treatments-xml/data/03/EC/DE/03ECDE0DFFDB7742F1049F8B77693A48.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.5411916" ID-GBIF-Dataset="03a6dd3f-a52a-4d60-bc0d-944c00931276" ID-GBIF-Taxon="123195636" ID-Zenodo-Dep="5411916" approvalRequired="73" approvalRequired_for_bibRefs="41" approvalRequired_for_document="1" approvalRequired_for_matCits="6" approvalRequired_for_taxonomicNames="3" approvalRequired_for_textStreams="19" approvalRequired_for_treatments="3" checkinTime="1468511911299" checkinUser="plazi" docAuthor="Lien Thi Phuong Nguyen &amp; James M. Carpenter" docDate="2016" docId="03ECDE0DFFDB7742F1049F8B77693A48" docLanguage="en" docName="entScience.12218.pdf" docOrigin="Entomological Science 2016" docStyle="DocumentStyle{}" docTitle="Zethus angulatus Nguyen &amp; Carpenter, sp. nov" docType="treatment" docVersion="6" lastPageId="4" lastPageNumber="5" masterDocId="FFD5A675FFD87746F23B995F74033D3E" masterDocTitle="Taxonomic review of the genus Zethus Fabricius (Hymenoptera: Vespidae: Eumeninae) from Vietnam with descriptions of four new species" masterLastPageNumber="9" masterPageNumber="1" pageId="3" pageNumber="4" updateTime="1643498771858" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Taxonomic review of the genus Zethus Fabricius (Hymenoptera: Vespidae: Eumeninae) from Vietnam with descriptions of four new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lien Thi Phuong Nguyen</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>James M. Carpenter</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Entomological Science</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>2016</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>9</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1111/ens.12218</mods:identifier>
<mods:identifier type="GBIF-Dataset">03a6dd3f-a52a-4d60-bc0d-944c00931276</mods:identifier>
<mods:identifier type="Zenodo-Dep">269807</mods:identifier>
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<heading bold="true" fontSize="11" level="3" pageId="3" pageNumber="4" reason="0">
<emphasis bold="true" pageId="3" pageNumber="4">
<subSubSection box="[831,1421,1748,1775]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph blockId="3.[831,1437,1748,1910]" box="[831,1421,1748,1775]" pageId="3" pageNumber="4">
<taxonomicName authority="Nguyen &amp; Carpenter" authorityName="Nguyen &amp; Carpenter" box="[831,1319,1748,1775]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="angulatus" status="sp. nov">
<emphasis bold="true" box="[831,1046,1748,1775]" italics="true" pageId="3" pageNumber="4">Zethus angulatus</emphasis>
Nguyen &amp; Carpenter
</taxonomicName>
, 
<taxonomicNameLabel box="[1331,1421,1749,1775]" pageId="3" pageNumber="4" rank="species">sp. nov</taxonomicNameLabel>
</paragraph>
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<subSubSection box="[831,987,1781,1807]" pageId="3" pageNumber="4" type="description">
<paragraph blockId="3.[831,1437,1748,1910]" box="[831,987,1781,1807]" pageId="3" pageNumber="4">
(
<figureCitation box="[838,977,1781,1807]" captionStart="Figures 13 – 18" captionStartId="4.[151,219,942,964]" captionTargetBox="[163,745,232,929]" captionTargetId="figure@4.[161,749,229,932]" captionTargetPageId="4" captionText="Figures 13 – 18 Zethus angulatus sp. nov. Holotype ♂ (13 – 17). 13 Head, frontal view. 14 Right antenna. 15 Propodeum, dorsal view. 16 TI &amp; II, dorsal view. 17 Metasomal segments I &amp; II, lateral view. 18 ♀, Head, frontal view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269810/files/figure.png" pageId="3" pageNumber="4">
Figs 
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</figureCitation>
)
</paragraph>
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</emphasis>
</heading>
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<paragraph blockId="3.[172,778,1835,1910]" pageId="3" pageNumber="4">
<emphasis bold="true" box="[172,500,1835,1857]" pageId="3" pageNumber="4">
Figures 8–12 
<taxonomicName box="[309,500,1835,1856]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="trimaculatus">
<emphasis bold="true" box="[309,500,1835,1856]" italics="true" pageId="3" pageNumber="4">
Zethus 
<taxonomicName box="[383,500,1835,1856]" pageId="3" pageNumber="4" rank="species" species="trimaculatus">trimaculatus</taxonomicName>
</emphasis>
</taxonomicName>
</emphasis>
, ♂. 
<emphasis bold="true" box="[550,563,1835,1857]" pageId="3" pageNumber="4">8</emphasis>
Head, frontal view. 
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Right antenna. 
<emphasis bold="true" box="[328,353,1862,1884]" pageId="3" pageNumber="4">10</emphasis>
Propodeum. 
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TI &amp; II, dorsal view. 
<emphasis bold="true" box="[753,778,1862,1884]" pageId="3" pageNumber="4">12</emphasis>
Metasomal segments I &amp; II, lateral view. Scale bar, 1 mm.
</paragraph>
</caption>
<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="materials_examined">
<paragraph blockId="3.[831,1437,1748,1910]" lastBlockId="4.[151,757,1123,1243]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<materialsCitation ID-GBIF-Occurrence="1275388310" collectingDate="2015-06-10" collectionCode="IEBR" collectorName="LTP Nguyen &amp; DD Nguyen &amp; LX Truong" country="Vietnam" elevation="120" lastPageId="4" lastPageNumber="5" latitude="22.347944" location="VIETNAM" longLatPrecision="14" longitude="105.430275" pageId="3" pageNumber="4" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[831,930,1823,1847]" pageId="3" pageNumber="4">
<emphasis bold="true" box="[831,930,1823,1847]" italics="true" pageId="3" pageNumber="4">Holotype</emphasis>
</typeStatus>
, 
<specimenCount box="[945,969,1823,1846]" pageId="3" pageNumber="4" type="male">♂</specimenCount>
, 
<emphasis bold="true" box="[983,997,1823,1846]" pageId="3" pageNumber="4">“</emphasis>
<collectingCountry box="[997,1121,1822,1846]" name="Vietnam" pageId="3" pageNumber="4">VIETNAM</collectingCountry>
, Bac Vang ranger station, Na Hang natural reserve, Na Hang, Tuyen Quang, 
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° 
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<emphasis bold="true" box="[858,865,1887,1910]" pageId="3" pageNumber="4">’</emphasis>
<number box="[865,917,1886,1910]" pageId="3" pageNumber="4" value="52.6">52.6</number>
&quot;N
</geoCoordinate>
<geoCoordinate box="[963,1107,1886,1910]" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="15" value="105.430275">
<number box="[963,1004,1886,1910]" pageId="3" pageNumber="4" value="105.0">105</number>
° 
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<emphasis bold="true" box="[1042,1049,1887,1910]" pageId="3" pageNumber="4">’</emphasis>
<number box="[1048,1079,1886,1910]" pageId="3" pageNumber="4" value="49.0">49</number>
&quot;E
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, ca 
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<quantity box="[1167,1235,1886,1910]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="4" unit="m" value="120.0">120 m</quantity>
</elevation>
, 
<collectingDate box="[1256,1373,1886,1910]" pageId="3" pageNumber="4" value="2015-06-10">
<date box="[1256,1373,1886,1910]" pageId="3" pageNumber="4" value="2015-06-10">
<number box="[1256,1283,1886,1910]" pageId="3" pageNumber="4" value="10.0">10</number>
.vi. 
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</date>
</collectingDate>
, 
<collectorName lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">LTP Nguyen</collectorName>
, 
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&amp; 
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<emphasis bold="true" box="[546,560,1124,1147]" pageId="4" pageNumber="5">”</emphasis>
[
<collectionCode box="[574,637,1124,1147]" pageId="4" pageNumber="5">IEBR</collectionCode>
].
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="1275388324" collectingDate="2004-07-18" collectionCode="IEBR" collectorName="Insect Systematic Department ’ s collectors" country="Vietnam" elevation="375" location="Huc Nghi, Dakrong, Quang Tri" pageId="4" pageNumber="5" specimenCount="1" typeStatus="paratype">
<typeStatus box="[656,750,1124,1148]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[656,750,1124,1148]" italics="true" pageId="4" pageNumber="5">Paratype</emphasis>
</typeStatus>
: 
<collectingCountry box="[151,276,1155,1179]" name="Vietnam" pageId="4" pageNumber="5">VIETNAM</collectingCountry>
: 
<quantity box="[299,332,1155,1179]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="4" pageNumber="5" unit="g" value="1.0">
<number box="[299,313,1155,1179]" pageId="4" pageNumber="5" value="1.0">
<specimenCount box="[299,313,1155,1179]" pageId="4" pageNumber="5" type="generic">1</specimenCount>
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♀
</quantity>
, 
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, 
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<quantity box="[151,271,1187,1211]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="4" pageNumber="5" unit="m" value="375.0" valueMax="400.0" valueMin="350.0">
350 
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400 m
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</elevation>
a.s.l., 
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<date box="[372,496,1187,1211]" pageId="4" pageNumber="5" value="2004-07-18">
<number box="[372,398,1187,1211]" pageId="4" pageNumber="5" value="18.0">18</number>
.vii. 
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</date>
</collectingDate>
, 
<collectorName pageId="4" pageNumber="5">
Insect Systematic Department 
<emphasis bold="true" box="[278,285,1220,1243]" pageId="4" pageNumber="5">’</emphasis>
s collectors
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[
<collectionCode box="[416,472,1220,1243]" collectionName="IEBR" pageId="4" pageNumber="5">IEBR</collectionCode>
].
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</paragraph>
</subSubSection>
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<emphasis bold="true" box="[151,544,942,964]" pageId="4" pageNumber="5">
Figures 13–18 
<taxonomicName box="[299,464,942,963]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="angulatus" status="sp. nov.">
<emphasis bold="true" box="[299,464,942,963]" italics="true" pageId="4" pageNumber="5">Zethus angulatus</emphasis>
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<taxonomicNameLabel box="[470,544,942,964]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype ♂ 
<emphasis bold="true" pageId="4" pageNumber="5">(13–17). 13</emphasis>
Head, frontal view. 
<emphasis bold="true" box="[370,395,969,991]" pageId="4" pageNumber="5">14</emphasis>
Right antenna. 
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Propodeum, dorsal view. 
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TI &amp; II, dorsal view. 
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Metasomal segments I &amp; II, lateral view. 
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♀, Head, frontal view. Scale bar, 1 mm.
</paragraph>
</caption>
<subSubSection pageId="4" pageNumber="5" type="description">
<paragraph blockId="4.[151,758,1301,1910]" box="[151,288,1301,1327]" pageId="4" pageNumber="5">
<heading bold="true" box="[151,288,1301,1327]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="0">
<emphasis bold="true" box="[151,288,1301,1327]" italics="true" pageId="4" pageNumber="5">Description</emphasis>
</heading>
</paragraph>
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<emphasis bold="true" box="[151,204,1343,1367]" italics="true" pageId="4" pageNumber="5">Male</emphasis>
. 
<typeStatus box="[225,328,1342,1366]" pageId="4" pageNumber="5">Holotype</typeStatus>
. Body length 
<quantity box="[491,567,1342,1366]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="5" unit="mm" value="15.0">15 mm</quantity>
; forewing length 
<quantity box="[151,228,1374,1398]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="5" unit="mm" value="12.0">12 mm</quantity>
. Structure as in 
<taxonomicName box="[421,589,1375,1399]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dolosus">
<emphasis bold="true" box="[421,589,1375,1399]" italics="true" pageId="4" pageNumber="5">Zethus dolosus</emphasis>
</taxonomicName>
, but differs as follows: body covered with shorter hairs. Head in frontal view wider than high, 
<number box="[400,433,1438,1462]" pageId="4" pageNumber="5" value="1.5">1.5</number>
× as wide as high (
<figureCitation box="[638,716,1438,1462]" captionStart="Figures 13 – 18" captionStartId="4.[151,219,942,964]" captionTargetBox="[163,745,232,929]" captionTargetId="figure@4.[161,749,229,932]" captionTargetPageId="4" captionText="Figures 13 – 18 Zethus angulatus sp. nov. Holotype ♂ (13 – 17). 13 Head, frontal view. 14 Right antenna. 15 Propodeum, dorsal view. 16 TI &amp; II, dorsal view. 17 Metasomal segments I &amp; II, lateral view. 18 ♀, Head, frontal view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269810/files/figure.png" pageId="4" pageNumber="5">
Fig. 
<number box="[687,716,1438,1462]" pageId="4" pageNumber="5" value="13.0">13</number>
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). A short longitudinal carina present at one-third of the length of the gena from base. Inner eye margins in frontal view 
<number box="[151,184,1534,1558]" pageId="4" pageNumber="5" value="1.1">1.1</number>
× further apart from each other at vertex than at clypeus. Clypeus in frontal view 
<number box="[522,555,1566,1590]" pageId="4" pageNumber="5" value="1.5">1.5</number>
× as wide as long, emarginated at basal margin and slightly concave medially, apical margin deeply emarginated medially with the emargination forming a semicircle (
<figureCitation box="[584,662,1662,1686]" captionStart="Figures 13 – 18" captionStartId="4.[151,219,942,964]" captionTargetBox="[163,745,232,929]" captionTargetId="figure@4.[161,749,229,932]" captionTargetPageId="4" captionText="Figures 13 – 18 Zethus angulatus sp. nov. Holotype ♂ (13 – 17). 13 Head, frontal view. 14 Right antenna. 15 Propodeum, dorsal view. 16 TI &amp; II, dorsal view. 17 Metasomal segments I &amp; II, lateral view. 18 ♀, Head, frontal view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269810/files/figure.png" pageId="4" pageNumber="5">
Fig. 
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), with a blunt teeth laterally, width of the emargination about 
<number box="[723,732,1694,1718]" pageId="4" pageNumber="5" value="1.0">1</number>
/ 
<number box="[742,757,1694,1718]" pageId="4" pageNumber="5" value="4.0">4</number>
width of clypeus between inner eye margins. Mandible with four teeth, the third tooth short and wide, with inner side round. Antennal scape 
<number box="[450,484,1790,1814]" pageId="4" pageNumber="5" value="2.6">2.6</number>
× as long as its maximum width; FI 
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× as long as its maximum width, FII 
<emphasis bold="true" box="[719,731,1823,1846]" pageId="4" pageNumber="5">–</emphasis>
IV longer than wide, FV 
<emphasis bold="true" box="[381,393,1855,1878]" pageId="4" pageNumber="5">–</emphasis>
VIII wider than long, FIX smaller than 
<collectionCode box="[212,271,1887,1910]" pageId="4" pageNumber="5">FVIII</collectionCode>
, as long as wide, FX small, FXI elongate, slightly curved, 
<number box="[988,1000,229,253]" pageId="4" pageNumber="5" value="3.0">3</number>
× as long as it basal width (
<figureCitation box="[1322,1404,229,253]" captionStart="Figures 13 – 18" captionStartId="4.[151,219,942,964]" captionTargetBox="[163,745,232,929]" captionTargetId="figure@4.[161,749,229,932]" captionTargetPageId="4" captionText="Figures 13 – 18 Zethus angulatus sp. nov. Holotype ♂ (13 – 17). 13 Head, frontal view. 14 Right antenna. 15 Propodeum, dorsal view. 16 TI &amp; II, dorsal view. 17 Metasomal segments I &amp; II, lateral view. 18 ♀, Head, frontal view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269810/files/figure.png" pageId="4" pageNumber="5">
Fig. 
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</figureCitation>
). Mesoscutum and scutellum in lateral view more convex than in 
<taxonomicName box="[904,1013,293,317]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dolosus">
<emphasis bold="true" box="[904,1013,293,317]" italics="true" pageId="4" pageNumber="5">Z. dolosus</emphasis>
</taxonomicName>
. Propodeum with some strong and slightly oblique carinae along median carina (
<figureCitation box="[1322,1403,325,349]" captionStart="Figures 13 – 18" captionStartId="4.[151,219,942,964]" captionTargetBox="[163,745,232,929]" captionTargetId="figure@4.[161,749,229,932]" captionTargetPageId="4" captionText="Figures 13 – 18 Zethus angulatus sp. nov. Holotype ♂ (13 – 17). 13 Head, frontal view. 14 Right antenna. 15 Propodeum, dorsal view. 16 TI &amp; II, dorsal view. 17 Metasomal segments I &amp; II, lateral view. 18 ♀, Head, frontal view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269810/files/figure.png" pageId="4" pageNumber="5">
Fig. 
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</figureCitation>
). Metasomal segment I proportionally shorter than in 
<taxonomicName class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dolosus">
<emphasis bold="true" italics="true" pageId="4" pageNumber="5">Z. dolosus</emphasis>
</taxonomicName>
, 
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× as long as wide, in dorsal view gradually widening from one-fifth of the distance from base, then slightly narrowly toward apical margin (
<figureCitation box="[1263,1344,453,477]" captionStart="Figures 13 – 18" captionStartId="4.[151,219,942,964]" captionTargetBox="[163,745,232,929]" captionTargetId="figure@4.[161,749,229,932]" captionTargetPageId="4" captionText="Figures 13 – 18 Zethus angulatus sp. nov. Holotype ♂ (13 – 17). 13 Head, frontal view. 14 Right antenna. 15 Propodeum, dorsal view. 16 TI &amp; II, dorsal view. 17 Metasomal segments I &amp; II, lateral view. 18 ♀, Head, frontal view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269810/files/figure.png" pageId="4" pageNumber="5">
Fig. 
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), with maximum width 
<number box="[990,1024,485,509]" pageId="4" pageNumber="5" value="2.6">2.6</number>
× its basal width, with a medial carina which runs from basal margin to one-third of the tergum, swollen part 
<number box="[958,971,549,573]" pageId="4" pageNumber="5" value="2.0">2</number>
× as long as wide in dorsal view. TII in lateral view strongly concave near base, then gradually swollen to midpoint and nearly straight to apical margin. SII in lateral view slightly concave near base, angular at basal half with obtuse angle, then nearly straight to apical margin (
<figureCitation box="[900,975,709,733]" captionStart="Figures 13 – 18" captionStartId="4.[151,219,942,964]" captionTargetBox="[163,745,232,929]" captionTargetId="figure@4.[161,749,229,932]" captionTargetPageId="4" captionText="Figures 13 – 18 Zethus angulatus sp. nov. Holotype ♂ (13 – 17). 13 Head, frontal view. 14 Right antenna. 15 Propodeum, dorsal view. 16 TI &amp; II, dorsal view. 17 Metasomal segments I &amp; II, lateral view. 18 ♀, Head, frontal view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269810/files/figure.png" pageId="4" pageNumber="5">
Fig. 
<number box="[946,975,709,733]" pageId="4" pageNumber="5" value="17.0">17</number>
</figureCitation>
), with apical lamella short.
</paragraph>
<paragraph blockId="4.[810,1417,229,1910]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[836,901,740,764]" italics="true" pageId="4" pageNumber="5">Color.</emphasis>
Black. A longitudinal stripe on mandible, two small spot between antenna and two marks at lateral margins of TI yellow. Propodeal valvulae dark brown.
</paragraph>
<paragraph blockId="4.[810,1417,229,1910]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[836,918,834,858]" italics="true" pageId="4" pageNumber="5">Female.</emphasis>
Body length 
<quantity box="[1058,1153,834,858]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="4" pageNumber="5" unit="mm" value="16.5">16.5 mm</quantity>
; forewing length 
<quantity box="[1336,1413,834,858]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="5" unit="mm" value="13.0">13 mm</quantity>
. Structure as in male, but differing as follows: head in frontal view 
<number box="[949,982,898,922]" pageId="4" pageNumber="5" value="1.3">1.3</number>
× as wide as high (
<figureCitation box="[1183,1261,898,922]" captionStart="Figures 13 – 18" captionStartId="4.[151,219,942,964]" captionTargetBox="[163,745,232,929]" captionTargetId="figure@4.[161,749,229,932]" captionTargetPageId="4" captionText="Figures 13 – 18 Zethus angulatus sp. nov. Holotype ♂ (13 – 17). 13 Head, frontal view. 14 Right antenna. 15 Propodeum, dorsal view. 16 TI &amp; II, dorsal view. 17 Metasomal segments I &amp; II, lateral view. 18 ♀, Head, frontal view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269810/files/figure.png" pageId="4" pageNumber="5">
Fig. 
<number box="[1232,1261,898,922]" pageId="4" pageNumber="5" value="18.0">18</number>
</figureCitation>
). Ocelli small, frontal ocellus with diameter nearly half diameter of antennal socket, area around ocelli strongly swollen, projecting over posterior ocelli. Distance from posterior ocelli to apical margin of the vertex 
<number box="[1183,1217,1026,1050]" pageId="4" pageNumber="5" value="2.7">2.7</number>
× the distance from posterior ocelli to inner eye margin. Inner eye margins in frontal view 
<number box="[947,980,1090,1114]" pageId="4" pageNumber="5" value="1.1">1.1</number>
× further apart from each other at vertex than at clypeus. Clypeus in frontal view 
<number box="[1252,1285,1122,1146]" pageId="4" pageNumber="5" value="1.5">1.5</number>
× as wide as high, apical margin thick, emarginated medially, forming short and sharp tooth each lateral side, apical part between two teeth slightly produced, forming a small tooth, width of emargination 
<number box="[1168,1177,1250,1274]" pageId="4" pageNumber="5" value="1.0">1</number>
/ 
<number box="[1187,1200,1250,1274]" pageId="4" pageNumber="5" value="3.0">3</number>
× width of clypeus between inner eye margins (
<figureCitation box="[1108,1185,1282,1306]" captionStart="Figures 13 – 18" captionStartId="4.[151,219,942,964]" captionTargetBox="[163,745,232,929]" captionTargetId="figure@4.[161,749,229,932]" captionTargetPageId="4" captionText="Figures 13 – 18 Zethus angulatus sp. nov. Holotype ♂ (13 – 17). 13 Head, frontal view. 14 Right antenna. 15 Propodeum, dorsal view. 16 TI &amp; II, dorsal view. 17 Metasomal segments I &amp; II, lateral view. 18 ♀, Head, frontal view. Scale bar, 1 mm." httpUri="https://zenodo.org/record/269810/files/figure.png" pageId="4" pageNumber="5">
Fig. 
<number box="[1156,1185,1282,1306]" pageId="4" pageNumber="5" value="18.0">18</number>
</figureCitation>
). Mandible with four distinct teeth on inner side, basal tooth short and pointed apically. Antennal scape 
<number box="[1067,1079,1346,1370]" pageId="4" pageNumber="5" value="3.0">3</number>
× as long as its maximum width; FI 
<number box="[839,873,1378,1402]" pageId="4" pageNumber="5" value="1.4">1.4</number>
× as long as its maximum width, FII 
<emphasis bold="true" box="[1254,1266,1379,1402]" pageId="4" pageNumber="5">–</emphasis>
IX wider than long, FX bullet-shaped, as long as its basal width.
</paragraph>
<paragraph blockId="4.[810,1417,229,1910]" box="[836,1400,1441,1465]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[836,901,1441,1465]" italics="true" pageId="4" pageNumber="5">Color.</emphasis>
As in male, except for mandible black entirely.
</paragraph>
</subSubSection>
<subSubSection box="[836,1360,1473,1497]" pageId="4" pageNumber="5" type="distribution">
<paragraph blockId="4.[810,1417,229,1910]" box="[836,1360,1473,1497]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[836,966,1473,1497]" italics="true" pageId="4" pageNumber="5">Distribution</emphasis>
. 
<collectingCountry box="[979,1072,1473,1497]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
: Tuyen Quang, Quang Tri.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="discussion">
<paragraph blockId="4.[810,1417,229,1910]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[836,937,1504,1528]" italics="true" pageId="4" pageNumber="5">Remarks.</emphasis>
This species can be distinguished from its congeners by the combination of the following characters: apical margin of clypeus strongly emarginated medially with the emargination semicirclular and width of the emargination 
<number box="[964,973,1631,1655]" pageId="4" pageNumber="5" value="1.0">1</number>
/ 
<number box="[983,998,1631,1655]" pageId="4" pageNumber="5" value="4.0">4</number>
width of clypeus between inner eye margin, TI 
<number box="[931,943,1663,1687]" pageId="4" pageNumber="5" value="3.0">3</number>
× as long as wide, swollen part 
<number box="[1279,1292,1663,1687]" pageId="4" pageNumber="5" value="2.0">2</number>
× as long as wide in dorsal view, TII in lateral view strongly concave near base, then gradually swollen to midpoint and nearly straight to apical margin, SII in lateral view slightly concave near base and angular at basal half with obtuse angle.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="etymology">
<paragraph blockId="4.[810,1417,229,1910]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[836,958,1854,1878]" italics="true" pageId="4" pageNumber="5">Etymology.</emphasis>
The specific name refers to the obtuse angle in SII.
</paragraph>
</subSubSection>
</treatment>
</document>