357 lines
55 KiB
XML
357 lines
55 KiB
XML
<document id="EF3A1D97B49A9F47171135F01DA5F562" ID-CLB-Dataset="24210" ID-DOI="10.11646/zootaxa.4729.3.5" ID-GBIF-Dataset="2a9de8f4-231e-4412-b987-f252b5c21cb0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3632801" ID-ZooBank="DC0128B0-1105-4D5C-B01B-49C7B5A6AF2B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580476580708" checkinUser="plazi" docAuthor="Dow, Rory A., Phan, Quoc Toan & Choong, Chee Yen" docDate="2020" docId="03E687F0FF96FF9CFF534827624BFE9D" docLanguage="en" docName="zootaxa.4729.3.5.pdf" docOrigin="Zootaxa 4729 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protosticta kinabaluensis Laidlaw 1915" docType="treatment" docVersion="9" lastPageNumber="385" masterDocId="FFDFFF88FF9AFF92FFC44B4E6171FFFA" masterDocTitle="Protosticta joepani sp. nov. from Borneo with notes on P. kinabaluensis Laidlaw 1915 (Odonata: Zygoptera: Platystictidae)" masterLastPageNumber="387" masterPageNumber="371" pageNumber="383" updateTime="1698779871450" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2D1AB6880C4EBB4B2EC7FEDDBC058BC7">Protosticta joepani sp. nov. from Borneo with notes on P. kinabaluensis Laidlaw 1915 (Odonata: Zygoptera: Platystictidae)</mods:title>
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<heading id="D0B8818AFF96FF9EFF534827630EFC79" bold="true" box="[151,639,873,900]" fontSize="11" level="1" pageId="12" pageNumber="383" reason="1">
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<taxonomicName id="4C4F4D65FF96FF9EFF534827630EFC79" ID-CoL="4N78K" authority="Laidlaw, 1915" authorityName="Laidlaw" authorityYear="1915" box="[151,639,873,900]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="kinabaluensis">
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<emphasis id="B93BEAF4FF96FF9EFF534827630EFC79" bold="true" box="[151,639,873,900]" pageId="12" pageNumber="383">
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<emphasis id="B93BEAF4FF96FF9EFF53482760B8FC7E" bold="true" box="[151,457,873,900]" italics="true" pageId="12" pageNumber="383">Protosticta kinabaluensis</emphasis>
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<bibRefCitation id="EFDE4B17FF96FF9EFE144827630EFC79" author="Laidlaw, F. F." box="[464,639,873,899]" pageId="12" pageNumber="383" pagination="25 - 39" refId="ref9753" refString="Laidlaw, F. F. (1915) Contributions to a study of the dragonfly fauna of Borneo. Part III. A collection made on Mount Kina Balu by Mr J. C. Moulton in September and October 1913. Proceedings of the Zoological Society of London, 1915, 25 - 39. https: // doi. org / 10.1111 / j. 1469 - 7998.1915.00025. x" type="book chapter" year="1915">Laidlaw, 1915</bibRefCitation>
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<subSubSection id="C355656DFF96FF9EFF5348C36008FC15" pageId="12" pageNumber="383" type="description">
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<paragraph id="8BF036E6FF96FF9EFF5348C362A5FC5D" blockId="12.[151,980,873,935]" box="[151,980,908,935]" pageId="12" pageNumber="383">
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(
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<figureCitation id="13742A63FF96FF9EFF5B48C36197FC5D" box="[159,230,909,935]" captionStart="FIGURE 1" captionStartId="1.[151,250,1119,1144]" captionTargetBox="[156,1435,235,1044]" captionTargetId="figure@1.[151,1435,179,1091]" captionTargetPageId="1" captionText="FIGURE 1. Neighbor joining COI gene tree for fifteen specimens of Protosticta joepani and four of P. kinabaluensis plus a Drepanosticta dentifera outgroup. RMNH collection codes are shown for each specimen. Male is indicated by “m” and female by “f”, the exceptionally small sized female referred to P. joepani with reservations (see under that species) is indicated by “small”." figureDoi="http://doi.org/10.5281/zenodo.3632803" httpUri="https://zenodo.org/record/3632803/files/figure.png" pageId="12" pageNumber="383">Figs 1</figureCitation>
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,
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<figureCitation id="13742A63FF96FF9EFF3048C36051FC5D" box="[244,288,908,935]" captionStart="FIGURES 2–5" captionStartId="4.[151,264,1331,1356]" captionTargetBox="[176,1411,345,1306]" captionTargetId="figure@4.[176,1411,345,1306]" captionTargetPageId="4" captionText="FIGURES 2–5. Heads of Protosticta species. Protosticta joepani: (2) holotype male, dorsal-frontal view; (4) paratype female used for description, dorsal-frontal view. Protosticta kinabaluensis: (3) male (SAB5_PST3), dorsal view; (5) female (SAB12_ PST69), dorsal frontal view." figureDoi="http://doi.org/10.5281/zenodo.3632805" httpUri="https://zenodo.org/record/3632805/files/figure.png" pageId="12" pageNumber="383">3, 5</figureCitation>
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,
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<figureCitation id="13742A63FF96FF9EFEE848C3604DFC5D" box="[300,316,909,935]" captionStart="FIGURES 6–7" captionStartId="5.[151,264,940,965]" captionTargetBox="[151,1436,273,916]" captionTargetId="figure@5.[151,1436,273,916]" captionTargetPageId="5" captionText="FIGURES 6–7. Prothorax of males of Protosticta species in lateral view. (6) Protosticta joepani holotype male. (7) Protosticta kinabaluensis (SAB12_PST71)." figureDoi="http://doi.org/10.5281/zenodo.3632807" httpUri="https://zenodo.org/record/3632807/files/figure.png" pageId="12" pageNumber="383">7</figureCitation>
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,
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<figureCitation id="13742A63FF96FF9EFE8C48C360F0FC5D" box="[328,385,909,935]" captionStart="FIGURES 8–11" captionStartId="6.[151,264,1634,1659]" captionTargetBox="[151,1436,181,1609]" captionTargetId="figure@6.[151,1436,181,1609]" captionTargetPageId="6" captionText="FIGURES 8–11. Prothorax of females of Protosticta species. Protosticta joepani, paratype female used for description: (8) dorsal; (10) lateral. Protosticta kinabaluensis (SAB12_PST69): (9) dorsal; (11) lateral." figureDoi="http://doi.org/10.5281/zenodo.3632809" httpUri="https://zenodo.org/record/3632809/files/figure.png" pageId="12" pageNumber="383">9, 11</figureCitation>
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,
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<figureCitation id="13742A63FF96FF9EFE4948C260A4FC5D" box="[397,469,908,935]" captionStart="FIGURES 15–16" captionStartId="8.[151,264,706,731]" captionTargetBox="[151,1435,181,681]" captionTargetId="figure@8.[151,1435,181,681]" captionTargetPageId="8" captionText="FIGURES 15–16. Synthorax of Protosticta kinabaluensis in lateral view: (15) male (SAB12_PST71); (16) female (SAB12_ PST69)." figureDoi="http://doi.org/10.5281/zenodo.3632813" httpUri="https://zenodo.org/record/3632813/files/figure.png" pageId="12" pageNumber="383">15, 16</figureCitation>
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,
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<figureCitation id="13742A63FF96FF9EFE2548C3608EFC5D" box="[481,511,909,935]" captionStart="FIGURES 17–18" captionStartId="8.[151,264,1089,1114]" captionTargetBox="[153,1428,794,1059]" captionTargetId="figure@8.[151,1436,789,1063]" captionTargetPageId="8" captionText="FIGURES 17–18. Genital ligula of Protosticta species in ventral view. (17) Protosticta joepani paratype, Mount Kinabalu, 4 iii 2008. (18) Protosticta kinabaluensis, Mount Kinabalu, 1 v 1979." figureDoi="http://doi.org/10.5281/zenodo.3632815" httpUri="https://zenodo.org/record/3632815/files/figure.png" pageId="12" pageNumber="383">18</figureCitation>
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,
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<figureCitation id="13742A63FF96FF9EFDCF48C363D6FC5D" box="[523,679,909,935]" captionStart="FIGURES 19–26" captionStartId="9.[151,264,1870,1895]" captionTargetBox="[151,1436,417,1845]" captionTargetId="figure@9.[151,1436,417,1845]" captionTargetPageId="9" captionText="FIGURES 19–26. Terminal abdominal segments of Protosticta species. Protosticta joepani: (19) holotype male, dorsal; (21) holotype male, lateral; (23) paratype female used for description, dorsal; (25) paratype female used for description, lateral. Protosticta kinabaluensis: (20) male (SAB12_PST71) dorsal; (22) male (SAB12_PST71) lateral; (24) female (SAB12_PST69), dorsal; (26) female (SAB12_PST69), lateral." figureDoi="http://doi.org/10.5281/zenodo.3632817" httpUri="https://zenodo.org/record/3632817/files/figure.png" pageId="12" pageNumber="383">20, 22, 24, 26</figureCitation>
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,
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<figureCitation id="13742A63FF96FF9EFD7748C3638AFC5D" box="[691,763,909,935]" captionStart="FIGURES 27–30" captionStartId="10.[151,264,1207,1232]" captionTargetBox="[151,1437,181,1181]" captionTargetId="figure@10.[151,1437,181,1181]" captionTargetPageId="10" captionText="FIGURES 27–30. Male anal appendages of Protosticta species. All scale bars 0.2mm. Protosticta joepani: (27) holotype male, lateral; (29) holotype male, paraprocts in ventral view. Protosticta kinabaluensis: (28) male (SAB12_PST71) lateral view; (30) male (SAB12_PST71) paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3632819" httpUri="https://zenodo.org/record/3632819/files/figure.png" pageId="12" pageNumber="383">28, 30</figureCitation>
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,
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<figureCitation id="13742A63FF96FF9EFCC348C3623EFC5D" box="[775,847,909,935]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1943,1968]" captionTargetBox="[151,1436,597,1918]" captionTargetId="figure@11.[151,1436,597,1918]" captionTargetPageId="11" captionText="FIGURES 31–34. Male anal appendages of Protosticta species, dorsal view. Protosticta joepani: (31) holotype male; (33) sketch of paratype, Poring Hot Springs, 4 iii 2008, right cercus removed. Protosticta kinabaluensis: (32) sketch of male, 1 v 1979; (34) male (SAB12_PST71), right cercus removed." figureDoi="http://doi.org/10.5281/zenodo.3632821" httpUri="https://zenodo.org/record/3632821/files/figure.png" pageId="12" pageNumber="383">32, 34</figureCitation>
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,
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<figureCitation id="13742A63FF96FF9EFC9F48C26208FC5D" box="[859,889,908,935]" captionStart="FIGURE 35" captionStartId="13.[151,250,1655,1680]" captionTargetBox="[151,1436,274,1646]" captionTargetId="figure@13.[151,1436,258,1646]" captionTargetPageId="13" captionText="FIGURE 35. Distribution of Protosticta joepani (red circle) and P. kinabaluensis (yellow square)." figureDoi="http://doi.org/10.5281/zenodo.3632823" httpUri="https://zenodo.org/record/3632823/files/figure.png" pageId="12" pageNumber="383">35</figureCitation>
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,
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<figureCitation id="13742A63FF96FF9EFC4148C362BDFC5D" box="[901,972,909,935]" captionStart="FIGURES 36–38" captionStartId="14.[151,264,1620,1645]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure@14.[151,1436,381,1595]" captionTargetPageId="14" captionText="FIGURES 36–38. Protosticta species in life. (36) Protosticta joepani, male, Kelabit Highlands, photo by G.T. Reels. (37) Protosticta kinabaluensis, male, photo by R.A. Dow; (38) Protosticta kinabaluensis, male, photo by R.A. Dow." figureDoi="http://doi.org/10.5281/zenodo.3632825" httpUri="https://zenodo.org/record/3632825/files/figure.png" pageId="12" pageNumber="383">37, 38</figureCitation>
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)
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</paragraph>
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<paragraph id="8BF036E6FF96FF9EFF53489B6008FC15" blockId="12.[151,377,981,1007]" box="[151,377,981,1007]" pageId="12" pageNumber="383">Selected references:</paragraph>
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<subSubSection id="C355656DFF96FF9EFF534F5462DCFB74" pageId="12" pageNumber="383" type="reference_group">
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<paragraph id="8BF036E6FF96FF9EFF534F54628DFB95" blockId="12.[151,1437,1049,1166]" pageId="12" pageNumber="383">
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<treatmentCitationGroup id="AB5F11C8FF96FF9EFF534F54628DFB95" pageId="12" pageNumber="383">
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<taxonomicName id="4C4F4D65FF96FF9EFF534F54635DFBC8" ID-CoL="4N78K" authority="Laidlaw, 1915" authorityName="Laidlaw" authorityYear="1915" box="[151,556,1049,1074]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="kinabaluensis">
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<emphasis id="B93BEAF4FF96FF9EFF534F5460E6FBCB" box="[151,407,1049,1073]" italics="true" pageId="12" pageNumber="383">Protosticta kinabaluensis</emphasis>
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<bibRefCitation id="EFDE4B17FF96FF9EFE594F57635DFBC8" author="Laidlaw, F. F." box="[413,556,1049,1074]" pageId="12" pageNumber="383" pagination="25 - 39" refId="ref9753" refString="Laidlaw, F. F. (1915) Contributions to a study of the dragonfly fauna of Borneo. Part III. A collection made on Mount Kina Balu by Mr J. C. Moulton in September and October 1913. Proceedings of the Zoological Society of London, 1915, 25 - 39. https: // doi. org / 10.1111 / j. 1469 - 7998.1915.00025. x" type="book chapter" year="1915">Laidlaw, 1915</bibRefCitation>
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</taxonomicName>
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;—
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<treatmentCitation id="0AEE10F7FF96FF9EFD8F4F576254FBCB" author="Laidlaw, F. F." box="[587,805,1049,1074]" page="37" pageId="12" pageNumber="383" year="1915">
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<bibRefCitation id="EFDE4B17FF96FF9EFD8F4F576254FBCB" author="Laidlaw, F. F." box="[587,805,1049,1074]" pageId="12" pageNumber="383" pagination="25 - 39" refId="ref9753" refString="Laidlaw, F. F. (1915) Contributions to a study of the dragonfly fauna of Borneo. Part III. A collection made on Mount Kina Balu by Mr J. C. Moulton in September and October 1913. Proceedings of the Zoological Society of London, 1915, 25 - 39. https: // doi. org / 10.1111 / j. 1469 - 7998.1915.00025. x" type="book chapter" year="1915">Laidlaw 1915: 37–38</bibRefCitation>
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</treatmentCitation>
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(original description male, Mount Kinabalu);—
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<treatmentCitation id="0AEE10F7FF96FF9EFACC4F57619CFBAA" author="Laidlaw, F. F." page="306" pageId="12" pageNumber="383" year="1924">
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<bibRefCitation id="EFDE4B17FF96FF9EFACC4F57619CFBAA" author="Laidlaw, F. F." pageId="12" pageNumber="383" pagination="296 - 308" refId="ref9826" refString="Laidlaw, F. F. (1924) A survey of the dragonfly fauna of the Malay Peninsula with notes on that of neighbouring countries, Part II. Journal of the Malayan Branch of the Royal Asiatic Society, 2 (3), 296 - 308, pls. III, IV and V." type="journal article" year="1924">Laidlaw 1924: 306</bibRefCitation>
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</treatmentCitation>
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(brief discussion);—
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<treatmentCitation id="0AEE10F7FF96FF9EFE064F7963E1FBAA" author="Lieftinck, M. A." box="[450,656,1079,1104]" page="285" pageId="12" pageNumber="383" year="1933">
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<bibRefCitation id="EFDE4B17FF96FF9EFE064F7963E1FBAA" author="Lieftinck, M. A." box="[450,656,1079,1104]" pageId="12" pageNumber="383" pagination="281 - 296" refId="ref9942" refString="Lieftinck, M. A. (1933) A synopsis of the Bornean Platystictinae, with descriptions of three new species (Odon.). Konowia, 11 (4), 281 - 296. [1932]" type="journal article" year="1933">Lieftinck 1933: 285</bibRefCitation>
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</treatmentCitation>
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(in key to Bornean species supposed to be in
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<taxonomicName id="4C4F4D65FF96FF9EFBA24F7765A7FBAA" ID-CoL="6XZ8" authorityName="Selys" authorityYear="1885" box="[1126,1238,1081,1104]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="383" phylum="Arthropoda" rank="genus">
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<emphasis id="B93BEAF4FF96FF9EFBA24F7765A7FBAA" box="[1126,1238,1081,1104]" italics="true" pageId="12" pageNumber="383">Protosticta</emphasis>
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</taxonomicName>
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);—
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<treatmentCitation id="0AEE10F7FF96FF9EFB3F4F796074FB95" author="Lieftinck, M. A." page="31" pageId="12" pageNumber="383" year="1954">
|
||
<bibRefCitation id="EFDE4B17FF96FF9EFB3F4F796074FB95" author="Lieftinck, M. A." pageId="12" pageNumber="383" pagination="1 - 202" refId="ref9983" refString="Lieftinck, M. A. (1954) Handlist of Malaysian Odonata. A catalogue of the dragonflies of the Malay Peninsula, Sumatra, Java and Borneo, including the adjacent small islands. Treubia, 22 (Supplement), i-xiii + 1 - 202, 1 folded map excl." type="journal article" year="1954">Lieftinck 1954: 31–32</bibRefCitation>
|
||
</treatmentCitation>
|
||
;—
|
||
<treatmentCitation id="0AEE10F7FF96FF9EFEE14F1960DDFB95" author="Orr, A. G." box="[293,428,1111,1135]" page="72" pageId="12" pageNumber="383" year="2003">
|
||
<bibRefCitation id="EFDE4B17FF96FF9EFEE14F1960DDFB95" author="Orr, A. G." box="[293,428,1111,1135]" pageId="12" pageNumber="383" refId="ref10453" refString="Orr, A. G. (2003) A guide to the dragonflies of Borneo: their identification and biology. Natural History Publications (Borneo), Kota Kinabalu, x + 195 pp." type="book" year="2003">Orr 2003: 72</bibRefCitation>
|
||
</treatmentCitation>
|
||
, Figs 91, 92 (brief discussion, photographs of both sexes).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BF036E6FF96FF9EFF534F3862DCFB74" blockId="12.[151,1437,1049,1166]" box="[151,941,1141,1166]" pageId="12" pageNumber="383">
|
||
<treatmentCitationGroup id="AB5F11C8FF96FF9EFF534F3862DCFB74" box="[151,941,1141,1166]" pageId="12" pageNumber="383">
|
||
<taxonomicName id="4C4F4D65FF96FF9EFF534F3860E2FB77" authority="Laidlaw 1934" authorityName="Laidlaw" authorityYear="1934" box="[151,403,1141,1165]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="kinabaluense">
|
||
<emphasis id="B93BEAF4FF96FF9EFF534F3860E2FB77" box="[151,403,1141,1165]" italics="true" pageId="12" pageNumber="383">Protosticta kinabaluense</emphasis>
|
||
</taxonomicName>
|
||
;—
|
||
<treatmentCitation id="0AEE10F7FF96FF9EFE754F3B6302FB74" author="Laidlaw, F. F." box="[433,627,1141,1166]" page="551" pageId="12" pageNumber="383" year="1934">
|
||
<bibRefCitation id="EFDE4B17FF96FF9EFE754F3B6302FB74" author="Laidlaw, F. F." box="[433,627,1141,1166]" pageId="12" pageNumber="383" pagination="549 - 561" refId="ref9884" refString="Laidlaw, F. F. (1934) A note of the dragonfly fauna (Odonata) of Mount Kinabalu and of some other mountain areas of Malaysia, with a description of some new or little known species. Journal of the Federated Malay States Museums, 17 (3), 549 - 561." type="journal article" year="1934">Laidlaw 1934: 551</bibRefCitation>
|
||
</treatmentCitation>
|
||
(listed from Mount Kinabalu).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C355656DFF96FF9CFF534FF565F1FF49" lastPageId="14" lastPageNumber="385" pageId="12" pageNumber="383" type="materials_examined">
|
||
<paragraph id="8BF036E6FF96FF9EFF534FF56342FA54" blockId="12.[151,1437,1211,2030]" pageId="12" pageNumber="383">
|
||
<emphasis id="B93BEAF4FF96FF9EFF534FF560F2FB2C" bold="true" box="[151,387,1211,1238]" pageId="12" pageNumber="383">Material examined.</emphasis>
|
||
All from
|
||
<collectingRegion id="498BF804FF96FF9EFE334FF2634AFB2C" box="[503,571,1212,1238]" country="Malaysia" name="Sabah" pageId="12" pageNumber="383">Sabah</collectingRegion>
|
||
in Malaysian Borneo.
|
||
<specimenCount id="9D49FD6FFF96FF9EFC874FF2621EFB2C" box="[835,879,1212,1238]" pageId="12" pageNumber="383" type="male">1 ♂</specimenCount>
|
||
(SAB5_PST3), Sungai Liwagu, ca
|
||
<quantity id="4CB79B03FF96FF9EFACE4FF56428FB2C" box="[1290,1369,1211,1238]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="383" unit="m" value="1500.0">1500m</quantity>
|
||
a.s.l., Headquarters area at Mount Kinabalu National Park, West Coast Division, (ca
|
||
<geoCoordinate id="EE7B5021FF96FF9EFC314F91653DFB03" box="[1013,1100,1247,1273]" degrees="6.008" direction="north" orientation="latitude" pageId="12" pageNumber="383" precision="55" value="6.008">6.008N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7B5021FF96FF9EFB914F9165B4FB00" box="[1109,1221,1247,1274]" degrees="116.549" direction="east" orientation="longitude" pageId="12" pageNumber="383" precision="55" value="116.549">116.549E</geoCoordinate>
|
||
), ca
|
||
<quantity id="4CB79B03FF96FF9EFB304F916433FB00" box="[1268,1346,1247,1274]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="383" unit="m" value="1500.0">1500m</quantity>
|
||
a.s.l.
|
||
<date id="FFF11026FF96FF9EFA444FAE619FFAE4" pageId="12" pageNumber="383" value="2005-04-30">30 iv 2005</date>
|
||
, leg. unknown, in coll. Dow;
|
||
<specimenCount id="9D49FD6FFF96FF9EFDF14E4A6302FAE4" box="[565,627,1284,1310]" pageId="12" pageNumber="383" type="male">3 ♂♂</specimenCount>
|
||
(SAB12_PST20; SAB12_PST71,
|
||
<collectionCode id="ED5EAE23FF96FF9EFC334E4D6520FAE4" box="[1015,1105,1283,1310]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="383" type="Museum">RMNH</collectionCode>
|
||
.INS.507705; SAB12_PST74,
|
||
<collectionCode id="ED5EAE23FF96FF9EFF534E696180FAB8" box="[151,241,1319,1346]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="383" type="Museum">RMNH</collectionCode>
|
||
.INS.507710),
|
||
<specimenCount id="9D49FD6FFF96FF9EFE504E6660A4FAB8" box="[404,469,1320,1346]" pageId="12" pageNumber="383" type="female">2 ♀♀</specimenCount>
|
||
(SAB12_PST21, SAB12_PST70;
|
||
<collectionCode id="ED5EAE23FF96FF9EFCA54E6962CDFAB8" box="[865,956,1319,1346]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="383" type="Museum">RMNH</collectionCode>
|
||
.INS.507701), Silau-Silau stream and trail, same area, (ca
|
||
<geoCoordinate id="EE7B5021FF96FF9EFEFA4E0560E5FA9F" box="[318,404,1355,1381]" degrees="6.007" direction="north" orientation="latitude" pageId="12" pageNumber="383" precision="55" value="6.007">6.007N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7B5021FF96FF9EFE5B4E05637EFA9C" box="[415,527,1355,1382]" degrees="116.545" direction="east" orientation="longitude" pageId="12" pageNumber="383" precision="55" value="116.545">116.545E</geoCoordinate>
|
||
), ca
|
||
<quantity id="4CB79B03FF96FF9EFD864E0563A4FA9C" box="[578,725,1355,1382]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="12" pageNumber="383" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">1500–1600m</quantity>
|
||
a.s.l.,
|
||
<date id="FFF11026FF96FF9EFCD04E0262E2FA9C" box="[788,915,1356,1382]" pageId="12" pageNumber="383" value="2012-09-13">13 ix 2012</date>
|
||
, leg.
|
||
<collectionCode id="ED5EAE23FF96FF9EFC0A4E026295FA9C" box="[974,996,1356,1382]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="383">R</collectionCode>
|
||
.
|
||
<materialsCitation id="3B273CBBFF96FF9EFC2C4E026416FA9C" ID-GBIF-Occurrence="2561485011" box="[1000,1383,1356,1382]" collectionCode="RMNH" collectorName="A. Dow & Dow" pageId="12" pageNumber="383" specimenCount="1">
|
||
<collectorName id="26BA5330FF96FF9EFC2C4E026530FA9C" box="[1000,1089,1356,1382]" pageId="12" pageNumber="383">A. Dow</collectorName>
|
||
, in coll.
|
||
<collectorName id="26BA5330FF96FF9EFB664E0265A9FA9C" box="[1186,1240,1356,1382]" pageId="12" pageNumber="383">Dow</collectorName>
|
||
and
|
||
<collectionCode id="ED5EAE23FF96FF9EFACB4E026416FA9C" box="[1295,1383,1356,1382]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="383" type="Museum">RMNH</collectionCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B273CBBFF96FF9EFAB64E02657BFA70" ID-GBIF-Occurrence="2561485022" collectionCode="RMNH" collectorName="J. van Tol" pageId="12" pageNumber="383" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="9D49FD6FFF96FF9EFAB64E0264EDFA9C" box="[1394,1436,1356,1382]" pageId="12" pageNumber="383" type="female">1 ♀</specimenCount>
|
||
(SAB12_PST69,
|
||
<collectionCode id="ED5EAE23FF96FF9EFE9E4E2160C5FA70" box="[346,436,1391,1418]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="383" type="Museum">RMNH</collectionCode>
|
||
.INS.507714), same data but leg.
|
||
<collectorName id="26BA5330FF96FF9EFCDB4E3E62FCFA70" box="[799,909,1392,1418]" pageId="12" pageNumber="383">J. van Tol</collectorName>
|
||
, in
|
||
<collectionCode id="ED5EAE23FF96FF9EFC764E3E657BFA70" box="[946,1034,1392,1418]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="383" type="Museum">RMNH</collectionCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B273CBBFF96FF9EFBD74E3E635EFA54" ID-GBIF-Occurrence="2561485020" collectingDate="1979-05-01" collectionCode="NMNS" collectorName="S. Asahina" elevation="1600" pageId="12" pageNumber="383" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="9D49FD6FFF96FF9EFBD74E3E654FFA70" box="[1043,1086,1392,1418]" pageId="12" pageNumber="383" type="male">1 ♂</specimenCount>
|
||
, same location, ca
|
||
<elevation id="0062D1D5FF96FF9EFACA4E2164E6FA70" box="[1294,1431,1391,1418]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="383" unit="m" value="1600.0">
|
||
<quantity id="4CB79B03FF96FF9EFACA4E21642DFA73" box="[1294,1372,1391,1417]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="383" unit="m" value="1600.0">1600m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFF11026FF96FF9EFF534EDA618DFA54" box="[151,252,1428,1454]" pageId="12" pageNumber="383" value="1979-05-01">
|
||
<collectingDate id="EFB5E9CEFF96FF9EFF534EDA618DFA54" box="[151,252,1428,1454]" pageId="12" pageNumber="383" value="1979-05-01">1 v 1979</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26BA5330FF96FF9EFEF34EDA60C3FA54" box="[311,434,1428,1454]" pageId="12" pageNumber="383">S. Asahina</collectorName>
|
||
, in
|
||
<collectionCode id="ED5EAE23FF96FF9EFE1E4EDA635EFA54" box="[474,559,1428,1454]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="12" pageNumber="383">NMNS</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF036E6FF96FF9EFF034EF964F0F998" blockId="12.[151,1437,1211,2030]" pageId="12" pageNumber="383">
|
||
<emphasis id="B93BEAF4FF96FF9EFF034EF96358FA28" bold="true" box="[199,553,1463,1490]" pageId="12" pageNumber="383">Descriptive notes on the male.</emphasis>
|
||
<bibRefCitation id="EFDE4B17FF96FF9EFDF44EF66240FA28" author="Laidlaw, F. F." box="[560,817,1463,1490]" pageId="12" pageNumber="383" pagination="25 - 39" refId="ref9753" refString="Laidlaw, F. F. (1915) Contributions to a study of the dragonfly fauna of Borneo. Part III. A collection made on Mount Kina Balu by Mr J. C. Moulton in September and October 1913. Proceedings of the Zoological Society of London, 1915, 25 - 39. https: // doi. org / 10.1111 / j. 1469 - 7998.1915.00025. x" type="book chapter" year="1915">Laidlaw (1915: 37–38)</bibRefCitation>
|
||
gave a fairly complete description of the
|
||
<typeStatus id="54F48844FF96FF9EFB394EF6642EFA28" box="[1277,1375,1464,1490]" pageId="12" pageNumber="383" type="holotype">holotype</typeStatus>
|
||
male and an essentially accurate sketch of the anal appendages in lateral view. Here we merely expand on some details useful for distinguishing
|
||
<taxonomicName id="4C4F4D65FF96FF9EFE724D4E631CF9E0" authorityName="Laidlaw" authorityYear="1915" box="[438,621,1536,1562]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="kinabaluensis">
|
||
<emphasis id="B93BEAF4FF96FF9EFE724D4E631CF9E0" box="[438,621,1536,1562]" italics="true" pageId="12" pageNumber="383">P. kinabaluensis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C4F4D65FF96FF9EFD714D4E6257F9E3" authorityName="Dow, Phan & Choong" authorityYear="2020" box="[693,806,1536,1561]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="joepani">
|
||
<emphasis id="B93BEAF4FF96FF9EFD714D4E6257F9E3" box="[693,806,1536,1561]" italics="true" pageId="12" pageNumber="383">P. joepani</emphasis>
|
||
</taxonomicName>
|
||
or for which Laidlaw’s description is lacking. Laidlaw only described the dorsum of the head and “a pale band on the anteclypeus … the third joint of the antennae which is light yellowish brown”. In fact, the anteclypeus, and much of the labrum and mandible bases are pale (
|
||
<figureCitation id="13742A63FF96FF9EFAEB4D066402F998" box="[1327,1395,1608,1634]" captionStart="FIGURES 2–5" captionStartId="4.[151,264,1331,1356]" captionTargetBox="[176,1411,345,1306]" captionTargetId="figure@4.[176,1411,345,1306]" captionTargetPageId="4" captionText="FIGURES 2–5. Heads of Protosticta species. Protosticta joepani: (2) holotype male, dorsal-frontal view; (4) paratype female used for description, dorsal-frontal view. Protosticta kinabaluensis: (3) male (SAB5_PST3), dorsal view; (5) female (SAB12_ PST69), dorsal frontal view." figureDoi="http://doi.org/10.5281/zenodo.3632805" httpUri="https://zenodo.org/record/3632805/files/figure.png" pageId="12" pageNumber="383">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF036E6FF96FF9EFF034D226530F8C0" blockId="12.[151,1437,1211,2030]" pageId="12" pageNumber="383">
|
||
Laidlaw’s description of the sides of the synthorax is vague, the reader is referred to
|
||
<figureCitation id="13742A63FF96FF9EFB424D2265A7F97C" box="[1158,1238,1643,1670]" captionStart="FIGURES 15–16" captionStartId="8.[151,264,706,731]" captionTargetBox="[151,1435,181,681]" captionTargetId="figure@8.[151,1435,181,681]" captionTargetPageId="8" captionText="FIGURES 15–16. Synthorax of Protosticta kinabaluensis in lateral view: (15) male (SAB12_PST71); (16) female (SAB12_ PST69)." figureDoi="http://doi.org/10.5281/zenodo.3632813" httpUri="https://zenodo.org/record/3632813/files/figure.png" pageId="12" pageNumber="383">Fig. 15</figureCitation>
|
||
here. The yellow bands on the metepisternum and of the metepimeron are both brighter and better defined than the corresponding markings in
|
||
<taxonomicName id="4C4F4D65FF96FF9EFEE64DFA60E0F937" authorityName="Dow, Phan & Choong" authorityYear="2020" box="[290,401,1716,1741]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="joepani">
|
||
<emphasis id="B93BEAF4FF96FF9EFEE64DFA60E0F937" box="[290,401,1716,1741]" italics="true" pageId="12" pageNumber="383">P. joepani</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13742A63FF96FF9EFE644DFA6350F934" box="[416,545,1716,1742]" captionStart="FIGURES 12–14" captionStartId="7.[151,264,1978,2003]" captionTargetBox="[480,1108,489,1953]" captionTargetId="figure@7.[480,1108,489,1953]" captionTargetPageId="7" captionText="FIGURES 12–14. Synthorax of Protosticta joepani in lateral view: (12) holotype male; (13) paratype female used for description; (14) dark male with purple crescentic area on mesepimeron, Mount Kinabalu, Sabah." figureDoi="http://doi.org/10.5281/zenodo.3632811" httpUri="https://zenodo.org/record/3632811/files/figure.png" pageId="12" pageNumber="383">Figs 12, 14</figureCitation>
|
||
). The “apical triangular mark on the ventral side” of abdominal segments 3–8 is basal in the convention employed here (where the end of the segment nearest to the thorax is considered basal). The mark on S8 is visible in
|
||
<figureCitation id="13742A63FF96FF9EFE6E4DB2608BF8EC" box="[426,506,1788,1814]" captionStart="FIGURES 19–26" captionStartId="9.[151,264,1870,1895]" captionTargetBox="[151,1436,417,1845]" captionTargetId="figure@9.[151,1436,417,1845]" captionTargetPageId="9" captionText="FIGURES 19–26. Terminal abdominal segments of Protosticta species. Protosticta joepani: (19) holotype male, dorsal; (21) holotype male, lateral; (23) paratype female used for description, dorsal; (25) paratype female used for description, lateral. Protosticta kinabaluensis: (20) male (SAB12_PST71) dorsal; (22) male (SAB12_PST71) lateral; (24) female (SAB12_PST69), dorsal; (26) female (SAB12_PST69), lateral." figureDoi="http://doi.org/10.5281/zenodo.3632817" httpUri="https://zenodo.org/record/3632817/files/figure.png" pageId="12" pageNumber="383">Fig. 22</figureCitation>
|
||
but on this segment it extends almost the entire length of the segment. The genital ligula is similar to that of
|
||
<taxonomicName id="4C4F4D65FF96FF9EFE7C4C6E6359F8C3" authorityName="Dow, Phan & Choong" authorityYear="2020" box="[440,552,1824,1849]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="joepani">
|
||
<emphasis id="B93BEAF4FF96FF9EFE7C4C6E6359F8C3" box="[440,552,1824,1849]" italics="true" pageId="12" pageNumber="383">P. joepani</emphasis>
|
||
</taxonomicName>
|
||
but with a much broader internal fold (
|
||
<figureCitation id="13742A63FF96FF9EFC254C6E6545F8C0" box="[993,1076,1824,1850]" captionStart="FIGURES 17–18" captionStartId="8.[151,264,1089,1114]" captionTargetBox="[153,1428,794,1059]" captionTargetId="figure@8.[151,1436,789,1063]" captionTargetPageId="8" captionText="FIGURES 17–18. Genital ligula of Protosticta species in ventral view. (17) Protosticta joepani paratype, Mount Kinabalu, 4 iii 2008. (18) Protosticta kinabaluensis, Mount Kinabalu, 1 v 1979." figureDoi="http://doi.org/10.5281/zenodo.3632815" httpUri="https://zenodo.org/record/3632815/files/figure.png" pageId="12" pageNumber="383">Fig. 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF036E6FF96FF9FFF034C0A650BFF2D" blockId="12.[151,1437,1211,2030]" lastBlockId="13.[151,1437,153,251]" lastPageId="13" lastPageNumber="384" pageId="12" pageNumber="383">
|
||
The anal appendages (
|
||
<figureCitation id="13742A63FF96FF9EFE7B4C0A634EF8A4" box="[447,575,1860,1886]" captionStart="FIGURES 27–30" captionStartId="10.[151,264,1207,1232]" captionTargetBox="[151,1437,181,1181]" captionTargetId="figure@10.[151,1437,181,1181]" captionTargetPageId="10" captionText="FIGURES 27–30. Male anal appendages of Protosticta species. All scale bars 0.2mm. Protosticta joepani: (27) holotype male, lateral; (29) holotype male, paraprocts in ventral view. Protosticta kinabaluensis: (28) male (SAB12_PST71) lateral view; (30) male (SAB12_PST71) paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3632819" httpUri="https://zenodo.org/record/3632819/files/figure.png" pageId="12" pageNumber="383">Figs 28, 30</figureCitation>
|
||
,
|
||
<figureCitation id="13742A63FF96FF9EFD8F4C0A63E0F8A4" box="[587,657,1860,1886]" captionStart="FIGURES 31–34" captionStartId="11.[151,264,1943,1968]" captionTargetBox="[151,1436,597,1918]" captionTargetId="figure@11.[151,1436,597,1918]" captionTargetPageId="11" captionText="FIGURES 31–34. Male anal appendages of Protosticta species, dorsal view. Protosticta joepani: (31) holotype male; (33) sketch of paratype, Poring Hot Springs, 4 iii 2008, right cercus removed. Protosticta kinabaluensis: (32) sketch of male, 1 v 1979; (34) male (SAB12_PST71), right cercus removed." figureDoi="http://doi.org/10.5281/zenodo.3632821" httpUri="https://zenodo.org/record/3632821/files/figure.png" pageId="12" pageNumber="383">32, 34</figureCitation>
|
||
) are similar to those of
|
||
<taxonomicName id="4C4F4D65FF96FF9EFC5D4C0A6579F8A7" authorityName="Dow, Phan & Choong" authorityYear="2020" box="[921,1032,1860,1885]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="joepani">
|
||
<emphasis id="B93BEAF4FF96FF9EFC5D4C0A6579F8A7" box="[921,1032,1860,1885]" italics="true" pageId="12" pageNumber="383">P. joepani</emphasis>
|
||
</taxonomicName>
|
||
and the following notes mostly deal with differences from that species. The cerci are lacking a rounded ventral spur basally on the blade-like section, which originates further apically than in
|
||
<taxonomicName id="4C4F4D65FF96FF9EFDAE4CC263ADF85F" authorityName="Dow, Phan & Choong" authorityYear="2020" box="[618,732,1932,1957]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="joepani">
|
||
<emphasis id="B93BEAF4FF96FF9EFDAE4CC263ADF85F" box="[618,732,1932,1957]" italics="true" pageId="12" pageNumber="383">P. joepani</emphasis>
|
||
</taxonomicName>
|
||
, the dorsal spine is situated after half-length and the rounded subapical projection is situated further apically (compare
|
||
<figureCitation id="13742A63FF96FF9EFCE74CFE6202F830" box="[803,883,1968,1994]" captionStart="FIGURES 27–30" captionStartId="10.[151,264,1207,1232]" captionTargetBox="[151,1437,181,1181]" captionTargetId="figure@10.[151,1437,181,1181]" captionTargetPageId="10" captionText="FIGURES 27–30. Male anal appendages of Protosticta species. All scale bars 0.2mm. Protosticta joepani: (27) holotype male, lateral; (29) holotype male, paraprocts in ventral view. Protosticta kinabaluensis: (28) male (SAB12_PST71) lateral view; (30) male (SAB12_PST71) paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3632819" httpUri="https://zenodo.org/record/3632819/files/figure.png" pageId="12" pageNumber="383">Fig. 27</figureCitation>
|
||
and
|
||
<figureCitation id="13742A63FF96FF9EFC6F4CFE6571F830" box="[939,1024,1968,1994]" captionStart="FIGURES 27–30" captionStartId="10.[151,264,1207,1232]" captionTargetBox="[151,1437,181,1181]" captionTargetId="figure@10.[151,1437,181,1181]" captionTargetPageId="10" captionText="FIGURES 27–30. Male anal appendages of Protosticta species. All scale bars 0.2mm. Protosticta joepani: (27) holotype male, lateral; (29) holotype male, paraprocts in ventral view. Protosticta kinabaluensis: (28) male (SAB12_PST71) lateral view; (30) male (SAB12_PST71) paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3632819" httpUri="https://zenodo.org/record/3632819/files/figure.png" pageId="12" pageNumber="383">Fig. 28</figureCitation>
|
||
). The paraprocts have a strong basal spur clearly visible in lateral view (
|
||
<figureCitation id="13742A63FF96FF9EFDD84C9A6301F814" box="[540,624,2004,2030]" captionStart="FIGURES 27–30" captionStartId="10.[151,264,1207,1232]" captionTargetBox="[151,1437,181,1181]" captionTargetId="figure@10.[151,1437,181,1181]" captionTargetPageId="10" captionText="FIGURES 27–30. Male anal appendages of Protosticta species. All scale bars 0.2mm. Protosticta joepani: (27) holotype male, lateral; (29) holotype male, paraprocts in ventral view. Protosticta kinabaluensis: (28) male (SAB12_PST71) lateral view; (30) male (SAB12_PST71) paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3632819" httpUri="https://zenodo.org/record/3632819/files/figure.png" pageId="12" pageNumber="383">Fig. 28</figureCitation>
|
||
). The internal spur originates only slightly subapically and the main body of each paraproct is expanded dorsally at this point, the spur is more expanded apically in lateral view and longer than that in
|
||
<taxonomicName id="4C4F4D65FF97FF9FFED84BF060FDFF2D" authorityName="Dow, Phan & Choong" authorityYear="2020" box="[284,396,190,215]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="384" phylum="Arthropoda" rank="species" species="joepani">
|
||
<emphasis id="B93BEAF4FF97FF9FFED84BF060FDFF2D" box="[284,396,190,215]" italics="true" pageId="13" pageNumber="384">P. joepani</emphasis>
|
||
</taxonomicName>
|
||
, and does not turn through a right angle (compare
|
||
<figureCitation id="13742A63FF97FF9FFC004BF3651FFF2D" box="[964,1134,189,215]" captionStart="FIGURES 27–30" captionStartId="10.[151,264,1207,1232]" captionTargetBox="[151,1437,181,1181]" captionTargetId="figure@10.[151,1437,181,1181]" captionTargetPageId="10" captionText="FIGURES 27–30. Male anal appendages of Protosticta species. All scale bars 0.2mm. Protosticta joepani: (27) holotype male, lateral; (29) holotype male, paraprocts in ventral view. Protosticta kinabaluensis: (28) male (SAB12_PST71) lateral view; (30) male (SAB12_PST71) paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3632819" httpUri="https://zenodo.org/record/3632819/files/figure.png" pageId="13" pageNumber="384">Figs 29 and 30</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF036E6FF97FF9FFF034BAF655DFF01" blockId="13.[151,1437,153,251]" box="[199,1068,225,251]" pageId="13" pageNumber="384">Measurements (mm). Hw 21–22, abdomen including anal appendages 33–36.</paragraph>
|
||
<caption id="DF30666EFF97FF9FFF534D39650DF96A" ID-DOI="http://doi.org/10.5281/zenodo.3632823" ID-Zenodo-Dep="3632823" box="[151,1148,1655,1680]" httpUri="https://zenodo.org/record/3632823/files/figure.png" pageId="13" pageNumber="384" startId="13.[151,250,1655,1680]" targetBox="[151,1436,274,1646]" targetPageId="13">
|
||
<paragraph id="8BF036E6FF97FF9FFF534D39650DF96A" blockId="13.[151,1148,1655,1680]" box="[151,1148,1655,1680]" pageId="13" pageNumber="384">
|
||
<emphasis id="B93BEAF4FF97FF9FFF534D396051F96A" bold="true" box="[151,288,1655,1680]" pageId="13" pageNumber="384">FIGURE 35.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4C4F4D65FF97FF9FFE064D3763F5F96A" authorityName="Dow, Phan & Choong" authorityYear="2020" box="[450,644,1657,1680]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="384" phylum="Arthropoda" rank="species" species="joepani">
|
||
<emphasis id="B93BEAF4FF97FF9FFE064D3763F5F96A" box="[450,644,1657,1680]" italics="true" pageId="13" pageNumber="384">Protosticta joepani</emphasis>
|
||
</taxonomicName>
|
||
(red circle) and
|
||
<taxonomicName id="4C4F4D65FF97FF9FFCE84D3762BEF96A" authorityName="Laidlaw" authorityYear="1915" box="[812,975,1656,1680]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="384" phylum="Arthropoda" rank="species" species="kinabaluensis">
|
||
<emphasis id="B93BEAF4FF97FF9FFCE84D3762BEF96A" box="[812,975,1656,1680]" italics="true" pageId="13" pageNumber="384">P. kinabaluensis</emphasis>
|
||
</taxonomicName>
|
||
(yellow square).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF036E6FF97FF9FFF034DF0627BF84B" blockId="13.[151,1437,1726,2041]" pageId="13" pageNumber="384">
|
||
<emphasis id="B93BEAF4FF97FF9FFF034DF0634EF923" bold="true" box="[199,575,1726,1753]" pageId="13" pageNumber="384">Descriptive notes on the female.</emphasis>
|
||
The female of
|
||
<taxonomicName id="4C4F4D65FF97FF9FFD224DF162EAF922" authorityName="Laidlaw" authorityYear="1915" box="[742,923,1726,1752]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="384" phylum="Arthropoda" rank="species" species="kinabaluensis">
|
||
<emphasis id="B93BEAF4FF97FF9FFD224DF162EAF922" box="[742,923,1726,1752]" italics="true" pageId="13" pageNumber="384">P. kinabaluensis</emphasis>
|
||
</taxonomicName>
|
||
has never been described, but it is sufficiently similar to the male that the illustrations and brief notes given here should suffice for identification. The female is generally similar to the male and the markings of head and thorax (
|
||
<figureCitation id="13742A63FF97FF9FFCBA4C4862B7F8DB" box="[894,966,1798,1825]" captionStart="FIGURES 2–5" captionStartId="4.[151,264,1331,1356]" captionTargetBox="[176,1411,345,1306]" captionTargetId="figure@4.[176,1411,345,1306]" captionTargetPageId="4" captionText="FIGURES 2–5. Heads of Protosticta species. Protosticta joepani: (2) holotype male, dorsal-frontal view; (4) paratype female used for description, dorsal-frontal view. Protosticta kinabaluensis: (3) male (SAB5_PST3), dorsal view; (5) female (SAB12_ PST69), dorsal frontal view." figureDoi="http://doi.org/10.5281/zenodo.3632805" httpUri="https://zenodo.org/record/3632805/files/figure.png" pageId="13" pageNumber="384">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="13742A63FF97FF9FFC164C49657BF8DB" box="[978,1034,1799,1825]" captionStart="FIGURES 8–11" captionStartId="6.[151,264,1634,1659]" captionTargetBox="[151,1436,181,1609]" captionTargetId="figure@6.[151,1436,181,1609]" captionTargetPageId="6" captionText="FIGURES 8–11. Prothorax of females of Protosticta species. Protosticta joepani, paratype female used for description: (8) dorsal; (10) lateral. Protosticta kinabaluensis (SAB12_PST69): (9) dorsal; (11) lateral." figureDoi="http://doi.org/10.5281/zenodo.3632809" httpUri="https://zenodo.org/record/3632809/files/figure.png" pageId="13" pageNumber="384">9, 11</figureCitation>
|
||
,
|
||
<figureCitation id="13742A63FF97FF9FFBD14C486543F8DB" box="[1045,1074,1798,1825]" captionStart="FIGURES 15–16" captionStartId="8.[151,264,706,731]" captionTargetBox="[151,1435,181,681]" captionTargetId="figure@8.[151,1435,181,681]" captionTargetPageId="8" captionText="FIGURES 15–16. Synthorax of Protosticta kinabaluensis in lateral view: (15) male (SAB12_PST71); (16) female (SAB12_ PST69)." figureDoi="http://doi.org/10.5281/zenodo.3632813" httpUri="https://zenodo.org/record/3632813/files/figure.png" pageId="13" pageNumber="384">16</figureCitation>
|
||
) are essentially the same in both sexes. The posterior pronotal lobe is simple (
|
||
<figureCitation id="13742A63FF97FF9FFD484C64638CF8BF" box="[652,765,1834,1861]" captionStart="FIGURES 8–11" captionStartId="6.[151,264,1634,1659]" captionTargetBox="[151,1436,181,1609]" captionTargetId="figure@6.[151,1436,181,1609]" captionTargetPageId="6" captionText="FIGURES 8–11. Prothorax of females of Protosticta species. Protosticta joepani, paratype female used for description: (8) dorsal; (10) lateral. Protosticta kinabaluensis (SAB12_PST69): (9) dorsal; (11) lateral." figureDoi="http://doi.org/10.5281/zenodo.3632809" httpUri="https://zenodo.org/record/3632809/files/figure.png" pageId="13" pageNumber="384">Figs 9, 11</figureCitation>
|
||
). The markings of the abdomen are also similar to the male, even those of the terminal segments (
|
||
<figureCitation id="13742A63FF97FF9FFDF34C0063C8F893" box="[567,697,1870,1897]" captionStart="FIGURES 19–26" captionStartId="9.[151,264,1870,1895]" captionTargetBox="[151,1436,417,1845]" captionTargetId="figure@9.[151,1436,417,1845]" captionTargetPageId="9" captionText="FIGURES 19–26. Terminal abdominal segments of Protosticta species. Protosticta joepani: (19) holotype male, dorsal; (21) holotype male, lateral; (23) paratype female used for description, dorsal; (25) paratype female used for description, lateral. Protosticta kinabaluensis: (20) male (SAB12_PST71) dorsal; (22) male (SAB12_PST71) lateral; (24) female (SAB12_PST69), dorsal; (26) female (SAB12_PST69), lateral." figureDoi="http://doi.org/10.5281/zenodo.3632817" httpUri="https://zenodo.org/record/3632817/files/figure.png" pageId="13" pageNumber="384">Figs 24, 26</figureCitation>
|
||
), however S7 is largely brown dorsally. The apex of the terebra of the ovipositor is visible outside the valves of S9, well developed, albeit less so than in
|
||
<taxonomicName id="4C4F4D65FF97FF9FFBA54C3D65A0F876" authorityName="Dow, Phan & Choong" authorityYear="2020" box="[1121,1233,1907,1932]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="384" phylum="Arthropoda" rank="species" species="joepani">
|
||
<emphasis id="B93BEAF4FF97FF9FFBA54C3D65A0F876" box="[1121,1233,1907,1932]" italics="true" pageId="13" pageNumber="384">P. joepani</emphasis>
|
||
</taxonomicName>
|
||
(compare
|
||
<figureCitation id="13742A63FF97FF9FFA8D4C3C6197F84B" captionStart="FIGURES 19–26" captionStartId="9.[151,264,1870,1895]" captionTargetBox="[151,1436,417,1845]" captionTargetId="figure@9.[151,1436,417,1845]" captionTargetPageId="9" captionText="FIGURES 19–26. Terminal abdominal segments of Protosticta species. Protosticta joepani: (19) holotype male, dorsal; (21) holotype male, lateral; (23) paratype female used for description, dorsal; (25) paratype female used for description, lateral. Protosticta kinabaluensis: (20) male (SAB12_PST71) dorsal; (22) male (SAB12_PST71) lateral; (24) female (SAB12_PST69), dorsal; (26) female (SAB12_PST69), lateral." figureDoi="http://doi.org/10.5281/zenodo.3632817" httpUri="https://zenodo.org/record/3632817/files/figure.png" pageId="13" pageNumber="384">Figs 25 and 26</figureCitation>
|
||
), blade-shaped, directed upwards and to the rear.
|
||
</paragraph>
|
||
<paragraph id="8BF036E6FF97FF9FFF034CF56285F802" blockId="13.[151,1437,1726,2041]" pageId="13" pageNumber="384">
|
||
<bibRefCitation id="EFDE4B17FF97FF9FFF034CF5601EF82F" author="Orr, A. G." box="[199,367,1979,2005]" pageId="13" pageNumber="384" refId="ref10453" refString="Orr, A. G. (2003) A guide to the dragonflies of Borneo: their identification and biology. Natural History Publications (Borneo), Kota Kinabalu, x + 195 pp." type="book" year="2003">Orr (2003: 72)</bibRefCitation>
|
||
notes that “In females there may be faint antehumeral bands”, this might be a relatively rare condition since it is not the case in the three females examined for this study.
|
||
</paragraph>
|
||
<paragraph id="8BF036E6FF94FF9CFF034BD765F1FF49" blockId="14.[151,1436,153,359]" box="[199,1152,153,179]" pageId="14" pageNumber="385">Measurements (mm). Hw 30, abdomen including anal appendages and ovipositor 30.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C355656DFF94FF9CFF034BF3624BFE9D" pageId="14" pageNumber="385" type="discussion">
|
||
<paragraph id="8BF036E6FF94FF9CFF034BF36425FF01" blockId="14.[151,1436,153,359]" pageId="14" pageNumber="385">
|
||
<emphasis id="B93BEAF4FF94FF9CFF034BF3604AFF2D" bold="true" box="[199,315,189,215]" pageId="14" pageNumber="385">Remarks.</emphasis>
|
||
Although not explicitly examined during this study, the
|
||
<typeStatus id="54F48844FF94FF9CFC6A4BF36561FF2D" box="[942,1040,189,215]" pageId="14" pageNumber="385" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4F4D65FF94FF9CFBF04BF06598FF2D" authorityName="Laidlaw" authorityYear="1915" box="[1076,1257,189,215]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="385" phylum="Arthropoda" rank="species" species="kinabaluensis">
|
||
<emphasis id="B93BEAF4FF94FF9CFBF04BF06598FF2D" box="[1076,1257,189,215]" italics="true" pageId="14" pageNumber="385">P. kinabaluensis</emphasis>
|
||
</taxonomicName>
|
||
has been examined previously by the first author. The recently collected material listed above is certainly the same species.
|
||
</paragraph>
|
||
<paragraph id="8BF036E6FF94FF9CFF034A4B624BFE9D" blockId="14.[151,1436,153,359]" pageId="14" pageNumber="385">
|
||
<bibRefCitation id="EFDE4B17FF94FF9CFF034A4B60CDFEDA" author="Donnelly, N." box="[199,444,261,288]" pageId="14" pageNumber="385" pagination="123 - 128" refId="ref9203" refString="Donnelly, N. [T. W.] (1997) Through darkest Borneo [and Malaysia] with net and camera. Malangpo, 14, 123 - 128." type="journal article" year="1997">Donnelly (1997: 126)</bibRefCitation>
|
||
recorded females of a
|
||
<taxonomicName id="4C4F4D65FF94FF9CFD074A48624FFEE5" authorityName="Selys" authorityYear="1885" box="[707,830,262,287]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="385" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93BEAF4FF94FF9CFD074A48624FFEE5" box="[707,830,262,287]" italics="true" pageId="14" pageNumber="385">Protosticta</emphasis>
|
||
</taxonomicName>
|
||
sp. from Mount Kinabalu, presumably from the park headquarters area. It is likely that this record refers to
|
||
<taxonomicName id="4C4F4D65FF94FF9CFCC64A6462C9FEB9" authorityName="Laidlaw" authorityYear="1915" box="[770,952,297,323]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="385" phylum="Arthropoda" rank="species" species="kinabaluensis">
|
||
<emphasis id="B93BEAF4FF94FF9CFCC64A6462C9FEB9" box="[770,952,297,323]" italics="true" pageId="14" pageNumber="385">P. kinabaluensis</emphasis>
|
||
</taxonomicName>
|
||
but it could possibly refer to
|
||
<taxonomicName id="4C4F4D65FF94FF9CFAC84A64640CFEB9" authorityName="Dow, Phan & Choong" authorityYear="2020" box="[1292,1405,298,323]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="385" phylum="Arthropoda" rank="species" species="joepani">
|
||
<emphasis id="B93BEAF4FF94FF9CFAC84A64640CFEB9" box="[1292,1405,298,323]" italics="true" pageId="14" pageNumber="385">P. joepani</emphasis>
|
||
</taxonomicName>
|
||
or some other species (for instance some species of
|
||
<taxonomicName id="4C4F4D65FF94FF9CFD7A4A03625BFE9D" authorityName="Dow & Orr" authorityYear="2012" box="[702,810,333,359]" class="Insecta" family="Platystictidae" genus="Telosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="385" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93BEAF4FF94FF9CFD7A4A03625BFE9D" box="[702,810,333,359]" italics="true" pageId="14" pageNumber="385">Telosticta</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |