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<document ID-DOI="10.37520/fi.2021.010" ID-ISSN="2533-4069" ID-Zenodo-Dep="7167243" approvalRequired="16" approvalRequired_for_taxonomicNames="3" approvalRequired_for_textStreams="11" approvalRequired_for_treatments="2" checkinTime="1665258969000" checkinUser="felipe" docAuthor="Pickford, Martin, Abdel Gawad, Mohamed, Hamdan, Mohamed, El-Barkooky, Ahmed N. &amp; Al Riaydh, Mohammed H." docDate="2021" docId="03E66B17D41B035ADBBFFB1EFE135790" docLanguage="en" docName="FossilImprint.77.1.111-125.pdf" docOrigin="Fossil Imprint 77 (1)" docSource="http://dx.doi.org/10.37520/fi.2021.010" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.2:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="2" docTitle="Diamantohyus africanus STROMER 1922" docType="treatment" docVersion="2" lastPageNumber="113" masterDocId="FFDF136FD41A0358D82AFFC8FFF4535F" masterDocTitle="New Suoid Fossils (Mammalia, Artiodactyla) From The Miocene Of Moghara, Egypt, And Gebel Zelten, Libya: Biochronological Implications" masterLastPageNumber="125" masterPageNumber="111" pageNumber="112" updateTime="1665272345592" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title>New Suoid Fossils (Mammalia, Artiodactyla) From The Miocene Of Moghara, Egypt, And Gebel Zelten, Libya: Biochronological Implications</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pickford, Martin</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Abdel Gawad, Mohamed</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hamdan, Mohamed</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>El-Barkooky, Ahmed N.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Al Riaydh, Mohammed H.</mods:namePart>
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<mods:date>2021</mods:date>
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<mods:number>2021-12-09</mods:number>
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<mods:number>77</mods:number>
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<mods:url>http://dx.doi.org/10.37520/fi.2021.010</mods:url>
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<mods:identifier type="DOI">10.37520/fi.2021.010</mods:identifier>
<mods:identifier type="ISSN">2533-4069</mods:identifier>
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<paragraph blockId="1.[917,1362,1238,1263]" box="[917,1362,1238,1263]" pageId="1" pageNumber="112">
<heading box="[917,1362,1238,1263]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="112" reason="2">
<taxonomicName authority="STROMER, 1922" authorityName="STROMER" authorityYear="1922" box="[917,1362,1238,1263]" class="Mammalia" family="Palaeochoeridae" genus="Diamantohyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="112" phylum="Chordata" rank="species" species="africanus">
<emphasis bold="true" box="[917,1362,1238,1263]" pageId="1" pageNumber="112">
<emphasis bold="true" box="[917,1190,1238,1262]" italics="true" pageId="1" pageNumber="112">Diamantohyus africanus</emphasis>
<bibRefCitation author="Stromer, E." box="[1197,1296,1238,1262]" pageId="1" pageNumber="112" pagination="331 - 340" refId="ref8306" refString="Stromer, E. (1922): Erste Mitteilung uber tertiare Wirbeltier- Reste aus Deutsch-Sudwestafrika. - Sitzungsberichte der mathematisch-physikalischen Klasse der Bayerischen Akademie der Wissenschaften zu Munchen, 1921: 331 - 340." type="journal article" year="1922">STROMER</bibRefCitation>
, 1922
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="1.[820,1459,1286,1406]" pageId="1" pageNumber="112">
H o l o t y p e: BSPG
<date box="[1117,1213,1286,1310]" pageId="1" pageNumber="112" value="1926-10">1926 X</date>
, right maxilla with P3/, M1/M2/, currently lost (Gertrud Rössner, personal communication 2009). Figured in
<bibRefCitation author="Stromer, E." box="[1207,1371,1350,1374]" pageId="1" pageNumber="112" pagination="107 - 153" refId="ref8342" refString="Stromer, E. (1926): Reste Land- und Susswasser-Bewohnender Wirbeltiere aus dem Diamantefeldern Deutsch- Sudwestafrikas. - In: Kaiser, E. (ed.), Die Diamantenwuste Sudwest-Afrikas, Bd. 2. Dietrich Reimer, Berlin, pp. 107 - 153." type="book chapter" year="1926">Stromer (1926</bibRefCitation>
: pl. 40, fig. 17); also figured in
<bibRefCitation author="Pickford, M." box="[1071,1235,1382,1406]" pageId="1" pageNumber="112" pagination="133 - 154" refId="ref7824" refString="Pickford, M. (1984): A revision of the Sanitheriidae (Suiformes, Mammalia). - Geobios, 17: 133 - 154. https: // doi. org / 10.1016 / S 0016 - 6995 (84) 80139 - 4" type="journal article" year="1984">Pickford (1984</bibRefCitation>
: fig. 1).
</paragraph>
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<paragraph blockId="1.[820,1459,1430,1646]" pageId="1" pageNumber="112">
D i a g n o s i s. Differs from species of
<taxonomicName authorityName="Meyer" authorityYear="1865" box="[1297,1427,1430,1454]" class="Mammalia" family="Palaeochoeridae" genus="Sanitherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="112" phylum="Chordata" rank="genus">
<emphasis box="[1297,1427,1430,1454]" italics="true" pageId="1" pageNumber="112">Sanitherium</emphasis>
</taxonomicName>
in having less molarized premolars, and in having anterior and postero-lingual cusps less developed; P4/ with two main cusps and two subsidiary ones; metastylid prominent on unworn specimens; m/1m/3 ca.
<quantity box="[1174,1248,1558,1582]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="112" unit="mm" value="40.0">40 mm</quantity>
long (range
<quantity metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.25" metricValueMin="3.75" pageId="1" pageNumber="112" unit="mm" value="40.0" valueMax="42.5" valueMin="37.5">37.5 to 42.5 mm</quantity>
) (after
<bibRefCitation author="Pickford, M." box="[992,1147,1590,1614]" pageId="1" pageNumber="112" pagination="133 - 154" refId="ref7824" refString="Pickford, M. (1984): A revision of the Sanitheriidae (Suiformes, Mammalia). - Geobios, 17: 133 - 154. https: // doi. org / 10.1016 / S 0016 - 6995 (84) 80139 - 4" type="journal article" year="1984">Pickford 1984</bibRefCitation>
). Smaller than
<taxonomicName authority="(WILKINSON, 1976)" baseAuthorityName="WILKINSON" baseAuthorityYear="1976" class="Mammalia" family="Palaeochoeridae" genus="Diamantohyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="112" phylum="Chordata" rank="species" species="nadirus">
<emphasis italics="true" pageId="1" pageNumber="112">Diamantohyus nadirus</emphasis>
(
<bibRefCitation author="Wilkinson, A." box="[917,1102,1622,1646]" pageId="1" pageNumber="112" pagination="173 - 282" refId="ref8445" refString="Wilkinson, A. (1976): The lower Miocene Suidae of Africa. - In: Savage, R. J. G., Coryndon, S. C. (eds), Fossil Vertebrates of Africa, vol. 4. Academic Press, London, New York, San Francisco, pp. 173 - 282, 10 pls." type="book chapter" year="1976">WILKINSON, 1976</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph blockId="1.[820,1459,1669,1726]" pageId="1" pageNumber="112">
T y p e l o c a l i t y a n d a g e. Langental,
<collectingCountry box="[1360,1455,1669,1693]" name="Namibia" pageId="1" pageNumber="112">Namibia</collectingCountry>
, early Miocene.
</paragraph>
<paragraph blockId="1.[820,1459,1749,1837]" pageId="1" pageNumber="112">N e w m a t e r i a l. CUWM 239, left m/3; CUWM 261, right mandible fragment with damaged m/2m/3; CUWM 360, right mandible fragment and associated m/3.</paragraph>
<paragraph blockId="1.[820,1459,1860,2077]" pageId="1" pageNumber="112">
D e s c r i p t i o n. CUWM 40 (palate), CUWM 57 (m/3), CUWM 59 (M2/), CUWM 63 (left juvenile mandible), CUWM 106 (left adult mandible), CUWM 121 (mandible with m/3) and CUWM 134 (m/1) have already been described (
<tableCitation box="[828,899,1989,2013]" captionStart="Table 1" captionStartId="2.[129,185,152,173]" captionText="Table 1. Old and new catalogue numbers of Diamantohyus africanus from Moghara, Egypt." pageId="1" pageNumber="112">Tab. 1</tableCitation>
) (
<bibRefCitation author="Pickford, M. &amp; Miller, E. &amp; El-Barkooky, A. N." box="[927,1158,1989,2013]" pageId="1" pageNumber="112" pagination="1 - 11" refId="ref8155" refString="Pickford, M., Miller, E., El-Barkooky, A. N. (2010): Suidae and Sanitheriidae from Wadi Moghra, Early Miocene, Egypt. - Acta Palaeontologica Polonica, 55 (1): 1 - 11. https: // doi. org / 10.4202 / app. 2009.0015" type="journal article" year="2010">Pickford et al. 2010</bibRefCitation>
). They are re-illustrated in stereo for convenience (
<figureCitation box="[1089,1236,2021,2045]" captionStart-0="Text-fig" captionStart-1="Text-fig" captionStart-2="Text-fig" captionStart-3="Text-fig" captionStart-4="Text-fig" captionStart-5="Text-fig" captionStartId-0="2.[147,228,2072,2093]" captionStartId-1="3.[125,206,1298,1319]" captionStartId-2="4.[147,228,1028,1049]" captionStartId-3="4.[147,228,1894,1915]" captionStartId-4="5.[125,206,764,785]" captionStartId-5="5.[125,206,1350,1371]" captionTargetBox-0="[172,1411,1300,2033]" captionTargetBox-1="[193,1392,189,1229]" captionTargetBox-2="[205,1395,167,978]" captionTargetBox-3="[171,1450,1158,1866]" captionTargetBox-4="[146,1412,162,740]" captionTargetBox-5="[128,1414,882,1328]" captionTargetId-0="figure-499@2.[147,1462,1271,2056]" captionTargetId-1="figure-382@3.[125,1440,151,1283]" captionTargetId-2="figure-69@4.[147,1462,113,1090]" captionTargetId-3="figure-100@4.[147,1462,1143,1878]" captionTargetId-4="figure-79@5.[125,1440,151,749]" captionTargetId-5="figure-111@5.[125,1440,871,1334]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="5" captionText-0="Text-fig. 1. CUWM 40, palate of Diamantohyus africanus from Moghara, Egypt, stereo occlusal view." captionText-1="Text-fig. 2. Diamantohyus africanus from Moghara, Egypt. a: CUWM 59, right upper molar, stereo occlusal view; b: CUWM 57, right m/3, b1 stereo occlusal views, b2 buccal view, b3 lingual view." captionText-2="Text-fig. 3. CUWM 134, right lower molar of Diamantohyus africanus from Moghara, Egypt. a: stereo occlusal view; b: lingual view; c: buccal view." captionText-3="Text-fig. 4. CUWM 106, left mandible with p/4m/3 of Diamantohyus africanus from Moghara, Egypt. a: buccal view; b: stereo occlusal views; c: lingual view." captionText-4="Text-fig. 5. CUWM 63, left mandible with d/4m/2 of Diamantohyus africanus from Moghara, Egypt. a: buccal view; b: stereo occlusal views; c: lingual view." captionText-5="Text-fig. 6. CUWM 121, right mandible with m/3 of Diamantohyus africanus from Moghara, Egypt. a: lingual view; b: stereo occlusal views; c: buccal view." figureDoi-0="http://doi.org/10.5281/zenodo.7167247" figureDoi-1="http://doi.org/10.5281/zenodo.7167249" figureDoi-2="http://doi.org/10.5281/zenodo.7167253" figureDoi-3="http://doi.org/10.5281/zenodo.7167255" figureDoi-4="http://doi.org/10.5281/zenodo.7167257" figureDoi-5="http://doi.org/10.5281/zenodo.7167259" httpUri-0="https://zenodo.org/record/7167247/files/figure.png" httpUri-1="https://zenodo.org/record/7167249/files/figure.png" httpUri-2="https://zenodo.org/record/7167253/files/figure.png" httpUri-3="https://zenodo.org/record/7167255/files/figure.png" httpUri-4="https://zenodo.org/record/7167257/files/figure.png" httpUri-5="https://zenodo.org/record/7167259/files/figure.png" pageId="1" pageNumber="112">Text-figs 16</figureCitation>
) and measurements are provided in
<tableCitation box="[988,1069,2053,2077]" captionStart="Table 2" captionStartId="7.[106,162,1476,1497]" captionText="Table 2. Measurements (in mm) of newly discovered teeth of Diamantohyus africanus from Moghara, Egypt (e estimated measurement)." pageId="1" pageNumber="112">Table 2</tableCitation>
.
</paragraph>
<caption box="[129,1060,152,173]" pageId="2" pageNumber="113" startId="2.[129,185,152,173]">
<paragraph blockId="2.[129,1060,152,173]" box="[129,1060,152,173]" pageId="2" pageNumber="113">
Table 1. Old and new catalogue numbers of
<taxonomicName authorityName="STROMER" authorityYear="1922" box="[578,824,152,173]" class="Mammalia" family="Palaeochoeridae" genus="Diamantohyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="africanus">
<emphasis bold="true" box="[578,824,152,173]" italics="true" pageId="2" pageNumber="113">Diamantohyus africanus</emphasis>
</taxonomicName>
from Moghara, Egypt.
</paragraph>
</caption>
<paragraph pageId="2" pageNumber="113">
<table box="[137,1318,220,517]" gridcols="3" gridrows="9" pageId="2" pageNumber="113">
<tr box="[137,1318,220,239]" gridrow="0" pageId="2" pageNumber="113">
<th box="[137,455,220,239]" gridcol="0" gridrow="0" pageId="2" pageNumber="113">
<emphasis bold="true" box="[251,455,220,239]" pageId="2" pageNumber="113">Old catalogue number</emphasis>
</th>
<th box="[583,904,220,239]" gridcol="1" gridrow="0" pageId="2" pageNumber="113">
<emphasis bold="true" box="[694,904,220,239]" pageId="2" pageNumber="113">New catalogue number</emphasis>
</th>
<th box="[1029,1318,220,239]" gridcol="2" gridrow="0" pageId="2" pageNumber="113">
<emphasis bold="true" box="[1205,1293,220,239]" pageId="2" pageNumber="113">Specimen</emphasis>
</th>
</tr>
<tr box="[137,1318,255,274]" gridrow="1" pageId="2" pageNumber="113">
<th box="[137,455,255,274]" gridcol="0" gridrow="1" pageId="2" pageNumber="113">WM 05-50</th>
<td box="[583,904,255,274]" gridcol="1" gridrow="1" pageId="2" pageNumber="113">CUWM 40</td>
<td box="[1029,1318,255,274]" gridcol="2" gridrow="1" pageId="2" pageNumber="113">Palate with both cheek tooth rows</td>
</tr>
<tr box="[137,1318,289,308]" gridrow="2" pageId="2" pageNumber="113">
<th box="[137,455,289,308]" gridcol="0" gridrow="2" pageId="2" pageNumber="113">WM 05-48 (DEC06-025)</th>
<td box="[583,904,289,308]" gridcol="1" gridrow="2" pageId="2" pageNumber="113">CUWM 57</td>
<td box="[1029,1318,289,308]" gridcol="2" gridrow="2" pageId="2" pageNumber="113">Right m/3</td>
</tr>
<tr box="[137,1318,324,343]" gridrow="3" pageId="2" pageNumber="113">
<th box="[137,455,324,343]" gridcol="0" gridrow="3" pageId="2" pageNumber="113">WM 05-21</th>
<td box="[583,904,324,343]" gridcol="1" gridrow="3" pageId="2" pageNumber="113">CUWM 59</td>
<td box="[1029,1318,324,343]" gridcol="2" gridrow="3" pageId="2" pageNumber="113">Right M2/</td>
</tr>
<tr box="[137,1318,359,378]" gridrow="4" pageId="2" pageNumber="113">
<th box="[137,455,359,378]" gridcol="0" gridrow="4" pageId="2" pageNumber="113">WM 06-14</th>
<td box="[583,904,359,378]" gridcol="1" gridrow="4" pageId="2" pageNumber="113">CUWM 63</td>
<td box="[1029,1318,359,378]" gridcol="2" gridrow="4" pageId="2" pageNumber="113">Left mandible d/4m/2</td>
</tr>
<tr box="[137,1318,394,413]" gridrow="5" pageId="2" pageNumber="113">
<th box="[137,455,394,413]" gridcol="0" gridrow="5" pageId="2" pageNumber="113">WM 06-49</th>
<td box="[583,904,394,413]" gridcol="1" gridrow="5" pageId="2" pageNumber="113">CUWM 106</td>
<td box="[1029,1318,394,413]" gridcol="2" gridrow="5" pageId="2" pageNumber="113">Left mandible, p/4m/3</td>
</tr>
<tr box="[137,1318,429,448]" gridrow="6" pageId="2" pageNumber="113">
<th box="[137,455,429,448]" gridcol="0" gridrow="6" pageId="2" pageNumber="113">WM 06-55</th>
<td box="[583,904,429,448]" gridcol="1" gridrow="6" pageId="2" pageNumber="113">CUWM 121</td>
<td box="[1029,1318,429,448]" gridcol="2" gridrow="6" pageId="2" pageNumber="113">Right mandible, m/3</td>
</tr>
<tr box="[137,1318,463,482]" gridrow="7" pageId="2" pageNumber="113">
<th box="[137,455,463,482]" gridcol="0" gridrow="7" pageId="2" pageNumber="113">WM DEC06-11</th>
<td box="[583,904,463,482]" gridcol="1" gridrow="7" pageId="2" pageNumber="113">CUWM 134</td>
<td box="[1029,1318,463,482]" gridcol="2" gridrow="7" pageId="2" pageNumber="113">Right m/1</td>
</tr>
<tr box="[137,1318,498,517]" gridrow="8" pageId="2" pageNumber="113">
<th box="[137,455,498,517]" gridcol="0" gridrow="8" pageId="2" pageNumber="113">97-697</th>
<td box="[583,904,498,517]" gridcol="1" gridrow="8" pageId="2" pageNumber="113">CGM 12612</td>
<td box="[1029,1318,498,517]" gridcol="2" gridrow="8" pageId="2" pageNumber="113">Right mandible, m/2m/3</td>
</tr>
</table>
</paragraph>
<paragraph blockId="2.[129,768,599,1231]" pageId="2" pageNumber="113">
CUWM 239 is a left m/
<quantity box="[422,461,599,624]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="2" pageNumber="113" unit="in" value="3.0">3 in</quantity>
light wear (
<figureCitation box="[589,705,599,624]" captionStart="Text-fig" captionStartId="5.[125,206,1873,1894]" captionTargetBox="[170,1383,1494,1834]" captionTargetId="figure-144@5.[125,1440,1462,1857]" captionTargetPageId="5" captionText="Text-fig. 7. CUWM 239, left m/3 of Diamantohyus africanus from Moghara, Egypt. a: buccal view; b: stereo occlusal views; c: lingual view." figureDoi="http://doi.org/10.5281/zenodo.7167261" httpUri="https://zenodo.org/record/7167261/files/figure.png" pageId="2" pageNumber="113">Text-fig. 7</figureCitation>
). Part of the protoconid is missing and there is a crack across the metaconid, but otherwise the specimen is well preserved. It has a small anterior accessory cusplet and larger median and posterior accessory cusplets. The talonid is comprised of a single cuspid which is in line with the protoconid and hypoconid and it has lingual and buccal precristids. The buccal cingulum is well developed and beaded.
</paragraph>
<paragraph blockId="2.[129,768,599,1231]" pageId="2" pageNumber="113">
Mandible CUWM 261 (
<figureCitation box="[428,543,855,880]" captionStart="Text-fig" captionStartId="6.[147,228,784,805]" captionTargetBox="[154,1431,153,753]" captionTargetId="figure-686@6.[147,1462,151,769]" captionTargetPageId="6" captionText="Text-fig. 8. CUWM 261, right mandible with damaged m/2m/3 of Diamantohyus africanus from Moghara, Egypt. a: lingual view; b: stereo occlusal views; c: buccal view." figureDoi="http://doi.org/10.5281/zenodo.7167263" httpUri="https://zenodo.org/record/7167263/files/figure.png" pageId="2" pageNumber="113">Text-fig. 8</figureCitation>
) is a right mandible preserved from the symphysis to the m/3 but the premolars and first molar are missing and m/2m/3 are damaged. The rear of the symphysis is beneath the p/3 and the mental foramen is in the lower third of the body of the mandible beneath the front of m/1. The body is deep beneath the m/3 but shallows markedly anteriorly.
</paragraph>
<paragraph blockId="2.[129,768,599,1231]" pageId="2" pageNumber="113">
CUWM 360 comprises an edentulous mandible fragment with the roots of m/1 and m/2 and the anterior alveoli of m/3, as well as an isolated m/3 missing part of the anterior lophid, found close by (
<figureCitation box="[303,420,1175,1200]" captionStart="Text-fig" captionStartId="7.[125,206,1354,1375]" captionTargetBox="[140,1427,160,1336]" captionTargetId="figure-170@7.[125,1440,110,1349]" captionTargetPageId="7" captionText="Text-fig. 9. CUWM 360, right mandible fragment and associated m/3 of Diamantohyus africanus from Moghara, Egypt. a: stereo occlusal views; b: buccal view; c: lingual view." figureDoi="http://doi.org/10.5281/zenodo.7167265" httpUri="https://zenodo.org/record/7167265/files/figure.png" pageId="2" pageNumber="113">Text-fig. 9</figureCitation>
). The m/3 is unworn and shows well the beaded buccal cingulum.
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