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<document ID-DOI="http://doi.org/10.5281/zenodo.3811499" ID-GBIF-Dataset="bfd41e0f-ff57-423f-910b-8fa72710ac7f" ID-GBIF-Taxon="163420833" ID-Zenodo-Dep="3811499" approvalRequired="4" approvalRequired_for_document="1" approvalRequired_for_matCits="1" approvalRequired_for_textStreams="2" checkinTime="1585803663184" checkinUser="plazi" docAuthor="Schmidt-Rhaesa, Andreas &amp; Vieler, Vera" docDate="2020" docId="03E4B37EFF8DFFF2ACBEFB0BFA96F97A" docLanguage="en" docName="zootaxa.4759.2.11.pdf" docOrigin="Zootaxa 4759 (2)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@7a34b7b8" docStyleName="zootaxa.2013.journal_article" docTitle="Spadella kappae Schmidt-Rhaesa &amp; Vieler 2018, sp. nov." docType="treatment" docVersion="5" lastPageNumber="300" masterDocId="FFDDCB06FF8DFFF2AC29FF86FFA7FFC4" masterDocTitle="Validation of Spadella kappae Schmidt-Rhaesa &amp; Vieler, a small benthic chaetognath from Roscoff, France (Chaetognatha)" masterLastPageNumber="300" masterPageNumber="300" pageId="0" pageNumber="300" updateTime="1643450110342" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Validation of Spadella kappae Schmidt-Rhaesa &amp; Vieler, a small benthic chaetognath from Roscoff, France (Chaetognatha)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schmidt-Rhaesa, Andreas</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vieler, Vera</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-04-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>4759</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>300</mods:start>
<mods:end>300</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">https://doi.org/10.11646/zootaxa.4759.2.12</mods:identifier>
<mods:identifier type="GBIF-Dataset">bfd41e0f-ff57-423f-910b-8fa72710ac7f</mods:identifier>
<mods:identifier type="Zenodo-Dep">3737391</mods:identifier>
<mods:identifier type="ZooBank">1BB61461-70FF-4945-AD67-8137A7F134D6</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3811499" ID-GBIF-Taxon="163420833" ID-Zenodo-Dep="3811499" LSID="urn:lsid:plazi:treatment:03E4B37EFF8DFFF2ACBEFB0BFA96F97A" httpUri="http://treatment.plazi.org/id/03E4B37EFF8DFFF2ACBEFB0BFA96F97A" lastPageNumber="300" pageId="0" pageNumber="300">
<subSubSection box="[151,450,1165,1192]" pageId="0" pageNumber="300" type="nomenclature">
<paragraph blockId="0.[151,1437,1130,1726]" box="[151,450,1165,1192]" pageId="0" pageNumber="300">
<heading bold="true" box="[151,450,1165,1192]" fontSize="11" level="1" pageId="0" pageNumber="300" reason="1">
<emphasis bold="true" box="[151,450,1165,1192]" pageId="0" pageNumber="300">
<taxonomicName authority="Schmidt-Rhaesa &amp; Vieler, 2020" authorityName="Schmidt-Rhaesa &amp; Vieler" authorityYear="2018" box="[151,349,1165,1192]" class="Sagittoidea" family="Spadellidae" genus="Spadella" kingdom="Animalia" order="Phragmophora" pageId="0" pageNumber="300" phylum="Chaetognatha" rank="species" species="kappae" status="sp. nov.">
<emphasis bold="true" box="[151,349,1165,1192]" italics="true" pageId="0" pageNumber="300">Spadella kappae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[357,450,1166,1192]" pageId="0" pageNumber="300" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,992,1197,1222]" pageId="0" pageNumber="300" type="reference_group">
<paragraph blockId="0.[151,1437,1130,1726]" box="[151,992,1197,1222]" pageId="0" pageNumber="300">
<taxonomicName authority="Schmidt-Rhaesa &amp; Vieler, 2018: 257 - 265" authorityName="Schmidt-Rhaesa &amp; Vieler" authorityPageNumber="257 - 265" authorityYear="2018" box="[151,753,1197,1222]" class="Sagittoidea" family="Spadellidae" genus="Spadella" kingdom="Animalia" order="Phragmophora" pageId="0" pageNumber="300" phylum="Chaetognatha" rank="species" species="kappae">
<emphasis box="[151,319,1198,1222]" italics="true" pageId="0" pageNumber="300">Spadella kappae</emphasis>
Schmidt-Rhaesa &amp; Vieler, 2018: 257–265
</taxonomicName>
, figs 1–5 [unavailable].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="300" type="materials_examined">
<paragraph blockId="0.[151,1437,1130,1726]" pageId="0" pageNumber="300">
<emphasis bold="true" box="[151,309,1234,1258]" pageId="0" pageNumber="300">Type material</emphasis>
.
<materialsCitation ID-GBIF-Occurrence="2594759836" box="[323,888,1234,1258]" collectionCode="ZMH" pageId="0" pageNumber="300" specimenCode="C861" specimenCount="1" typeStatus="holotype">
<typeStatus box="[323,426,1234,1258]" pageId="0" pageNumber="300">Holotype</typeStatus>
: adult specimen, on SEM stub,
<collectionCode box="[762,821,1234,1257]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="0" pageNumber="300" type="Museum">ZMH</collectionCode>
<specimenCode box="[827,888,1234,1258]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="0" pageNumber="300" type="Museum">C861</specimenCode>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2594759835" box="[898,1432,1234,1258]" collectionCode="ZMH" pageId="0" pageNumber="300" specimenCode="C859" specimenCount="3" typeStatus="paratype">
<typeStatus box="[898,1006,1234,1258]" pageId="0" pageNumber="300">Paratypes</typeStatus>
:
<specimenCount box="[1016,1147,1234,1258]" pageId="0" pageNumber="300" type="generic">3 specimens</specimenCount>
on SEM stub,
<collectionCode box="[1305,1364,1234,1257]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="0" pageNumber="300" type="Museum">ZMH</collectionCode>
<specimenCode box="[1371,1432,1234,1258]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="0" pageNumber="300" type="Museum">C859</specimenCode>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2594759834" box="[151,569,1270,1294]" collectionCode="ZMH" pageId="0" pageNumber="300" specimenCode="C860" specimenCount="5" typeStatus="paratype">
<specimenCount box="[151,282,1270,1294]" pageId="0" pageNumber="300" type="generic">5 specimens</specimenCount>
on SEM stub,
<collectionCode box="[442,501,1270,1293]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="0" pageNumber="300" type="Museum">ZMH</collectionCode>
<specimenCode box="[507,569,1270,1294]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="0" pageNumber="300" type="Museum">C860</specimenCode>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2594759832" box="[579,985,1270,1294]" collectionCode="ZMH" pageId="0" pageNumber="300" specimenCode="C862" specimenCount="1" typeStatus="paratype">
<specimenCount box="[579,700,1270,1294]" pageId="0" pageNumber="300" type="generic">1 specimen</specimenCount>
on SEM stub,
<collectionCode box="[859,918,1270,1293]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="0" pageNumber="300" type="Museum">ZMH</collectionCode>
<specimenCode box="[925,985,1270,1294]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="0" pageNumber="300" type="Museum">C862</specimenCode>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2594759837" collectionCode="ZMH" pageId="0" pageNumber="300" specimenCode="C863" specimenCount="1" typeStatus="paratype">
<specimenCount box="[996,1117,1270,1294]" pageId="0" pageNumber="300" type="generic">1 specimen</specimenCount>
on microscopical slide,
<collectionCode box="[1377,1436,1270,1293]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="0" pageNumber="300" type="Museum">ZMH</collectionCode>
<specimenCode box="[151,212,1306,1330]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="0" pageNumber="300" type="Museum">C863</specimenCode>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2594759833" collectorName="All" elevation="20" latitude="48.82453" location="Marine Biological Station in Roscoff" longLatPrecision="1" longitude="-4.1622167" pageId="0" pageNumber="300" specimenCount="1" typeStatus="paratype">
<collectorName box="[222,256,1306,1330]" pageId="0" pageNumber="300">All</collectorName>
specimens from same locations: “amphioxus-sand” at
<elevation box="[853,975,1306,1330]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="300" unit="m" value="20.0">
<quantity box="[853,909,1306,1330]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="300" unit="m" value="20.0">20 m</quantity>
depth
</elevation>
(
<geoCoordinate box="[991,1147,1306,1330]" degrees="48" direction="north" minutes="43.3450" orientation="latitude" pageId="0" pageNumber="300" precision="1" value="48.722416">48°43.3450´N</geoCoordinate>
,
<geoCoordinate box="[1160,1319,1306,1330]" degrees="003" direction="west" minutes="50.802" orientation="longitude" pageId="0" pageNumber="300" precision="1" value="-3.8467">003°50.802´W</geoCoordinate>
) and “trou aux Raies” at
<elevation box="[300,420,1342,1366]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="0" pageNumber="300" unit="m" value="70.0">
<quantity box="[300,354,1342,1366]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="0" pageNumber="300" unit="m" value="70.0">70 m</quantity>
depth
</elevation>
(
<geoCoordinate box="[434,577,1342,1366]" degrees="48" direction="north" minutes="49.472" orientation="latitude" pageId="0" pageNumber="300" precision="1" value="48.82453">48°49.472´N</geoCoordinate>
,
<geoCoordinate box="[588,750,1342,1366]" degrees="004" direction="west" minutes="09.733" orientation="longitude" pageId="0" pageNumber="300" precision="1" value="-4.1622167">004°09.733´W</geoCoordinate>
), both off the
<location LSID="urn:lsid:plazi:treatment:03E4B37EFF8DFFF2ACBEFB0BFA96F97A:8E9254B3FF8DFFF2AFAAFAB8FAB5FA92" box="[899,1298,1342,1366]" latitude="48.82453" longLatPrecision="1" longitude="-4.1622167" name="Marine Biological Station in Roscoff" pageId="0" pageNumber="300">Marine Biological Station in Roscoff</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="300" type="diagnosis">
<paragraph blockId="0.[151,1437,1130,1726]" pageId="0" pageNumber="300">
<emphasis bold="true" box="[199,309,1378,1402]" pageId="0" pageNumber="300">Diagnosis</emphasis>
. Length
<quantity box="[406,560,1378,1402]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="3.5" metricValueMin="1.7" pageId="0" pageNumber="300" unit="mm" value="2.6" valueMax="3.5" valueMin="1.7">1.7 to 3.5 mm</quantity>
. Septum at approximately 50% of total body length. Paired and completely rayed lateral fins and unpaired tail fin, lateral fins continuing ventral to seminal vesicles. Collarette tissue abundant. 11 to 12 grasping spines per side. Fine structure with ridges and denticles. 3 to 4 teeth in anterior region of vestibulum, each tooth with four longitudinal serrated ridges and tip with four peaks. Ventral ganglion 40.2–63.1% of trunk length. Dorsal and lateral side covered by groups of cilia, adhesive papillae on ventral side. Kidney-shaped corona ciliata. Intestine without diverticula. Testes are not visible. Ovary 60% of trunk length, receptacula seminis present, paired seminal vesicles oval, in the last third of tail.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="300" type="etymology">
<paragraph blockId="0.[151,1437,1130,1726]" pageId="0" pageNumber="300">
<emphasis bold="true" box="[199,322,1630,1654]" pageId="0" pageNumber="300">Etymology</emphasis>
. Named
<taxonomicName authorityName="Schmidt-Rhaesa &amp; Vieler" authorityYear="2018" box="[421,498,1630,1654]" class="Sagittoidea" family="Spadellidae" genus="Spadella" kingdom="Animalia" order="Phragmophora" pageId="0" pageNumber="300" phylum="Chaetognatha" rank="species" species="kappae">
<emphasis box="[421,498,1630,1654]" italics="true" pageId="0" pageNumber="300">kappae</emphasis>
</taxonomicName>
, after Helga Kapp, to acknowledge her contributions to the study of Chaetognatha and for giving valuable advice for the description of this species.
</paragraph>
</subSubSection>
<subSubSection box="[199,1329,1702,1726]" pageId="0" pageNumber="300" type="discussion">
<paragraph blockId="0.[151,1437,1130,1726]" box="[199,1329,1702,1726]" pageId="0" pageNumber="300">
<emphasis bold="true" box="[199,303,1702,1726]" pageId="0" pageNumber="300">Remarks</emphasis>
. A full description and figures of
<taxonomicName authorityName="Schmidt-Rhaesa &amp; Vieler" authorityYear="2018" box="[663,840,1702,1726]" class="Sagittoidea" family="Spadellidae" genus="Spadella" kingdom="Animalia" order="Phragmophora" pageId="0" pageNumber="300" phylum="Chaetognatha" rank="species" species="kappae">
<emphasis box="[663,840,1702,1726]" italics="true" pageId="0" pageNumber="300">Spadella kappae</emphasis>
</taxonomicName>
is given by Schmidt-Rhaesa &amp; Vieler (2018).
</paragraph>
</subSubSection>
</treatment>
</document>
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