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<mods:title id="3338622D8D1F985073DE5F94BF1AAC81">Taxonomic revision of the Menticirrhus americanus (Linnaeus, 1758) and M littoralis (Holbrook, 1847) (Percomorphacea: Sciaenidae) species complexes from the western Atlantic</mods:title>
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<mods:namePart id="70756B7461AF7AC080FC71F9696DFF15">Cerqueira, Najila Nolie Catarine Dantas</mods:namePart>
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<mods:namePart id="28800DFC42E98B2DA6B1759D25F5EE40">Siccha-Ramirez, Raquel</mods:namePart>
<mods:affiliation id="F240A83BB8509E668D37B8D4239E22CE">Laboratorio Costero de Tumbes, Instituto del Mar, Tumbes, Peru.</mods:affiliation>
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<treatment id="03E687FFFFF87E21BA831F57955BABA3" ID-DOI="http://doi.org/10.5281/zenodo.4401556" ID-GBIF-Taxon="177013298" ID-Zenodo-Dep="4401556" LSID="urn:lsid:plazi:treatment:03E687FFFFF87E21BA831F57955BABA3" httpUri="http://treatment.plazi.org/id/03E687FFFFF87E21BA831F57955BABA3" lastPageId="26" lastPageNumber="326" pageId="24" pageNumber="325">
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<taxonomicName id="4C4F4D6AFFF87E23BA831F579628A9DE" ID-CoL="72ZKJ" authority="(Holbrook, 1847)" baseAuthorityName="Holbrook" baseAuthorityYear="1847" box="[151,639,618,645]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="littoralis">
<emphasis id="B93BEAFBFFF87E23BA831F579628A9DE" bold="true" box="[151,639,618,645]" pageId="24" pageNumber="325">
<emphasis id="B93BEAFBFFF87E23BA831F5795F4A9DF" bold="true" box="[151,419,618,645]" italics="true" pageId="24" pageNumber="325">Menticirrhus littoralis</emphasis>
(
<bibRefCitation id="EFDE4B18FFF87E23BBA71F579620A9DE" author="Holbrook, J. E." box="[435,631,618,644]" pageId="24" pageNumber="325" refId="ref26882" refString="Holbrook, J. E. (1847) Southern ichthyology: or, a description of the fishes inhabiting the waters of South Carolina Georgia and Florida. No. II. Wiley, New York and Putnam, London, 32 pp., 4 pls." type="book" year="1847">Holbrook, 1847</bibRefCitation>
)
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<figureCitation id="13742A6CFFF87E23BA831FB294A0A9F3" box="[151,247,655,681]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="24" pageNumber="325">Figure 2</figureCitation>
,
<tableCitation id="C6CD0352FFF87E23BB171FB29536A9F3" box="[259,353,655,681]" captionStart="TABLE 3" captionStartId="11.[151,239,404,428]" captionText="TABLE 3. Meristic data of Menticirrhus americanus, M. martinicensis, M. cuiaranensis, M. gracilis and M. littoralis. A. Dorsal-fin rays. B. Pectoral-fin rays. C. Gill rakers on the first gill arch. D. Scales above the lateral line." pageId="24" pageNumber="325">Tables 3</tableCitation>
,
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<emphasis id="B93BEAFBFFF87E23BA831FEE9503A9B1" box="[151,340,723,747]" italics="true" pageId="24" pageNumber="325">Umbrina littoralis</emphasis>
<bibRefCitation id="EFDE4B18FFF87E23BB4B1FEE967CA9B1" author="Holbrook, J. E." box="[351,555,723,747]" pageId="24" pageNumber="325" refId="ref26882" refString="Holbrook, J. E. (1847) Southern ichthyology: or, a description of the fishes inhabiting the waters of South Carolina Georgia and Florida. No. II. Wiley, New York and Putnam, London, 32 pp., 4 pls." type="book" year="1847">Holbrook 1847: 10</bibRefCitation>
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, pl. 1 (original description; type locality:
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,
<collectingCountry id="F3587679FFF87E23BE891FEE90B4A9B1" box="[1181,1251,723,747]" name="United States of America" pageId="24" pageNumber="325">U.S.A.</collectingCountry>
<typeStatus id="54F4884BFFF87E23BEFA1FEE9107A9B1" box="[1262,1360,723,747]" pageId="24" pageNumber="325" type="holotype">Holotype</typeStatus>
: ANSP 11576).—
<treatmentCitation id="0AEE10F8FFF87E23BB3E1FCF95ABA850" author="Holbrook, J. E." box="[298,508,754,778]" page="142" pageId="24" pageNumber="325" year="1855">
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(fishes of
<collectingRegion id="498BF80BFFF87E23B87C1FCF9753A850" box="[616,772,754,778]" country="United States of America" name="South Carolina" pageId="24" pageNumber="325">South Carolina</collectingRegion>
; short description).—
<treatmentCitation id="0AEE10F8FFF87E23B9CA1FCF90F7A850" author="Gunther, A." box="[990,1184,754,778]" page="276" pageId="24" pageNumber="325" year="1860">
<bibRefCitation id="EFDE4B18FFF87E23B9CA1FCF90F7A850" author="Gunther, A." box="[990,1184,754,778]" pageId="24" pageNumber="325" refId="ref26590" refString="Gunther, A. (1860) Catalogue of the fishes in the British Museum. Catalogue of the acanthopterygian fishes in the collection of the British Museum. Squamipinnes, Cirrhitidae, Triglidae, Trachinidae, Sciaenidae, Polynemidae, Sphyraenidae, Trichiuridae, Scombridae, Carangidae, Xiphiidae. Vol. 2. British Museum, London, xxi + 548 pp." type="book" year="1860">Günther 1860: 276</bibRefCitation>
</treatmentCitation>
(Catalog of fishes of the British Museum; short description).—
<bibRefCitation id="EFDE4B18FFF87E23B85D1E2D96AEA872" author="Bohlke, E. B." box="[585,761,784,808]" pageId="24" pageNumber="325" pagination="1 - 216" refId="ref24614" refString="Bohlke, E. B. (1984) Catalog of type specimens in the ichthyological collection of the Academy of Natural Sciences of Philadelphia. Special Publication, Academy of Natural Sciences of Philadelphia, 14, 1 - 216." type="journal article" year="1984">Böhlke 1984:152</bibRefCitation>
(type catalog).
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</paragraph>
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<taxonomicName id="4C4F4D6AFFF87E23BA831E12952BA81D" ID-CoL="72ZKJ" baseAuthorityName="Holbrook" baseAuthorityYear="1847" box="[151,380,815,839]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="littoralis">
<emphasis id="B93BEAFBFFF87E23BA831E12952BA81D" box="[151,380,815,839]" italics="true" pageId="24" pageNumber="325">Menticirrhus littoralis</emphasis>
</taxonomicName>
<treatmentCitation id="0AEE10F8FFF87E23BB911E1296FCA81D" author="Jordan, D. S. &amp; Gilbert, C. H." box="[389,683,815,839]" page="378" pageId="24" pageNumber="325" year="1879">
<bibRefCitation id="EFDE4B18FFF87E23BB911E1296FCA81D" author="Jordan, D. S. &amp; Gilbert, C. H." box="[389,683,815,839]" pageId="24" pageNumber="325" pagination="365 - 388" refId="ref27418" refString="Jordan, D. S. &amp; Gilbert, C. H. (1879) Notes on the fishes of Beaufort Harbor, North Carolina. Proceedings of the United States National Museum, 1 (55), 365 - 388. https: // doi. org / 10.5479 / si. 00963801.1 - 55.365" type="journal article" year="1879">
<collectingCountry id="F3587679FFF87E23BB911E12959EA81D" box="[389,457,815,839]" name="Jordan" pageId="24" pageNumber="325">Jordan</collectingCountry>
&amp; Gilbert 1879: 378
</bibRefCitation>
</treatmentCitation>
(Beaufort; listed).—
<treatmentCitation id="0AEE10F8FFF87E23B9901E129048A81D" author="Bean, T. H." box="[900,1055,815,839]" page="93" pageId="24" pageNumber="325" year="1880">
<bibRefCitation id="EFDE4B18FFF87E23B9901E129048A81D" author="Bean, T. H." box="[900,1055,815,839]" pageId="24" pageNumber="325" pagination="75 - 116" refId="ref24249" refString="Bean, T. H. (1880) Check-list of duplicates of North American fishes distributed by the Smithsonian Institution in behalf of the United States National Museum, 1877 - 1880. Proceedings of the United States National Museum, 3 (127), 75 - 116. https: // doi. org / 10.5479 / si. 00963801.3 - 127.75" type="journal article" year="1880">Bean 1880: 93</bibRefCitation>
</treatmentCitation>
(Florida; listed).—
<treatmentCitation id="0AEE10F8FFF87E23BEFC1E12957AA83C" author="Jordan, D. S. &amp; Gilbert, C. H." page="283" pageId="24" pageNumber="325" year="1882">
<bibRefCitation id="EFDE4B18FFF87E23BEFC1E12957AA83C" author="Jordan, D. S. &amp; Gilbert, C. H." pageId="24" pageNumber="325" pagination="241 - 307" refId="ref27476" refString="Jordan, D. S. &amp; Gilbert, C. H. (1882) Notes on fishes observed about Pensacola, Florida, and Galveston, Texas, with description of new species. Proceedings of the United States National Museum, 5 (282), 241 - 307. https: // doi. org / 10.5479 / si. 00963801.5 - 282.241" type="journal article" year="1882">
<collectingCountry id="F3587679FFF87E23BEFC1E129179A81D" box="[1256,1326,815,839]" name="Jordan" pageId="24" pageNumber="325">Jordan</collectingCountry>
&amp; Gilbert 1882: 283
</bibRefCitation>
</treatmentCitation>
(Pensacola, Galveston; listed).—
<treatmentCitation id="0AEE10F8FFF87E23B86B1E7397CDA83C" author="Jordan, D. S. &amp; Gilbert, C. H." box="[639,922,846,870]" page="606" pageId="24" pageNumber="325" year="1882">
<bibRefCitation id="EFDE4B18FFF87E23B86B1E7397CDA83C" author="Jordan, D. S. &amp; Gilbert, C. H." box="[639,922,846,870]" pageId="24" pageNumber="325" pagination="241 - 307" refId="ref27476" refString="Jordan, D. S. &amp; Gilbert, C. H. (1882) Notes on fishes observed about Pensacola, Florida, and Galveston, Texas, with description of new species. Proceedings of the United States National Museum, 5 (282), 241 - 307. https: // doi. org / 10.5479 / si. 00963801.5 - 282.241" type="journal article" year="1882">
<collectingCountry id="F3587679FFF87E23B86B1E739692A83C" box="[639,709,846,870]" name="Jordan" pageId="24" pageNumber="325">Jordan</collectingCountry>
&amp; Gilbert 1882: 606
</bibRefCitation>
</treatmentCitation>
(Charleston; listed).—
<treatmentCitation id="0AEE10F8FFF87E23BE951E7391CBA83C" author="Jordan, D. S. &amp; Gilbert, C. H." box="[1153,1436,846,870]" page="933" pageId="24" pageNumber="325" year="1883">
<bibRefCitation id="EFDE4B18FFF87E23BE951E7391CBA83C" author="Jordan, D. S. &amp; Gilbert, C. H." box="[1153,1436,846,870]" pageId="24" pageNumber="325" pagination="1 - 1018" refId="ref27543" refString="Jordan, D. S. &amp; Gilbert, C. H. (1883) Synopsis of the fishes of North America. Bulletin of the United States National Museum, 16, 1 - 1018. https: // doi. org / 10.5479 / si. 03629236.16. i" type="journal article" year="1883">
<collectingCountry id="F3587679FFF87E23BE951E739090A83C" box="[1153,1223,846,870]" name="Jordan" pageId="24" pageNumber="325">Jordan</collectingCountry>
&amp; Gilbert 1883: 933
</bibRefCitation>
</treatmentCitation>
(synopsis North American fishes; listed).—
<collectingCountry id="F3587679FFF87E23B8941E519691A8DE" box="[640,710,876,900]" name="Jordan" pageId="24" pageNumber="325">Jordan</collectingCountry>
1885: (name only).—
<collectingCountry id="F3587679FFF87E23B9B31E5197B9A8DE" box="[935,1006,876,900]" name="Jordan" pageId="24" pageNumber="325">Jordan</collectingCountry>
&amp; Evermann 1896: 1474 (fishes of North America; description; synonymy).—
<treatmentCitation id="0AEE10F8FFF87E23B82E1EB69686A8F9" author="Chao, L. N." box="[570,721,907,931]" page="29" pageId="24" pageNumber="325" year="1977">
<bibRefCitation id="EFDE4B18FFF87E23B82E1EB69686A8F9" author="Chao, L. N." box="[570,721,907,931]" pageId="24" pageNumber="325" pagination="1 - 48" refId="ref25569" refString="Chao, L. N. (1977) Sciaenidae. In: Fischer, W. (Ed.), Identification sheets of Central West Atlantic, fishing area 30 and 31. FAO, Rome, sheets 1 - 48." type="book chapter" year="1977">Chao 1977: 29</bibRefCitation>
</treatmentCitation>
(in part; FAO fact sheets; description).—
<treatmentCitation id="0AEE10F8FFF87E23BE621EB6915CA8F9" author="Chao, L. N." box="[1142,1291,907,931]" page="31" pageId="24" pageNumber="325" year="1978">
<bibRefCitation id="EFDE4B18FFF87E23BE621EB6915CA8F9" author="Chao, L. N." box="[1142,1291,907,931]" pageId="24" pageNumber="325" pagination="1 - 64" refId="ref25613" refString="Chao, L. N. (1978) A basis for classifying western Atlantic Sciaenidae (Teleostei: Perciformes). NOAA (National Oceanic and Atmospheric Administration) Technical Report NMFS (National Marine Fisheries Service) Circular, 415, 1 - 64." type="journal article" year="1978">Chao 1978: 31</bibRefCitation>
</treatmentCitation>
(in part; basis for classifying western Atlantic sciaenids; description).—
<bibRefCitation id="EFDE4B18FFF87E23B91B1E9797EFA898" author="Chao, L. N." box="[783,952,938,962]" pageId="24" pageNumber="325" pagination="1583 - 1653" refId="ref25660" refString="Chao, L. N. (2003) Sciaenidae. In: Carpenter, K. E. (Ed.), The living marine resources of the Western Central Atlantic. Vol. 3. Bony fishes part 2. Opistognathidae to Molidae. FAO species identification guide for fishery purposes and American Society of Ichthyologist and Herpetologists Special Publication No. 5. FAO, Rome, pp. 1583 - 1653." type="book chapter" year="2003">Chao 2003:1627</bibRefCitation>
(in part; FAO species identification guide; central Western Atlantic; description).—
<treatmentCitation id="0AEE10F8FFF87E23B8291EF5976CA8BB" author="Robins, C. R. &amp; Ray, G. C." box="[573,827,968,993]" page="187" pageId="24" pageNumber="325" year="1986">
<bibRefCitation id="EFDE4B18FFF87E23B8291EF5976CA8BB" author="Robins, C. R. &amp; Ray, G. C." box="[573,827,968,993]" pageId="24" pageNumber="325" refId="ref29080" refString="Robins, C. R. &amp; Ray, G. C. (1986) A field guide to Atlantic coast fishes of North America. The Peterson Field Guide Series, Boston, Massachusetts, 354 pp." type="book" year="1986">Robins &amp; Ray 1986: 187</bibRefCitation>
</treatmentCitation>
(Atlantic coast fishes; list, short description, illustration).—
<treatmentCitation id="0AEE10F8FFF87E23BAD31EDA95C1A8A5" author="Boschung, H. T." box="[199,406,999,1023]" page="146" pageId="24" pageNumber="325" year="1992">
<bibRefCitation id="EFDE4B18FFF87E23BAD31EDA95C1A8A5" author="Boschung, H. T." box="[199,406,999,1023]" pageId="24" pageNumber="325" pagination="1 - 266" refId="ref24747" refString="Boschung, H. T. (1992) Catalogue of freshwater and marine fishes of Alabama. Alabama Museum of Natural History Bulletin, 14, 1 - 266." type="journal article" year="1992">Boschung 1992: 146</bibRefCitation>
</treatmentCitation>
(catalog of fishes of Alabama; listed).—
<bibRefCitation id="EFDE4B18FFF87E23B93E1EDA905BA8A5" author="Murdy, E. O. &amp; Birdsong, R. S. &amp; Musick, J. A." box="[810,1036,999,1023]" pageId="24" pageNumber="325" refId="ref28388" refString="Murdy, E. O., Birdsong, R. S. &amp; Musick, J. A. (1997) Fishes of Chesapeake Bay. Smithsonian Institution Press, Washington, 324 pp." type="book" year="1997">
Murdy
<emphasis id="B93BEAFBFFF87E23B9621ED597F6A8A5" box="[886,929,999,1023]" italics="true" pageId="24" pageNumber="325">et al</emphasis>
. 1997:196
</bibRefCitation>
(fishes of Chesapeake Bay; listed, short description).—
<treatmentCitation id="0AEE10F8FFF87E23BB74193B96F4AF44" author="Castro-Aguirre, J. L. &amp; Espinosa, P. H. &amp; Schmitter-Soto, J. J." box="[352,675,1030,1054]" page="371" pageId="24" pageNumber="325" year="1999">
<bibRefCitation id="EFDE4B18FFF87E23BB74193B96F4AF44" author="Castro-Aguirre, J. L. &amp; Espinosa, P. H. &amp; Schmitter-Soto, J. J." box="[352,675,1030,1054]" pageId="24" pageNumber="325" refId="ref25287" refString="Castro-Aguirre, J. L., Espinosa, P. H. &amp; Schmitter-Soto, J. J. (1999) Ictiofauna estuarino-lagunar y vicaria de Mexico. Noriega- Limusa, IPN. Mexico City, 711 pp." type="book" year="1999">
Castro-Aguirre
<emphasis id="B93BEAFBFFF87E23B815193A9679AF44" box="[513,558,1030,1054]" italics="true" pageId="24" pageNumber="325">et al</emphasis>
. 1999: 371
</bibRefCitation>
</treatmentCitation>
(ichthyofauna of
<collectingCountry id="F3587679FFF87E23B94F193B97FBAF44" box="[859,940,1030,1054]" name="Mexico" pageId="24" pageNumber="325">Mexico</collectingCountry>
; listed).—
<treatmentCitation id="0AEE10F8FFF87E23BE00193B915FAF44" author="Nelson, J. S. &amp; Crossman, E. J. &amp; Espinosa Perez, H. &amp; Findley, L. T. &amp; Gilbert, C. R. &amp; Lea, R. N. &amp; Williams, J. D." box="[1044,1288,1030,1054]" page="147" pageId="24" pageNumber="325" year="2004">
<bibRefCitation id="EFDE4B18FFF87E23BE00193B915FAF44" author="Nelson, J. S. &amp; Crossman, E. J. &amp; Espinosa Perez, H. &amp; Findley, L. T. &amp; Gilbert, C. R. &amp; Lea, R. N. &amp; Williams, J. D." box="[1044,1288,1030,1054]" pageId="24" pageNumber="325" refId="ref28425" refString="Nelson, J. S., Crossman, E. J., Espinosa Perez, H., Findley, L. T., Gilbert, C. R., Lea, R. N. &amp; Williams, J. D. (2004) Common and scientific names of fishes from the United States, Canada, and Mexico. 6 th Edition. American Fisheries Society Special Publication 29. American Fisheries Society, Bethesda, Maryland, 386 pp." type="book" year="2004">
Nelson
<emphasis id="B93BEAFBFFF87E23BE71193A90C4AF44" box="[1125,1171,1030,1054]" italics="true" pageId="24" pageNumber="325">et al</emphasis>
. 2004: 147
</bibRefCitation>
</treatmentCitation>
(common and scientific names of the fishes from the
<collectingCountry id="F3587679FFF87E23B85819199681AF67" box="[588,726,1060,1085]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
,
<collectingCountry id="F3587679FFF87E23B8CB19199779AF66" box="[735,814,1060,1084]" name="Canada" pageId="24" pageNumber="325">Canada</collectingCountry>
,
<collectingCountry id="F3587679FFF87E23B923191997DFAF66" box="[823,904,1060,1084]" name="Mexico" pageId="24" pageNumber="325">Mexico</collectingCountry>
; listed).—
<treatmentCitation id="0AEE10F8FFF87E23B9F919199104AF67" author="McEachran, J. D. &amp; Fechhelm, J. D." box="[1005,1363,1060,1085]" page="432" pageId="24" pageNumber="325" year="2005">
<bibRefCitation id="EFDE4B18FFF87E23B9F919199104AF67" author="McEachran, J. D. &amp; Fechhelm, J. D." box="[1005,1363,1060,1085]" pageId="24" pageNumber="325" refId="ref28236" refString="McEachran, J. D. &amp; Fechhelm, J. D. (2005) F ishes of the Gulf of Mexico. Vol. 2. Scorpaeniformes to Tetraodontiformes. University of Texas Press, Austin, Texas, viii + 1004 pp." type="book" year="2005">McEachran &amp; Fechhelm 2005: 432</bibRefCitation>
</treatmentCitation>
(fishes of the Gulf of
<collectingCountry id="F3587679FFF87E23BB4E197E95FCAF01" box="[346,427,1091,1115]" name="Mexico" pageId="24" pageNumber="325">Mexico</collectingCountry>
; description; illustration).—
<treatmentCitation id="0AEE10F8FFF87E23B8DD197E97F5AF01" author="Page, L. M. &amp; Espinosa-Perez, H. &amp; Findley, L. D. &amp; Gilbert, C. R. &amp; Lea, R. N. &amp; Mandrak, N. E. &amp; Mayden, R. L. &amp; Nelson, J. S." box="[713,930,1091,1115]" page="152" pageId="24" pageNumber="325" year="2013">
<bibRefCitation id="EFDE4B18FFF87E23B8DD197E97F5AF01" author="Page, L. M. &amp; Espinosa-Perez, H. &amp; Findley, L. D. &amp; Gilbert, C. R. &amp; Lea, R. N. &amp; Mandrak, N. E. &amp; Mayden, R. L. &amp; Nelson, J. S." box="[713,930,1091,1115]" pageId="24" pageNumber="325" refId="ref28600" refString="Page, L. M., Espinosa-Perez, H., Findley, L. D., Gilbert, C. R., Lea, R. N., Mandrak, N. E., Mayden, R. L. &amp; Nelson, J. S. (2013) Common and scientific names of fishes from the United States, Canada, and Mexico. 7 th Edition. American Fisheries Society Special Publication 34. American Fisheries Society, Bethesda, Maryland, 384 pp." type="book" year="2013">
Page
<emphasis id="B93BEAFBFFF87E23B91619799778AF01" box="[770,815,1091,1115]" italics="true" pageId="24" pageNumber="325">et al</emphasis>
. 2013: 152
</bibRefCitation>
</treatmentCitation>
(common and scientific names of the fishes from the
<collectingCountry id="F3587679FFF87E23BAF8195F9520AF20" box="[236,375,1122,1146]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
,
<collectingCountry id="F3587679FFF87E23BB95195F9587AF20" box="[385,464,1122,1146]" name="Canada" pageId="24" pageNumber="325">Canada</collectingCountry>
,
<collectingCountry id="F3587679FFF87E23BBCE195F967CAF20" box="[474,555,1122,1146]" name="Mexico" pageId="24" pageNumber="325">Mexico</collectingCountry>
; listed).—
<bibRefCitation id="EFDE4B18FFF87E23B884195F97ECAF20" author="Raz-Guzman, A. &amp; Huidobro, L. &amp; Padilla, V." box="[656,955,1122,1146]" pageId="24" pageNumber="325" pagination="341 - 362" refId="ref28924" refString="Raz-Guzman, A., Huidobro, L. &amp; Padilla, V. (2018) An updated checklist and characterisation of the ichthyofauna (Elasmobranchii and Actinopterygii) of the Laguna de Tamiahua, Veracruz, Mexico. Acta Ichthyologica et Piscatoria, 48 (4), 341 - 362. https: // doi. org / 10.3750 / AIEP / 02451" type="journal article" year="2018">
Raz-Guzmán
<emphasis id="B93BEAFBFFF87E23B90B195E971BAF20" box="[799,844,1122,1146]" italics="true" pageId="24" pageNumber="325">et al</emphasis>
. 2018:346
</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3556562FFF87E23BA83199590D7AEB8" pageId="24" pageNumber="325" type="materials_examined">
<paragraph id="8BF036E9FFF87E23BA83199590D7AEB8" blockId="24.[151,1437,1192,2011]" pageId="24" pageNumber="325">
<emphasis id="B93BEAFBFFF87E23BA83199595D7AF98" bold="true" box="[151,384,1192,1218]" pageId="24" pageNumber="325">Material examined.</emphasis>
<materialsCitation id="3B273CB4FFF87E23BB93199590F0AF98" ID-GBIF-Occurrence="3022747304" box="[391,1191,1192,1219]" collectionCode="ANSP" country="United States of America" location="South Carolina" pageId="24" pageNumber="325" specimenCode="ANSP 11576" specimenCount="1" stateProvince="South Carolina" typeStatus="holotype">
<emphasis id="B93BEAFBFFF87E23BB93199595A0AF98" box="[391,503,1192,1218]" italics="true" pageId="24" pageNumber="325">
<typeStatus id="54F4884BFFF87E23BB93199595A6AF98" box="[391,497,1192,1218]" pageId="24" pageNumber="325" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DBE99E92FFF87E23BBEA199596C6AF99" box="[510,657,1192,1219]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="24" pageNumber="325">ANSP 11576</specimenCode>
(
<quantity id="4CB79B0CFFF87E23B8B71995974EAF99" box="[675,793,1192,1219]" metricMagnitude="0" metricUnit="m" metricValue="1.1949999999999998" pageId="24" pageNumber="325" unit="mm" value="1195.0">1, 195 mm</quantity>
SL),
<collectingRegion id="498BF80BFFF87E23B94C19959054AF98" box="[856,1027,1192,1218]" inferredBySuffix="true" pageId="24" pageNumber="325">South Carolina</collectingRegion>
,
<collectingCountry id="F3587679FFF87E23BE1A199590F0AF98" box="[1038,1191,1192,1218]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
</materialsCitation>
.
<emphasis id="B93BEAFBFFF87E23BEA6199491CBAF98" box="[1202,1436,1193,1218]" italics="true" pageId="24" pageNumber="325">Non-type specimens:</emphasis>
<materialsCitation id="3B273CB4FFF87E23BA8319F1972EAFBC" ID-GBIF-Occurrence="3022747362" box="[151,889,1228,1255]" collectionCode="ANSP" country="United States of America" county="Charleston Co." location="Charleston Co." pageId="24" pageNumber="325" specimenCode="ANSP 151380" specimenCount="1" stateProvince="South Carolina">
<specimenCode id="DBE99E92FFF87E23BA8319F1956EAFBD" box="[151,313,1228,1255]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="24" pageNumber="325">ANSP 151380</specimenCode>
(8),
<collectingCounty id="62914E65FFF87E23BB6419F1964CAFBC" box="[368,539,1228,1254]" pageId="24" pageNumber="325">Charleston Co.</collectingCounty>
,
<collectingRegion id="498BF80BFFF87E23B83319F19684AFBC" box="[551,723,1228,1254]" country="United States of America" name="South Carolina" pageId="24" pageNumber="325">South Carolina</collectingRegion>
,
<collectingCountry id="F3587679FFF87E23B8CB19F1972EAFBC" box="[735,889,1228,1254]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B273CB4FFF87E23B99019F1953EAE50" ID-GBIF-Occurrence="3022747319" collectionCode="ANSP" country="United States of America" county="Charleston Co." location="Charleston Co." pageId="24" pageNumber="325" specimenCode="ANSP 151385" specimenCount="1" stateProvince="South Carolina">
<specimenCode id="DBE99E92FFF87E23B99019F19071AFBD" box="[900,1062,1228,1255]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="24" pageNumber="325">ANSP 151385</specimenCode>
(4),
<collectingCounty id="62914E65FFF87E23BE4819F19150AFBC" box="[1116,1287,1228,1254]" pageId="24" pageNumber="325">Charleston Co.</collectingCounty>
,
<collectingRegion id="498BF80BFFF87E23BF0719F19493AE50" country="United States of America" name="South Carolina" pageId="24" pageNumber="325">South Carolina</collectingRegion>
,
<collectingCountry id="F3587679FFF87E23BAC419CD953EAE50" box="[208,361,1264,1290]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B273CB4FFF87E23BB6019CD9036AE50" ID-GBIF-Occurrence="3022747340" box="[372,1121,1264,1291]" collectionCode="ANSP" country="United States of America" county="Charleston Co." location="Charleston Co." pageId="24" pageNumber="325" specimenCode="ANSP 151386" specimenCount="1" stateProvince="South Carolina">
<specimenCode id="DBE99E92FFF87E23BB6019CD9642AE51" box="[372,533,1264,1291]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="24" pageNumber="325">ANSP 151386</specimenCode>
(15),
<collectingCounty id="62914E65FFF87E23B84E19CD9753AE50" box="[602,772,1264,1290]" pageId="24" pageNumber="325">Charleston Co.</collectingCounty>
,
<collectingRegion id="498BF80BFFF87E23B90419CD97EBAE50" box="[784,956,1264,1290]" country="United States of America" name="South Carolina" pageId="24" pageNumber="325">South Carolina</collectingRegion>
,
<collectingCountry id="F3587679FFF87E23B9DC19CD9036AE50" box="[968,1121,1264,1290]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B273CB4FFF87E23BE7919CD958BAE74" ID-GBIF-Occurrence="3022747301" collectionCode="USNM" country="United States of America" location="Brunswick" pageId="24" pageNumber="325" specimenCode="USNM 203280" specimenCount="1" stateProvince="Georgia">
<specimenCode id="DBE99E92FFF87E23BE7919CD914DAE50" box="[1133,1306,1264,1290]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="325" type="Museum">USNM 203280</specimenCode>
(3),
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,
<collectingRegion id="498BF80BFFF87E23BACF1829956FAE74" box="[219,312,1300,1326]" country="United States of America" name="Georgia" pageId="24" pageNumber="325">Georgia</collectingRegion>
,
<collectingCountry id="F3587679FFF87E23BB571829958BAE74" box="[323,476,1300,1326]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B273CB4FFF87E23BBF3182991CFAE74" ID-GBIF-Occurrence="3022747361" box="[487,1432,1300,1327]" collectionCode="MZUSP" country="United States of America" county="Charlotte Co." location="Gasparilla" municipality="Florida" pageId="24" pageNumber="325" specimenCode="MZUSP 69846" specimenCount="1">
<specimenCode id="DBE99E92FFF87E23BBF3182996C7AE74" box="[487,656,1300,1326]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="24" pageNumber="325">MZUSP 69846</specimenCode>
(4, 96-
<quantity id="4CB79B0CFFF87E23B8F51829976AAE75" box="[737,829,1300,1327]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" pageId="24" pageNumber="325" unit="mm" value="113.0">113 mm</quantity>
SL),
<collectingCounty id="62914E65FFF87E23B96F18299043AE74" box="[891,1044,1300,1326]" pageId="24" pageNumber="325">Charlotte Co.</collectingCounty>
,
<location id="8E906032FFF87E23BE0B182990C2AE74" LSID="urn:lsid:plazi:treatment:03E687FFFFF87E21BA831F57955BABA3:8E906032FFF87E23BE0B182990C2AE74" box="[1055,1173,1300,1326]" country="United States of America" county="Charlotte Co." municipality="Florida" name="Gasparilla" pageId="24" pageNumber="325">Gasparilla</location>
,
<collectingMunicipality id="6B94AC93FFF87E23BEB4182990A3AE74" box="[1184,1268,1300,1326]" pageId="24" pageNumber="325">Florida</collectingMunicipality>
,
<collectingCountry id="F3587679FFF87E23BEEB182991CFAE74" box="[1279,1432,1300,1326]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B273CB4FFF87E23BA8318059745AE08" ID-GBIF-Occurrence="3022747310" box="[151,786,1336,1363]" collectionCode="ANSP" country="United States of America" location="Boca Grande" municipality="Florida" pageId="24" pageNumber="325" specimenCode="ANSP 79851" specimenCount="1">
<specimenCode id="DBE99E92FFF87E23BA8318059579AE09" box="[151,302,1336,1363]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="24" pageNumber="325">ANSP 79851</specimenCode>
(3),
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,
<collectingMunicipality id="6B94AC93FFF87E23B80718059630AE08" box="[531,615,1336,1362]" pageId="24" pageNumber="325">Florida</collectingMunicipality>
,
<collectingCountry id="F3587679FFF87E23B86118059745AE08" box="[629,786,1336,1362]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
</materialsCitation>
;
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(1),
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,
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,
<collectingCountry id="F3587679FFF87E23BEEF180591CFAE08" box="[1275,1432,1336,1362]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
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;
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(2),
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back at
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,
<collectingCounty id="62914E65FFF87E23B8621861969DAE2C" box="[630,714,1372,1398]" pageId="24" pageNumber="325">Florida</collectingCounty>
,
<collectingCountry id="F3587679FFF87E23B8C01861973BAE2C" box="[724,876,1372,1398]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
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;
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<specimenCode id="DBE99E92FFF87E23B96218619076AE2C" box="[886,1057,1372,1398]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="325" type="Museum">USNM 126784</specimenCode>
(4),
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,
<collectingMunicipality id="6B94AC93FFF87E23BEE618619111AE2C" box="[1266,1350,1372,1398]" pageId="24" pageNumber="325">Florida</collectingMunicipality>
,
<collectingCountry id="F3587679FFF87E23BF441861948AAEC0" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
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;
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<specimenCode id="DBE99E92FFF87E23BAFE18BD95DEAEC1" box="[234,393,1408,1435]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="325" type="Museum">USNM 30815</specimenCode>
(1),
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,
<collectingMunicipality id="6B94AC93FFF87E23B85418BD96C3AEC0" box="[576,660,1408,1434]" pageId="24" pageNumber="325">Florida</collectingMunicipality>
,
<collectingCountry id="F3587679FFF87E23B8B418BD976DAEC0" box="[672,826,1408,1434]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
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;
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(1),
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,
<collectingMunicipality id="6B94AC93FFF87E23BE8918BD90A6AEC0" box="[1181,1265,1408,1434]" pageId="24" pageNumber="325">Florida</collectingMunicipality>
,
<collectingCountry id="F3587679FFF87E23BEE918BD91CFAEC0" box="[1277,1432,1408,1434]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
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;
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<specimenCode id="DBE99E92FFF87E23BA8318999515AEE4" box="[151,322,1444,1470]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="325" type="Museum">USNM 127347</specimenCode>
(1),
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,
<collectingRegion id="498BF80BFFF87E23B81C1899962EAEE4" box="[520,633,1444,1470]" country="United States of America" name="Louisiana" pageId="24" pageNumber="325">Louisiana</collectingRegion>
,
<collectingCountry id="F3587679FFF87E23B8971899974CAEE4" box="[643,795,1444,1470]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
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;
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<specimenCode id="DBE99E92FFF87E23B93118999787AEE5" box="[805,976,1444,1471]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="325" type="Museum">USNM 125775</specimenCode>
(1),
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,
<collectingRegion id="498BF80BFFF87E23BEA6189890A1AEE5" box="[1202,1270,1445,1471]" country="United States of America" name="Texas" pageId="24" pageNumber="325">Texas</collectingRegion>
,
<collectingCountry id="F3587679FFF87E23BF14189991CFAEE4" box="[1280,1432,1444,1470]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
</materialsCitation>
;
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<specimenCode id="DBE99E92FFF87E23BA8318F59562AEB8" box="[151,309,1480,1506]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="325" type="Museum">USNM 31048</specimenCode>
(1),
<location id="8E906032FFF87E23BB7F18F59588AEB8" LSID="urn:lsid:plazi:treatment:03E687FFFFF87E21BA831F57955BABA3:8E906032FFF87E23BB7F18F59588AEB8" box="[363,479,1480,1506]" country="United States of America" name="Galveston" pageId="24" pageNumber="325" stateProvince="Texas">Galveston</location>
,
<collectingRegion id="498BF80BFFF87E23BBFE18F49679AEB9" box="[490,558,1481,1507]" country="United States of America" name="Texas" pageId="24" pageNumber="325">Texas</collectingRegion>
,
<collectingCountry id="F3587679FFF87E23B82D18F59685AEB8" box="[569,722,1480,1506]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
</materialsCitation>
;
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<specimenCode id="DBE99E92FFF87E23B8CA18F5972CAEB8" box="[734,891,1480,1506]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="325" type="Museum">USNM 61229</specimenCode>
(1),
<location id="8E906032FFF87E23B9A518F49040AEB9" LSID="urn:lsid:plazi:treatment:03E687FFFFF87E21BA831F57955BABA3:8E906032FFF87E23B9A518F49040AEB9" box="[945,1047,1481,1507]" country="Mexico" name="Progreso" pageId="24" pageNumber="325">Progreso</location>
,
<collectingCountry id="F3587679FFF87E23BE3718F5902BAEB8" box="[1059,1148,1480,1506]" name="Mexico" pageId="24" pageNumber="325">Mexico</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3556562FFF87E22BAD318D19177A964" lastPageId="25" lastPageNumber="326" pageId="24" pageNumber="325" type="diagnosis">
<paragraph id="8BF036E9FFF87E23BAD318D197EAAC34" blockId="24.[151,1437,1192,2011]" pageId="24" pageNumber="325">
<emphasis id="B93BEAFBFFF87E23BAD318D1956DAD5C" bold="true" box="[199,314,1516,1542]" pageId="24" pageNumber="325">Diagnosis</emphasis>
.
<taxonomicName id="4C4F4D6AFFF87E23BB5D18D19614AD5C" baseAuthorityName="Holbrook" baseAuthorityYear="1847" box="[329,579,1516,1542]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="littoralis">
<emphasis id="B93BEAFBFFF87E23BB5D18D19614AD5C" box="[329,579,1516,1542]" italics="true" pageId="24" pageNumber="325">Menticirrhus littoralis</emphasis>
</taxonomicName>
, which is found in the
<collectingCountry id="F3587679FFF87E23B95F18D197B5AD5C" box="[843,994,1516,1542]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
and the Gulf of Mexico, can be differentiated readily from its congeners due to the black blotch or hue on the tip of the upper caudal lobe in live or fresh specimens (
<emphasis id="B93BEAFBFFF87E23BB0F1B08956EAD14" box="[283,313,1589,1614]" italics="true" pageId="24" pageNumber="325">vs.</emphasis>
caudal fin without dark hue,
<figureCitation id="13742A6CFFF87E23B86E1B0996E3AD14" box="[634,692,1588,1614]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="24" pageNumber="325">Fig 2</figureCitation>
)
<taxonomicName id="4C4F4D6AFFF87E23B8D61B089710AD14" baseAuthorityName="Holbrook" baseAuthorityYear="1847" box="[706,839,1588,1614]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="littoralis">
<emphasis id="B93BEAFBFFF87E23B8D61B089710AD14" box="[706,839,1588,1614]" italics="true" pageId="24" pageNumber="325">M. littoralis</emphasis>
</taxonomicName>
can also be differentiated from
<taxonomicName id="4C4F4D6AFFF87E23BEB01B08911CAD14" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1188,1355,1589,1614]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="americanus">
<emphasis id="B93BEAFBFFF87E23BEB01B08911CAD14" box="[1188,1355,1589,1614]" italics="true" pageId="24" pageNumber="325">M. americanus</emphasis>
</taxonomicName>
, which is found in the
<collectingCountry id="F3587679FFF87E23BB541B659581AD28" box="[320,470,1624,1650]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
and the Gulf of Mexico, by pectoral-fin tip barely reaching tip of depressed pelvic fin tip of depressed pelvic fin (
<emphasis id="B93BEAFBFFF87E23BBDA1B4095B2ADCC" box="[462,485,1661,1686]" italics="true" pageId="24" pageNumber="325">vs</emphasis>
. surpassing tip of pelvic fin,
<figureCitation id="13742A6CFFF87E23B9251B419721ADCC" box="[817,886,1660,1686]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="24" pageNumber="325">Fig. 2</figureCitation>
), body without dark bars (
<emphasis id="B93BEAFBFFF87E23BEB61B4090EEADCC" box="[1186,1209,1661,1686]" italics="true" pageId="24" pageNumber="325">vs</emphasis>
. with irregular dark bars,
<figureCitation id="13742A6CFFF87E23BAC11B9D954EADE0" box="[213,281,1696,1722]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="24" pageNumber="325">Fig. 2</figureCitation>
); from
<taxonomicName id="4C4F4D6AFFF87E23BB791B9C95BEADE0" authorityName="Cuvier in Cuvier &amp; Valenciennes" authorityYear="1830" box="[365,489,1696,1722]" class="Actinopterygii" family="Sciaenidae" genus="Umbrina" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B93BEAFBFFF87E23BB791B9C95BEADE0" box="[365,489,1696,1722]" italics="true" pageId="24" pageNumber="325">M. gracilis</emphasis>
</taxonomicName>
, which is found on the southern and southeastern coasts of
<collectingCountry id="F3587679FFF87E23BE981B9D9084ADE0" box="[1164,1235,1696,1722]" name="Brazil" pageId="24" pageNumber="325">Brazil</collectingCountry>
, by having 2225 dorsal-fin rays (
<emphasis id="B93BEAFBFFF87E23BB5B1BF8953AAD84" box="[335,365,1733,1758]" italics="true" pageId="24" pageNumber="325">vs.</emphasis>
1821,
<tableCitation id="C6CD0352FFF87E23BBD81BF99674AD84" box="[460,547,1732,1758]" captionStart="TABLE 3" captionStartId="11.[151,239,404,428]" captionText="TABLE 3. Meristic data of Menticirrhus americanus, M. martinicensis, M. cuiaranensis, M. gracilis and M. littoralis. A. Dorsal-fin rays. B. Pectoral-fin rays. C. Gill rakers on the first gill arch. D. Scales above the lateral line." pageId="24" pageNumber="325">Table 3</tableCitation>
); from
<taxonomicName id="4C4F4D6AFFF87E23B8681BF8976CAD84" baseAuthorityName="Cuvier" baseAuthorityYear="1830" box="[636,827,1733,1758]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="martinicensis">
<emphasis id="B93BEAFBFFF87E23B8681BF8976CAD84" box="[636,827,1733,1758]" italics="true" pageId="24" pageNumber="325">M. martinicensis</emphasis>
</taxonomicName>
, which is found in the Caribbean and Atlantic South America, by pectoral-fin tip barely reaching tip of depressed pelvic fin (
<emphasis id="B93BEAFBFFF87E23B9A11BD4979BAC58" box="[949,972,1769,1794]" italics="true" pageId="24" pageNumber="325">vs</emphasis>
. surpassing tip of pelvic fin,
<figureCitation id="13742A6CFFF87E23BF191BD59107AC58" box="[1293,1360,1768,1794]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="24" pageNumber="325">Fig. 2</figureCitation>
), body without dark bars (
<emphasis id="B93BEAFBFFF87E23BB7B1A3095D1AC7C" box="[367,390,1805,1830]" italics="true" pageId="24" pageNumber="325">vs</emphasis>
. with irregular dark bars,
<figureCitation id="13742A6CFFF87E23B8BB1A3196A3AC7C" box="[687,756,1804,1830]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="24" pageNumber="325">Fig. 2</figureCitation>
), and from
<taxonomicName id="4C4F4D6AFFF87E23B96D1A3097AAAC7C" box="[889,1021,1804,1830]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="saxatilis">
<emphasis id="B93BEAFBFFF87E23B96D1A3097AAAC7C" box="[889,1021,1804,1830]" italics="true" pageId="24" pageNumber="325">M. saxatilis</emphasis>
</taxonomicName>
, which is found in the
<collectingCountry id="F3587679FFF87E23BF111A3191CBAC7C" box="[1285,1436,1804,1830]" name="United States of America" pageId="24" pageNumber="325">United States</collectingCountry>
and the Gulf of Mexico, by pectoral-fin tip barely reaching tip of depressed pelvic fin (
<emphasis id="B93BEAFBFFF87E23BE7B1A0C90D1AC10" box="[1135,1158,1841,1866]" italics="true" pageId="24" pageNumber="325">vs</emphasis>
. surpassing tip of pelvic fin,
<figureCitation id="13742A6CFFF87E23BAD01A69955FAC34" box="[196,264,1876,1902]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="24" pageNumber="325">Fig. 2</figureCitation>
), body without dark bars (
<emphasis id="B93BEAFBFFF87E23B8241A689610AC34" box="[560,583,1877,1902]" italics="true" pageId="24" pageNumber="325">vs</emphasis>
. with irregular dark bars,
<figureCitation id="13742A6CFFF87E23B97F1A6997E7AC34" box="[875,944,1876,1902]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="24" pageNumber="325">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BF036E9FFF87E22BAD31A459177A964" blockId="24.[151,1437,1192,2011]" lastBlockId="25.[151,1437,150,2013]" lastPageId="25" lastPageNumber="326" pageId="24" pageNumber="325">
<taxonomicName id="4C4F4D6AFFF87E23BAD31A459596ACC8" baseAuthorityName="Holbrook" baseAuthorityYear="1847" box="[199,449,1912,1938]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="littoralis">
<emphasis id="B93BEAFBFFF87E23BAD31A459596ACC8" box="[199,449,1912,1938]" italics="true" pageId="24" pageNumber="325">Menticirrhus littoralis</emphasis>
</taxonomicName>
can also be distinguished from its congeners in the eastern Pacific as follows: from
<taxonomicName id="4C4F4D6AFFF87E23BF6A1A449553ACEC" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="325" phylum="Chordata" rank="species" species="elongatus">
<emphasis id="B93BEAFBFFF87E23BF6A1A449553ACEC" italics="true" pageId="24" pageNumber="325">M. elongatus</emphasis>
</taxonomicName>
, which occurs from the Gulf of California to
<collectingCountry id="F3587679FFF87E23B90C1AA09719ACED" box="[792,846,1949,1975]" name="Peru" pageId="24" pageNumber="325">Peru</collectingCountry>
, by having 1215 gill rakers in the first arches (
<emphasis id="B93BEAFBFFF87E23BF6A1AA091CBACEC" box="[1406,1436,1949,1974]" italics="true" pageId="24" pageNumber="325">vs.</emphasis>
710,
<tableCitation id="C6CD0352FFF87E23BAC91AFD9565AC80" box="[221,306,1984,2010]" captionStart="TABLE 3" captionStartId="11.[151,239,404,428]" captionText="TABLE 3. Meristic data of Menticirrhus americanus, M. martinicensis, M. cuiaranensis, M. gracilis and M. littoralis. A. Dorsal-fin rays. B. Pectoral-fin rays. C. Gill rakers on the first gill arch. D. Scales above the lateral line." pageId="24" pageNumber="325">Table 3</tableCitation>
), pectoral-fin tip barely reaching tip of depressed pelvic fin (
<emphasis id="B93BEAFBFFF87E23B9F21AFC97AAAC80" box="[998,1021,1985,2010]" italics="true" pageId="24" pageNumber="325">vs</emphasis>
. surpassing tip of pelvic fin,
<figureCitation id="13742A6CFFF87E23BF5D1AFD91DAAC80" box="[1353,1421,1984,2010]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="24" pageNumber="325">Fig. 2</figureCitation>
); from
<taxonomicName id="4C4F4D6AFFF97E22BAC11DAA956DABEA" box="[213,314,151,176]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="326" phylum="Chordata" rank="species" species="nasus">
<emphasis id="B93BEAFBFFF97E22BAC11DAA956DABEA" box="[213,314,151,176]" italics="true" pageId="25" pageNumber="326">M. nasus</emphasis>
</taxonomicName>
, ranging from the Gulf of California to
<collectingCountry id="F3587679FFF97E22B8EC1DAA9779ABEB" box="[760,814,151,177]" name="Peru" pageId="25" pageNumber="326">Peru</collectingCountry>
, by having 1215 gill rakers in the first arches (
<emphasis id="B93BEAFBFFF97E22BF521DAA9133ABEA" box="[1350,1380,151,176]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
49,
<tableCitation id="C6CD0352FFF97E22BA831D8694BBAB8F" box="[151,236,187,213]" captionStart="TABLE 3" captionStartId="11.[151,239,404,428]" captionText="TABLE 3. Meristic data of Menticirrhus americanus, M. martinicensis, M. cuiaranensis, M. gracilis and M. littoralis. A. Dorsal-fin rays. B. Pectoral-fin rays. C. Gill rakers on the first gill arch. D. Scales above the lateral line." pageId="25" pageNumber="326">Table 3</tableCitation>
), pectoral-fin tip barely reaching tip of depressed pelvic fin (
<emphasis id="B93BEAFBFFF97E22B9B11D8697EBAB8E" box="[933,956,187,212]" italics="true" pageId="25" pageNumber="326">vs</emphasis>
. surpassing tip of pelvic fin,
<figureCitation id="13742A6CFFF97E22BF1E1D869119AB8F" box="[1290,1358,187,213]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="25" pageNumber="326">Fig. 2</figureCitation>
); from
<taxonomicName id="4C4F4D6AFFF97E22BA831DE29506ABA3" box="[151,337,223,249]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="326" phylum="Chordata" rank="species" species="ophicephalus">
<emphasis id="B93BEAFBFFF97E22BA831DE29506ABA3" box="[151,337,223,249]" italics="true" pageId="25" pageNumber="326">M. ophicephalus</emphasis>
</taxonomicName>
, ranging from
<collectingCountry id="F3587679FFF97E22BBED1DE29601ABA3" box="[505,598,223,249]" name="Ecuador" pageId="25" pageNumber="326">Ecuador</collectingCountry>
to
<collectingCountry id="F3587679FFF97E22B86D1DE296EFABA3" box="[633,696,223,249]" name="Chile" pageId="25" pageNumber="326">Chile</collectingCountry>
, by having 5060 scales with pores along lateral line to caudal-fin base (
<emphasis id="B93BEAFBFFF97E22BACD1C3E94AFAA46" box="[217,248,259,284]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
6366), 1215 gill rakers in the first arches (
<emphasis id="B93BEAFBFFF97E22B8E61C3E9747AA46" box="[754,784,259,284]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
69,
<tableCitation id="C6CD0352FFF97E22B95A1C3E97F5AA47" box="[846,930,259,285]" captionStart="TABLE 3" captionStartId="11.[151,239,404,428]" captionText="TABLE 3. Meristic data of Menticirrhus americanus, M. martinicensis, M. cuiaranensis, M. gracilis and M. littoralis. A. Dorsal-fin rays. B. Pectoral-fin rays. C. Gill rakers on the first gill arch. D. Scales above the lateral line." pageId="25" pageNumber="326">Table 3</tableCitation>
), S-shaped posterior margin of the caudal fin (
<emphasis id="B93BEAFBFFF97E22BAB51C1A94E8AA1A" box="[161,191,295,320]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
concave), scales present along base of soft dorsal fin (
<emphasis id="B93BEAFBFFF97E22B90E1C1A976FAA1A" box="[794,824,295,320]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
soft dorsal fin naked); from
<taxonomicName id="4C4F4D6AFFF97E22BE611C1A90AAAA1A" box="[1141,1277,295,320]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="326" phylum="Chordata" rank="species" species="paitensis">
<emphasis id="B93BEAFBFFF97E22BE611C1A90AAAA1A" box="[1141,1277,295,320]" italics="true" pageId="25" pageNumber="326">M. paitensis</emphasis>
</taxonomicName>
, ranging from the Gulf of California to
<collectingCountry id="F3587679FFF97E22BBA61C7695BFAA3F" box="[434,488,331,357]" name="Peru" pageId="25" pageNumber="326">Peru</collectingCountry>
, by having 5060 scales with pores along lateral line to caudal-fin base (
<emphasis id="B93BEAFBFFF97E22BF341C769169AA3E" box="[1312,1342,331,356]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
7698), 1820 pectoral-fin rays (
<emphasis id="B93BEAFBFFF97E22BBBD1C529590AAD2" box="[425,455,367,392]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
2123,
<tableCitation id="C6CD0352FFF97E22B80B1C529626AAD3" box="[543,625,367,393]" captionStart="TABLE 3" captionStartId="11.[151,239,404,428]" captionText="TABLE 3. Meristic data of Menticirrhus americanus, M. martinicensis, M. cuiaranensis, M. gracilis and M. littoralis. A. Dorsal-fin rays. B. Pectoral-fin rays. C. Gill rakers on the first gill arch. D. Scales above the lateral line." pageId="25" pageNumber="326">Table 3</tableCitation>
), and 1215 gill rakers in the first arches (
<emphasis id="B93BEAFBFFF97E22BE551C529037AAD2" box="[1089,1120,367,392]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
69,
<tableCitation id="C6CD0352FFF97E22BE8F1C5290B8AAD3" box="[1179,1263,367,393]" captionStart="TABLE 2" captionStartId="5.[151,239,508,534]" captionText="TABLE 2. Kimura-2-parameter (K2P) genetic distance among genetic lineages of Menticirrhus (below diagonal) and values of standard error (above diagonal). Numbers in bold represent intraespecific K2P genetic distance. Western Pacific (EP); Western Atlantic (WA)." pageId="25" pageNumber="326">Table 2</tableCitation>
), scales present along base of soft dorsal fin (
<emphasis id="B93BEAFBFFF97E22BBCC1CAE95A1AAF6" box="[472,502,403,428]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
soft dorsal fin naked), pectoral-fin tip barely reaching tip of depressed pelvic fin (
<emphasis id="B93BEAFBFFF97E22BF6A1CAE91C2AAF6" box="[1406,1429,403,428]" italics="true" pageId="25" pageNumber="326">vs</emphasis>
. surpassing tip of pelvic fin,
<figureCitation id="13742A6CFFF97E22BBC11C8A964EAA8B" box="[469,537,439,465]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="25" pageNumber="326">Fig. 2</figureCitation>
); from
<taxonomicName id="4C4F4D6AFFF97E22B87B1C8A974EAA8A" box="[623,793,439,464]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="326" phylum="Chordata" rank="species" species="panamensis">
<emphasis id="B93BEAFBFFF97E22B87B1C8A974EAA8A" box="[623,793,439,464]" italics="true" pageId="25" pageNumber="326">M. panamensis</emphasis>
</taxonomicName>
, ranging from the Gulf of California to
<collectingCountry id="F3587679FFF97E22BEF41C8A9148AA8B" box="[1248,1311,439,465]" name="Chile" pageId="25" pageNumber="326">Chile</collectingCountry>
, by having seven anal-fin rays (
<emphasis id="B93BEAFBFFF97E22BB6B1CE695CAAAAE" box="[383,413,475,500]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
eight or nine), 1215 gill rakers in the first arches (
<emphasis id="B93BEAFBFFF97E22B9FA1CE6905BAAAE" box="[1006,1036,475,500]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
38,
<tableCitation id="C6CD0352FFF97E22BE591CE690F5AAAF" box="[1101,1186,475,501]" captionStart="TABLE 3" captionStartId="11.[151,239,404,428]" captionText="TABLE 3. Meristic data of Menticirrhus americanus, M. martinicensis, M. cuiaranensis, M. gracilis and M. littoralis. A. Dorsal-fin rays. B. Pectoral-fin rays. C. Gill rakers on the first gill arch. D. Scales above the lateral line." pageId="25" pageNumber="326">Table 3</tableCitation>
), scales present along base of soft dorsal fin (
<emphasis id="B93BEAFBFFF97E22BB8C1CC295E1A942" box="[408,438,511,536]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
soft dorsal fin naked); from
<taxonomicName id="4C4F4D6AFFF97E22B8E11CC297DFA943" box="[757,904,511,537]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="326" phylum="Chordata" rank="species" species="undulatus">
<emphasis id="B93BEAFBFFF97E22B8E11CC297DFA943" box="[757,904,511,537]" italics="true" pageId="25" pageNumber="326">M. undulatus</emphasis>
</taxonomicName>
, which is found in
<collectingRegion id="498BF80BFFF97E22BE4A1CC29085A943" box="[1118,1234,511,537]" country="United States of America" name="California" pageId="25" pageNumber="326">California</collectingRegion>
,
<collectingCountry id="F3587679FFF97E22BEC91CC29121A943" box="[1245,1398,511,537]" name="United States of America" pageId="25" pageNumber="326">United States</collectingCountry>
, by having seven anal-fin rays (
<emphasis id="B93BEAFBFFF97E22BBDA1F1E95BAA966" box="[462,493,547,572]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
eight or nine), scales along base of soft dorsal (
<emphasis id="B93BEAFBFFF97E22BE101F1E9074A966" box="[1028,1059,547,572]" italics="true" pageId="25" pageNumber="326">vs.</emphasis>
soft dorsal fin naked).
</paragraph>
</subSubSection>
<subSubSection id="C3556562FFF97E22BAD31F7B95FAAC5F" pageId="25" pageNumber="326" type="description">
<paragraph id="8BF036E9FFF97E22BAD31F7B962DAE9B" blockId="25.[151,1437,150,2013]" pageId="25" pageNumber="326">
<emphasis id="B93BEAFBFFF97E22BAD31F7B9506A93A" bold="true" box="[199,337,582,608]" pageId="25" pageNumber="326">Description</emphasis>
: D. X+2225; A. I+7; P. 1819; C. 17; GR 1215; LL. 5060; SA 89; SB. 1118 (
<tableCitation id="C6CD0352FFF97E22BF1A1F7A9127A93B" box="[1294,1392,583,609]" captionStart="TABLE 3" captionStartId="11.[151,239,404,428]" captionText="TABLE 3. Meristic data of Menticirrhus americanus, M. martinicensis, M. cuiaranensis, M. gracilis and M. littoralis. A. Dorsal-fin rays. B. Pectoral-fin rays. C. Gill rakers on the first gill arch. D. Scales above the lateral line." pageId="25" pageNumber="326">Tables 3</tableCitation>
,
<tableCitation id="C6CD0352FFF97E22BF6A1F7A91D8A93B" box="[1406,1423,583,609]" captionStart="TABLE 4" captionStartId="12.[151,239,472,498]" captionText="TABLE 4. Measurements of Menticirrhus americanus, M. martinicensis, M. cuiaranensis, M. gracilis and M. littoralis, expressed as percentages of standard length. T = neotype specimen; H = holotype; N = number of non-type specimens examined." pageId="25" pageNumber="326">4</tableCitation>
). Body slender in lateral view, not compressed, ventral profile flat; maximum depth at first dorsal-fin origin. Dorsal profile ascending steeply from snout to eye-level, ascending, nearly straight, over eye, gently convex to first dorsal-fin base, straight, slightly descending along second dorsal-fin base, concave in caudal peduncle. Ventral profile convex on snout, nearly straight on head to anal fin origin, ascending, nearly concave, along anal-fin base to caudal-fin base. Snout deep, with blunt to pug-nosed tip, triangular in lateral view. Mouth subterminal, upper jaw slender, rear tip slightly surpassing a vertical line through middle of eye. Premaxilla with 47 rows of acicular teeth, external most (about 2030) larger, caniniform; dentary with 47 tooth rows. Eye slightly oval, nearly two times snout length, not adjoining dorsal profile. Interorbital space smaller than orbital diameter, convex, covered with large ctenoid scales. Nostrils small, anterior nearly round, posterior larger, oval, nearly on horizontal line through ventral pupil border. Head with five sensory pores on snout, in semicircle around mouth angle, followed by larger central pore, lower jaw with four pores, two on each side encircling mental barbel, pores otherwise absent; mental barbel short, rigid, with blunt tip without pore. Lateral line slightly arched to second dorsal-fin origin, then straight, descending to caudal peduncle, nearly horizontal elsewhere to caudal-fin tip. Preopercle margin rough, with about ten subtle spines. Opercle tip fleshy, blunt, over vertical line that passes through pectoral-fin base. Gill rakers short. Ctenoid scales on trunk, belly, pectoral-fin base, opercle, preopercle, infraorbital, and interorbital region to snout, cycloid in gular area and encircling nostrils. Dorsal fins without scales on interradial membranes, second dorsal fin with basal sheath of scales (one row); pectoral-fin base densely covered by small ctenoid scales, rows of cycloid scales along upper pectoral-fin rays (pectoral tip naked), caudal-fin base covered with large ctenoid scales, rows of cycloid scales along rays, except at tip. Spinous dorsal fin short, first spine shortest, second and third spines longest; second dorsal fin adjoining first one. Origin of second dorsal fin in front of vertical line through pectoral-fin tip, second dorsal soft rays as long as longest dorsal spines. Anal-fin origin on vertical line passing through sixty-seventh rays of the second dorsal fin; first spine very slender. Pectoral fin very large, falcate, reaching pelvic fin tip. Pelvic-fin origin behind pectoral-fin base, shorter than pectoral fin. Caudal peduncle short, deep, depth larger than orbital diameter. Caudal fin short, truncate.
</paragraph>
<paragraph id="8BF036E9FFF97E22BAD318F79152AD77" blockId="25.[151,1437,150,2013]" pageId="25" pageNumber="326">
<emphasis id="B93BEAFBFFF97E22BAD318F795D0AEBF" bold="true" box="[199,391,1482,1509]" pageId="25" pageNumber="326">Color in alcohol</emphasis>
.
<typeStatus id="54F4884BFFF97E22BB8018F6959BAEBF" box="[404,460,1483,1509]" pageId="25" pageNumber="326">Type</typeStatus>
material with straw-beige pigmentation, mostly faded due to the age of the specimens. Other specimens: dorsal and anal fins beige, slender dark line along distal margin of spines and rays; pectoral fins beige, caudal fin straw-colored, dark area in distal region, more conspicuous on upper rays (
<figureCitation id="13742A6CFFF97E22BE8A1B2E90AFAD77" box="[1182,1272,1555,1581]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="25" pageNumber="326">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="8BF036E9FFF97E22BAD31B0B95FAAC5F" blockId="25.[151,1437,150,2013]" pageId="25" pageNumber="326">
<emphasis id="B93BEAFBFFF97E22BAD31B0B958CAD0A" bold="true" box="[199,475,1590,1616]" pageId="25" pageNumber="326">Color of fresh material</emphasis>
. Background color silvery gray dorsally, silvery white ventrally; head silver to silver gray, fainter below eyes, darker dorsally. First dorsal fin plain gray to dusky, tip of longest spines sometimes darkened; second dorsal fin rusty gray, lighter distally; anal fin light gray to dusky. Pectoral fin dusky, a darkened line along upper half; pelvic fin gray to dusky, sometimes with a dark hue along spine and neighboring two rays. Caudal fin plain gray or dusky, a dark area at base and lower rays; evident black spot on upper caudal-fin lobe tip, over 34 upper rays (
<figureCitation id="13742A6CFFF97E22BB081BD695F7AC5F" box="[284,416,1771,1797]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="25" pageNumber="326">Figs. 2B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3556562FFF97E22BAD31A3390B9AC17" pageId="25" pageNumber="326" type="distribution">
<paragraph id="8BF036E9FFF97E22BAD31A3390B9AC17" blockId="25.[151,1437,150,2013]" pageId="25" pageNumber="326">
<emphasis id="B93BEAFBFFF97E22BAD31A3395BAAC72" bold="true" box="[199,493,1806,1833]" pageId="25" pageNumber="326">Distribution and habitat</emphasis>
. Western North Atlantic from
<collectingRegion id="498BF80BFFF97E22B9521A3297A7AC73" box="[838,1008,1807,1833]" country="United States of America" name="North Carolina" pageId="25" pageNumber="326">North Carolina</collectingRegion>
to the Gulf of
<collectingRegion id="498BF80BFFF97E22BE8F1A3290A6AC73" box="[1179,1265,1807,1833]" country="Mexico" name="Mexico" pageId="25" pageNumber="326">Mexico</collectingRegion>
(
<figureCitation id="13742A6CFFF97E22BF151A329110AC73" box="[1281,1351,1807,1833]" captionStart="FIGURE 4" captionStartId="7.[151,250,1941,1965]" captionTargetBox="[176,1410,1004,1918]" captionTargetId="figure-51@7.[176,1410,1004,1918]" captionTargetPageId="7" captionText="FIGURE 4. The Atlantic coast of North, Central, and South America showing geographic distribution. A. Menticirrhus americanus (red) and M. martinicensis (green). B. M. littoralis (yellow), M. cuiaranensis (light blue) and M. gracilis (dark blue). Some symbols represent more than one locality or a large number of specimens." figureDoi="http://doi.org/10.5281/zenodo.4401574" httpUri="https://zenodo.org/record/4401574/files/figure.png" pageId="25" pageNumber="326">Fig. 4</figureCitation>
). Common in shallow waters, mainly in the vicinity of estuaries and in the surf zone, over sandy bottoms.
</paragraph>
</subSubSection>
<subSubSection id="C3556562FFF97E21BAD31A6A955BABA3" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326" type="discussion">
<paragraph id="8BF036E9FFF97E21BAD31A6A955BABA3" blockId="25.[151,1437,150,2013]" lastBlockId="26.[151,1436,151,249]" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326">
<emphasis id="B93BEAFBFFF97E22BAD31A6A9563AC2B" bold="true" box="[199,308,1879,1905]" pageId="25" pageNumber="326">Remarks</emphasis>
.
<taxonomicName id="4C4F4D6AFFF97E22BB501A6A9645AC2B" authorityName="Holbrook" authorityYear="1847" box="[324,530,1879,1905]" class="Actinopterygii" family="Sciaenidae" genus="Umbrina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="326" phylum="Chordata" rank="species" species="littoralis">
<emphasis id="B93BEAFBFFF97E22BB501A6A9645AC2B" box="[324,530,1879,1905]" italics="true" pageId="25" pageNumber="326">Umbrina littoralis</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFDE4B18FFF97E22B8FE1A6A97FEAC2B" author="Holbrook, J. E." box="[746,937,1879,1905]" pageId="25" pageNumber="326" refId="ref26882" refString="Holbrook, J. E. (1847) Southern ichthyology: or, a description of the fishes inhabiting the waters of South Carolina Georgia and Florida. No. II. Wiley, New York and Putnam, London, 32 pp., 4 pls." type="book" year="1847">Holbrook (1847)</bibRefCitation>
from the
<typeStatus id="54F4884BFFF97E22BE091A6A90D5AC2B" box="[1053,1154,1879,1905]" pageId="25" pageNumber="326" type="holotype">holotype</typeStatus>
, ANSP 11576 (
<figureCitation id="13742A6CFFF97E22BF271A6A91D8AC2B" box="[1331,1423,1879,1905]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="25" pageNumber="326">Fig. 2A</figureCitation>
), collected in
<collectingRegion id="498BF80BFFF97E22BB341A46959BACCF" box="[288,460,1915,1941]" country="United States of America" name="South Carolina" pageId="25" pageNumber="326">South Carolina</collectingRegion>
, in the
<collectingCountry id="F3587679FFF97E22B80A1A4696E0ACCF" box="[542,695,1915,1941]" name="United States of America" pageId="25" pageNumber="326">United States</collectingCountry>
.
<taxonomicName id="4C4F4D6AFFF97E22B8D61A4697ECACCF" baseAuthorityName="Holbrook" baseAuthorityYear="1847" box="[706,955,1915,1941]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="326" phylum="Chordata" rank="species" species="littoralis">
<emphasis id="B93BEAFBFFF97E22B8D61A4697ECACCF" box="[706,955,1915,1941]" italics="true" pageId="25" pageNumber="326">Menticirrhus littoralis</emphasis>
</taxonomicName>
is recognized here as being closely related to
<taxonomicName id="4C4F4D6AFFF97E22BAA11AA2953EACE2" authorityName="Marceniuk &amp; Caires &amp; Rotundo &amp; Cerqueira &amp; Siccha-Ramirez &amp; Wosiacki &amp; Oliveira" authorityYear="2020" box="[181,361,1951,1976]" class="Actinopterygii" family="Sciaenidae" genus="Menticirrhus" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="326" phylum="Chordata" rank="species" species="cuiaranensis">
<emphasis id="B93BEAFBFFF97E22BAA11AA2953EACE2" box="[181,361,1951,1976]" italics="true" pageId="25" pageNumber="326">M. cuiaranensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4F4D6AFFF97E22BBB51AA2964AACE3" authorityName="Cuvier in Cuvier &amp; Valenciennes" authorityYear="1830" box="[417,541,1951,1977]" class="Actinopterygii" family="Sciaenidae" genus="Umbrina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="326" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B93BEAFBFFF97E22BBB51AA2964AACE3" box="[417,541,1951,1977]" italics="true" pageId="25" pageNumber="326">M. gracilis</emphasis>
</taxonomicName>
, but can be differentiated from these species based on its geographic range, the number of rays in the pectoral and dorsal fins, as well as the black blotch or hue on the tip of the upper caudal lobe (
<figureCitation id="13742A6CFFFA7E21BA8B1DAA9572ABEB" box="[159,293,151,177]" captionStart="FIGURE 2" captionStartId="6.[151,250,1945,1969]" captionTargetBox="[183,1405,808,1921]" captionTargetId="figure-288@6.[183,1405,808,1921]" captionTargetPageId="6" captionText="FIGURE 2. Menticirrhus littoralis in lateral view. A. Holotype, ANSP 11576, 195 mm SL, South Carolina, United States. B. Live specimen captured in the Gulf of Mexico by Luiz Malabarba. C. Fresh specimen, Mexico, photographed by Eduardo Perusquia. Scale bar equals 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4401570" httpUri="https://zenodo.org/record/4401570/files/figure.png" pageId="26" pageNumber="327">Figs. 2B, C</figureCitation>
). It is important to note that, while
<bibRefCitation id="EFDE4B18FFFA7E21B8A11DAA9714ABEB" author="Irwin, R. J." box="[693,835,151,177]" pageId="26" pageNumber="327" refId="ref27032" refString="Irwin, R. J. (1971) Geographical variation systematics and general biology of shore fishes of the genus Menticirrhus, family Sciaendae. Ph. D. dissertation, Tulane University, New Orleans, Louisiana, 295 pp." type="book" year="1971">Irwin (1971)</bibRefCitation>
reported that “the upper tip of the caudal fin is black in most specimens longer than
<quantity id="4CB79B0CFFFA7E21BBE01D869602AB8F" box="[500,597,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="327" unit="mm" value="100.0">100 mm</quantity>
”, he did not attribute any taxonomic significance to this trait in the species definition.
</paragraph>
</subSubSection>
</treatment>
</document>