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<mods:title id="6F73FCD2F94FF093B5575C0807926427">Cryptic Species of a Cascade Frog from Southeast Asia: Taxonomic Revisions and Descriptions of Six New Species</mods:title>
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<mods:namePart id="105B247B96621F43BBEB6AF9301EC1D5">BAIN, RAOUL H.</mods:namePart>
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<treatment id="03EF403E0054FFCBFA7FDF8EFBC01F4C" ID-DOI="http://doi.org/10.5281/zenodo.5818463" ID-GBIF-Taxon="190301398" ID-Zenodo-Dep="5818463" LSID="urn:lsid:plazi:treatment:03EF403E0054FFCBFA7FDF8EFBC01F4C" httpUri="http://treatment.plazi.org/id/03EF403E0054FFCBFA7FDF8EFBC01F4C" lastPageId="37" lastPageNumber="38" pageId="31" pageNumber="32">
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<heading id="D0B146440054FFF1FA7FDF8EFBED185F" box="[781,1145,1513,1537]" centered="true" fontSize="36" level="2" pageId="31" pageNumber="32" reason="8">
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<taxonomicName id="4C468AAB0054FFF1FA7FDF8EFC4E185E" ID-CoL="4RDHY" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[781,986,1513,1537]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="bacboensis" status="sp. nov.">
<emphasis id="B9322D3A0054FFF1FA7FDF8EFC4E185E" bold="true" box="[781,986,1513,1537]" italics="true" pageId="31" pageNumber="32">Rana bacboensis</emphasis>
</taxonomicName>
,
</emphasis>
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</paragraph>
<paragraph id="8BF9F1280054FFF1FBCCDC72FC771816" blockId="31.[702,1224,1557,1639]" pageId="31" pageNumber="32">(Previously referred to as species 2, Black Egg)</paragraph>
</subSubSection>
<subSubSection id="C35CA2A30054FFF1FA31DC34FBD61839" box="[835,1090,1616,1639]" pageId="31" pageNumber="32" type="description">
<paragraph id="8BF9F1280054FFF1FA31DC34FBD61839" blockId="31.[702,1224,1557,1639]" box="[835,1090,1616,1639]" pageId="31" pageNumber="32">
<figureCitation id="137DEDAD0054FFF1FA31DC34FC2C1839" box="[835,952,1619,1638]" captionStart="Fig" captionStartId="32.[150,188,1523,1544]" captionTargetBox="[129,1225,187,1501]" captionTargetId="figure-15@32.[129,1225,187,1501]" captionTargetPageId="32" captionText="Fig. 11. Members of the Rana chloronota complex. (A) Male and female and (B) male R. chloronota, Nghe An Province, Vietnam. C. R. graminea, male, Hainan Island, China, (NLO field series 26375). D. R. bacboensis, new species, holotype, a female from Nghe An Province, Vietnam. (E) Female holotype and male paratype in amplexus, and (F) juvenile paratype of R. daorum, new species, Lao Cai Province, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.4734904" httpUri="https://zenodo.org/record/4734904/files/figure.png" pageId="31" pageNumber="32">Figures 11D</figureCitation>
,
<figureCitation id="137DEDAD0054FFF1FAB8DC34FB9E1839" box="[970,1034,1619,1638]" captionStart="Fig" captionStartId="33.[149,187,1380,1401]" captionTargetBox="[131,1668,442,1357]" captionTargetId="figure-5@33.[131,1668,439,1358]" captionTargetPageId="33" captionText="Fig. 12. Heads of members of the Rana chloronota complex in lateral and dorsal views: (A, B) Rana chloronota, male (ROM 26402), and (C, D) female (ROM 26406); paratypes of (E, F) Rana bacboensis, new species (ROM 29529); (G, H) Rana daorum, new species (ROM 38538); (I, J) Rana hmongorum, new species (ROM 39891); (K, L) Rana morafkai, new species (ROM 39906); (M, N) Rana banaorum, new species (ROM39941); and (O, P) Rana megatympanum, new species (ROM 26398)." figureDoi="http://doi.org/10.5281/zenodo.4734907" httpUri="https://zenodo.org/record/4734907/files/figure.png" pageId="31" pageNumber="32">12E, F</figureCitation>
,
<figureCitation id="137DEDAD0054FFF1FD69DC34FBD61839" box="[1051,1090,1619,1638]" captionStart="Fig" captionStartId="34.[110,148,1407,1428]" captionTargetBox="[92,1628,413,1387]" captionTargetId="figure-5@34.[92,1629,412,1387]" captionTargetPageId="34" captionText="Fig. 13. Hands and feet of members of the Rana chloronota complex: (A) Rana chloronota (ROM 26402); paratypes of (B) Rana bacboensis, new species (ROM 29529); (C) Rana daorum, new species (ROM 38538); (D) Rana hmongorum, new species (ROM 26379); (E) Rana morafkai, new species (ROM 39906); (F) Rana banaorum, new species (ROM 39941); and (G) Rana megatympanum, new species (ROM 26398). Scale equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4734911" httpUri="https://zenodo.org/record/4734911/files/figure.png" pageId="31" pageNumber="32">13B</figureCitation>
</paragraph>
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<paragraph id="8BF9F1280054FFCDFBAADC1BFE771F34" blockId="31.[699,1227,1660,1741]" lastBlockId="35.[123,652,193,1741]" lastPageId="35" lastPageNumber="36" pageId="31" pageNumber="32">
<materialsCitation id="3B2EFB750054FFCDFBAADC1BFE771F34" ID-GBIF-Occurrence="3408947319" accessionNumber="ROM 29534" collectingDate="1994-10-24" collectorName="I. S. Darevsky &amp; L. A. Lowcock &amp; R. W. Murphy &amp; N. L. Orlov" country="Vietnam" county="Con Cuong District" lastPageId="35" lastPageNumber="36" location="Khe Moi" municipality="Con Cuong village" pageId="31" pageNumber="32" specimenCount="2" stateProvince="Nghe An" typeStatus="holotype">
<typeStatus id="54FD4F8A0054FFF1FBAADC1BFCDF18CD" box="[728,843,1660,1683]" pageId="31" pageNumber="32" type="holotype">HOLOTYPE</typeStatus>
: (ROM field no. 13171) ROM 29534 an adult female from the
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River, approximately
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west of
<collectingMunicipality id="6B9D6B520054FFCDFD36DCD1FF5F1E87" lastPageId="35" lastPageNumber="36" pageId="31" pageNumber="32">Con Cuong village</collectingMunicipality>
,
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,
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,
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(18°56̍30̎N, 104°48̍35̎E) collected
<date id="FFF8D7E80068FFCDF9BDDA9BFE361F4C" box="[207,418,252,275]" pageId="35" pageNumber="36" value="1994-10-24">
<collectingDate id="EFBC2E000068FFCDF9BDDA9BFE361F4C" box="[207,418,252,275]" pageId="35" pageNumber="36" value="1994-10-24">24 October 1994</collectingDate>
</date>
by
<collectorName id="26B394FE0068FFCDF892DA9BFD1C1F4C" box="[480,648,252,275]" pageId="35" pageNumber="36">I.S. Darevsky</collectorName>
,
<collectorName id="26B394FE0068FFCDF909DB7EFEB71F6F" box="[123,291,281,304]" pageId="35" pageNumber="36">L.A. Lowcock</collectorName>
,
<collectorName id="26B394FE0068FFCDF85CDB7EFE581F6F" box="[302,460,281,304]" pageId="35" pageNumber="36">R.W. Murphy</collectorName>
, and
<collectorName id="26B394FE0068FFCDFB75DB7EFF3C1F12" pageId="35" pageNumber="36">N.L. Orlov. The</collectorName>
<typeStatus id="54FD4F8A0068FFCDF9DDDB51FE871F12" box="[175,275,310,333]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
had leg and liver tissue removed shortly after it was euthanised.
</materialsCitation>
</paragraph>
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<paragraph id="8BF9F128006BFFCEF9E4DF94FEDD182D" blockId="32.[124,1229,1522,1650]" pageId="32" pageNumber="33">
Fig. 11. Members of the
<taxonomicName id="4C468AAB006BFFCEF8C6DF94FDF31857" box="[436,615,1523,1544]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="chloronota">
<emphasis id="B9322D3A006BFFCEF8C6DF94FDF31857" box="[436,615,1523,1544]" italics="true" pageId="32" pageNumber="33">Rana chloronota</emphasis>
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complex. (
<emphasis id="B9322D3A006BFFCEFB93DF95FD601858" bold="true" box="[737,756,1522,1543]" pageId="32" pageNumber="33">A</emphasis>
) Male and female and (
<emphasis id="B9322D3A006BFFCEFA85DF95FB9C1858" bold="true" box="[1015,1032,1522,1543]" pageId="32" pageNumber="33">B</emphasis>
) male
<taxonomicName id="4C468AAB006BFFCEFD26DF94FF09187D" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="chloronota">
<emphasis id="B9322D3A006BFFCEFD26DF94FF09187D" italics="true" pageId="32" pageNumber="33">R. chloronota</emphasis>
</taxonomicName>
, Nghe An Province, Vietnam.
<emphasis id="B9322D3A006BFFCEF896DC6AFE69187D" bold="true" box="[484,509,1549,1570]" pageId="32" pageNumber="33">C.</emphasis>
<taxonomicName id="4C468AAB006BFFCEFB76DC6AFD12187D" authorityName=": Boulenger" authorityYear="1899" box="[516,646,1549,1570]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="graminea">
<emphasis id="B9322D3A006BFFCEFB76DC6AFD12187D" box="[516,646,1549,1570]" italics="true" pageId="32" pageNumber="33">R. graminea</emphasis>
</taxonomicName>
, male, Hainan Island, China, (NLO field series 26375).
<emphasis id="B9322D3A006BFFCEF90EDC4FFF021862" bold="true" box="[124,150,1576,1597]" pageId="32" pageNumber="33">D.</emphasis>
<taxonomicName id="4C468AAB006BFFCEF9D1DC4FFEAE1862" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[163,314,1576,1597]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="bacboensis" status="sp. nov.">
<emphasis id="B9322D3A006BFFCEF9D1DC4FFEAE1862" box="[163,314,1576,1597]" italics="true" pageId="32" pageNumber="33">R. bacboensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2019041006BFFCEF83CDC4FFE401862" box="[334,468,1576,1597]" pageId="32" pageNumber="33" rank="species">new species</taxonomicNameLabel>
, holotype, a female from Nghe An Province, Vietnam. (
<emphasis id="B9322D3A006BFFCEFD2BDC4FFBFE1862" bold="true" box="[1113,1130,1576,1597]" pageId="32" pageNumber="33">E</emphasis>
) Female holotype and male paratype in amplexus, and (
<emphasis id="B9322D3A006BFFCEFB02DC25FD141808" bold="true" box="[624,640,1602,1623]" pageId="32" pageNumber="33">F</emphasis>
) juvenile paratype of
<taxonomicName id="4C468AAB006BFFCEFA1FDC24FC481807" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[877,988,1603,1624]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="daorum" status="sp. nov.">
<emphasis id="B9322D3A006BFFCEFA1FDC24FC481807" box="[877,988,1603,1624]" italics="true" pageId="32" pageNumber="33">R. daorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2019041006BFFCEFA99DC24FBF81807" box="[1003,1132,1603,1624]" pageId="32" pageNumber="33" rank="species">new species</taxonomicNameLabel>
, Lao Cai Province, Vietnam.
</paragraph>
</caption>
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Fig. 12. Heads of members of the
<taxonomicName id="4C468AAB006AFFCFFB6DDF03FD471B26" box="[543,723,1380,1401]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="chloronota">
<emphasis id="B9322D3A006AFFCFFB6DDF03FD471B26" box="[543,723,1380,1401]" italics="true" pageId="33" pageNumber="34">Rana chloronota</emphasis>
</taxonomicName>
complex in lateral and dorsal views: (
<emphasis id="B9322D3A006AFFCFFD05DF04FB3F1B27" bold="true" box="[1143,1195,1379,1400]" pageId="33" pageNumber="34">A, B</emphasis>
)
<taxonomicName id="4C468AAB006AFFCFFDCCDF03FAE51B26" box="[1214,1393,1380,1401]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="chloronota">
<emphasis id="B9322D3A006AFFCFFDCCDF03FAE51B26" box="[1214,1393,1380,1401]" italics="true" pageId="33" pageNumber="34">Rana chloronota</emphasis>
</taxonomicName>
, male (ROM 26402), and (
<emphasis id="B9322D3A006AFFCFF9F1DF19FF2C1BCC" bold="true" box="[131,184,1406,1427]" pageId="33" pageNumber="34">C, D</emphasis>
) female (ROM 26406); paratypes of (
<emphasis id="B9322D3A006AFFCFFB3CDF19FDEA1BCC" bold="true" box="[590,638,1406,1427]" pageId="33" pageNumber="34">E, F</emphasis>
)
<taxonomicName id="4C468AAB006AFFCFFBFCDF19FCD51BCC" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[654,833,1406,1427]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="bacboensis" status="sp. nov.">
<emphasis id="B9322D3A006AFFCFFBFCDF19FCD51BCC" box="[654,833,1406,1427]" italics="true" pageId="33" pageNumber="34">Rana bacboensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2019041006AFFCFFA3DDF19FC591BCC" box="[847,973,1406,1427]" pageId="33" pageNumber="34" rank="species">new species</taxonomicNameLabel>
(ROM 29529); (
<emphasis id="B9322D3A006AFFCFFDF1DF19FB2E1BCC" bold="true" box="[1155,1210,1406,1427]" pageId="33" pageNumber="34">G, H</emphasis>
)
<taxonomicName id="4C468AAB006AFFCFFDB8DF19FACE1BCC" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[1226,1370,1406,1427]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="daorum" status="sp. nov.">
<emphasis id="B9322D3A006AFFCFFDB8DF19FACE1BCC" box="[1226,1370,1406,1427]" italics="true" pageId="33" pageNumber="34">Rana daorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2019041006AFFCFFC1ADF19FA721BCC" box="[1384,1510,1406,1427]" pageId="33" pageNumber="34" rank="species">new species</taxonomicNameLabel>
(ROM 38538); (
<emphasis id="B9322D3A006AFFCFF9F1DFFEFF3F1BF1" bold="true" box="[131,171,1433,1454]" pageId="33" pageNumber="34">I, J</emphasis>
)
<taxonomicName id="4C468AAB006AFFCFF9CFDFFEFEE91BF1" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[189,381,1433,1454]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="hmongorum" status="sp. nov.">
<emphasis id="B9322D3A006AFFCFF9CFDFFEFEE91BF1" box="[189,381,1433,1454]" italics="true" pageId="33" pageNumber="34">Rana hmongorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2019041006AFFCFF8FFDFFEFD991BF1" box="[397,525,1433,1454]" pageId="33" pageNumber="34" rank="species">new species</taxonomicNameLabel>
(ROM 39891); (
<emphasis id="B9322D3A006AFFCFFBB8DFFEFD6B1BF1" bold="true" box="[714,767,1433,1454]" pageId="33" pageNumber="34">K, L</emphasis>
)
<taxonomicName id="4C468AAB006AFFCFFA60DFFEFC251BF1" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[786,945,1433,1454]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="morafkai" status="sp. nov.">
<emphasis id="B9322D3A006AFFCFFA60DFFEFC251BF1" box="[786,945,1433,1454]" italics="true" pageId="33" pageNumber="34">Rana morafkai</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2019041006AFFCFFAB0DFFEFBD71BF1" box="[962,1091,1433,1454]" pageId="33" pageNumber="34" rank="species">new species</taxonomicNameLabel>
(ROM 39906); (
<emphasis id="B9322D3A006AFFCFFD8DDFFEFAAF1BF1" bold="true" box="[1279,1339,1433,1454]" pageId="33" pageNumber="34">M, N</emphasis>
)
<taxonomicName id="4C468AAB006AFFCFFC3FDFFEFA6D1BF1" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[1357,1529,1433,1454]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="banaorum" status="sp. nov.">
<emphasis id="B9322D3A006AFFCFFC3FDFFEFA6D1BF1" box="[1357,1529,1433,1454]" italics="true" pageId="33" pageNumber="34">Rana banaorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2019041006AFFCFFF78DFFEF91E1BF1" box="[1546,1674,1433,1454]" pageId="33" pageNumber="34" rank="species">new species</taxonomicNameLabel>
(ROM39941); and (
<emphasis id="B9322D3A006AFFCFF822DFD4FE101B97" bold="true" box="[336,388,1459,1480]" pageId="33" pageNumber="34">O, P</emphasis>
)
<taxonomicName id="4C468AAB006AFFCFF8E4DFD3FDEF1B96" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[406,635,1460,1481]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="megatympanum" status="sp. nov.">
<emphasis id="B9322D3A006AFFCFF8E4DFD3FDEF1B96" box="[406,635,1460,1481]" italics="true" pageId="33" pageNumber="34">Rana megatympanum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2019041006AFFCFFBF9DFD3FC9E1B96" box="[651,778,1460,1481]" pageId="33" pageNumber="34" rank="species">new species</taxonomicNameLabel>
(ROM 26398).
</paragraph>
</caption>
<caption id="DF39A1A00069FFCCF91CDF18FF721BBB" ID-DOI="http://doi.org/10.5281/zenodo.4734911" ID-Zenodo-Dep="4734911" httpUri="https://zenodo.org/record/4734911/files/figure.png" pageId="34" pageNumber="35" startId="34.[110,148,1407,1428]" targetBox="[92,1628,413,1387]" targetPageId="34">
<paragraph id="8BF9F1280069FFCCF91CDF18FF721BBB" blockId="34.[84,1637,1407,1508]" pageId="34" pageNumber="35">
Fig. 13. Hands and feet of members of the
<taxonomicName id="4C468AAB0069FFCCFB3EDF18FD6A1BCB" box="[588,766,1407,1428]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="chloronota">
<emphasis id="B9322D3A0069FFCCFB3EDF18FD6A1BCB" box="[588,766,1407,1428]" italics="true" pageId="34" pageNumber="35">Rana chloronota</emphasis>
</taxonomicName>
complex: (
<emphasis id="B9322D3A0069FFCCFA05DF18FC1E1BCB" bold="true" box="[887,906,1407,1428]" pageId="34" pageNumber="35">A</emphasis>
)
<taxonomicName id="4C468AAB0069FFCCFAE8DF18FBD81BCB" box="[922,1100,1407,1428]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="chloronota">
<emphasis id="B9322D3A0069FFCCFAE8DF18FBD81BCB" box="[922,1100,1407,1428]" italics="true" pageId="34" pageNumber="35">Rana chloronota</emphasis>
</taxonomicName>
(ROM 26402); paratypes of (
<emphasis id="B9322D3A0069FFCCFCF8DF18FA0F1BCB" bold="true" box="[1418,1435,1407,1428]" pageId="34" pageNumber="35">B</emphasis>
)
<taxonomicName id="4C468AAB0069FFCCFCD9DF18F9CA1BCB" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[1451,1630,1407,1428]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="bacboensis" status="sp. nov.">
<emphasis id="B9322D3A0069FFCCFCD9DF18F9CA1BCB" box="[1451,1630,1407,1428]" italics="true" pageId="34" pageNumber="35">Rana bacboensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20190410069FFCCF926DFFDFF471BF0" box="[84,211,1434,1455]" pageId="34" pageNumber="35" rank="species">new species</taxonomicNameLabel>
(ROM 29529); (
<emphasis id="B9322D3A0069FFCCF8FADFFEFE0F1BF1" bold="true" box="[392,411,1433,1454]" pageId="34" pageNumber="35">C</emphasis>
)
<taxonomicName id="4C468AAB0069FFCCF8DEDFFDFDA81BF0" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[428,572,1434,1455]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="daorum" status="sp. nov.">
<emphasis id="B9322D3A0069FFCCF8DEDFFDFDA81BF0" box="[428,572,1434,1455]" italics="true" pageId="34" pageNumber="35">Rana daorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20190410069FFCCFB38DFFDFD5C1BF0" box="[586,712,1434,1455]" pageId="34" pageNumber="35" rank="species">new species</taxonomicNameLabel>
(ROM 38538); (
<emphasis id="B9322D3A0069FFCCFA0CDFFEFC051BF1" bold="true" box="[894,913,1433,1454]" pageId="34" pageNumber="35">D</emphasis>
)
<taxonomicName id="4C468AAB0069FFCCFAD3DFFDFBCA1BF0" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[929,1118,1434,1455]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="hmongorum" status="sp. nov.">
<emphasis id="B9322D3A0069FFCCFAD3DFFDFBCA1BF0" box="[929,1118,1434,1455]" italics="true" pageId="34" pageNumber="35">Rana hmongorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20190410069FFCCFD1EDFFDFB7E1BF0" box="[1132,1258,1434,1455]" pageId="34" pageNumber="35" rank="species">new species</taxonomicNameLabel>
(ROM 26379); (
<emphasis id="B9322D3A0069FFCCFCD2DFFEFA251BF1" bold="true" box="[1440,1457,1433,1454]" pageId="34" pageNumber="35">E</emphasis>
)
<taxonomicName id="4C468AAB0069FFCCFCB3DFFDF9CA1BF0" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[1473,1630,1434,1455]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="morafkai" status="sp. nov.">
<emphasis id="B9322D3A0069FFCCFCB3DFFDF9CA1BF0" box="[1473,1630,1434,1455]" italics="true" pageId="34" pageNumber="35">Rana morafkai</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20190410069FFCCF926DFD3FF411B96" box="[84,213,1460,1481]" pageId="34" pageNumber="35" rank="species">new species</taxonomicNameLabel>
(ROM 39906); (
<emphasis id="B9322D3A0069FFCCF8E3DFD3FE351B96" bold="true" box="[401,417,1460,1481]" pageId="34" pageNumber="35">F</emphasis>
)
<taxonomicName id="4C468AAB0069FFCCF8C1DFD2FDCB1B95" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[435,607,1461,1482]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="banaorum" status="sp. nov.">
<emphasis id="B9322D3A0069FFCCF8C1DFD2FDCB1B95" box="[435,607,1461,1482]" italics="true" pageId="34" pageNumber="35">Rana banaorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20190410069FFCCFB02DFD3FD641B96" box="[624,752,1460,1481]" pageId="34" pageNumber="35" rank="species">new species</taxonomicNameLabel>
(ROM 39941); and (
<emphasis id="B9322D3A0069FFCCFAAEDFD3FC641B96" bold="true" box="[988,1008,1460,1481]" pageId="34" pageNumber="35">G</emphasis>
)
<taxonomicName id="4C468AAB0069FFCCFD71DFD2FB7D1B95" authority="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC, 2003" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[1027,1257,1461,1482]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="megatympanum" status="sp. nov.">
<emphasis id="B9322D3A0069FFCCFD71DFD2FB7D1B95" box="[1027,1257,1461,1482]" italics="true" pageId="34" pageNumber="35">Rana megatympanum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20190410069FFCCFD8BDFD3FAED1B96" box="[1273,1401,1460,1481]" pageId="34" pageNumber="35" rank="species">new species</taxonomicNameLabel>
(ROM 26398). Scale equals 5 mm.
</paragraph>
</caption>
<paragraph id="8BF9F1280068FFCDF9EADB16FDBB1AA7" blockId="35.[123,652,193,1741]" pageId="35" pageNumber="36">
<materialsCitation id="3B2EFB750068FFCDF9EADB16FE321FBF" ID-GBIF-Occurrence="3408947312" accessionNumber="ROM 29531" collectingDate="1994-10-24" collectorName="I. S. Darevsky &amp; L. A. Lowcock &amp; R. W. Murphy &amp; N. L. Orlov" country="Vietnam" county="Con Cuong District" location="Khe Moi" municipality="Con Cuong village" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Nghe An" typeStatus="paratype">
<typeStatus id="54FD4F8A0068FFCDF9EADB16FE821FD8" box="[152,278,369,392]" pageId="35" pageNumber="36" type="paratype">PARATYPES</typeStatus>
: ROM 2953129533, all females, collected with holotype on
<date id="FFF8D7E80068FFCDFB7BDBE9FF211F9C" pageId="35" pageNumber="36" value="1994-10-24">
<collectingDate id="EFBC2E000068FFCDFB7BDBE9FF211F9C" pageId="35" pageNumber="36" value="1994-10-24">24 October 1994</collectingDate>
</date>
by I.S. Darevsky, L.A. Lowcock, R.W. Murphy, and N.L. Orlov;
</materialsCitation>
<materialsCitation id="3B2EFB750068FFCDF8C2DBAEFF2D1C67" ID-GBIF-Occurrence="3408947329" accessionNumber="ROM 26404" collectingDate="1995-06-05" collectorName="A. Lathrop &amp; R. W. Murphy" country="Vietnam" location="Orlov" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Nghe An" typeStatus="paratype">
ROM 26404, a female collected at the type locality by
<collectorName id="26B394FE0068FFCDFB4FDB81FF2F1C44" pageId="35" pageNumber="36">A. Lathrop</collectorName>
,
<collectorName id="26B394FE0068FFCDF9BAD863FEFE1C44" box="[200,362,516,539]" pageId="35" pageNumber="36">R.W. Murphy</collectorName>
, and N.
<location id="8E99A7F30068FFCDF8BCD863FD861C44" LSID="urn:lsid:plazi:treatment:03EF403E0054FFCBFA7FDF8EFBC01F4C:8E99A7F30068FFCDF8BCD863FD861C44" box="[462,530,516,539]" country="Vietnam" name="Orlov" pageId="35" pageNumber="36" stateProvince="Nghe An">Orlov</location>
on
<date id="FFF8D7E80068FFCDFB4DD863FF2D1C67" pageId="35" pageNumber="36" value="1995-06-05">
<collectingDate id="EFBC2E000068FFCDFB4DD863FF2D1C67" pageId="35" pageNumber="36" value="1995-06-05">5 June 1995</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2EFB750068FFCDF9B6D846FD1F1C0A" ID-GBIF-Occurrence="3408947336" accessionNumber="ROM 26357" collectingDate="1995-06-05" country="Vietnam" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Nghe An" typeStatus="paratype">
ROM 2635726358, adult females collected from the type locality on
<date id="FFF8D7E80068FFCDF889D859FD1F1C0A" box="[507,651,574,597]" pageId="35" pageNumber="36" value="1995-06-05">
<collectingDate id="EFBC2E000068FFCDF889D859FD1F1C0A" box="[507,651,574,597]" pageId="35" pageNumber="36" value="1995-06-05">5 June 1995</collectingDate>
</date>
</materialsCitation>
by A. Lathrop, R.W. Murphy, and N.L. Orlov;
<materialsCitation id="3B2EFB750068FFCDF9CBD81EFE751D5A" ID-GBIF-Occurrence="3408947333" accessionNumber="ROM 29359" collectingDate="1995-05-24" collectorName="R. H. Bain" country="Vietnam" location="Ba Be Lake" municipality="Ba Be Lake National Park" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Bac Kan" typeStatus="paratype">
ROM 29359, a female, from
<location id="8E99A7F30068FFCDFB48D81EFF2C1CF2" LSID="urn:lsid:plazi:treatment:03EF403E0054FFCBFA7FDF8EFBC01F4C:8E99A7F30068FFCDFB48D81EFF2C1CF2" country="Vietnam" municipality="Ba Be Lake National Park" name="Ba Be Lake" pageId="35" pageNumber="36" stateProvince="Bac Kan">Ba Be Lake</location>
,
<collectingMunicipality id="6B9D6B520068FFCDF9BAD8F1FD841CF2" box="[200,528,662,685]" pageId="35" pageNumber="36">Ba Be Lake National Park</collectingMunicipality>
,
<collectingRegion id="49823FCA0068FFCDFB53D8F1FF761C94" country="Vietnam" name="Bac Kan" pageId="35" pageNumber="36">Bac Kan Province</collectingRegion>
(formerly
<collectingRegion id="49823FCA0068FFCDF817D8D3FDC41C94" box="[357,592,692,715]" country="Vietnam" name="Cao Bang" pageId="35" pageNumber="36">Cao Bang Province</collectingRegion>
),
<collectingCountry id="F351B1B80068FFCDFB17D8D3FF561CB7" name="Vietnam" pageId="35" pageNumber="36">Vietnam</collectingCountry>
(22°25̍05̎N, 105°38̍05̎E), collected by
<collectorName id="26B394FE0068FFCDF9D2D889FE821D5A" box="[160,278,750,773]" pageId="35" pageNumber="36">R.H. Bain</collectorName>
on
<date id="FFF8D7E80068FFCDF836D889FE751D5A" box="[324,481,750,773]" pageId="35" pageNumber="36" value="1995-05-24">
<collectingDate id="EFBC2E000068FFCDF836D889FE751D5A" box="[324,481,750,773]" pageId="35" pageNumber="36" value="1995-05-24">24 May 1995</collectingDate>
</date>
</materialsCitation>
<materialsCitation id="3B2EFB750068FFCDF898D889FDA01D7C" ID-GBIF-Occurrence="3408947320" country="Vietnam" location="Ba Be Lake" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Bac Kan" typeStatus="paratype">
at the outflow on the south side of
<location id="8E99A7F30068FFCDF8E6D96BFDA01D7C" LSID="urn:lsid:plazi:treatment:03EF403E0054FFCBFA7FDF8EFBC01F4C:8E99A7F30068FFCDF8E6D96BFDA01D7C" box="[404,564,780,803]" country="Vietnam" name="Ba Be Lake" pageId="35" pageNumber="36" stateProvince="Bac Kan">Ba Be Lake</location>
</materialsCitation>
;
<materialsCitation id="3B2EFB750068FFCDFB3BD96BFEC31D8C" ID-GBIF-Occurrence="3408947327" accessionNumber="ROM 29526" collectingDate="1996-05-25" collectingDateMax="1996-05-30" collectingDateMin="1996-05-25" collectorName="R. W. Murphy &amp; A. Lathrop" country="Vietnam" location="Na Hang Nature Reserve" municipality="Pac Ban village" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Tuyen Quang" typeStatus="paratype">
ROM 2952629530, all females, from
<location id="8E99A7F30068FFCDFB68D94EFEA21D02" LSID="urn:lsid:plazi:treatment:03EF403E0054FFCBFA7FDF8EFBC01F4C:8E99A7F30068FFCDFB68D94EFEA21D02" country="Vietnam" municipality="Pac Ban village" name="Na Hang Nature Reserve" pageId="35" pageNumber="36" stateProvince="Tuyen Quang">Na Hang Nature Reserve</location>
,
<collectingRegion id="49823FCA0068FFCDF836D921FDC31D02" box="[324,599,838,861]" country="Vietnam" name="Tuyen Quang" pageId="35" pageNumber="36">Tuyen Quang Province</collectingRegion>
,
<collectingCountry id="F351B1B80068FFCDFB17D921FF561D24" name="Vietnam" pageId="35" pageNumber="36">Vietnam</collectingCountry>
(22°21̍54̎N, 105°25̍40̎E) approximately
<quantity id="4CBE5CCD0068FFCDF9A0D9E6FE881DC7" box="[210,284,897,920]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="35" pageNumber="36" unit="km" value="15.0">15 km</quantity>
from
<collectingMunicipality id="6B9D6B520068FFCDF816D9E6FD8F1DC7" box="[356,539,897,920]" pageId="35" pageNumber="36">Pac Ban village</collectingMunicipality>
collected by
<collectorName id="26B394FE0068FFCDF9D3D9F9FED51DEA" box="[161,321,926,949]" pageId="35" pageNumber="36">R.W. Murphy</collectorName>
and
<collectorName id="26B394FE0068FFCDF80ED9F9FE681DEA" box="[380,508,926,949]" pageId="35" pageNumber="36">A. Lathrop</collectorName>
between
<date id="FFF8D7E80068FFCDFB1CD9F9FEC31D8C" pageId="35" pageNumber="36" value="1996-05-25" valueMax="1996-05-30" valueMin="1996-05-25">
<collectingDate id="EFBC2E000068FFCDFB1CD9F9FEC31D8C" pageId="35" pageNumber="36" value="1996-05-25" valueMax="1996-05-30" valueMin="1996-05-25">25 and 30 May 1996</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2EFB750068FFCDF815D9DBFD1F1A17" ID-GBIF-Occurrence="3408947306" collectingDate="1998-04-29" country="Vietnam" county="Con Cuong District" elevation="300" location="Hieng Stream" municipality="Chau Khe Commune" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Nghe An" typeStatus="paratype">
AMNH A­161248, a female, from
<location id="8E99A7F30068FFCDF879D9BEFE271DAF" LSID="urn:lsid:plazi:treatment:03EF403E0054FFCBFA7FDF8EFBC01F4C:8E99A7F30068FFCDF879D9BEFE271DAF" box="[267,435,985,1008]" country="Vietnam" county="Con Cuong District" municipality="Chau Khe Commune" name="Hieng Stream" pageId="35" pageNumber="36" stateProvince="Nghe An">Hieng Stream</location>
,
<collectingMunicipality id="6B9D6B520068FFCDF8B1D9BEFF2B1A52" pageId="35" pageNumber="36">Chau Khe Commune</collectingMunicipality>
,
<collectingCounty id="629889A40068FFCDF9BDD991FE2B1A52" box="[207,447,1014,1037]" pageId="35" pageNumber="36">Con Cuong District</collectingCounty>
,
<collectingRegion id="49823FCA0068FFCDF8A2D991FF3A1A74" country="Vietnam" name="Nghe An" pageId="35" pageNumber="36">Nghe An Province</collectingRegion>
,
<collectingCountry id="F351B1B80068FFCDF9CEDE73FE8B1A74" box="[188,287,1044,1067]" name="Vietnam" pageId="35" pageNumber="36">Vietnam</collectingCountry>
(19°02̍17̎N, 104°42̍06̎E,
<elevation id="006B161B0068FFCDFB1EDE73FEB81A17" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="36" unit="m" value="300.0">
elevation
<quantity id="4CBE5CCD0068FFCDF9AEDE56FEB81A17" box="[220,300,1073,1096]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="36" unit="m" value="300.0">300 m</quantity>
</elevation>
) collected on
<date id="FFF8D7E80068FFCDF8ADDE56FD1F1A17" box="[479,651,1073,1096]" pageId="35" pageNumber="36" value="1998-04-29">
<collectingDate id="EFBC2E000068FFCDF8ADDE56FD1F1A17" box="[479,651,1073,1096]" pageId="35" pageNumber="36" value="1998-04-29">29 April 1998</collectingDate>
</date>
</materialsCitation>
<materialsCitation id="3B2EFB750068FFCDF909DE29FEAF1AA7" ID-GBIF-Occurrence="3408947317" collectingDate="1998-09-08" collectorName="N. Q. Truong" country="Vietnam" county="Con Cuong District" elevation="300" location="Khe Chat Stream" municipality="Pu Mat Nature Reserve" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Nghe An" typeStatus="paratype">
by
<collectorName id="26B394FE0068FFCDF9D7DE29FED61A3A" box="[165,322,1102,1125]" pageId="35" pageNumber="36">N.Q. Truong</collectorName>
; FMNH 255611 (adult female), 255612 (adult male) along the
<location id="8E99A7F30068FFCDFB28DE0BFE801AFF" LSID="urn:lsid:plazi:treatment:03EF403E0054FFCBFA7FDF8EFBC01F4C:8E99A7F30068FFCDFB28DE0BFE801AFF" country="Vietnam" county="Con Cuong District" municipality="Pu Mat Nature Reserve" name="Khe Chat Stream" pageId="35" pageNumber="36" stateProvince="Nghe An">Khe Chat Stream</location>
,
<collectingMunicipality id="6B9D6B520068FFCDF857DEEEFDDE1AFF" box="[293,586,1161,1184]" pageId="35" pageNumber="36">Pu Mat Nature Reserve</collectingMunicipality>
,
<collectingCounty id="629889A40068FFCDFB28DEEEFEB81AE2" pageId="35" pageNumber="36">Con Cuong District</collectingCounty>
,
<collectingRegion id="49823FCA0068FFCDF848DEC1FD8E1AE2" box="[314,538,1190,1213]" country="Vietnam" name="Nghe An" pageId="35" pageNumber="36">Nghe An Province</collectingRegion>
,
<collectingCountry id="F351B1B80068FFCDFB5ADEC1FD1F1AE2" box="[552,651,1190,1213]" name="Vietnam" pageId="35" pageNumber="36">Vietnam</collectingCountry>
(18°56̍N, 104°45̍E,
<elevation id="006B161B0068FFCDF80ADEA3FDAB1A84" box="[376,575,1220,1243]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="36" unit="m" value="300.0">
elevation
<quantity id="4CBE5CCD0068FFCDF89DDEA3FDAB1A84" box="[495,575,1220,1243]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="36" unit="m" value="300.0">300 m</quantity>
</elevation>
) on
<date id="FFF8D7E80068FFCDFB0EDEA3FEAF1AA7" pageId="35" pageNumber="36" value="1998-09-08">
<collectingDate id="EFBC2E000068FFCDFB0EDEA3FEAF1AA7" pageId="35" pageNumber="36" value="1998-09-08">8 September 1998</collectingDate>
</date>
</materialsCitation>
by Bryan L. Stuart.
</paragraph>
</subSubSection>
<subSubSection id="C35CA2A30068FFCAF9EADE99FF5C1AA7" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="8BF9F1280068FFCDF9EADE99FB581CCF" blockId="35.[123,652,193,1741]" lastBlockId="35.[699,1228,193,1742]" pageId="35" pageNumber="36">
DIAGNOSIS:
<taxonomicName id="4C468AAB0068FFCDF853DE98FE731B49" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[289,487,1279,1302]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A0068FFCDF853DE98FE731B49" box="[289,487,1279,1302]" italics="true" pageId="35" pageNumber="36">Rana bacboensis</emphasis>
</taxonomicName>
, a member of the subgenus
<taxonomicName id="4C468AAB0068FFCDF85EDF7BFE341B6C" authorityName="Fei, Ye, and Huang" authorityYear="1990" box="[300,416,1308,1331]" class="Amphibia" family="Ranidae" genus="Odorrana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="genus">
<emphasis id="B9322D3A0068FFCDF85EDF7BFE341B6C" box="[300,416,1308,1331]" italics="true" pageId="35" pageNumber="36">Odorrana</emphasis>
</taxonomicName>
(sensu
<bibRefCitation id="EFD78CD90068FFCDFB7EDF7BFF2D1B0F" author="Fei, L." pageId="35" pageNumber="36" refId="ref37298" refString="Fei, L., C. - y. Ye, and Y. - z. Huang. 1990. Keys to Chinese Amphibians. Chongqing: Chongqing Branch, Science and Technology Materials Press, 364 pp. [in Chinese]" type="book" year="1990">Fei et al., 1990</bibRefCitation>
), is characterized by a combination of the following attributes: (1) body dorsoventrally compressed; (2) SVL of males
<quantity id="4CBE5CCD0068FFCDFB2ADF13FF3F1BF7" metricMagnitude="-2" metricUnit="m" metricValue="5.49" pageId="35" pageNumber="36" unit="mm" value="54.9">54.9 mm</quantity>
, means of females
<quantity id="4CBE5CCD0068FFCDF8F9DFF6FE481BF7" box="[395,476,1425,1448]" metricMagnitude="-2" metricUnit="m" metricValue="9.6" pageId="35" pageNumber="36" unit="mm" value="96.0">96 mm</quantity>
(
<quantity id="4CBE5CCD0068FFCDF89EDFF6FDEA1BF7" box="[492,638,1425,1448]" metricMagnitude="-2" metricUnit="m" metricValue="9.35" metricValueMax="10.5" metricValueMin="8.2" pageId="35" pageNumber="36" unit="mm" value="93.5" valueMax="105.0" valueMin="82.0">82105 mm</quantity>
); (3) vomerine teeth present in rows oblique to choanae; (4) vertical black stripes on upper lip (especially under eye), light colored glandule above insertion of arm; (5) head broad, bluntly rounded in profile; (6) tympanum circular, distinct, TMP:EYE
<quantity id="4CBE5CCD0068FFCDF8B0DC26FD8F1807" box="[450,539,1601,1624]" metricMagnitude="-2" metricUnit="m" metricValue="1.0922" pageId="35" pageNumber="36" unit="in" value="0.43">0.43 in</quantity>
females,
<quantity id="4CBE5CCD0068FFCDF909DC39FF42182A" box="[123,214,1630,1653]" metricMagnitude="-2" metricUnit="m" metricValue="1.6764" pageId="35" pageNumber="36" unit="in" value="0.66">0.66 in</quantity>
the male; (7) supratympanic fold weak; (8) dorsal skin shagreened, becoming granular laterally, dorsolateral fold absent; venter smooth; (9) dorsum brown with black blotches; forelimbs and hindlimbs with transverse bands of distinct blotches to tips of digits; (10) median callous pad on base of fingers II and III to proximal tubercle; (11) disks on fingers and toes enlarged (&lt;2× base of phalanges); (12) feet fully webbed to disks, but as a fringe from distal subarticular tubercle of IV, slight lateral fringes on toes I and V to terminal phalanges, webbing marbled brown on white; (13) subarticular tubercles distinct, conical; inner metatarsal tubercle distinct, ovoid; (14) terminal phalanges T­ shaped; (15) xiphisternum large, deeply notched posteriorly; (16) male with velvety nuptial pad on thumb, paired gular pouches, no pectoral spines; (17) eggs black.
</paragraph>
<paragraph id="8BF9F1280068FFCAFBAAD8F1FF5C1AA7" blockId="35.[699,1228,193,1742]" lastBlockId="36.[124,653,193,1741]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
COMPARISONS:
<taxonomicName id="4C468AAB0068FFCDFAFAD8F0FBC61CF1" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[904,1106,663,686]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A0068FFCDFAFAD8F0FBC61CF1" box="[904,1106,663,686]" italics="true" pageId="35" pageNumber="36">Rana bacboensis</emphasis>
</taxonomicName>
is one of the larger species of cascade ranids (SVL female
<quantity id="4CBE5CCD0068FFCDFA73D8B6FC0C1CB7" box="[769,920,721,744]" metricMagnitude="-2" metricUnit="m" metricValue="9.3" metricValueMax="10.5" metricValueMin="8.1" pageId="35" pageNumber="36" unit="mm" value="93.0" valueMax="105.0" valueMin="81.0">81105 mm</quantity>
). It can be distinguished from all other cascade ranids of Southeast Asia by its dark, pigmented eggs (immaculate white or white with melanic pole in other species) (table 12). Its black vertical lip­bars distinguish it from
<taxonomicName id="4C468AAB0068FFCDFAD6D903FBCD1D24" authorityName="Inger &amp; Chan-ard" authorityYear="1997" box="[932,1113,868,891]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="archotaphus">
<emphasis id="B9322D3A0068FFCDFAD6D903FBCD1D24" box="[932,1113,868,891]" italics="true" pageId="35" pageNumber="36">R. archotaphus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFD19D903FC9E1DC7" baseAuthorityName="Schlegel" baseAuthorityYear="1837" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A0068FFCDFD19D903FC9E1DC7" italics="true" pageId="35" pageNumber="36">R. chalconota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFA6CD9E6FC501DC7" box="[798,964,897,920]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="chloronota">
<emphasis id="B9322D3A0068FFCDFA6CD9E6FC501DC7" box="[798,964,897,920]" italics="true" pageId="35" pageNumber="36">R. chloronota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFAA5D9E6FBF41DC7" authorityName="Boulenger" authorityYear="1917" box="[983,1120,897,920]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="grahami">
<emphasis id="B9322D3A0068FFCDFAA5D9E6FBF41DC7" box="[983,1120,897,920]" italics="true" pageId="35" pageNumber="36">R. grahami</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFD01D9E6FC961DE9" authorityName=": Boulenger" authorityYear="1899" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="graminea">
<emphasis id="B9322D3A0068FFCDFD01D9E6FC961DE9" italics="true" pageId="35" pageNumber="36">R. graminea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFA61D9F8FC531DE9" authorityName="Y.u.Deng" authorityYear="1992" box="[787,967,927,950]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="hejiangensis">
<emphasis id="B9322D3A0068FFCDFA61D9F8FC531DE9" box="[787,967,927,950]" italics="true" pageId="35" pageNumber="36">R. hejiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFAABD9F8FBA11DE9" authorityName="Boulenger" authorityYear="1891" box="[985,1077,927,950]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="hosii">
<emphasis id="B9322D3A0068FFCDFAABD9F8FBA11DE9" box="[985,1077,927,950]" italics="true" pageId="35" pageNumber="36">R. hosii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFD34D9F8FC901D8C" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A0068FFCDFD34D9F8FC901D8C" italics="true" pageId="35" pageNumber="36">R. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFA60D9DBFC4F1D8C" box="[786,987,956,979]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="kwangwuensis">
<emphasis id="B9322D3A0068FFCDFA60D9DBFC4F1D8C" box="[786,987,956,979]" italics="true" pageId="35" pageNumber="36">R. kwangwuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFA9BD9DBFBE21D8C" authorityName=": Werner" authorityYear="1930" box="[1001,1142,956,979]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="leporipes">
<emphasis id="B9322D3A0068FFCDFA9BD9DBFBE21D8C" box="[1001,1142,956,979]" italics="true" pageId="35" pageNumber="36">R. leporipes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFDF6D9DBFD741DAF" baseAuthorityName="Boulenger" baseAuthorityYear="1887" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="livida">
<emphasis id="B9322D3A0068FFCDFDF6D9DBFD741DAF" italics="true" pageId="35" pageNumber="36">R. livida</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFB80D9BEFC0F1DAF" authorityName="Boettger" authorityYear="1892" box="[754,923,985,1008]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="schmackeri">
<emphasis id="B9322D3A0068FFCDFB80D9BEFC0F1DAF" box="[754,923,985,1008]" italics="true" pageId="35" pageNumber="36">R. schmackeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFADCD9BEFB831DAF" authorityName=": Ahl" authorityYear="1925" box="[942,1047,985,1008]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="sinica">
<emphasis id="B9322D3A0068FFCDFADCD9BEFB831DAF" box="[942,1047,985,1008]" italics="true" pageId="35" pageNumber="36">R. sinica</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C468AAB0068FFCDFD2DD9BEFD7E1A51" baseAuthorityName="Yang" baseAuthorityYear="1991" class="Amphibia" family="Ranidae" genus="Huia" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="nasica">
<emphasis id="B9322D3A0068FFCDFD2DD9BEFD7E1A51" italics="true" pageId="35" pageNumber="36">Huia nasica</emphasis>
</taxonomicName>
. The brown dorsum with dark spots distinguishes it from
<taxonomicName id="4C468AAB0068FFCDFAAADE73FB021A74" authorityName="Inger &amp; Chan-ard" authorityYear="1997" box="[984,1174,1044,1067]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="archotaphus">
<emphasis id="B9322D3A0068FFCDFAAADE73FB021A74" box="[984,1174,1044,1067]" italics="true" pageId="35" pageNumber="36">R. archotaphus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFDC0DE73FCA81A17" baseAuthorityName="Schlegel" baseAuthorityYear="1837" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A0068FFCDFDC0DE73FCA81A17" italics="true" pageId="35" pageNumber="36">R. chalconota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFA20DE56FC6E1A17" box="[850,1018,1073,1096]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="chloronota">
<emphasis id="B9322D3A0068FFCDFA20DE56FC6E1A17" box="[850,1018,1073,1096]" italics="true" pageId="35" pageNumber="36">R. chloronota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFD63DE56FB0F1A17" authorityName="Boulenger" authorityYear="1917" box="[1041,1179,1073,1096]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="grahami">
<emphasis id="B9322D3A0068FFCDFD63DE56FB0F1A17" box="[1041,1179,1073,1096]" italics="true" pageId="35" pageNumber="36">R. grahami</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFDC0DE56FCBE1A39" authorityName=": Boulenger" authorityYear="1899" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="graminea">
<emphasis id="B9322D3A0068FFCDFDC0DE56FCBE1A39" italics="true" pageId="35" pageNumber="36">R. graminea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFA4EDE28FC651A39" authorityName="Y.u.Deng" authorityYear="1992" box="[828,1009,1103,1126]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="hejiangensis">
<emphasis id="B9322D3A0068FFCDFA4EDE28FC651A39" box="[828,1009,1103,1126]" italics="true" pageId="35" pageNumber="36">R. hejiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFD71DE28FBCB1A39" authorityName="Boulenger" authorityYear="1891" box="[1027,1119,1103,1126]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="hosii">
<emphasis id="B9322D3A0068FFCDFD71DE28FBCB1A39" box="[1027,1119,1103,1126]" italics="true" pageId="35" pageNumber="36">R. hosii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFD03DE28FCBB1ADC" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A0068FFCDFD03DE28FCBB1ADC" italics="true" pageId="35" pageNumber="36">R. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFA4CDE0BFC791ADC" baseAuthorityName="Huang, Fei &amp; Ye" baseAuthorityYear="2001" box="[830,1005,1132,1155]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="junlianensis">
<emphasis id="B9322D3A0068FFCDFA4CDE0BFC791ADC" box="[830,1005,1132,1155]" italics="true" pageId="35" pageNumber="36">R. junlianensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFA89DE0BFB501ADC" box="[1019,1220,1132,1155]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="kwangwuensis">
<emphasis id="B9322D3A0068FFCDFA89DE0BFB501ADC" box="[1019,1220,1132,1155]" italics="true" pageId="35" pageNumber="36">R. kwangwuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFBC9DEEEFCDF1AFF" authorityName=": Werner" authorityYear="1930" box="[699,843,1161,1184]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="leporipes">
<emphasis id="B9322D3A0068FFCDFBC9DEEEFCDF1AFF" box="[699,843,1161,1184]" italics="true" pageId="35" pageNumber="36">R. leporipes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFA2CDEEEFB9E1AFF" authorityName="Liu" authorityYear="1950" box="[862,1034,1161,1184]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="margaretae">
<emphasis id="B9322D3A0068FFCDFA2CDEEEFB9E1AFF" box="[862,1034,1161,1184]" italics="true" pageId="35" pageNumber="36">R. margaretae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFD6EDEEEFB501AFF" authorityName="Boettger" authorityYear="1892" box="[1052,1220,1161,1184]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="schmackeri">
<emphasis id="B9322D3A0068FFCDFD6EDEEEFB501AFF" box="[1052,1220,1161,1184]" italics="true" pageId="35" pageNumber="36">R. schmackeri</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C468AAB0068FFCDFB9DDEC0FCCC1AE1" authorityName=": Ahl" authorityYear="1925" box="[751,856,1191,1214]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="sinica">
<emphasis id="B9322D3A0068FFCDFB9DDEC0FCCC1AE1" box="[751,856,1191,1214]" italics="true" pageId="35" pageNumber="36">R. sinica</emphasis>
</taxonomicName>
, all of which have a predominantly green dorsum. The presence of gular pouches in males differentiates
<taxonomicName id="4C468AAB0068FFCDFD58DE86FB5F1AA7" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[1066,1227,1249,1272]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A0068FFCDFD58DE86FB5F1AA7" box="[1066,1227,1249,1272]" italics="true" pageId="35" pageNumber="36">R. bacboensis</emphasis>
</taxonomicName>
from
<taxonomicName id="4C468AAB0068FFCDFA72DE98FC361B49" authorityName="Boulenger" authorityYear="1882" box="[768,930,1279,1302]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A0068FFCDFA72DE98FC361B49" box="[768,930,1279,1302]" italics="true" pageId="35" pageNumber="36">R. andersonii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFAC5DE98FBCA1B49" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[951,1118,1279,1302]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A0068FFCDFAC5DE98FBCA1B49" box="[951,1118,1279,1302]" italics="true" pageId="35" pageNumber="36">R. chalconota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFD01DE98FD611B6C" authorityName="Boulenger" authorityYear="1917" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="grahami">
<emphasis id="B9322D3A0068FFCDFD01DE98FD611B6C" italics="true" pageId="35" pageNumber="36">R. grahami</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFA7BDF7BFC2F1B6C" box="[777,955,1308,1331]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B9322D3A0068FFCDFA7BDF7BFC2F1B6C" box="[777,955,1308,1331]" italics="true" pageId="35" pageNumber="36">R. hainanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFABDDF7BFBBA1B6C" authorityName="Boulenger" authorityYear="1891" box="[975,1070,1308,1331]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="hosii">
<emphasis id="B9322D3A0068FFCDFABDDF7BFBBA1B6C" box="[975,1070,1308,1331]" italics="true" pageId="35" pageNumber="36">R. hosii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFD30DF7BFC901B0F" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A0068FFCDFD30DF7BFC901B0F" italics="true" pageId="35" pageNumber="36">R. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFA6CDF5EFC431B0F" baseAuthorityName="Huang, Fei &amp; Ye" baseAuthorityYear="2001" box="[798,983,1337,1360]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="junlianensis">
<emphasis id="B9322D3A0068FFCDFA6CDF5EFC431B0F" box="[798,983,1337,1360]" italics="true" pageId="35" pageNumber="36">R. junlianensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB0068FFCDFA82DF5EFB501B0F" box="[1008,1220,1337,1360]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="kwangwuensis">
<emphasis id="B9322D3A0068FFCDFA82DF5EFB501B0F" box="[1008,1220,1337,1360]" italics="true" pageId="35" pageNumber="36">R. kwangwuensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C468AAB0068FFCDFB8BDF30FC221B31" authorityName="Liu" authorityYear="1950" box="[761,950,1367,1390]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="margaretae">
<emphasis id="B9322D3A0068FFCDFB8BDF30FC221B31" box="[761,950,1367,1390]" italics="true" pageId="35" pageNumber="36">R. margaretae</emphasis>
</taxonomicName>
.
<taxonomicName id="4C468AAB0068FFCDFAA3DF30FBE51B31" baseAuthorityName="Yang" baseAuthorityYear="1991" box="[977,1137,1367,1390]" class="Amphibia" family="Ranidae" genus="Huia" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="nasica">
<emphasis id="B9322D3A0068FFCDFAA3DF30FBE51B31" box="[977,1137,1367,1390]" italics="true" pageId="35" pageNumber="36">Huia nasica</emphasis>
</taxonomicName>
has a smooth, olive­brown dorsum with lighter flanks (
<taxonomicName id="4C468AAB0068FFCDFA61DFF6FC231BF7" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[787,951,1425,1448]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A0068FFCDFA61DFF6FC231BF7" box="[787,951,1425,1448]" italics="true" pageId="35" pageNumber="36">R. bacboensis</emphasis>
</taxonomicName>
is shagreened and uniformly brown with black spots both on the dorsum and flanks), and adult females are smaller than those of
<taxonomicName id="4C468AAB0068FFCDFAB2DF8EFBF0185F" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[960,1124,1513,1536]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A0068FFCDFAB2DF8EFBF0185F" box="[960,1124,1513,1536]" italics="true" pageId="35" pageNumber="36">R. bacboensis</emphasis>
</taxonomicName>
(
<quantity id="4CBE5CCD0068FFCDFD04DF8EFB5F185F" box="[1142,1227,1513,1536]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="35" pageNumber="36" unit="mm" value="67.0">67 mm</quantity>
vs.&gt;
<quantity id="4CBE5CCD0068FFCDFB88DC61FCC61842" box="[762,850,1542,1565]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="35" pageNumber="36" unit="mm" value="80.0">80 mm</quantity>
). The absence of an outer metatarsal tubercle and the large female SVL also distinguishes
<taxonomicName id="4C468AAB0068FFCDFA2DDC26FB921807" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[863,1030,1601,1624]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A0068FFCDFA2DDC26FB921807" box="[863,1030,1601,1624]" italics="true" pageId="35" pageNumber="36">R. bacboensis</emphasis>
</taxonomicName>
from
<taxonomicName id="4C468AAB0068FFCDFD2ADC26FC9D1829" authorityName="Inger &amp; Chan-ard" authorityYear="1997" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="archotaphus">
<emphasis id="B9322D3A0068FFCDFD2ADC26FC9D1829" italics="true" pageId="35" pageNumber="36">R. archotaphus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C468AAB0068FFCDFA3BDC38FC7A1829" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[841,1006,1631,1654]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A0068FFCDFA3BDC38FC7A1829" box="[841,1006,1631,1654]" italics="true" pageId="35" pageNumber="36">R. chalconota</emphasis>
</taxonomicName>
(female SVL
<quantity id="4CBE5CCD0068FFCDFDEDDC39FC8B18CC" metricMagnitude="-2" metricUnit="m" metricValue="9.3" metricValueMax="10.5" metricValueMin="8.1" pageId="35" pageNumber="36" unit="mm" value="93.0" valueMax="105.0" valueMin="81.0">81 105 mm</quantity>
for
<taxonomicName id="4C468AAB0068FFCDFA2ADC1BFC6A18CC" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[856,1022,1660,1683]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A0068FFCDFA2ADC1BFC6A18CC" box="[856,1022,1660,1683]" italics="true" pageId="35" pageNumber="36">R. bacboensis</emphasis>
</taxonomicName>
,
<quantity id="4CBE5CCD0068FFCDFD63DC1BFB0118CC" box="[1041,1173,1660,1683]" metricMagnitude="-2" metricUnit="m" metricValue="6.05" metricValueMax="6.2" metricValueMin="5.9" pageId="35" pageNumber="36" unit="mm" value="60.5" valueMax="62.0" valueMin="59.0">5962 mm</quantity>
and
<quantity id="4CBE5CCD0068FFCDFBC9DCFEFCA818EF" box="[699,828,1689,1712]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="5.9" metricValueMin="4.6" pageId="35" pageNumber="36" unit="mm" value="52.5" valueMax="59.0" valueMin="46.0">4659 mm</quantity>
for
<taxonomicName id="4C468AAB0068FFCDFA07DCFEFBBE18EF" authorityName="Inger &amp; Chan-ard" authorityYear="1997" box="[885,1066,1689,1712]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="archotaphus">
<emphasis id="B9322D3A0068FFCDFA07DCFEFBBE18EF" box="[885,1066,1689,1712]" italics="true" pageId="35" pageNumber="36">R. archotaphus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C468AAB0068FFCDFD19DCFEFC9E1891" baseAuthorityName="Schlegel" baseAuthorityYear="1837" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A0068FFCDFD19DCFEFC9E1891" italics="true" pageId="35" pageNumber="36">R. chalconota</emphasis>
</taxonomicName>
, respectively). The absence of dorsolateral folds distinguishes
<taxonomicName id="4C468AAB006FFFCAF8DEDAA6FDD91E87" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[428,589,193,216]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A006FFFCAF8DEDAA6FDD91E87" box="[428,589,193,216]" italics="true" pageId="36" pageNumber="37">R. bacboensis</emphasis>
</taxonomicName>
from
<taxonomicName id="4C468AAB006FFFCAF90EDAB8FEB41EA9" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[124,288,223,246]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="chalconota">
<emphasis id="B9322D3A006FFFCAF90EDAB8FEB41EA9" box="[124,288,223,246]" italics="true" pageId="36" pageNumber="37">R. chalconota</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB006FFFCAF843DAB8FE571EA9" authorityName=": Boulenger" authorityYear="1899" box="[305,451,223,246]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="graminea">
<emphasis id="B9322D3A006FFFCAF843DAB8FE571EA9" box="[305,451,223,246]" italics="true" pageId="36" pageNumber="37">R. graminea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB006FFFCAF8A6DAB8FDA41EA9" authorityName="Boulenger" authorityYear="1891" box="[468,560,223,246]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="hosii">
<emphasis id="B9322D3A006FFFCAF8A6DAB8FDA41EA9" box="[468,560,223,246]" italics="true" pageId="36" pageNumber="37">R. hosii</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C468AAB006FFFCAFB01DAB8FF7C1F4C" authorityName=": Werner" authorityYear="1930" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="leporipes">
<emphasis id="B9322D3A006FFFCAFB01DAB8FF7C1F4C" italics="true" pageId="36" pageNumber="37">R. leporipes</emphasis>
</taxonomicName>
(pustules on the dorsum of
<taxonomicName id="4C468AAB006FFFCAFB4ADA9BFF221F6F" authorityName="Boulenger" authorityYear="1917" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="grahami">
<emphasis id="B9322D3A006FFFCAFB4ADA9BFF221F6F" italics="true" pageId="36" pageNumber="37">R. grahami</emphasis>
</taxonomicName>
sometimes form a dorsolateral fold).
<taxonomicName id="4C468AAB006FFFCAF90EDB50FED01F11" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[124,324,311,334]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A006FFFCAF90EDB50FED01F11" box="[124,324,311,334]" italics="true" pageId="36" pageNumber="37">Rana bacboensis</emphasis>
</taxonomicName>
has webbing to the base of the toe pad distinguishing it from
<taxonomicName id="4C468AAB006FFFCAFB68DB33FF371FD7" authorityName=": Werner" authorityYear="1930" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="leporipes">
<emphasis id="B9322D3A006FFFCAFB68DB33FF371FD7" italics="true" pageId="36" pageNumber="37">R. leporipes</emphasis>
</taxonomicName>
(webbing to distal phalanx).
<taxonomicName id="4C468AAB006FFFCAFB73DB16FD181FD7" authorityName=": Ahl" authorityYear="1925" box="[513,652,369,392]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="sinica">
<emphasis id="B9322D3A006FFFCAFB73DB16FD181FD7" box="[513,652,369,392]" italics="true" pageId="36" pageNumber="37">Rana sinica</emphasis>
</taxonomicName>
can further be distinguished from
<taxonomicName id="4C468AAB006FFFCAFB60DBE8FF471F9C" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A006FFFCAFB60DBE8FF471F9C" italics="true" pageId="36" pageNumber="37">Rana bacboensis</emphasis>
</taxonomicName>
by its indistinct, skin­covered tympanum, and different finger formula (I &lt;II &lt;IV for
<taxonomicName id="4C468AAB006FFFCAF981DB80FE141FA2" authority=", II" authorityName="II" box="[243,384,486,510]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="sinica">
<emphasis id="B9322D3A006FFFCAF981DB80FECF1FA1" box="[243,347,487,510]" italics="true" pageId="36" pageNumber="37">R. sinica</emphasis>
, II
</taxonomicName>
&lt;I &lt;IV for
<taxonomicName id="4C468AAB006FFFCAFB47DB80FF471C44" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A006FFFCAFB47DB80FF471C44" italics="true" pageId="36" pageNumber="37">R. bacboensis</emphasis>
</taxonomicName>
).
<taxonomicName id="4C468AAB006FFFCAF982D863FE2F1C44" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[240,443,516,539]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A006FFFCAF982D863FE2F1C44" box="[240,443,516,539]" italics="true" pageId="36" pageNumber="37">Rana bacboensis</emphasis>
</taxonomicName>
shares a superficial resemblance to
<taxonomicName id="4C468AAB006FFFCAF80DD846FD8B1C67" box="[383,543,545,568]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="tiannensis">
<emphasis id="B9322D3A006FFFCAF80DD846FD8B1C67" box="[383,543,545,568]" italics="true" pageId="36" pageNumber="37">R. tiannensis</emphasis>
</taxonomicName>
, another large brown cascade ranid, but differs in having shagreened dorsal skin with small lateral granulations (dorsum of
<taxonomicName id="4C468AAB006FFFCAF8B0D81EFDF11CCF" box="[450,613,633,656]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="tiannensis">
<emphasis id="B9322D3A006FFFCAF8B0D81EFDF11CCF" box="[450,613,633,656]" italics="true" pageId="36" pageNumber="37">R. tiannensis</emphasis>
</taxonomicName>
is rough with large, prominent lateral granulations) and smaller toe disk than finger disk (the opposite condition of
<taxonomicName id="4C468AAB006FFFCAF8A9D8B6FDE81CB7" box="[475,636,721,744]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="tiannensis">
<emphasis id="B9322D3A006FFFCAF8A9D8B6FDE81CB7" box="[475,636,721,744]" italics="true" pageId="36" pageNumber="37">R. tiannensis</emphasis>
</taxonomicName>
).
<taxonomicName id="4C468AAB006FFFCAF90ED888FED21D59" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[124,326,751,774]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A006FFFCAF90ED888FED21D59" box="[124,326,751,774]" italics="true" pageId="36" pageNumber="37">Rana bacboensis</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="4C468AAB006FFFCAFB01D888FE931D7C" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B9322D3A006FFFCAFB01D888FE931D7C" italics="true" pageId="36" pageNumber="37">R. hainanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C468AAB006FFFCAF865D96BFE4E1D7C" box="[279,474,780,803]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A006FFFCAF865D96BFE4E1D7C" box="[279,474,780,803]" italics="true" pageId="36" pageNumber="37">R. jingdongensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C468AAB006FFFCAFB6CD96BFF211D1F" authorityName="Boulenger" authorityYear="1882" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A006FFFCAFB6CD96BFF211D1F" italics="true" pageId="36" pageNumber="37">R. andersonii</emphasis>
</taxonomicName>
.
<taxonomicName id="4C468AAB006FFFCAF9BBD94EFE001D1F" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[201,404,809,832]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A006FFFCAF9BBD94EFE001D1F" box="[201,404,809,832]" italics="true" pageId="36" pageNumber="37">Rana bacboensis</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="4C468AAB006FFFCAF90ED920FEB81D01" box="[124,300,839,862]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B9322D3A006FFFCAF90ED920FEB81D01" box="[124,300,839,862]" italics="true" pageId="36" pageNumber="37">R. hainanensis</emphasis>
</taxonomicName>
in its relative lengths of fingers (II &lt;IV &lt;I &lt;III for
<taxonomicName id="4C468AAB006FFFCAF8A6D903FD171D24" box="[468,643,868,891]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B9322D3A006FFFCAF8A6D903FD171D24" box="[468,643,868,891]" italics="true" pageId="36" pageNumber="37">R. hainanensis</emphasis>
</taxonomicName>
) and by its shagreened skin (smooth for
<taxonomicName id="4C468AAB006FFFCAFB01D9E6FE931DE9" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="B9322D3A006FFFCAFB01D9E6FE931DE9" italics="true" pageId="36" pageNumber="37">R. hainanensis</emphasis>
</taxonomicName>
).
<taxonomicName id="4C468AAB006FFFCAF855D9F8FE601DE9" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[295,500,927,950]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A006FFFCAF855D9F8FE601DE9" box="[295,500,927,950]" italics="true" pageId="36" pageNumber="37">Rana bacboensis</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="4C468AAB006FFFCAF9CDD9DBFE171D8C" box="[191,387,956,979]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A006FFFCAF9CDD9DBFE171D8C" box="[191,387,956,979]" italics="true" pageId="36" pageNumber="37">R. jingdongensis</emphasis>
</taxonomicName>
in profile of its snout shape (rounded or obtusely pointed in
<taxonomicName id="4C468AAB006FFFCAFB01D9BEFE891A51" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A006FFFCAFB01D9BEFE891A51" italics="true" pageId="36" pageNumber="37">R. jingdongensis</emphasis>
</taxonomicName>
versus rounded in
<taxonomicName id="4C468AAB006FFFCAF88DD990FF0F1A74" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A006FFFCAF88DD990FF0F1A74" italics="true" pageId="36" pageNumber="37">R. bacboensis</emphasis>
</taxonomicName>
) and texture of skin (
<taxonomicName id="4C468AAB006FFFCAF8B0DE73FD181A74" box="[450,652,1044,1067]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B9322D3A006FFFCAF8B0DE73FD181A74" box="[450,652,1044,1067]" italics="true" pageId="36" pageNumber="37">R. jingdongensis</emphasis>
</taxonomicName>
dorsum scattered with tubercles and large warts, lips and sides of heads with white spines, all absent in
<taxonomicName id="4C468AAB006FFFCAF8F4DE0BFDBB1ADC" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" box="[390,559,1132,1155]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A006FFFCAF8F4DE0BFDBB1ADC" box="[390,559,1132,1155]" italics="true" pageId="36" pageNumber="37">R. bacboensis</emphasis>
</taxonomicName>
).
<taxonomicName id="4C468AAB006FFFCAFB3DDE0BFF691AFF" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A006FFFCAFB3DDE0BFF691AFF" italics="true" pageId="36" pageNumber="37">Rana bacboensis</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="4C468AAB006FFFCAF8A3DEEEFDFA1AFF" authorityName="Boulenger" authorityYear="1882" box="[465,622,1161,1184]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A006FFFCAF8A3DEEEFDFA1AFF" box="[465,622,1161,1184]" italics="true" pageId="36" pageNumber="37">R. andersonii</emphasis>
</taxonomicName>
in its head shape (obtusely pointed in
<taxonomicName id="4C468AAB006FFFCAFB6CDEC0FF211A84" authorityName="Boulenger" authorityYear="1882" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="B9322D3A006FFFCAFB6CDEC0FF211A84" italics="true" pageId="36" pageNumber="37">R. andersonii</emphasis>
</taxonomicName>
) and absence of ventral spines in the males.
</paragraph>
</subSubSection>
<subSubSection id="C35CA2A3006FFFCBF9EBDE99FF7A1DAF" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
<paragraph id="8BF9F128006FFFCAF9EBDE99FBA71F34" blockId="36.[124,653,193,1741]" lastBlockId="36.[700,1230,193,1741]" pageId="36" pageNumber="37">
DESCRIPTION OF
<typeStatus id="54FD4F8A006FFFCAF811DE99FE4D1B4A" box="[355,473,1278,1301]" pageId="36" pageNumber="37" type="holotype">HOLOTYPE</typeStatus>
: A gravid female (ROM 29534), head length greater than width (127%), head width 34% of SVL, length 43% of SVL; snout short, protruding beyond margin of lower jaw, rounded in dorsal view, bluntly rounded in profile; eye large, prominent, 81% of snout length; eyelid broader than interorbital distance. Top of head flat; canthus rostralis rounded; loreal region concave; lip flared just anterior to orbit; nostril about three­fourths distance from eye to tip of snout; supratympanic fold barely evident, curving posteroventrally from posterior corner of eye to a level above insertion of arm; tympanum round, distinctly visible, separated from eye by distance equal to TMP:EYE 0.62. Choanae ovoid; vomerine dentigerous processes prominent, slightly oblique, posteromedial to choanae, each bearing numerous teeth. Tongue cordiform, distinctly notched posteriorly, free for approximately one­half its length.
</paragraph>
<paragraph id="8BF9F128006FFFCAFBABDB16FB591A52" blockId="36.[700,1230,193,1741]" pageId="36" pageNumber="37">
Forearms moderately robust; fingers moderately short, slender; hands 27% of SVL, relative lengths of fingers II &lt;I &lt;IV &lt;III; ventromedial callous ridge on fingers II and III prominent, extending to proximal tubercle; disks expanded (&lt;2× base of phalanges), relative pad size II &lt;I &lt;IV &lt;III, pad length (III) 75% of pad width; ventral circummarginal grooves present; terminal phalanges T­ shaped; subarticular tubercles conical. Hindlimbs moderately robust; tibia length 60% of SVL; foot length 62% of SVL; relative toe lengths I &lt;II &lt;III &lt;V &lt;IV; inner tarsal fold absent; feet fully webbed to disks, but as a fringe from distal subarticular tubercle of IV, lateral fringes on toes I and V to terminal phalanges; toes long, slender, with large, rounded triangular disks, relative pad size I = II = III&gt; IV
<emphasis id="B9322D3A006FFFCAFD2CD9E5FBED1DC6" bold="true" box="[1118,1145,898,921]" italics="true" pageId="36" pageNumber="37">k</emphasis>
V, pad width (IV) 85% of pad length, circummarginal grooves present; subarticular tubercles prominent, conical; inner metatarsal tubercle ovoid, long; outer metatarsal tubercle absent.
</paragraph>
<paragraph id="8BF9F128006FFFCAFBABDE73FC8D1A17" blockId="36.[700,1230,193,1741]" pageId="36" pageNumber="37">Xiphisternum large, deeply notched posteriorly.</paragraph>
<paragraph id="8BF9F128006FFFCAFBABDE29FC9F1B0F" blockId="36.[700,1230,193,1741]" pageId="36" pageNumber="37">Skin on dorsum shagreened with heavy granulations, leathery in alcohol preservation; dorsolateral folds absent; small tubercles anterior and posterior to tympanum; flanks with small granulations and large pustules; distinct granules on posterior thighs and around cloaca; cloacal opening unmodified, directed posteriorly, at upper level of thighs.</paragraph>
<paragraph id="8BF9F128006FFFCAFBABDF31FB3C18CC" blockId="36.[700,1230,193,1741]" pageId="36" pageNumber="37">COLOR IN LIFE (in preservative): Dorsum, flanks, and loreal region brown (brownish gray) with small black spots, becoming larger on the flanks; upper and lower lips creamy yellow with vertical black bars; dorsal limbs and digits brown with black transverse bands; webbing on feet marbled white and dark brown (black); venter creamy white, sometimes with light spotting on belly, chest, and chin; iris golden, margin of pupil outlined in a striking yellow and red border.</paragraph>
<paragraph id="8BF9F128006FFFCBFBABDCFEFE861F4C" blockId="36.[700,1230,193,1741]" lastBlockId="37.[123,651,193,1742]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
SECONDARY SEXUAL CHARACTERS: The
<typeStatus id="54FD4F8A006FFFCAFDD4DCFEFC911892" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
possesses large, black eggs (
<quantity id="4CBE5CCD006FFFCAFD17DCD1FB381892" box="[1125,1196,1718,1741]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="mm" value="2.0">2 mm</quantity>
in diameter). The lone male
<typeStatus id="54FD4F8A006EFFCBF8C3DAA6FD811E87" box="[433,533,193,216]" pageId="37" pageNumber="38" type="paratype">paratype</typeStatus>
has gular pouches, thickened forearms, and thick white nuptial pads.
</paragraph>
<paragraph id="8BF9F128006EFFCBF9EADB7EFDBE1FFA" blockId="37.[123,651,193,1742]" pageId="37" pageNumber="38">
MEASUREMENTS OF
<typeStatus id="54FD4F8A006EFFCBF8FDDB7EFD911F70" box="[399,517,281,304]" pageId="37" pageNumber="38" type="holotype">HOLOTYPE</typeStatus>
(in mm): SVL 95.1; SNT 12.0; HDL 41.0; HDW 32.2; EYE 9.7; IOD 6.4; TMP 6.0; TEY 5.0; HND 25.6; FGR 21.8; FPL 2.8; FPW 3.7; TIB 56.8; FTL 59.6; TPL 2.4; TPW 2.0.
</paragraph>
<paragraph id="8BF9F128006EFFCBF9EADBCBFD9A1FBF" blockId="37.[123,651,193,1742]" pageId="37" pageNumber="38">
VARIATION OF
<typeStatus id="54FD4F8A006EFFCBF837DBCBFE571F9D" box="[325,451,428,451]" pageId="37" pageNumber="38" type="paratype">PARATYPES</typeStatus>
: Variation in all type material is given in
<tableCitation id="C6C4C493006EFFCBF8DADBAEFD9A1FBF" box="[424,526,457,480]" captionStart="TABLE 15" captionStartId="46.[620,694,194,213]" captionTargetId="figure-50@46.[121,1231,304,1673]" captionTargetPageId="46" captionText="TABLE 15 Variation in Body Proportions for Rana chloronota sensu stricto and Vietnamese Species of the Rana chloronota Complex Given as Means (in mm) ± 1 SD and ranges See Materials and Methods for abbreviations." pageId="37" pageNumber="38">table 15.</tableCitation>
</paragraph>
<paragraph id="8BF9F128006EFFCBF9EADB81FE9E1D1F" blockId="37.[123,651,193,1742]" pageId="37" pageNumber="38">
MEASUREMENTS OF FEMALE
<typeStatus id="54FD4F8A006EFFCBF890DB81FDF41FA2" box="[482,608,486,509]" pageId="37" pageNumber="38" type="paratype">PARATYPES</typeStatus>
(in mm, n = 6, ROM 29359, 2952629530): SVL
<quantity id="4CBE5CCD006EFFCBF9C8D846FEB51C67" box="[186,289,545,568]" metricMagnitude="-2" metricUnit="m" metricValue="9.58" pageId="37" pageNumber="38" unit="mm" value="95.8">95.8 mm</quantity>
± 6.4 (81.8105.1); SNT 13.8 ± 1.4 (10.815.6); HDL 46.4 ± 3.5 (43.5 51.2); HDW 35.4 ± 2.9 (34.139.6); EYE 9.8 ± 0.5 (9.410.8); IOD 7.9 ± 1.7 (5.8 11.0); TMP 5.3 ± 0.6 (4.06.0); TEY 4.9 ± 0.7 (3.66.0); HND 25.5 ± 2.8 (18.630.3); FGR 20.8 ± 2.3 (14.024.1); FPL 3.0 ± 0.3 (2.43.6); FPW 3.5 ± 0.6 (2.64.3); TIB 58.6 ± 3.7 (50.566.2); FTL 70.2 ± 7.8 (55.879.1).
</paragraph>
<paragraph id="8BF9F128006EFFCBF9EAD921FF7A1DAF" blockId="37.[123,651,193,1742]" pageId="37" pageNumber="38">
MEASUREMENTS OF MALE
<typeStatus id="54FD4F8A006EFFCBF897D921FDC31D02" box="[485,599,838,861]" pageId="37" pageNumber="38" type="paratype">PARATYPE</typeStatus>
(in mm, FMNH 255611): SVL 54.9, SNT 8.8, HDL 28.0, HDW 18.1, EYE 6.6, IOD 6.1, TMP 4.4, TEY 1.8, HND 16.5, FPL 2.4, FPW 1.8, TIB 32.4, FTL 45.4, TPL 2.4, TPW 1.6.
</paragraph>
</subSubSection>
<subSubSection id="C35CA2A3006EFFCBF9EAD991FDD51A3A" pageId="37" pageNumber="38" type="etymology">
<paragraph id="8BF9F128006EFFCBF9EAD991FDD51A3A" blockId="37.[123,651,193,1742]" pageId="37" pageNumber="38">
ETYMOLOGY: The specific name, derived from Bac Bo, the Vietnamese name for northern
<collectingCountry id="F351B1B8006EFFCBF99BDE56FED81A17" box="[233,332,1073,1096]" name="Vietnam" pageId="37" pageNumber="38">Vietnam</collectingCountry>
(often referred to as
<collectingCountry id="F351B1B8006EFFCBFB27DE56FF301A3A" name="Vietnam" pageId="37" pageNumber="38">Tonkin</collectingCountry>
), reflects this species distribution.
</paragraph>
</subSubSection>
<subSubSection id="C35CA2A3006EFFCBF9EADE0BFE681864" pageId="37" pageNumber="38" type="distribution">
<paragraph id="8BF9F128006EFFCBF9EADE0BFE681864" blockId="37.[123,651,193,1742]" pageId="37" pageNumber="38">
<materialsCitation id="3B2EFB75006EFFCBF9EADE0BFE6C1864" ID-GBIF-Occurrence="3408947307" accessionNumber="ROM 26358" collectionCode="DISTRIBUTION" country="Vietnam" county="October" location="These" municipality="April - May" pageId="37" pageNumber="38" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectionCode id="ED5769ED006EFFCBF9EADE0BFEA71ADD" box="[152,307,1132,1155]" pageId="37" pageNumber="38">DISTRIBUTION</collectionCode>
AND ECOLOGY:
<taxonomicName id="4C468AAB006EFFCBFB7ADE0BFF461AFF" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A006EFFCBFB7ADE0BFF461AFF" italics="true" pageId="37" pageNumber="38">Rana bacboensis</emphasis>
</taxonomicName>
occurs in forested montane river systems across northern
<collectingCountry id="F351B1B8006EFFCBF8F3DEC1FE7C1AE2" box="[385,488,1190,1213]" name="Vietnam" pageId="37" pageNumber="38">Vietnam</collectingCountry>
.
<location id="8E99A7F3006EFFCBF88ADEC1FDA91AE2" LSID="urn:lsid:plazi:treatment:03EF403E0054FFCBFA7FDF8EFBC01F4C:8E99A7F3006EFFCBF88ADEC1FDA91AE2" box="[504,573,1190,1213]" country="Vietnam" county="October" municipality="April - May" name="These" pageId="37" pageNumber="38">These</location>
rivers vary from shallow and slow moving to torrential and deep.
<location id="8E99A7F3006EFFCBF827DE86FE471AA7" LSID="urn:lsid:plazi:treatment:03EF403E0054FFCBFA7FDF8EFBC01F4C:8E99A7F3006EFFCBF827DE86FE471AA7" box="[341,467,1249,1272]" country="Vietnam" county="October" municipality="April - May" name="Specimens" pageId="37" pageNumber="38">Specimens</location>
may be found on boulders and logs, both in and around the water and in the adjacent forest.
<location id="8E99A7F3006EFFCBF88ADF7BFD1F1B6C" LSID="urn:lsid:plazi:treatment:03EF403E0054FFCBFA7FDF8EFBC01F4C:8E99A7F3006EFFCBF88ADF7BFD1F1B6C" box="[504,651,1308,1331]" country="Vietnam" county="October" municipality="April - May" name="Radiographs" pageId="37" pageNumber="38">Radiographs</location>
revealed that females feed on large invertebrates, including small freshwater crabs.
<location id="8E99A7F3006EFFCBFB16DF31FF2B1BD4" LSID="urn:lsid:plazi:treatment:03EF403E0054FFCBFA7FDF8EFBC01F4C:8E99A7F3006EFFCBFB16DF31FF2B1BD4" country="Vietnam" county="October" municipality="April - May" name="Females" pageId="37" pageNumber="38">Females</location>
were collected in
<collectingMunicipality id="6B9D6B52006EFFCBF8D2DF13FDB61BD4" box="[416,546,1396,1419]" pageId="37" pageNumber="38">AprilMay</collectingMunicipality>
and October. The
<typeStatus id="54FD4F8A006EFFCBF98ADFF6FEF51BF7" box="[248,353,1425,1448]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
, collected in
<collectingCounty id="629889A4006EFFCBF889DFF6FDCD1BF7" box="[507,601,1425,1448]" pageId="37" pageNumber="38">October</collectingCounty>
, has fully developed ova, and
<specimenCount id="9D403AA1006EFFCBF8DADFC9FDA31B9A" box="[424,567,1454,1477]" pageId="37" pageNumber="38" type="female">two females</specimenCount>
(ROM 26358, 29529) collected in the spring have undeveloped ova suggesting a fall breeding season. No calls are associated with this species. The tadpoles are unknown
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35CA2A3006EFFCBF9EADC26FBC01F4C" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8BF9F128006EFFCBF9EADC26FBC01F4C" blockId="37.[123,651,193,1742]" lastBlockId="37.[699,1227,193,275]" pageId="37" pageNumber="38">
REMARKS: Cascade ranids bearing white eggs lay them under rocks, sheltered from the sunlight (
<bibRefCitation id="EFD78CD9006EFFCBF86EDC1BFE3C18CC" author="Pope, C. H." box="[284,424,1660,1683]" pageId="37" pageNumber="38" pagination="397 - 611" refId="ref38937" refString="Pope, C. H. 1931. Notes on amphibians from Fukien, Hainan, and other parts of China. Bulleetin of the American Museum of Natural History 61 (8): 397 - 611, 10 pls." type="book chapter" year="1931">Pope, 1931</bibRefCitation>
; ROM field notes, 1996). In contrast, the black eggs of
<taxonomicName id="4C468AAB006EFFCBFB47DCFEFF461891" authorityName="BAIN &amp; LATHROP &amp; MURPHY &amp; ORLOV &amp; CUC" authorityYear="2003" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="bacboensis">
<emphasis id="B9322D3A006EFFCBFB47DCFEFF461891" italics="true" pageId="37" pageNumber="38">R. bacboensis</emphasis>
</taxonomicName>
might be found where they are exposed to sunlight to promote development, a requirement for some species with melanic eggs (
<bibRefCitation id="EFD78CD9006EFFCBFA70DA9BFBD31F4C" author="Duellman, W. E. &amp; L. Trueb" box="[770,1095,252,275]" pageId="37" pageNumber="38" refId="ref37074" refString="Duellman, W. E. and L. Trueb. 1986. Biology of the Amphibians. Baltimore, Md: Johns Hopkins Press, 114 pp." type="book" year="1986">Duellman and Trueb, 1986</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>