason="1">
<emphasis id="B9320D60E467FFEE6AEA063CFDBF7408" bold="true" box="[151,564,556,582]" pageId="12" pageNumber="313">
<taxonomicName id="4C46AAF1E467FFEE6AEA063CFE447408" ID-CoL="7X635" box="[151,463,556,582]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B9320D60E467FFEE6AEA063CFE447408" bold="true" box="[151,463,556,582]" italics="true" pageId="12" pageNumber="313">Tyrphonothrus maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201B01BE467FFEE6BAB063CFDBF7408" box="[470,564,556,582]" pageId="12" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D172E467FFEE6AEA065FFE9A7429" blockId="12.[151,564,556,615]" box="[151,273,591,615]" pageId="12" pageNumber="313">(Figs. 7, 8)</paragraph>
<paragraph id="8BF9D172E467FFEE6AEA0686FAAC74E1" blockId="12.[151,1437,662,2025]" box="[151,1319,662,687]" pageId="12" pageNumber="313">
<emphasis id="B9320D60E467FFEE6AEA0686FEA274E1" bold="true" box="[151,297,662,687]" pageId="12" pageNumber="313">Dimensions.</emphasis>
<typeStatus id="54FD6FD0E467FFEE6B520687FE1C74E1" box="[303,407,663,687]" pageId="12" pageNumber="313" type="holotype">Holotype</typeStatus>
female length 596 μm, breadth 293 μm. Ratio of prodorsum to total length: 0.24.
</paragraph>
</subSubSection>
<subSubSection id="C35C82F9E467FFEF6ABA06ABFB3573E1" lastPageId="13" lastPageNumber="314" pageId="12" pageNumber="313" type="description">
<paragraph id="8BF9D172E467FFEE6ABA06ABFAC0752D" blockId="12.[151,1437,662,2025]" pageId="12" pageNumber="313">
<emphasis id="B9320D60E467FFEE6ABA06ABFE95749D" bold="true" box="[199,286,699,723]" pageId="12" pageNumber="313">Female</emphasis>
.
<emphasis id="B9320D60E467FFEE6B5306ABFE27749D" box="[302,428,699,723]" italics="true" pageId="12" pageNumber="313">Prodorsum</emphasis>
. Rostrum acute, projecting apically beyond margin of prodorsum, giving impression of partial, transverse rostral ridge (Fig. 7a). Rostral seta (
<emphasis id="B9320D60E467FFEE688506F0FC9A74B9" box="[760,785,736,759]" italics="true" pageId="12" pageNumber="313">ro</emphasis>
) smooth, curved, setiform, thick, 57 μm long, longer than mutual distance, emerging from well-developed tubercles. Lamellar seta (
<emphasis id="B9320D60E467FFEE69AA0713FC607555" box="[983,1003,771,795]" italics="true" pageId="12" pageNumber="313">le</emphasis>
) barbed, flagelliform apically, 97 μm, longer than their mutual distance, enclosed by incurved, S-shaped lateral carina. Interlamellar seta (
<emphasis id="B9320D60E467FFEE6F640738FAA47571" box="[1305,1327,808,831]" italics="true" pageId="12" pageNumber="313">in</emphasis>
) smooth, setiform, short (92 μm), 3 × longer than smooth seta
<emphasis id="B9320D60E467FFEE689A075CFC8B752D" box="[743,768,844,867]" italics="true" pageId="12" pageNumber="313">ex</emphasis>
1. Prodorsum granular with scattered small foveolae.
</paragraph>
<paragraph id="8BF9D172E467FFEE6ABA0763FB0A737F" blockId="12.[151,1437,662,2025]" pageId="12" pageNumber="313">
<emphasis id="B9320D60E467FFEE6ABA0763FEC875C4" box="[199,323,883,906]" italics="true" pageId="12" pageNumber="313">Notogaster</emphasis>
. Dorsosejugal suture slightly convex (Fig. 7a). Humeral region obtusely angled, almost at right angles; with short ridge between setae
<emphasis id="B9320D60E467FFEE68270787FDED75E0" box="[602,614,919,942]" italics="true" pageId="12" pageNumber="313">c</emphasis>
2 and
<emphasis id="B9320D60E467FFEE68D60787FD3C75E0" box="[683,695,919,942]" italics="true" pageId="12" pageNumber="313">c</emphasis>
3 extending towards
<emphasis id="B9320D60E467FFEE69D90787FC3475E0" box="[932,959,919,942]" italics="true" pageId="12" pageNumber="313">cp</emphasis>
. With 15 pairs of barbed, thick, setiform setae. Setae
<emphasis id="B9320D60E467FFEE6B5E07AEFEA4759B" box="[291,303,958,981]" italics="true" pageId="12" pageNumber="313">e</emphasis>
2,
<emphasis id="B9320D60E467FFEE6B3E07ADFEDA759B" box="[323,337,957,981]" italics="true" pageId="12" pageNumber="313">h</emphasis>
2 and
<emphasis id="B9320D60E467FFEE6BEC07AEFE14759B" box="[401,415,958,981]" italics="true" pageId="12" pageNumber="313">p</emphasis>
2 much longer than others. Seta
<emphasis id="B9320D60E467FFEE697407AEFC9E759B" box="[777,789,958,981]" italics="true" pageId="12" pageNumber="313">c</emphasis>
1 51 μm long, almost equal distance to
<emphasis id="B9320D60E467FFEE6EAD07ADFB55759B" box="[1232,1246,957,981]" italics="true" pageId="12" pageNumber="313">d</emphasis>
1;
<emphasis id="B9320D60E467FFEE6E8B07AEFA89759B" box="[1270,1282,958,981]" italics="true" pageId="12" pageNumber="313">c</emphasis>
2 52 μm long, shorter than distance to
<emphasis id="B9320D60E467FFEE6BD807F4FE4B75B5" box="[421,448,996,1019]" italics="true" pageId="12" pageNumber="313">cp</emphasis>
;
<emphasis id="B9320D60E467FFEE6BB207F4FE5075B5" box="[463,475,996,1019]" italics="true" pageId="12" pageNumber="313">c</emphasis>
3 65 μm long, almost equal distance to
<emphasis id="B9320D60E467FFEE69EE07F4FC2675B5" box="[915,941,996,1019]" italics="true" pageId="12" pageNumber="313">cp</emphasis>
;
<emphasis id="B9320D60E467FFEE69C107F4FC5D75B5" box="[956,982,996,1019]" italics="true" pageId="12" pageNumber="313">cp</emphasis>
62 μm, longer than distance to
<emphasis id="B9320D60E467FFEE6F4007F3FAC075B5" box="[1341,1355,995,1019]" italics="true" pageId="12" pageNumber="313">d</emphasis>
2;
<emphasis id="B9320D60E467FFEE6F1E07F3FAFA75B5" box="[1379,1393,995,1019]" italics="true" pageId="12" pageNumber="313">d</emphasis>
1 48 μm, almost equal their mutual distance;
<emphasis id="B9320D60E467FFEE681E001AFDFA726C" box="[611,625,1034,1058]" italics="true" pageId="12" pageNumber="313">d</emphasis>
2 59 μm, equal distance to
<emphasis id="B9320D60E467FFEE69D9001BFC3B726C" box="[932,944,1035,1058]" italics="true" pageId="12" pageNumber="313">e</emphasis>
2;
<emphasis id="B9320D60E467FFEE69B5001BFC5F726C" box="[968,980,1035,1058]" italics="true" pageId="12" pageNumber="313">e</emphasis>
1 54 μm long, almost equal their mutual distance;
<emphasis id="B9320D60E467FFEE6B7C0022FE867207" box="[257,269,1074,1097]" italics="true" pageId="12" pageNumber="313">e</emphasis>
2 124 μm, overlapping
<emphasis id="B9320D60E467FFEE68710021FD9F7207" box="[524,532,1073,1097]" italics="true" pageId="12" pageNumber="313">f</emphasis>
2;
<emphasis id="B9320D60E467FFEE68570021FDB97207" box="[554,562,1073,1097]" italics="true" pageId="12" pageNumber="313">f</emphasis>
2 78 μm long, almost equal distance to
<emphasis id="B9320D60E467FFEE699F0021FC7B7207" box="[994,1008,1073,1097]" italics="true" pageId="12" pageNumber="313">h</emphasis>
2;
<emphasis id="B9320D60E467FFEE6E7B0021FB9F7207" box="[1030,1044,1073,1097]" italics="true" pageId="12" pageNumber="313">h</emphasis>
1 65 μm long, half distance to
<emphasis id="B9320D60E467FFEE6F1F0022FAFB7207" box="[1378,1392,1074,1097]" italics="true" pageId="12" pageNumber="313">p</emphasis>
1;
<emphasis id="B9320D60E467FFEE6FFB0021FA1F7207" box="[1414,1428,1073,1097]" italics="true" pageId="12" pageNumber="313">h</emphasis>
2 105 μm long, half distance to
<emphasis id="B9320D60E467FFEE6B980049FE78723E" box="[485,499,1113,1136]" italics="true" pageId="12" pageNumber="313">p</emphasis>
1;
<emphasis id="B9320D60E467FFEE68770048FD93723E" box="[522,536,1112,1136]" italics="true" pageId="12" pageNumber="313">h</emphasis>
3 95 μm long;
<emphasis id="B9320D60E467FFEE68CB0049FD4F723E" box="[694,708,1113,1136]" italics="true" pageId="12" pageNumber="313">p</emphasis>
1 43 μm long, half mutual distance, positioned dorsally;
<emphasis id="B9320D60E467FFEE6F4A0049FACE723E" box="[1335,1349,1113,1136]" italics="true" pageId="12" pageNumber="313">p</emphasis>
3 86 μm long, extending as far as
<emphasis id="B9320D60E467FFEE6BAF0090FE6B72D9" box="[466,480,1152,1175]" italics="true" pageId="12" pageNumber="313">p</emphasis>
2; seta
<emphasis id="B9320D60E467FFEE684B0090FDCF72D9" box="[566,580,1152,1175]" italics="true" pageId="12" pageNumber="313">p</emphasis>
2 121 μm long). Notogaster with complex series of median and lateral longitudinal ridges, converging anterior of
<emphasis id="B9320D60E467FFEE68FD00B5FD0572F3" box="[640,654,1189,1213]" italics="true" pageId="12" pageNumber="313">h</emphasis>
1; then with posterior M-shaped ridge diverging medially, anterior of
<emphasis id="B9320D60E467FFEE6AEA00DDFF2E72AA" box="[151,165,1229,1252]" italics="true" pageId="12" pageNumber="313">p</emphasis>
1. Lateral ridges diverging anteriorly. Margins narrowing anteriorly, broadest between
<emphasis id="B9320D60E467FFEE6EDB00DDFB3972AA" box="[1190,1202,1229,1252]" italics="true" pageId="12" pageNumber="313">e</emphasis>
2 and
<emphasis id="B9320D60E467FFEE6E8100DCFA8F72AA" box="[1276,1284,1228,1252]" italics="true" pageId="12" pageNumber="313">f</emphasis>
2. With short concave ridges between
<emphasis id="B9320D60E467FFEE6BCD00E3FE357345" box="[432,446,1267,1291]" italics="true" pageId="12" pageNumber="313">h</emphasis>
3 and
<emphasis id="B9320D60E467FFEE687C00E4FD847345" box="[513,527,1268,1291]" italics="true" pageId="12" pageNumber="313">p</emphasis>
2 and between
<emphasis id="B9320D60E467FFEE68C500E4FD4D7345" box="[696,710,1268,1291]" italics="true" pageId="12" pageNumber="313">p</emphasis>
2 and posterior margin of notogaster. Ratio of length to breadth 1.53. Caudal margin V-shaped, indented at level of seta
<emphasis id="B9320D60E467FFEE6971010AFC91737C" box="[780,794,1306,1330]" italics="true" pageId="12" pageNumber="313">h</emphasis>
2. Notogaster with small foveae.
</paragraph>
<paragraph id="8BF9D172E467FFEE6ABA0151FC06738A" blockId="12.[151,1437,662,2025]" pageId="12" pageNumber="313">
<emphasis id="B9320D60E467FFEE6ABA0151FEDD7316" box="[199,342,1345,1368]" italics="true" pageId="12" pageNumber="313">Coxisternum</emphasis>
. Epimeral setal formula 3-1-3-2 (Fig. 7b). Epimeral setae smooth, setiform;
<emphasis id="B9320D60E467FFEE6EC10151FB5C7317" box="[1212,1239,1345,1369]" italics="true" pageId="12" pageNumber="313">1b</emphasis>
,
<emphasis id="B9320D60E467FFEE6E9A0151FA967317" box="[1255,1309,1345,1369]" italics="true" pageId="12" pageNumber="313">3ac</emphasis>
,
<emphasis id="B9320D60E467FFEE6F500151FAC27317" box="[1325,1353,1345,1369]" italics="true" pageId="12" pageNumber="313">4a</emphasis>
and
<emphasis id="B9320D60E467FFEE6FFC0151FA107317" box="[1409,1435,1345,1369]" italics="true" pageId="12" pageNumber="313">4c</emphasis>
longer than others (2231 μm);
<emphasis id="B9320D60E467FFEE68700175FDA17333" box="[525,554,1381,1405]" italics="true" pageId="12" pageNumber="313">3b</emphasis>
in posteriolateral position;
<emphasis id="B9320D60E467FFEE69110175FC0C7333" box="[876,903,1381,1405]" italics="true" pageId="12" pageNumber="313">4b</emphasis>
absent. Epimeral plates I not fused medially, rounded anteriolaterally. Epimeral plates II acute posteriolaterally, not fused medially. Epimeral plates III fused medially. Apodeme IV arched. Epimeral plate IV rounded laterally.
</paragraph>
<paragraph id="8BF9D172E467FFEE6ABA01C1FF5B70EC" blockId="12.[151,1437,662,2025]" pageId="12" pageNumber="313">
<emphasis id="B9320D60E467FFEE6ABA01C1FE1273A7" box="[199,409,1489,1513]" italics="true" pageId="12" pageNumber="313">Anogenital Region</emphasis>
. With five pairs of smooth, flagelliform genital setae;
<emphasis id="B9320D60E467FFEE6E7601C2FB9273A7" box="[1035,1049,1490,1513]" italics="true" pageId="12" pageNumber="313">g</emphasis>
1-4 subequal in length (5164 μm),
<emphasis id="B9320D60E467FFEE6AEA01E8FF2E7041" box="[151,165,1528,1551]" italics="true" pageId="12" pageNumber="313">g</emphasis>
1-4 closely spaced,
<emphasis id="B9320D60E467FFEE6B0701E8FE037041" box="[378,392,1528,1551]" italics="true" pageId="12" pageNumber="313">g</emphasis>
5 shorter (38 μm) and spaced more broadly apart (Fig. 7b). Genital plates sub-rectangular, rounded apically; each plate 151 μm long, 48 μm broad. Posterior margin of genital plate transverse, overlapping adanal plate. Anterior margin of adanal plate acute, with sclerotised transverse region. Each adanal plate 156 μm long, 35 μm broad; three pairs of barbed, setiform adanal setae, 5465 μm long. One pair of minute anal setae 5 μm long.
</paragraph>
<paragraph id="8BF9D172E467FFEE6ABA02BFFE8971A7" blockId="12.[151,1437,662,2025]" pageId="12" pageNumber="313">
<emphasis id="B9320D60E467FFEE6ABA02BFFF767088" box="[199,253,1711,1734]" italics="true" pageId="12" pageNumber="313">Legs</emphasis>
. Pre-tarsi homotridactylous; claws smooth (
<figureCitation id="137DCDF7E467FFEE688202BEFCCD7088" box="[767,838,1710,1734]" captionStart="FIGURE 8" captionStartId="14.[151,250,1723,1746]" captionTargetBox="[353,1251,198,1701]" captionTargetId="figure@14.[285,1269,194,1709]" captionTargetPageId="14" captionText="FIGURE 8. Tyrphonothrus maritimus sp. nov., holotype female, left legs, antiaxial view; a) Leg I; b) Leg II; c) Leg III; d) Leg IV." httpUri="https://zenodo.org/record/222805/files/figure.png" pageId="12" pageNumber="313">Fig. 8</figureCitation>
). Tarsi long and thin; ratio of lengths to breadth of tarsi: I 2.69; II 2.67; III 3.02; IV 3.91. Setal formula: Leg
<date id="FFF8F7B2E467FFEE694C02C3FCED70A4" box="[817,870,1746,1770]" pageId="12" pageNumber="313">I 1-5</date>
-3(1)-4(1)-11(3); Leg
<date id="FFF8F7B2E467FFEE6E1802C3FB2F70A4" box="[1125,1188,1746,1770]" pageId="12" pageNumber="313">II 1-5</date>
-2(1)-4(1)-10(1); Leg
<date id="FFF8F7B2E467FFEE6AEA02E7FF6F7140" box="[151,228,1782,1806]" pageId="12" pageNumber="313">III 2-3</date>
-1-2(1)-10; Leg
<date id="FFF8F7B2E467FFEE6BE102E7FE6C7140" box="[412,487,1782,1806]" pageId="12" pageNumber="313">IV 1-2</date>
-1-2-10. Most setae of tibiae, genuae and femora barbed; most tarsal setae smooth. Proral setae (
<emphasis id="B9320D60E467FFEE6B47030BFEC3717C" box="[314,328,1819,1842]" italics="true" pageId="12" pageNumber="313">p</emphasis>
) on tarsi I-IV short, broad, blunt (
<figureCitation id="137DCDF7E467FFEE6886030AFCCD717C" box="[763,838,1818,1842]" captionStart="FIGURE 8" captionStartId="14.[151,250,1723,1746]" captionTargetBox="[353,1251,198,1701]" captionTargetId="figure@14.[285,1269,194,1709]" captionTargetPageId="14" captionText="FIGURE 8. Tyrphonothrus maritimus sp. nov., holotype female, left legs, antiaxial view; a) Leg I; b) Leg II; c) Leg III; d) Leg IV." httpUri="https://zenodo.org/record/222805/files/figure.png" pageId="12" pageNumber="313">Fig. 8</figureCitation>
a): Tarsus I with very long, broad, blunt, barbed primiventral setae (
<emphasis id="B9320D60E467FFEE6B04032FFE1F7118" box="[377,404,1855,1878]" italics="true" pageId="12" pageNumber="313">pv</emphasis>
). Solenidion w1 separate from w1-2. Famulus (e) thin, setiform 4 μm long. Fastigial setae (
<emphasis id="B9320D60E467FFEE6ADD0375FF3B7133" box="[160,176,1893,1917]" italics="true" pageId="12" pageNumber="313">ft</emphasis>
) heteromorphic:
<emphasis id="B9320D60E467FFEE6B050375FE037133" box="[376,392,1893,1917]" italics="true" pageId="12" pageNumber="313">ft</emphasis>
long, curved, smooth and pointed;
<emphasis id="B9320D60E467FFEE69540375FCB27133" box="[809,825,1893,1917]" italics="true" pageId="12" pageNumber="313">ft</emphasis>
” large, thick, spine-like, blunt apically. Setae
<emphasis id="B9320D60E467FFEE6F330375FAD57133" box="[1358,1374,1893,1917]" italics="true" pageId="12" pageNumber="313">ft</emphasis>
” and
<emphasis id="B9320D60E467FFEE6AEA0399FF2C71EF" box="[151,167,1929,1953]" italics="true" pageId="12" pageNumber="313">ft</emphasis>
on tarsi II similarly shaped (
<figureCitation id="137DCDF7E467FFEE6B860399FDB471EF" box="[507,575,1929,1953]" captionStart="FIGURE 8" captionStartId="14.[151,250,1723,1746]" captionTargetBox="[353,1251,198,1701]" captionTargetId="figure@14.[285,1269,194,1709]" captionTargetPageId="14" captionText="FIGURE 8. Tyrphonothrus maritimus sp. nov., holotype female, left legs, antiaxial view; a) Leg I; b) Leg II; c) Leg III; d) Leg IV." httpUri="https://zenodo.org/record/222805/files/figure.png" pageId="12" pageNumber="313">Fig. 8</figureCitation>
b); those on tarsi II homeomorphic; long, curved, smooth and pointed (
<figureCitation id="137DCDF7E467FFEE6F100399FF2D718B" captionStart="FIGURE 8" captionStartId="14.[151,250,1723,1746]" captionTargetBox="[353,1251,198,1701]" captionTargetId="figure@14.[285,1269,194,1709]" captionTargetPageId="14" captionText="FIGURE 8. Tyrphonothrus maritimus sp. nov., holotype female, left legs, antiaxial view; a) Leg I; b) Leg II; c) Leg III; d) Leg IV." httpUri="https://zenodo.org/record/222805/files/figure.png" pageId="12" pageNumber="313">Fig. 8</figureCitation>
c). Tarsus IV with seta
<emphasis id="B9320D60E467FFEE6BCA03BDFE4C718B" box="[439,455,1965,1989]" italics="true" pageId="12" pageNumber="313">ft</emphasis>
” curved, pointed, smooth; seta
<emphasis id="B9320D60E467FFEE694403BEFCCF718B" box="[825,836,1966,1989]" italics="true" pageId="12" pageNumber="313">s</emphasis>
same shape and length as paired primiventral setae (
<figureCitation id="137DCDF7E467FFEE6AE203C1FF6871A7" box="[159,227,2001,2025]" captionStart="FIGURE 8" captionStartId="14.[151,250,1723,1746]" captionTargetBox="[353,1251,198,1701]" captionTargetId="figure@14.[285,1269,194,1709]" captionTargetPageId="14" captionText="FIGURE 8. Tyrphonothrus maritimus sp. nov., holotype female, left legs, antiaxial view; a) Leg I; b) Leg II; c) Leg III; d) Leg IV." httpUri="https://zenodo.org/record/222805/files/figure.png" pageId="12" pageNumber="313">Fig. 8</figureCitation>
d).
</paragraph>
<paragraph id="8BF9D172E466FFEF6AEA0188FB3573E1" blockId="13.[151,1214,1432,1459]" box="[151,1214,1432,1459]" pageId="13" pageNumber="314">
<emphasis id="B9320D60E466FFEF6AEA0188FE9873E0" bold="true" box="[151,275,1432,1455]" pageId="13" pageNumber="314">FIGURE 7.</emphasis>
<taxonomicName id="4C46AAF1E466FFEF6B640189FD9073E1" box="[281,539,1433,1455]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B9320D60E466FFEF6B640189FD9073E1" box="[281,539,1433,1455]" italics="true" pageId="13" pageNumber="314">Tyrphonothrus maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201B01BE466FFEF685D0189FDFA73E0" box="[544,625,1433,1454]" pageId="13" pageNumber="314" rank="species">
<emphasis id="B9320D60E466FFEF685D0189FDFA73E0" bold="true" box="[544,625,1433,1454]" pageId="13" pageNumber="314">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="54FD6FD0E466FFEF68030189FD5C73E1" box="[638,727,1433,1455]" pageId="13" pageNumber="314" type="holotype">holotype</typeStatus>
female; a) dorsal; b) ventral; c) detail of seta
<emphasis id="B9320D60E466FFEF6ED8018AFB3B73E1" box="[1189,1200,1434,1455]" italics="true" pageId="13" pageNumber="314">e</emphasis>
1.
</paragraph>
</subSubSection>
<subSubSection id="C35C82F9E466FFEF6ABA01CDFEFE7070" pageId="13" pageNumber="314" type="materials_examined">
<paragraph id="8BF9D172E466FFEF6ABA01CDFEFE7070" blockId="13.[151,1436,1501,1997]" pageId="13" pageNumber="314">
<emphasis id="B9320D60E466FFEF6ABA01CDFE3B73B8" bold="true" box="[199,432,1501,1526]" pageId="13" pageNumber="314">Material examined.</emphasis>
moss, Point Perpendicular, Beecroft Peninsula, New South
<collectingCountry id="F35191E2E466FFEF6E3101CEFB1E73B8" box="[1100,1173,1502,1526]" name="United Kingdom" pageId="13" pageNumber="314">Wales</collectingCountry>
,
<geoCoordinate id="EE72B7B5E466FFEF6EE201CEFB6773B8" box="[1183,1260,1502,1526]" direction="south" orientation="latitude" pageId="13" pageNumber="314" precision="925" value="-36.1">36°6'S</geoCoordinate>
,
<geoCoordinate id="EE72B7B5E466FFEF6E8B01CEFAEB73B8" box="[1270,1376,1502,1526]" direction="east" orientation="longitude" pageId="13" pageNumber="314" precision="925" value="150.8">150°48'E</geoCoordinate>
, coll. P. Greenslade,
<date id="FFF8F7B2E466FFEF6B3A0212FE4F7054" box="[327,452,1538,1562]" pageId="13" pageNumber="314" value="1995-03-21">21.iii.1995</date>
.
<typeStatus id="54FD6FD0E466FFEF6BAF0212FDB17054" box="[466,570,1538,1562]" pageId="13" pageNumber="314" type="holotype">Holotype</typeStatus>
deposited in the Australian National Insect Collection,
<collectionCode id="ED5749B7E466FFEF6EB80212FA937054" box="[1221,1304,1538,1562]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="13" pageNumber="314">CSIRO</collectionCode>
Ecosystem Sciences, Canberra.
</paragraph>
</subSubSection>
<subSubSection id="C35C82F9E466FFEF6ABA025BFBC8702C" box="[199,1091,1610,1635]" pageId="13" pageNumber="314" type="etymology">
<paragraph id="8BF9D172E466FFEF6ABA025BFBC8702C" blockId="13.[151,1436,1501,1997]" box="[199,1091,1610,1635]" pageId="13" pageNumber="314">
<emphasis id="B9320D60E466FFEF6ABA025BFEC4702D" bold="true" box="[199,335,1611,1635]" pageId="13" pageNumber="314">Etymology.</emphasis>
This species is named for the maritime location of its
<typeStatus id="54FD6FD0E466FFEF69CD025BFC6B702D" box="[944,992,1611,1635]" pageId="13" pageNumber="314">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C35C82F9E466FFEF6ABA027EFBB97183" pageId="13" pageNumber="314" type="discussion">
<paragraph id="8BF9D172E466FFEF6ABA027EFA177174" blockId="13.[151,1436,1501,1997]" pageId="13" pageNumber="314">
<emphasis id="B9320D60E466FFEF6ABA027EFEBF70C8" bold="true" box="[199,308,1646,1670]" pageId="13" pageNumber="314">Remarks</emphasis>
.
<taxonomicName id="4C46AAF1E466FFEF6B3A027EFDE770C8" box="[327,620,1646,1670]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B9320D60E466FFEF6B3A027EFDE770C8" box="[327,620,1646,1670]" italics="true" pageId="13" pageNumber="314">Tyrphonothrus maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201B01BE466FFEF6805027EFD5270C8" box="[632,729,1646,1670]" pageId="13" pageNumber="314" rank="species">
<emphasis id="B9320D60E466FFEF6805027EFD5A70C8" bold="true" box="[632,721,1646,1670]" pageId="13" pageNumber="314">sp. nov</emphasis>
.
</taxonomicNameLabel>
differs from other members of the genus by the following combination of characters: 1) the acute rostrum; 2) the rostral setae on well-developed tubercles; 3) the lamellar setae are as longas the interlamellar setae; 4) of the prodorsal setae, the lamellar setae are barbed and the rest smooth; 5) the interlamellar setae are only three times the length of the exobothridial setae; 6) the minute foveolae on the cuticle; 7) the complex series of median and lateral longitudinal notogastral ridges; 8) the pointed caudal region; 9) the barbed notogastral and adanal setae; 10) the five pairs of genital setae with
<emphasis id="B9320D60E466FFEF6E020333FB067174" box="[1151,1165,1827,1850]" italics="true" pageId="13" pageNumber="314">g</emphasis>
5 shorter than the others.
</paragraph>
<paragraph id="8BF9D172E466FFEF6ABA0359FBB97183" blockId="13.[151,1436,1501,1997]" pageId="13" pageNumber="314">
<taxonomicName id="4C46AAF1E466FFEF6ABA0359FE63712F" box="[199,488,1865,1889]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B9320D60E466FFEF6ABA0359FE63712F" box="[199,488,1865,1889]" italics="true" pageId="13" pageNumber="314">Tyrphonothrus maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201B01BE466FFEF6B8C0359FDC5712F" box="[497,590,1865,1889]" pageId="13" pageNumber="314" rank="species">
<emphasis id="B9320D60E466FFEF6B8C0359FDCD712F" bold="true" box="[497,582,1865,1889]" pageId="13" pageNumber="314">sp. nov</emphasis>
.
</taxonomicNameLabel>
shares with
<taxonomicName id="4C46AAF1E466FFEF689D035AFCE5712F" box="[736,878,1865,1889]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="binodulus">
<emphasis id="B9320D60E466FFEF689D035AFCE5712F" box="[736,878,1865,1889]" italics="true" pageId="13" pageNumber="314">T. binodulus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C46AAF1E466FFEF6903035AFC7E712F" box="[894,1013,1865,1889]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="fijiensis">
<emphasis id="B9320D60E466FFEF6903035AFC7E712F" box="[894,1013,1865,1889]" italics="true" pageId="13" pageNumber="314">T. fijiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C46AAF1E466FFEF6E78035AFBFD712F" box="[1029,1142,1866,1889]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="gringai">
<emphasis id="B9320D60E466FFEF6E78035AFBFD712F" box="[1029,1142,1866,1889]" italics="true" pageId="13" pageNumber="314">T. gringai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C46AAF1E466FFEF6ECC035AFA97712F" box="[1201,1308,1865,1889]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="taylori">
<emphasis id="B9320D60E466FFEF6ECC035AFA97712F" box="[1201,1308,1865,1889]" italics="true" pageId="13" pageNumber="314">T. taylori</emphasis>
</taxonomicName>
the barbed notogastral setae, five pairs of genital setae and the pattern of median, lateral and M-shaped posterior notogastral ridges. Only
<taxonomicName id="4C46AAF1E466FFEF6B510382FE2971E7" box="[300,418,1937,1961]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="fijiensis">
<emphasis id="B9320D60E466FFEF6B510382FE2971E7" box="[300,418,1937,1961]" italics="true" pageId="13" pageNumber="314">T. fijiensis</emphasis>
</taxonomicName>
also has the M-shaped ridge diverging medially, but differs from
<taxonomicName id="4C46AAF1E466FFEF6EE50382FAA371E7" box="[1176,1320,1938,1961]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B9320D60E466FFEF6EE50382FAA371E7" box="[1176,1320,1938,1961]" italics="true" pageId="13" pageNumber="314">T. maritimus</emphasis>
</taxonomicName>
in having barbed interlamellar setae and a broad rostrum without a partial inter-rostral ridge.
</paragraph>
</subSubSection>
</treatment>
</document>