329 lines
49 KiB
XML
329 lines
49 KiB
XML
<document id="B99461CC57A326CEA8117094FAB2C8D6" ID-DOI="10.5281/zenodo.278442" ID-GBIF-Dataset="6b9505e0-ac6e-4baa-9066-6f729d38e9e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="278442" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460341493798" checkinUser="plazi" docAuthor="Hershler, Robert, Liu, Hsiu-Ping & Landye, Jerry" docDate="2011" docId="03E687C5BF78831FFF2E1576FBE51B73" docLanguage="en" docName="zt03001p032.pdf" docOrigin="Zootaxa 3001" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tryonia contrerasi Hershler, Liu & Landye, 2011, new species" docType="treatment" docVersion="8" lastPageNumber="18" masterDocId="FFDFFFBDBF77830EFFB9171AFFA4194F" masterDocTitle="New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot" masterLastPageNumber="32" masterPageNumber="1" pageNumber="16" updateTime="1698261240372" updateUser="plazi">
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<mods:title id="88A9CBF4E53EF6481B66F764682A0D48">New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot</mods:title>
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<mods:namePart id="D14C43660F95132C72F034F2EDBA4D34">Hershler, Robert</mods:namePart>
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<mods:namePart id="6EB9850B7F4F176296C34192A670DABF">Liu, Hsiu-Ping</mods:namePart>
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<mods:namePart id="AF1401292DD0160B8F400D03D3D474B3">Landye, Jerry</mods:namePart>
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<mods:title id="2412680914FD114C450A244CBF83B45A">Zootaxa</mods:title>
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<mods:part id="E7125AF0B778C30F4749FDDD9EA8CAE7">
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<mods:date id="D28AF55D33C728D1684DB775B54F6154">2011</mods:date>
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<mods:identifier id="D53F7A1EAB3D99D836D49AF876182995" type="DOI">10.5281/zenodo.278442</mods:identifier>
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<mods:identifier id="518C39F56A8B2659622A425B0344542A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E687C5BF78831FFF2E1576FBE51B73" ID-DOI="http://doi.org/10.5281/zenodo.5629201" ID-GBIF-Taxon="119406527" ID-Zenodo-Dep="5629201" LSID="urn:lsid:plazi:treatment:03E687C5BF78831FFF2E1576FBE51B73" httpUri="http://treatment.plazi.org/id/03E687C5BF78831FFF2E1576FBE51B73" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
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<subSubSection id="C3556558BF788301FF2E1576FB981A5D" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph id="8BF036D3BF788301FF2E1576FDAA1BCA" blockId="15.[151,526,619,678]" box="[151,526,619,646]" pageId="15" pageNumber="16">
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<heading id="D0B881BFBF788301FF2E1576FDAA1BCA" bold="true" box="[151,526,619,646]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
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<emphasis id="B93BEAC1BF788301FF2E1576FDAA1BCA" bold="true" box="[151,526,619,646]" pageId="15" pageNumber="16">
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<taxonomicName id="4C4F4D50BF788301FF2E1576FED31BC9" box="[151,375,620,646]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="contrerasi" status="sp. nov.">
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<emphasis id="B93BEAC1BF788301FF2E1576FED31BC9" bold="true" box="[151,375,620,646]" italics="true" pageId="15" pageNumber="16">Tryonia contrerasi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20857BABF788301FEC71571FDAA1BCA" box="[382,526,619,645]" pageId="15" pageNumber="16" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF036D3BF788301FF2E1594FE1B1BE9" blockId="15.[151,526,619,678]" box="[151,447,654,678]" pageId="15" pageNumber="16">
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(
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<figureCitation id="13742A56BF788301FF261594FF4F1BE9" box="[159,235,654,678]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="15" pageNumber="16">Figs. 5</figureCitation>
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F, 8D–F, I–J, N–P)
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</paragraph>
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<paragraph id="8BF036D3BF788301FF2E15CDFB981A5D" blockId="15.[151,1436,726,1974]" pageId="15" pageNumber="16">
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<emphasis id="B93BEAC1BF788301FF2E15CDFF7E1BA0" bold="true" box="[151,218,727,751]" pageId="15" pageNumber="16">
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<typeStatus id="54F48871BF788301FF2E15CDFF7E1BA0" box="[151,218,727,751]" pageId="15" pageNumber="16">Types</typeStatus>
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</emphasis>
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.
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<typeStatus id="54F48871BF788301FF5515CCFEF31BA1" box="[236,343,726,750]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
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, USNM 874121, Ojo del Apache, south of Rancho Nuevo, Chihuahua,
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<geoCoordinate id="EE7B5014BF788301FB3115CCFAB01BA1" box="[1160,1300,726,750]" direction="north" orientation="latitude" pageId="15" pageNumber="16" precision="15" value="30.574167">30º34'27" N</geoCoordinate>
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,
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<geoCoordinate id="EE7B5014BF788301FA9815CCFF161A5D" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="15" value="-106.9075">106º54'27" W</geoCoordinate>
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, leg. J.J.L. and D. Wong,
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<date id="FFF11013BF788301FE6B15E0FDE51A5D" box="[466,577,762,786]" pageId="15" pageNumber="16" value="1991-04-02">2/iv/1991</date>
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.
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<typeStatus id="54F48871BF788301FDF215E1FD1C1A5C" box="[587,696,763,787]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
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(from same lot), USNM 1153670.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF788301FF7E1405FD4E1A31" pageId="15" pageNumber="16" type="etymology">
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<paragraph id="8BF036D3BF788301FF7E1405FD4E1A31" blockId="15.[151,1436,726,1974]" pageId="15" pageNumber="16">
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<emphasis id="B93BEAC1BF788301FF7E1405FEE21A78" bold="true" box="[199,326,799,823]" pageId="15" pageNumber="16">Etymology</emphasis>
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. A patronym honoring
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<collectingCountry id="F3587643BF788301FDEA1404FD121A79" box="[595,694,798,822]" name="El Salvador" pageId="15" pageNumber="16">Salvador</collectingCountry>
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Contreras-Balderas (Professor Emeritus, Department of Biology, Universidad Autonoma de Nuevo Leon, Monterrey,
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<collectingCountry id="F3587643BF788301FD5C1458FC981A15" box="[741,828,834,858]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
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) for his dedicated efforts over many years to promote the conservation of Chihuahuan Desert aquatic biota.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF788301FF7E1490FC6D1D41" pageId="15" pageNumber="16" type="materials_examined">
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<paragraph id="8BF036D3BF788301FF7E1490FC6D1D41" blockId="15.[151,1436,726,1974]" pageId="15" pageNumber="16">
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<emphasis id="B93BEAC1BF788301FF7E1490FE3A1AEC" bold="true" box="[199,414,906,931]" pageId="15" pageNumber="16">Referred material</emphasis>
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. CHIHUAHUA.
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<collectionCode id="ED5EAE16BF788301FDDC1490FD121AED" LSID="urn:lsid:biocol.org:col:34871" box="[613,694,906,930]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="16">USNM</collectionCode>
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854951, topotypes, R.H. and J.J.L.,
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<date id="FFF11013BF788301FBE81490FB721AED" box="[1105,1238,906,930]" pageId="15" pageNumber="16" value="1998-12-14">14/xii/1998</date>
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.
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<collectionCode id="ED5EAE16BF788301FB581491FA831AED" box="[1249,1319,907,930]" pageId="15" pageNumber="16">UTEP</collectionCode>
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881, ca.
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<quantity id="4CB79B36BF788301FA371490FF1F1A89" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="km" value="4.0">4 km</quantity>
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southwest of Hacienda Santo
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<collectingCountry id="F3587643BF788301FD9E14B4FD301A89" box="[551,660,942,966]" name="Dominican Republic" pageId="15" pageNumber="16">Domingo</collectingCountry>
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, leg. W. Strain, no date.
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||
<collectionCode id="ED5EAE16BF788301FC7A14B4FBB01A89" LSID="urn:lsid:biocol.org:col:34871" box="[963,1044,942,966]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="16">USNM</collectionCode>
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854124,
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<collectionCode id="ED5EAE16BF788301FB3E14B4FB7C1A89" LSID="urn:lsid:biocol.org:col:34871" box="[1159,1240,942,966]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="16">USNM</collectionCode>
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874124,
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<collectionCode id="ED5EAE16BF788301FAF214B4FA381A89" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,942,966]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="16">USNM</collectionCode>
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874128, Ojo Carbonero, southeast of Rancho Nuevo, Chihuahua,
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<geoCoordinate id="EE7B5014BF788301FC3B14C8FB801AA5" box="[898,1060,978,1002]" direction="north" orientation="latitude" pageId="15" pageNumber="16" precision="1" value="30.579863">30º34'47.5" N</geoCoordinate>
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,
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<geoCoordinate id="EE7B5014BF788301FB8914C8FB6A1AA5" box="[1072,1230,978,1002]" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="15" value="-106.87389">106º52'26" W</geoCoordinate>
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, leg. J.J.L. and D. Wong,
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<date id="FFF11013BF788301FF5E14ECFEF21D41" box="[231,342,1014,1038]" pageId="15" pageNumber="16" value="1991-04-02">2/iv/1991</date>
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.
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<collectionCode id="ED5EAE16BF788301FED914ECFE151D41" LSID="urn:lsid:biocol.org:col:34871" box="[352,433,1014,1038]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="16">USNM</collectionCode>
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854952,
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<emphasis id="B93BEAC1BF788301FDA314ECFDE91D41" box="[538,589,1014,1038]" italics="true" pageId="15" pageNumber="16">ibid.</emphasis>
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, leg. R.H. and J.J.L.,
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<date id="FFF11013BF788301FCF914ECFC611D41" box="[832,965,1014,1038]" pageId="15" pageNumber="16" value="1998-12-14">14/xii/1998</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF788301FF7E1300FA9B1D8D" pageId="15" pageNumber="16" type="diagnosis">
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<paragraph id="8BF036D3BF788301FF7E1300FA9B1D8D" blockId="15.[151,1436,726,1974]" pageId="15" pageNumber="16">
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<emphasis id="B93BEAC1BF788301FF7E1300FE9E1D7C" bold="true" box="[199,314,1050,1075]" pageId="15" pageNumber="16">Diagnosis</emphasis>
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. Shell medium-to large-sized, conic, rather strongly sculptured; penis having two distal and one basal papillae on the inner edge. Distinguished from other regional congeners by its more strongly sculptured shell. Further differentiated from conchologically similar
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<taxonomicName id="4C4F4D50BF788301FD60137EFCEA1D34" box="[729,846,1123,1147]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="molinae">
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<emphasis id="B93BEAC1BF788301FD60137EFCEA1D34" box="[729,846,1123,1147]" italics="true" pageId="15" pageNumber="16">T. molinae</emphasis>
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||
</taxonomicName>
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by its larger size (t-test for shell height, t = 8.892, df = 8.892,
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<emphasis id="B93BEAC1BF788301FF451392FEA91DD0" box="[252,269,1160,1183]" italics="true" pageId="15" pageNumber="16">P</emphasis>
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<0.01) and evenly rounded distal bulb of the penis; and from
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<taxonomicName id="4C4F4D50BF788301FC7E1392FB861DD0" box="[967,1058,1160,1183]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="ovata">
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<emphasis id="B93BEAC1BF788301FC7E1392FB861DD0" box="[967,1058,1160,1183]" italics="true" pageId="15" pageNumber="16">T. ovata</emphasis>
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</taxonomicName>
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, which has closely similar penial morphology, by its less convex teleoconch whorls, typically incomplete parietal lip and smaller umbilicus.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3556558BF788301FF7E13D4FEF31EF9" pageId="15" pageNumber="16" type="description">
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<paragraph id="8BF036D3BF788301FF7E13D4FB431CD5" blockId="15.[151,1436,726,1974]" pageId="15" pageNumber="16">
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<emphasis id="B93BEAC1BF788301FF7E13D4FEF51DA8" bold="true" box="[199,337,1230,1255]" pageId="15" pageNumber="16">Description</emphasis>
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. Shell (
|
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<figureCitation id="13742A56BF788301FE0D13D4FE5F1DA9" box="[436,507,1230,1254]" captionStart="FIGURE 8" captionStartId="16.[151,244,1854,1875]" captionTargetBox="[151,1436,151,1835]" captionTargetId="figure@16.[151,1436,149,1835]" captionTargetPageId="16" captionText="FIGURE 8. Tryonia molinae sp. nov. and Tryonia contrerasi sp. nov. A. Holotype, T. molinae, USNM 1139278. B – C. Shells, T. molinae, USNM 1153674 (C is a male). D. Holotype, T. contrerasi, USNM 874121. E – F. Shells, T. contrerasi, USNM 874128 (F is a male). G – H. Operculum (outer, inner sides), T. molinae, USNM 1153674. I – J. Operculum (outer, inner sides), T. contrerasi, USNM 1153670. K. Portion of radular ribbon, T. molinae, USNM 1153674. L. Central radular teeth, T. molinae, USNM 1153670. M. Lateral and inner marginal teeth, T. molinae, USNM 1153670. N. Portion of radular ribbon, T. contrerasi, USNM 1153670. Q. Central radular teeth, T. contrerasi, USNM 1153670. R. Lateral tooth, T. contrerasi, USNM 1153670. Scale bars A – F = 1.0 mm; G – J = 250 µm; K, N = 25 µm, L – M, O – P = 10 µm." httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
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||
D–F) up to
|
||
<quantity id="4CB79B36BF788301FD2B13D4FD491DA9" box="[658,749,1230,1254]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="15" pageNumber="16" unit="mm" value="6.2">6.2 mm</quantity>
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||
tall, large females having 5.75–8.75 whorls, spire height 140–200% width of shell, male shells smaller than those of females. Teleoconch whorls weak to medium convex, evenly rounded with impressed sutures. Sculpture of widely spaced collabral riblets or cords and more numerous spiral threads or cords, sometimes resulting in a cancellate appearance. Aperture strongly angled adapically, parietal lip usually incomplete or a thin glaze across the parietal wall, rarely complete and adnate or slightly disjunct, umbilicus narrow or absent. Outer lip slightly orthocline or prosocline. Periostracum brown or tan.
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||
</paragraph>
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||
<paragraph id="8BF036D3BF788301FF7E12BCFA881F49" blockId="15.[151,1436,726,1974]" pageId="15" pageNumber="16">
|
||
<emphasis id="B93BEAC1BF788301FF7E12BCFE6D1CF0" bold="true" box="[199,457,1446,1471]" pageId="15" pageNumber="16">Shell measurements (</emphasis>
|
||
mean in parentheses): height
|
||
<quantity id="4CB79B36BF788301FCA012BCFC1A1CF1" box="[793,958,1446,1470]" metricMagnitude="-3" metricUnit="m" metricValue="5.26" metricValueMax="6.13" metricValueMin="4.39" pageId="15" pageNumber="16" unit="mm" value="5.26" valueMax="6.13" valueMin="4.39">4.39–6.13 mm</quantity>
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||
(4.92), width
|
||
<quantity id="4CB79B36BF788301FBDA12BCFAAC1CF1" box="[1123,1288,1446,1470]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.25" metricValueMin="1.85" pageId="15" pageNumber="16" unit="mm" value="2.05" valueMax="2.25" valueMin="1.85">1.85–2.25 mm</quantity>
|
||
(2.02), body whorl height
|
||
<quantity id="4CB79B36BF788301FE9412D0FE741CAD" box="[301,464,1482,1506]" metricMagnitude="-3" metricUnit="m" metricValue="2.68" metricValueMax="2.97" metricValueMin="2.39" pageId="15" pageNumber="16" unit="mm" value="2.68" valueMax="2.97" valueMin="2.39">2.39–2.97 mm</quantity>
|
||
(2.62), body whorl width
|
||
<quantity id="4CB79B36BF788301FD4012D0FC391CAD" box="[761,925,1482,1506]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.09" metricValueMin="1.71" pageId="15" pageNumber="16" unit="mm" value="1.9" valueMax="2.09" valueMin="1.71">1.71–2.09 mm</quantity>
|
||
(1.89), aperture height
|
||
<quantity id="4CB79B36BF788301FB1E12D0FAEE1CAD" box="[1191,1354,1482,1506]" metricMagnitude="-3" metricUnit="m" metricValue="1.565" metricValueMax="1.71" metricValueMin="1.42" pageId="15" pageNumber="16" unit="mm" value="1.565" valueMax="1.71" valueMin="1.42">1.42–1.71 mm</quantity>
|
||
(1.56), aperture width
|
||
<quantity id="4CB79B36BF788301FEF912F4FE471F49" box="[320,483,1518,1542]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" metricValueMax="1.26" metricValueMin="1.08" pageId="15" pageNumber="16" unit="mm" value="1.17" valueMax="1.26" valueMin="1.08">1.08–1.26 mm</quantity>
|
||
(1.15), total number of whorls 5.75–7.75 (6.54) (USNM 1153670,
|
||
<emphasis id="B93BEAC1BF788301FB6D12EAFB461F48" box="[1236,1250,1520,1543]" italics="true" pageId="15" pageNumber="16">n</emphasis>
|
||
= 13).
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF788301FF7E1109FC671F01" blockId="15.[151,1436,726,1974]" pageId="15" pageNumber="16">
|
||
<emphasis id="B93BEAC1BF788301FF7E1109FDA71F64" bold="true" box="[199,515,1554,1579]" pageId="15" pageNumber="16">
|
||
Measurements of
|
||
<typeStatus id="54F48871BF788301FE251109FDA71F64" box="[412,515,1555,1579]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
: height
|
||
<quantity id="4CB79B36BF788301FDD91108FD631F65" box="[608,711,1554,1578]" metricMagnitude="-3" metricUnit="m" metricValue="5.22" pageId="15" pageNumber="16" unit="mm" value="5.22">5.22 mm</quantity>
|
||
, width
|
||
<quantity id="4CB79B36BF788301FCA01108FC241F65" box="[793,896,1554,1578]" metricMagnitude="-3" metricUnit="m" metricValue="2.21" pageId="15" pageNumber="16" unit="mm" value="2.21">2.21 mm</quantity>
|
||
, body whorl height
|
||
<quantity id="4CB79B36BF788301FBD81108FB6D1F65" box="[1121,1225,1554,1578]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="15" pageNumber="16" unit="mm" value="2.9">2.90 mm</quantity>
|
||
, body whorl width
|
||
<quantity id="4CB79B36BF788301FF2E112CFF591F01" box="[151,253,1590,1614]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="15" pageNumber="16" unit="mm" value="2.03">2.03 mm</quantity>
|
||
, aperture height
|
||
<quantity id="4CB79B36BF788301FE01112CFDBA1F01" box="[440,542,1590,1614]" metricMagnitude="-3" metricUnit="m" metricValue="1.69" pageId="15" pageNumber="16" unit="mm" value="1.69">1.69 mm</quantity>
|
||
, aperture width
|
||
<quantity id="4CB79B36BF788301FD6A112CFC9D1F01" box="[723,825,1590,1614]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" pageId="15" pageNumber="16" unit="mm" value="1.27">1.27 mm</quantity>
|
||
, 6.5 whorls.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF788301FF7E1140FB1A1E69" blockId="15.[151,1436,726,1974]" pageId="15" pageNumber="16">
|
||
Edges of last 0.5 operculum whorl sometimes frilled on outer side (
|
||
<figureCitation id="13742A56BF788301FC721140FBAB1F3D" box="[971,1039,1626,1650]" captionStart="FIGURE 8" captionStartId="16.[151,244,1854,1875]" captionTargetBox="[151,1436,151,1835]" captionTargetId="figure@16.[151,1436,149,1835]" captionTargetPageId="16" captionText="FIGURE 8. Tryonia molinae sp. nov. and Tryonia contrerasi sp. nov. A. Holotype, T. molinae, USNM 1139278. B – C. Shells, T. molinae, USNM 1153674 (C is a male). D. Holotype, T. contrerasi, USNM 874121. E – F. Shells, T. contrerasi, USNM 874128 (F is a male). G – H. Operculum (outer, inner sides), T. molinae, USNM 1153674. I – J. Operculum (outer, inner sides), T. contrerasi, USNM 1153670. K. Portion of radular ribbon, T. molinae, USNM 1153674. L. Central radular teeth, T. molinae, USNM 1153670. M. Lateral and inner marginal teeth, T. molinae, USNM 1153670. N. Portion of radular ribbon, T. contrerasi, USNM 1153670. Q. Central radular teeth, T. contrerasi, USNM 1153670. R. Lateral tooth, T. contrerasi, USNM 1153670. Scale bars A – F = 1.0 mm; G – J = 250 µm; K, N = 25 µm, L – M, O – P = 10 µm." httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="15" pageNumber="16">
|
||
Fig.
|
||
<date id="FFF11013BF788301FBB81140FBAB1F3D" box="[1025,1039,1626,1650]" pageId="15" pageNumber="16">8</date>
|
||
</figureCitation>
|
||
I), inner side of operculum smooth (
|
||
<figureCitation id="13742A56BF788301FF261164FF401FD9" box="[159,228,1662,1686]" captionStart="FIGURE 8" captionStartId="16.[151,244,1854,1875]" captionTargetBox="[151,1436,151,1835]" captionTargetId="figure@16.[151,1436,149,1835]" captionTargetPageId="16" captionText="FIGURE 8. Tryonia molinae sp. nov. and Tryonia contrerasi sp. nov. A. Holotype, T. molinae, USNM 1139278. B – C. Shells, T. molinae, USNM 1153674 (C is a male). D. Holotype, T. contrerasi, USNM 874121. E – F. Shells, T. contrerasi, USNM 874128 (F is a male). G – H. Operculum (outer, inner sides), T. molinae, USNM 1153674. I – J. Operculum (outer, inner sides), T. contrerasi, USNM 1153670. K. Portion of radular ribbon, T. molinae, USNM 1153674. L. Central radular teeth, T. molinae, USNM 1153670. M. Lateral and inner marginal teeth, T. molinae, USNM 1153670. N. Portion of radular ribbon, T. contrerasi, USNM 1153670. Q. Central radular teeth, T. contrerasi, USNM 1153670. R. Lateral tooth, T. contrerasi, USNM 1153670. Scale bars A – F = 1.0 mm; G – J = 250 µm; K, N = 25 µm, L – M, O – P = 10 µm." httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
J). Radula (
|
||
<figureCitation id="13742A56BF788301FED31164FE091FD9" box="[362,429,1662,1686]" captionStart="FIGURE 8" captionStartId="16.[151,244,1854,1875]" captionTargetBox="[151,1436,151,1835]" captionTargetId="figure@16.[151,1436,149,1835]" captionTargetPageId="16" captionText="FIGURE 8. Tryonia molinae sp. nov. and Tryonia contrerasi sp. nov. A. Holotype, T. molinae, USNM 1139278. B – C. Shells, T. molinae, USNM 1153674 (C is a male). D. Holotype, T. contrerasi, USNM 874121. E – F. Shells, T. contrerasi, USNM 874128 (F is a male). G – H. Operculum (outer, inner sides), T. molinae, USNM 1153674. I – J. Operculum (outer, inner sides), T. contrerasi, USNM 1153670. K. Portion of radular ribbon, T. molinae, USNM 1153674. L. Central radular teeth, T. molinae, USNM 1153670. M. Lateral and inner marginal teeth, T. molinae, USNM 1153670. N. Portion of radular ribbon, T. contrerasi, USNM 1153670. Q. Central radular teeth, T. contrerasi, USNM 1153670. R. Lateral tooth, T. contrerasi, USNM 1153670. Scale bars A – F = 1.0 mm; G – J = 250 µm; K, N = 25 µm, L – M, O – P = 10 µm." httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
N-P): dorsal edge of central radular teeth concave, basal tongue rounded, median cusps elongate, distally pointed, parallel-sided proximally, lateral cusps four–seven, basal cusps one–three (innermost larger) (
|
||
<figureCitation id="13742A56BF788301FF4811DCFE901F91" box="[241,308,1734,1758]" captionStart="FIGURE 8" captionStartId="16.[151,244,1854,1875]" captionTargetBox="[151,1436,151,1835]" captionTargetId="figure@16.[151,1436,149,1835]" captionTargetPageId="16" captionText="FIGURE 8. Tryonia molinae sp. nov. and Tryonia contrerasi sp. nov. A. Holotype, T. molinae, USNM 1139278. B – C. Shells, T. molinae, USNM 1153674 (C is a male). D. Holotype, T. contrerasi, USNM 874121. E – F. Shells, T. contrerasi, USNM 874128 (F is a male). G – H. Operculum (outer, inner sides), T. molinae, USNM 1153674. I – J. Operculum (outer, inner sides), T. contrerasi, USNM 1153670. K. Portion of radular ribbon, T. molinae, USNM 1153674. L. Central radular teeth, T. molinae, USNM 1153670. M. Lateral and inner marginal teeth, T. molinae, USNM 1153670. N. Portion of radular ribbon, T. contrerasi, USNM 1153670. Q. Central radular teeth, T. contrerasi, USNM 1153670. R. Lateral tooth, T. contrerasi, USNM 1153670. Scale bars A – F = 1.0 mm; G – J = 250 µm; K, N = 25 µm, L – M, O – P = 10 µm." httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
<collectingCountry id="F3587643BF788301FE8D11DCFEEF1F91" box="[308,331,1734,1758]" name="Iceland" pageId="15" pageNumber="16">O</collectingCountry>
|
||
). Lateral teeth having two–four cusps on inner and three–five cusps on outer side, length of outer wing about 180% width of cutting edge, central cusp hoe-shaped (
|
||
<figureCitation id="13742A56BF788301FCC411F0FC641E4D" box="[893,960,1770,1794]" captionStart="FIGURE 8" captionStartId="16.[151,244,1854,1875]" captionTargetBox="[151,1436,151,1835]" captionTargetId="figure@16.[151,1436,149,1835]" captionTargetPageId="16" captionText="FIGURE 8. Tryonia molinae sp. nov. and Tryonia contrerasi sp. nov. A. Holotype, T. molinae, USNM 1139278. B – C. Shells, T. molinae, USNM 1153674 (C is a male). D. Holotype, T. contrerasi, USNM 874121. E – F. Shells, T. contrerasi, USNM 874128 (F is a male). G – H. Operculum (outer, inner sides), T. molinae, USNM 1153674. I – J. Operculum (outer, inner sides), T. contrerasi, USNM 1153670. K. Portion of radular ribbon, T. molinae, USNM 1153674. L. Central radular teeth, T. molinae, USNM 1153670. M. Lateral and inner marginal teeth, T. molinae, USNM 1153670. N. Portion of radular ribbon, T. contrerasi, USNM 1153670. Q. Central radular teeth, T. contrerasi, USNM 1153670. R. Lateral tooth, T. contrerasi, USNM 1153670. Scale bars A – F = 1.0 mm; G – J = 250 µm; K, N = 25 µm, L – M, O – P = 10 µm." httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
P). Inner marginal teeth with 16–25 cusps, outer marginal teeth with 30–40 cusps. Radula data are from USNM 874128, USNM 1153670.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF788301FF7E1028FEF31EF9" blockId="15.[151,1436,726,1974]" pageId="15" pageNumber="16">
|
||
Animal darkly pigmented. Penis (
|
||
<figureCitation id="13742A56BF788301FDF91028FD251E05" box="[576,641,1842,1866]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
|
||
F) having two distal papillae and one basal papilla on inner edge (56 of
|
||
<specimenCount id="9D49FD5ABF788301FF2E104CFE961E21" box="[151,306,1878,1902]" pageId="15" pageNumber="16" type="generic">57 specimens</specimenCount>
|
||
from two samples); one specimen differed in lacking a basal papilla. Distal bulb of penis evenly rounded; stylet small. Penial duct weakly undulating along most of length. Penial data are from USNM 874128, USNM 1153670.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF30665BBF67831EFF2E1024FE1E1EBC" httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="16" pageNumber="17" targetBox="[151,1436,151,1835]" targetPageId="16">
|
||
<paragraph id="8BF036D3BF67831EFF2E1024FE1E1EBC" blockId="16.[151,1436,1854,2035]" pageId="16" pageNumber="17">
|
||
<emphasis id="B93BEAC1BF67831EFF2E1024FEA21E1C" bold="true" box="[151,262,1854,1875]" pageId="16" pageNumber="17">FIGURE 8</emphasis>
|
||
.
|
||
<taxonomicName id="4C4F4D50BF67831EFEA81024FE0B1E1C" box="[273,431,1854,1875]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="molinae" status="sp. nov.">
|
||
<emphasis id="B93BEAC1BF67831EFEA81024FE0B1E1C" box="[273,431,1854,1875]" italics="true" pageId="16" pageNumber="17">Tryonia molinae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20857BABF67831EFE0D1024FDA51E1C" box="[436,513,1854,1875]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B93BEAC1BF67831EFE0D1024FE5F1E1C" bold="true" box="[436,507,1854,1875]" pageId="16" pageNumber="17">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C4F4D50BF67831EFD961024FD441E1C" box="[559,736,1854,1875]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="contrerasi" status="sp. nov.">
|
||
<emphasis id="B93BEAC1BF67831EFD961024FD441E1C" box="[559,736,1854,1875]" italics="true" pageId="16" pageNumber="17">Tryonia contrerasi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20857BABF67831EFD5F1024FC961E1C" box="[742,818,1854,1875]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B93BEAC1BF67831EFD5F1024FC891E1C" bold="true" box="[742,813,1854,1875]" pageId="16" pageNumber="17">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
A. Holotype,
|
||
<taxonomicName id="4C4F4D50BF67831EFC031024FBBB1E1C" box="[954,1055,1854,1875]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="molinae">
|
||
<emphasis id="B93BEAC1BF67831EFC031024FBBB1E1C" box="[954,1055,1854,1875]" italics="true" pageId="16" pageNumber="17">T. molinae</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1139278. B–C. Shells,
|
||
<taxonomicName id="4C4F4D50BF67831EFAEB1024FF1E1E21" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="molinae">
|
||
<emphasis id="B93BEAC1BF67831EFAEB1024FF1E1E21" italics="true" pageId="16" pageNumber="17">T. molinae</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153674 (C is a male). D. Holotype,
|
||
<taxonomicName id="4C4F4D50BF67831EFD311043FCA01E21" box="[648,772,1881,1902]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="contrerasi">
|
||
<emphasis id="B93BEAC1BF67831EFD311043FCA01E21" box="[648,772,1881,1902]" italics="true" pageId="16" pageNumber="17">T. contrerasi</emphasis>
|
||
</taxonomicName>
|
||
, USNM 874121. E–F. Shells,
|
||
<taxonomicName id="4C4F4D50BF67831EFB8D1043FB091E21" box="[1076,1197,1881,1902]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="contrerasi">
|
||
<emphasis id="B93BEAC1BF67831EFB8D1043FB091E21" box="[1076,1197,1881,1902]" italics="true" pageId="16" pageNumber="17">T. contrerasi</emphasis>
|
||
</taxonomicName>
|
||
, USNM 874128 (F is a male). G–H. Operculum (outer, inner sides),
|
||
<taxonomicName id="4C4F4D50BF67831EFDF4106EFD151EC6" box="[589,689,1908,1929]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="molinae">
|
||
<emphasis id="B93BEAC1BF67831EFDF4106EFD151EC6" box="[589,689,1908,1929]" italics="true" pageId="16" pageNumber="17">T. molinae</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153674. I–J. Operculum (outer, inner sides),
|
||
<taxonomicName id="4C4F4D50BF67831EFB76106EFAED1EC6" box="[1231,1353,1908,1929]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="contrerasi">
|
||
<emphasis id="B93BEAC1BF67831EFB76106EFAED1EC6" box="[1231,1353,1908,1929]" italics="true" pageId="16" pageNumber="17">T. contrerasi</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153670. K. Portion of radular ribbon,
|
||
<taxonomicName id="4C4F4D50BF67831EFDA91094FDD11EEC" box="[528,629,1934,1955]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="molinae">
|
||
<emphasis id="B93BEAC1BF67831EFDA91094FDD11EEC" box="[528,629,1934,1955]" italics="true" pageId="16" pageNumber="17">T. molinae</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153674. L. Central radular teeth,
|
||
<taxonomicName id="4C4F4D50BF67831EFBA31094FBDB1EEC" box="[1050,1151,1934,1955]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="molinae">
|
||
<emphasis id="B93BEAC1BF67831EFBA31094FBDB1EEC" box="[1050,1151,1934,1955]" italics="true" pageId="16" pageNumber="17">T. molinae</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153670. M. Lateral and inner marginal teeth,
|
||
<taxonomicName id="4C4F4D50BF67831EFE3510B3FE551EF1" box="[396,497,1961,1982]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="molinae">
|
||
<emphasis id="B93BEAC1BF67831EFE3510B3FE551EF1" box="[396,497,1961,1982]" italics="true" pageId="16" pageNumber="17">T. molinae</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153670. N. Portion of radular ribbon,
|
||
<taxonomicName id="4C4F4D50BF67831EFC0610B3FB931EF1" box="[959,1079,1961,1982]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="contrerasi">
|
||
<emphasis id="B93BEAC1BF67831EFC0610B3FB931EF1" box="[959,1079,1961,1982]" italics="true" pageId="16" pageNumber="17">T. contrerasi</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153670. Q. Central radular teeth,
|
||
<taxonomicName id="4C4F4D50BF67831EFF6810DEFEEE1E96" box="[209,330,1988,2009]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="contrerasi">
|
||
<emphasis id="B93BEAC1BF67831EFF6810DEFEEE1E96" box="[209,330,1988,2009]" italics="true" pageId="16" pageNumber="17">T. contrerasi</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153670. R. Lateral tooth,
|
||
<taxonomicName id="4C4F4D50BF67831EFD1D10DEFCB81E96" box="[676,796,1988,2009]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="contrerasi">
|
||
<emphasis id="B93BEAC1BF67831EFD1D10DEFCB81E96" box="[676,796,1988,2009]" italics="true" pageId="16" pageNumber="17">T. contrerasi</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153670. Scale bars A–F = 1.0 mm; G–J = 250 µm; K, N = 25 µm, L–M, O–P = 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3556558BF66831FFF7E178DFD421880" pageId="17" pageNumber="18" type="distribution">
|
||
<paragraph id="8BF036D3BF66831FFF7E178DFD421880" blockId="17.[151,1437,151,573]" pageId="17" pageNumber="18">
|
||
<emphasis id="B93BEAC1BF66831FFF7E178DFE5719FF" bold="true" box="[199,499,151,176]" pageId="17" pageNumber="18">Distribution and habitat.</emphasis>
|
||
<taxonomicName id="4C4F4D50BF66831FFE451783FD6919FF" box="[508,717,153,176]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="contrerasi">
|
||
<emphasis id="B93BEAC1BF66831FFE451783FD6919FF" box="[508,717,153,176]" italics="true" pageId="17" pageNumber="18">Tryonia contrerasi</emphasis>
|
||
</taxonomicName>
|
||
is distributed in two closely proximal (ca.
|
||
<quantity id="4CB79B36BF66831FFB0E178DFB5719E0" box="[1207,1267,151,175]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="km" value="3.0">3 km</quantity>
|
||
) springs in the (endorheic) Bolson de los Muertos (
|
||
<figureCitation id="13742A56BF66831FFD9117A6FDCE199B" box="[552,618,188,212]" captionStart="FIGURE 1" captionStartId="2.[151,244,1795,1816]" captionTargetBox="[179,1402,157,1789]" captionTargetId="figure@2.[179,1423,125,1806]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of Tryonia species in the Chihuahuan Desert. 1. Tryonia hertleini, Las Palomas. 2. Tryonia peregrina sp. nov., Ojo Vareleño, northwest of Casas Grandes. 3. Tryonia peregrina sp. nov., Yepomera. 4. Tryonia zaragozae sp. nov., northeast of Ignacio Zaragoza. 5. Tryonia angostura sp. nov., T. ovata sp. nov., south of Galeana. 6. Tryonia molinae sp. nov., El Molino. 7. Tryonia taylori sp. nov., vicinity of Rancho Ojo Caliente. 8. Tryonia contrerasi sp. nov., vicinity of Rancho Nuevo. 9. Tryonia chuviscarae sp. nov., T. minckleyi sp. nov., Balneario de San Diego de Alcala. 10. Tryonia julimesensis sp. nov., south of Julimes. 11. Tryonia allendae sp. nov., southwest of Talamantes. 12. Tryonia metcalfi sp. nov., La Cienega, south of Candelaria. 13. Tryonia oasiensis sp. nov., Caroline Spring, Oasis Ranch. 14. Tryonia cheatumi, Phantom Lake Spring. 15. Tryonia circumstriata, Diamond Y Draw. 16. Tryonia seemani, Ojo de Agua del Obispo, Durango City. 17. Tryonia seemani, Nombre de Dios. 18. Tryonia seemani, Amado Nervo." httpUri="https://zenodo.org/record/278443/files/figure.png" pageId="17" pageNumber="18">Fig. 1</figureCitation>
|
||
, locality 8). These springs are part of a complex to the west of Villa Ahumada near Rancho Nuevo. All of these springs with the exception of Ojo Carbonero (referred to as Ojo Carbonera by
|
||
<bibRefCitation id="EFDE4B22BF66831FFF00161EFEDA1853" author="Miller" box="[185,382,260,284]" pageId="17" pageNumber="18" refString="Miller, R. R., Minckley, W. L. & Norris, S. M. (2005) Freshwater Fishes of Mexico. The University of Chicago Press, Chicago, pp. 1 - 490, plates 1 - 99." type="book chapter" year="2005">
|
||
Miller
|
||
<emphasis id="B93BEAC1BF66831FFEBF161FFE931853" box="[262,311,260,284]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
|
||
. 2005
|
||
</bibRefCitation>
|
||
) have been highly modified. The
|
||
<typeStatus id="54F48871BF66831FFD4F161FFC821852" box="[758,806,261,285]" pageId="17" pageNumber="18">type</typeStatus>
|
||
locality, Ojo del Apache, is a thermal spring (27°C) that has been severely degraded (trampled) by horses (
|
||
<figureCitation id="13742A56BF66831FFD75163DFCB41870" box="[716,784,295,319]" captionStart="FIGURE 6" captionStartId="11.[151,244,1710,1731]" captionTargetBox="[221,1371,156,1678]" captionTargetId="figure@11.[212,1381,149,1685]" captionTargetPageId="11" captionText="FIGURE 6. Photographs of habitats of species treated in this paper. A. Ojo Vareleño, Chihuahua, type locality of T. peregrina sp. nov. (photograph taken on 16 / ix / 1990). B. Main spring in Yepomera, Chihuahua, T. peregrina habitat (18 / ix / 1990). C. Springs northeast of Ignacio Zaragoza, Chihuaua, type locality of T. zaragozae sp. nov. (8 / xii / 1998). D. Western-most head spring, Ojos de Arrey, Chihuahua, type locality of T. angosturae sp. nov. and T. ovata sp. nov. (7 / xii / 1998). E. Spring in northern part of El Molino, Chihuahua, type locality of T. molinae sp. nov. (26 / iv / 2010). F. Ojo del Apache, Chihuahua, type locality of T. contrerasi sp. nov. (14 / xii / 1998). G. Ojo Carbonero, Chihuahua, T. contrerasi habitat (2 / iv / 1991). H. Outflow of Ojo Caliente, Chihuahua, type locality of T. taylori sp. nov. (7 / xii / 1998). I – J. El Ojo de Servin, Chihuahua, T. taylori habitat (27 - viii- 1971, 7 / xii / 1998, respectively). K. Seeps in La Cienega, Texas, type locality of T. metcalfi sp. nov. (6 / xii / 2001). L – M. Balneario de San Diego de Alcala, Chihuahua, type locality of T. chuviscarae sp. nov. and T. minckleyi sp. nov. (4 / xii / 2001). N. Springs southeast of Julimes, Chihuahua, type locality of T. julimesensis sp. nov. (5 / iv / 1991). O. El Ojito (outflow), southwest of Talamantes, Chihuahua, type locality of T. allendae sp. nov. (10 / xii / 1998). P. Caroline Spring, Oasis Ranch, Texas, type locality of T. oasiensis sp. nov. (30 / iii / 2009). Q. Spring in Nombre de Dios, Durango, T. seemani habitat (2 / ix / 1971). R. Spring in Amado Nervo, Durango, T. seemani habitat (7 / iv / 1991)." httpUri="https://zenodo.org/record/278448/files/figure.png" pageId="17" pageNumber="18">Fig. 6</figureCitation>
|
||
F). Snails were found in this spring on aquatic vegetation. Ojo Carbonero contains at least eight discrete sources that coalesce into a spring run for about
|
||
<quantity id="4CB79B36BF66831FFB661656FA8C182B" box="[1247,1320,332,356]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="100.0">100 m</quantity>
|
||
(
|
||
<figureCitation id="13742A56BF66831FFA801656FADA182B" box="[1337,1406,332,356]" captionStart="FIGURE 6" captionStartId="11.[151,244,1710,1731]" captionTargetBox="[221,1371,156,1678]" captionTargetId="figure@11.[212,1381,149,1685]" captionTargetPageId="11" captionText="FIGURE 6. Photographs of habitats of species treated in this paper. A. Ojo Vareleño, Chihuahua, type locality of T. peregrina sp. nov. (photograph taken on 16 / ix / 1990). B. Main spring in Yepomera, Chihuahua, T. peregrina habitat (18 / ix / 1990). C. Springs northeast of Ignacio Zaragoza, Chihuaua, type locality of T. zaragozae sp. nov. (8 / xii / 1998). D. Western-most head spring, Ojos de Arrey, Chihuahua, type locality of T. angosturae sp. nov. and T. ovata sp. nov. (7 / xii / 1998). E. Spring in northern part of El Molino, Chihuahua, type locality of T. molinae sp. nov. (26 / iv / 2010). F. Ojo del Apache, Chihuahua, type locality of T. contrerasi sp. nov. (14 / xii / 1998). G. Ojo Carbonero, Chihuahua, T. contrerasi habitat (2 / iv / 1991). H. Outflow of Ojo Caliente, Chihuahua, type locality of T. taylori sp. nov. (7 / xii / 1998). I – J. El Ojo de Servin, Chihuahua, T. taylori habitat (27 - viii- 1971, 7 / xii / 1998, respectively). K. Seeps in La Cienega, Texas, type locality of T. metcalfi sp. nov. (6 / xii / 2001). L – M. Balneario de San Diego de Alcala, Chihuahua, type locality of T. chuviscarae sp. nov. and T. minckleyi sp. nov. (4 / xii / 2001). N. Springs southeast of Julimes, Chihuahua, type locality of T. julimesensis sp. nov. (5 / iv / 1991). O. El Ojito (outflow), southwest of Talamantes, Chihuahua, type locality of T. allendae sp. nov. (10 / xii / 1998). P. Caroline Spring, Oasis Ranch, Texas, type locality of T. oasiensis sp. nov. (30 / iii / 2009). Q. Spring in Nombre de Dios, Durango, T. seemani habitat (2 / ix / 1971). R. Spring in Amado Nervo, Durango, T. seemani habitat (7 / iv / 1991)." httpUri="https://zenodo.org/record/278448/files/figure.png" pageId="17" pageNumber="18">Fig. 6</figureCitation>
|
||
G) before flowing into a canal. Snails were common in this thermal (26–27°C) spring both in detritus and on sand. UTEP 881, which consists of old, worn shells, may have been collected from a playa lake (Laguna Colorada) along the highway southwest of Hacienda Santo
|
||
<collectingCountry id="F3587643BF66831FFDCD16ADFD461880" box="[628,738,439,463]" name="Dominican Republic" pageId="17" pageNumber="18">Domingo</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF66831FFF7E16C6FBE51B73" pageId="17" pageNumber="18" type="discussion">
|
||
<paragraph id="8BF036D3BF66831FFF7E16C6FBE51B73" blockId="17.[151,1437,151,573]" pageId="17" pageNumber="18">
|
||
<emphasis id="B93BEAC1BF66831FFF7E16C6FE9018BB" bold="true" box="[199,308,476,500]" pageId="17" pageNumber="18">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C4F4D50BF66831FFEFC16C7FDBC18BB" box="[325,536,477,500]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="contrerasi">
|
||
<emphasis id="B93BEAC1BF66831FFEFC16C7FDBC18BB" box="[325,536,477,500]" italics="true" pageId="17" pageNumber="18">Tryonia contrerasi</emphasis>
|
||
</taxonomicName>
|
||
was delineated in the Bayesian analysis as a well supported lineage (
|
||
<figureCitation id="13742A56BF66831FFAFF16C6FA2A18BB" box="[1350,1422,476,500]" captionStart="FIGURE 2" captionStartId="5.[151,244,1938,1959]" captionTargetBox="[244,1345,828,1915]" captionTargetId="figure@5.[244,1346,828,1920]" captionTargetPageId="5" captionText="FIGURE 2. Bayesian tree based on the COI dataset. Posterior probabilities are provided when> 95 %. The names of the species treated herein are in bold face. Terminals and specimen codes are labeled as in Table 1. The two well supported clades composed of new species described herein are labeled A – B." httpUri="https://zenodo.org/record/278444/files/figure.png" pageId="17" pageNumber="18">Fig. 2</figureCitation>
|
||
). Snails from Ojo Carbonero (
|
||
<figureCitation id="13742A56BF66831FFE6C16E5FDB11B58" box="[469,533,511,535]" captionStart="FIGURE 8" captionStartId="16.[151,244,1854,1875]" captionTargetBox="[151,1436,151,1835]" captionTargetId="figure@16.[151,1436,149,1835]" captionTargetPageId="16" captionText="FIGURE 8. Tryonia molinae sp. nov. and Tryonia contrerasi sp. nov. A. Holotype, T. molinae, USNM 1139278. B – C. Shells, T. molinae, USNM 1153674 (C is a male). D. Holotype, T. contrerasi, USNM 874121. E – F. Shells, T. contrerasi, USNM 874128 (F is a male). G – H. Operculum (outer, inner sides), T. molinae, USNM 1153674. I – J. Operculum (outer, inner sides), T. contrerasi, USNM 1153670. K. Portion of radular ribbon, T. molinae, USNM 1153674. L. Central radular teeth, T. molinae, USNM 1153670. M. Lateral and inner marginal teeth, T. molinae, USNM 1153670. N. Portion of radular ribbon, T. contrerasi, USNM 1153670. Q. Central radular teeth, T. contrerasi, USNM 1153670. R. Lateral tooth, T. contrerasi, USNM 1153670. Scale bars A – F = 1.0 mm; G – J = 250 µm; K, N = 25 µm, L – M, O – P = 10 µm." httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="17" pageNumber="18">Fig. 8</figureCitation>
|
||
E–F) are slightly smaller and have more highly sculptured shells than those from the
|
||
<typeStatus id="54F48871BF66831FFF79153FFF541B72" box="[192,240,549,573]" pageId="17" pageNumber="18">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13742A56BF66831FFEE3153EFE381B73" box="[346,412,548,572]" captionStart="FIGURE 8" captionStartId="16.[151,244,1854,1875]" captionTargetBox="[151,1436,151,1835]" captionTargetId="figure@16.[151,1436,149,1835]" captionTargetPageId="16" captionText="FIGURE 8. Tryonia molinae sp. nov. and Tryonia contrerasi sp. nov. A. Holotype, T. molinae, USNM 1139278. B – C. Shells, T. molinae, USNM 1153674 (C is a male). D. Holotype, T. contrerasi, USNM 874121. E – F. Shells, T. contrerasi, USNM 874128 (F is a male). G – H. Operculum (outer, inner sides), T. molinae, USNM 1153674. I – J. Operculum (outer, inner sides), T. contrerasi, USNM 1153670. K. Portion of radular ribbon, T. molinae, USNM 1153674. L. Central radular teeth, T. molinae, USNM 1153670. M. Lateral and inner marginal teeth, T. molinae, USNM 1153670. N. Portion of radular ribbon, T. contrerasi, USNM 1153670. Q. Central radular teeth, T. contrerasi, USNM 1153670. R. Lateral tooth, T. contrerasi, USNM 1153670. Scale bars A – F = 1.0 mm; G – J = 250 µm; K, N = 25 µm, L – M, O – P = 10 µm." httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="17" pageNumber="18">Fig. 8</figureCitation>
|
||
D); these two populations differed by 0.6% COI divergence.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |