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<document id="568B9D242307BFF7AC39D23A905FBE83" ID-CLB-Dataset="6332" ID-DOI="10.11646/phytotaxa.7.1.7" ID-GBIF-Dataset="a402e118-9c07-463b-9281-fcb13c7745a8" ID-ISSN="1179-3163" ID-Zenodo-Dep="4893512" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621603564933" checkinUser="felipe" docAuthor="Yesilyurt, Jovita C. &amp; Schneider, Harald" docDate="2010" docId="03DF1874FF8BFFCCFF5081ADFA75FBF0" docLanguage="en" docName="Phytotaxa.7.52-59.pdf" docOrigin="Phytotaxa 7" docStyle="DocumentStyle:E765CC7F3D29AC31B7D5F874DD99C2DD.4:Phytotaxa.2009-2010.journal_article" docStyleId="E765CC7F3D29AC31B7D5F874DD99C2DD" docStyleName="Phytotaxa.2009-2010.journal_article" docStyleVersion="4" docTitle="Calciphilopteris Yesilyurt &amp; H. Schneider 2010, gen. nov." docType="treatment" docVersion="6" lastPageNumber="55" masterDocId="FFE6600CFF8AFFCFFFC78231FFE9FFD4" masterDocTitle="The new fern genus Calciphilopteris (Pteridaceae)" masterLastPageNumber="59" masterPageNumber="52" pageNumber="53" updateTime="1698940832469" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E20F1FEFD2B60FAB4C66B5A791318A59">The new fern genus Calciphilopteris (Pteridaceae)</mods:title>
</mods:titleInfo>
<mods:name id="1A6EC957D9AB2C7B8F245C79BC7960E2" type="personal">
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<mods:namePart id="BA8006A9AF9908F02D362A7F99B2CE05">Yesilyurt, Jovita C.</mods:namePart>
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<mods:namePart id="2557BB5CAFA1ADEE0B0404AE20B72F7A">Schneider, Harald</mods:namePart>
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<mods:title id="6B119FC65ECA23F9CD5417C11B0CD3AE">Phytotaxa</mods:title>
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<mods:date id="D86B6318F5E0102CC4C08C7C24794B05">2010</mods:date>
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<mods:number id="A521AADE38B673378EBA0D9ABAD37CAE">2010-07-15</mods:number>
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<treatment id="03DF1874FF8BFFCCFF5081ADFA75FBF0" ID-DOI="http://doi.org/10.5281/zenodo.4893474" ID-GBIF-Taxon="182484943" ID-Zenodo-Dep="4893474" LSID="urn:lsid:plazi:treatment:03DF1874FF8BFFCCFF5081ADFA75FBF0" httpUri="http://treatment.plazi.org/id/03DF1874FF8BFFCCFF5081ADFA75FBF0" lastPageId="3" lastPageNumber="55" pageId="1" pageNumber="53">
<subSubSection id="C36CFAE9FF8BFFCEFF5081ADFC4FFC62" box="[151,934,923,950]" pageId="1" pageNumber="53" type="nomenclature">
<paragraph id="8BC9A962FF8BFFCEFF5081ADFC4FFC62" blockId="1.[151,934,923,950]" box="[151,934,923,950]" pageId="1" pageNumber="53">
<heading id="D0811E0EFF8BFFCEFF5081ADFC4FFC62" box="[151,934,923,950]" fontSize="11" level="2" pageId="1" pageNumber="53" reason="3">
<emphasis id="B9027570FF8BFFCEFF5081ADFCEDFC62" bold="true" box="[151,772,923,950]" pageId="1" pageNumber="53">
<taxonomicName id="4C76D2E1FF8BFFCEFF5081ADFD65FC61" ID-CoL="3F44" authority="Yesilyurt &amp; H.Schneider" authorityName="Yesilyurt &amp; H. Schneider" authorityYear="2010" box="[151,652,923,950]" class="Polypodiopsida" family="Pteridaceae" genus="Calciphilopteris" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="53" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="B9027570FF8BFFCEFF5081ADFEBAFC62" bold="true" box="[151,339,924,950]" italics="true" pageId="1" pageNumber="53">Calciphilopteris</emphasis>
Yesilyurt &amp; H.Schneider
</taxonomicName>
,
<emphasis id="B9027570FF8BFFCEFD5E81ADFCEDFC62" bold="true" box="[665,772,924,950]" italics="true" pageId="1" pageNumber="53">
<taxonomicNameLabel id="A231C80BFF8BFFCEFD5E81ADFCEDFC62" box="[665,772,924,950]" pageId="1" pageNumber="53" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</emphasis>
<figureCitation id="134DB5E7FF8BFFCEFCE381ADFC4FFC62" box="[804,934,924,950]" captionStart="FIGURE 1" captionStartId="2.[151,255,1674,1698]" captionTargetBox="[195,1395,307,1652]" captionTargetId="figure-134@2.[164,1433,269,1705]" captionTargetPageId="2" captionText="FIGURE 1. Sketches of fertile leaves showing their general attributes and morphological variation: A. Calciphilopteris ludens. B. Calciphilopteris papuana. C. Calciphilopteris wallichii." figureDoi="http://doi.org/10.5281/zenodo.4893514" httpUri="https://zenodo.org/record/4893514/files/figure.png" pageId="1" pageNumber="53">Figs. 1AC</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36CFAE9FF8BFFCEFF5081D8FE70FB54" pageId="1" pageNumber="53" type="description">
<paragraph id="8BC9A962FF8BFFCEFF5081D8FE70FB54" blockId="1.[151,1438,1001,1216]" pageId="1" pageNumber="53">
<emphasis id="B9027570FF8BFFCEFF5081D8FE70FB54" italics="true" pageId="1" pageNumber="53">
<taxonomicName id="4C76D2E1FF8BFFCEFF5081D8FEACFBD4" authority="Yesilyurt &amp; Schneider, 2010" authorityName="Yesilyurt &amp; H. Schneider" authorityYear="2010" box="[151,325,1001,1024]" class="Polypodiopsida" family="Pteridaceae" genus="Calciphilopteris" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="53" phylum="Tracheophyta" rank="genus" status="gen. nov.">Calciphilopteris</taxonomicName>
<taxonomicNameLabel id="A231C80BFF8BFFCEFE8B81D8FE44FBD4" box="[332,429,1001,1024]" pageId="1" pageNumber="53" rank="genus">gen. nov.</taxonomicNameLabel>
<taxonomicName id="4C76D2E1FF8BFFCEFE7381D8FD7CFBD4" box="[436,661,1001,1024]" class="Polypodiopsida" family="Pteridaceae" genus="Doryopteridis" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="53" phylum="Tracheophyta" rank="species" species="affinis">Doryopteridis affinis</taxonomicName>
sed rhizomate breviter vel longe repenti, paleis rhizomatum lanceolatis vel anguste lanceolatis clathratis vel semiclathratis, petiolo tereti fasciculum vascularem unum tantum continenti, petiolo laminaque fibrillas (pilos squamiformes) atque squamas non clathratas vel semiclathratas ferentibus, sporangio plerumque longipedicellato 32 sporas continenti, plantis plerumque rupicola locus calcareos incolentibus differt.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36CFAE9FF8BFFCEFF5086B8FD28FB14" pageId="1" pageNumber="53" type="type_taxon">
<paragraph id="8BC9A962FF8BFFCEFF5086B8FD28FB14" blockId="1.[151,1438,1001,1216]" pageId="1" pageNumber="53">
<emphasis id="B9027570FF8BFFCEFF5086B8FECCFB75" bold="true" box="[151,293,1161,1185]" pageId="1" pageNumber="53">Type species</emphasis>
:—
<taxonomicName id="4C76D2E1FF8BFFCEFE9686B8FBFBFB74" ID-CoL="PHN5" authority="(Wall. ex Hook.) Yesilyurt &amp; H. Schneid." authorityName="Yesilyurt &amp; H. Schneid." baseAuthorityName="Wall. ex Hook." box="[337,1042,1161,1184]" class="Polypodiopsida" family="Pteridaceae" genus="Calciphilopteris" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="53" phylum="Tracheophyta" rank="species" species="ludens">
<emphasis id="B9027570FF8BFFCEFE9686B8FDA5FB74" box="[337,588,1161,1184]" italics="true" pageId="1" pageNumber="53">Calciphilopteris ludens</emphasis>
(Wall. ex Hook.) Yesilyurt &amp; H.Schneid.
</taxonomicName>
=
<taxonomicName id="4C76D2E1FF8BFFCEFBF386B8FAF3FB74" ID-CoL="78KSW" authority="Wallich" authorityName="Wallich" box="[1076,1306,1161,1184]" class="Polypodiopsida" family="Pteridaceae" genus="Pteris" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="53" phylum="Tracheophyta" rank="species" species="ludens">
<emphasis id="B9027570FF8BFFCEFBF386B8FB2BFB74" box="[1076,1218,1161,1184]" italics="true" pageId="1" pageNumber="53">Pteris ludens</emphasis>
Wallich
</taxonomicName>
ex.
<bibRefCitation id="EFE7D493FF8BFFCEFA8986B8FEAEFB14" author="Hooker, W. J." pageId="1" pageNumber="53" refId="ref5115" refString="Hooker, W. J. (1858) Species Filicum: being descriptions of the known ferns, particularly of such as exist in the author's herbarium, or are with sufficient accuracy described in work to which he has access, vol. 2. William Pamplin, London, 250 pp." type="book" year="1858">Hooker (1858: 210)</bibRefCitation>
.
<typeStatus id="54CD17C0FF8BFFCEFE968698FE51FB14" box="[337,440,1193,1216]" pageId="1" pageNumber="53" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F361E9F2FF8BFFCEFE048698FDFAFB14" box="[451,531,1193,1216]" name="India" pageId="1" pageNumber="53">INDIA</collectingCountry>
:
<emphasis id="B9027570FF8BFFCEFDDB8698FD67FB14" box="[540,654,1193,1216]" italics="true" pageId="1" pageNumber="53">
<location id="8EA9FFB9FF8BFFCEFDDB8698FD85FB14" LSID="urn:lsid:plazi:treatment:03DF1874FF8BFFCCFF5081ADFA75FBF0:8EA9FFB9FF8BFFCEFDDB8698FD85FB14" box="[540,620,1193,1216]" country="India" name="Wallich" pageId="1" pageNumber="53">Wallich</location>
88
</emphasis>
(K).
</paragraph>
</subSubSection>
<subSubSection id="C36CFAE9FF8BFFCDFF5086C1FBD6FF2B" lastPageId="2" lastPageNumber="54" pageId="1" pageNumber="53" type="description">
<paragraph id="8BC9A962FF8BFFCDFF5086C1FBD6FF2B" blockId="1.[151,1437,1264,2024]" lastBlockId="2.[151,1436,152,255]" lastPageId="2" lastPageNumber="54" pageId="1" pageNumber="53">
Plants terrestrial or rupicolous; essentially growing on limestone.
<emphasis id="B9027570FF8BFFCEFC6986C1FBFDFADD" box="[942,1044,1264,1289]" italics="true" pageId="1" pageNumber="53">Rhizome</emphasis>
slender, short- to long-creeping, scales up to
<quantity id="4C8E0487FF8BFFCEFEEF8727FE99FAE4" box="[296,368,1302,1328]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="53" unit="mm" value="3.0">3 mm</quantity>
, linear to ovate-lanceolate, acuminate to caudate, predominantly bicoloured, with clathrate to semi-clathrate centre, and paler, narrow to broad border, margin entire to dentate towards the apex.
<emphasis id="B9027570FF8BFFCEFA8E870CFA75FA82" box="[1353,1436,1341,1366]" italics="true" pageId="1" pageNumber="53">Petiole</emphasis>
terete to slightly sulcate (towards) at the base of the lamina; one vascular bundle; dark brown, atropurpureous to black, base with scales as those of the rhizome, higher up the petiole with indumentum (generally dense towards the base of the lamina) usually scaly, scales semi- to non-clathrate, with fibrils, rarely with white hairs or glabrous.
<emphasis id="B9027570FF8BFFCEFEED87E9FE9CFA25" box="[298,373,1496,1521]" italics="true" pageId="1" pageNumber="53">Sterile</emphasis>
and
<emphasis id="B9027570FF8BFFCEFE6987E9FDACFA25" box="[430,581,1496,1521]" italics="true" pageId="1" pageNumber="53">fertile fronds</emphasis>
dimorphic (architecture of fertile lamina more complex), sub-coriaceous to coriaceous; glabrescent to glabrous, indumentum on the abaxial side with glandular hairs and/or fibrils and non-clathrate scales at the base of the lamina; veins anastomosing, without included free veinlets.
<emphasis id="B9027570FF8BFFCEFA8A8414FF0DF9B1" italics="true" pageId="1" pageNumber="53">Sterile fronds</emphasis>
up to
<quantity id="4C8E0487FF8BFFCEFEF2847DFE6CF9B2" box="[309,389,1612,1638]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="53" unit="cm" value="45.0">45 cm</quantity>
; lamina up to
<quantity id="4C8E0487FF8BFFCEFDFF847DFD61F9B2" box="[568,648,1612,1638]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="53" unit="cm" value="17.0">17 cm</quantity>
, ranging from simple, entire or 3-lobed to pedate; oblong-oval, cordate, sagittate, hastate, lobed to pentagonal in outline; pinnae/segments12(4) pairs; the margins usually with a whitish or brown cartilagineous border, entire; hydathodes generally present on the upper surface; veins ending free near the margin (occasionally confluent in
<taxonomicName id="4C76D2E1FF8BFFCEFCE384F1FC76F90D" box="[804,927,1728,1753]" class="Polypodiopsida" family="Pteridaceae" genus="Calciphilopteris" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="53" phylum="Tracheophyta" rank="species" species="allenae">
<emphasis id="B9027570FF8BFFCEFCE384F1FC76F90D" box="[804,927,1728,1753]" italics="true" pageId="1" pageNumber="53">C. allenae</emphasis>
</taxonomicName>
); basal pinnae lobed basiscopically or not.
<emphasis id="B9027570FF8BFFCEFF5084D6FEABF8D4" box="[151,322,1767,1792]" italics="true" pageId="1" pageNumber="53">Fertile fronds</emphasis>
up to
<quantity id="4C8E0487FF8BFFCEFE5284D7FE0CF8D4" box="[405,485,1766,1792]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="53" unit="cm" value="65.0">65 cm</quantity>
; lamina up to 20(25) cm; sagittate, hastate, pentagonal to broad-cordate or suborbicular in outline, 3-lobed to pedate, pedate-pinnatifid (pedate-2-pinnatifid); pinnae/segments 25(6) pairs; ascending to patent, deltoid to long-lanceolate, acute or acuminate, predominantly with entire margins; basal pinna/segment with acroscopic and/or with basiscopic segments/lobes; first segment/lobe divided or not, falcate to lanceolate.
<emphasis id="B9027570FF8BFFCEFE5485B0FE2DF84E" box="[403,452,1921,1946]" italics="true" pageId="1" pageNumber="53">Sori</emphasis>
marginal, usually on a continuous vascular commissure (connecting the vein-tips), rarely interrupted at sinuses, covered by a continuous modified marginal indusium, without paraphyses; receptacle and indusium continuous, including in the sinuses; indusium with margin entire to slightly eroded.
<emphasis id="B9027570FF88FFCDFF5082A9FECEFF65" box="[151,295,152,177]" italics="true" pageId="2" pageNumber="54">Sporangium</emphasis>
with pedicel 12(2.5) times the capsule length; capsule without basal cell; annulus with 1734 indurate cells; with 32 spores per sporangium.
<emphasis id="B9027570FF88FFCDFD1C828EFCC5FF0C" box="[731,812,191,216]" italics="true" pageId="2" pageNumber="54">Spores</emphasis>
trilete, tetrahedral to globose, echinate or cristate, without equatorial flange. Chromosome number n = 29, 30 or multiples thereof.
</paragraph>
</subSubSection>
<caption id="DF09F9EAFF88FFCDFF5084BBFC88F910" ID-DOI="http://doi.org/10.5281/zenodo.4893514" ID-Zenodo-Dep="4893514" httpUri="https://zenodo.org/record/4893514/files/figure.png" pageId="2" pageNumber="54" startId="2.[151,255,1674,1698]" targetBox="[195,1395,307,1652]" targetPageId="2">
<paragraph id="8BC9A962FF88FFCDFF5084BBFC88F910" blockId="2.[151,1435,1674,1733]" pageId="2" pageNumber="54">
<emphasis id="B9027570FF88FFCDFF5084BBFEF3F976" bold="true" box="[151,282,1674,1698]" pageId="2" pageNumber="54">FIGURE 1.</emphasis>
Sketches of fertile leaves showing their general attributes and morphological variation:
<emphasis id="B9027570FF88FFCDFB0A84BBFB0EF976" bold="true" box="[1229,1255,1674,1698]" pageId="2" pageNumber="54">A.</emphasis>
<taxonomicName id="4C76D2E1FF88FFCDFB2984BAFF34F910" authorityName="Yesilyurt &amp; H. Schneider" authorityYear="2010" baseAuthorityName="Yesilyurt &amp; H. Schneider" baseAuthorityYear="2010" class="Polypodiopsida" family="Pteridaceae" genus="Calciphilopteris" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="54" phylum="Tracheophyta" rank="species" species="ludens">
<emphasis id="B9027570FF88FFCDFB2984BAFF34F910" italics="true" pageId="2" pageNumber="54">Calciphilopteris ludens</emphasis>
</taxonomicName>
.
<emphasis id="B9027570FF88FFCDFF2D849CFF15F911" bold="true" box="[234,252,1709,1733]" pageId="2" pageNumber="54">B</emphasis>
.
<taxonomicName id="4C76D2E1FF88FFCDFECE849CFDF3F910" authorityName="Yesilyurt &amp; H. Schneider" authorityYear="2010" baseAuthorityName="Yesilyurt &amp; H. Schneider" baseAuthorityYear="2010" box="[265,538,1709,1732]" class="Polypodiopsida" family="Pteridaceae" genus="Calciphilopteris" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="54" phylum="Tracheophyta" rank="species" species="papuana">
<emphasis id="B9027570FF88FFCDFECE849CFDF3F910" box="[265,538,1709,1732]" italics="true" pageId="2" pageNumber="54">Calciphilopteris papuana</emphasis>
</taxonomicName>
.
<emphasis id="B9027570FF88FFCDFDE0849CFDD3F911" bold="true" box="[551,570,1709,1733]" pageId="2" pageNumber="54">C</emphasis>
.
<taxonomicName id="4C76D2E1FF88FFCDFD8F849CFCB4F910" authorityName="Yesilyurt &amp; H. Schneider" authorityYear="2010" baseAuthorityName="Yesilyurt &amp; H. Schneider" baseAuthorityYear="2010" box="[584,861,1709,1732]" class="Polypodiopsida" family="Pteridaceae" genus="Calciphilopteris" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="54" phylum="Tracheophyta" rank="species" species="wallichii">Calciphilopteris wallichii</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C36CFAE9FF88FFCDFF01852CFB69F8E3" box="[198,1152,1821,1847]" pageId="2" pageNumber="54" type="distribution">
<paragraph id="8BC9A962FF88FFCDFF01852CFB69F8E3" blockId="2.[151,1435,1821,1963]" box="[198,1152,1821,1847]" pageId="2" pageNumber="54">
<emphasis id="B9027570FF88FFCDFF01852CFEB6F8E3" bold="true" box="[198,351,1821,1847]" pageId="2" pageNumber="54">Distribution</emphasis>
:—Southeast Asia, from Indo-China to New
<collectingCountry id="F361E9F2FF88FFCDFCAB852CFC29F8E3" box="[876,960,1821,1847]" name="Guinea-Bissau" pageId="2" pageNumber="54">Guinea</collectingCountry>
(and
<collectingCountry id="F361E9F2FF88FFCDFBC4852CFB9DF8E3" box="[1027,1140,1821,1847]" name="Australia" pageId="2" pageNumber="54">Australia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C36CFAE9FF88FFCDFF018572FC87F88A" box="[198,878,1859,1886]" pageId="2" pageNumber="54" type="biology_ecology">
<paragraph id="8BC9A962FF88FFCDFF018572FC87F88A" blockId="2.[151,1435,1821,1963]" box="[198,878,1859,1886]" pageId="2" pageNumber="54">
<emphasis id="B9027570FF88FFCDFF018572FE58F889" bold="true" box="[198,433,1859,1885]" pageId="2" pageNumber="54">Habitat &amp; Ecology</emphasis>
:—exclusively on limestone outcrops.
</paragraph>
</subSubSection>
<subSubSection id="C36CFAE9FF88FFCDFF01855BFB0FF87F" pageId="2" pageNumber="54" type="etymology">
<paragraph id="8BC9A962FF88FFCDFF01855BFB0FF87F" blockId="2.[151,1435,1821,1963]" pageId="2" pageNumber="54">
<emphasis id="B9027570FF88FFCDFF01855BFEA4F850" bold="true" box="[198,333,1898,1924]" pageId="2" pageNumber="54">Etymology</emphasis>
:—the genus name is a composite from the Greek
<emphasis id="B9027570FF88FFCDFC65855AFC3AF850" box="[930,979,1899,1924]" italics="true" pageId="2" pageNumber="54">calx</emphasis>
(limestone),
<emphasis id="B9027570FF88FFCDFBB7855AFB51F850" box="[1136,1208,1899,1924]" italics="true" pageId="2" pageNumber="54">philus</emphasis>
(loving) and
<emphasis id="B9027570FF88FFCDFA9E855AFA72F850" box="[1369,1435,1899,1924]" italics="true" pageId="2" pageNumber="54">pteris</emphasis>
(fern) and emphasizes the exclusivity of limestone outcrops as the natural habitat for this fern.
</paragraph>
</subSubSection>
<subSection id="E2F9B089FF89FFCCFF5082A6FA75FBF0" pageId="3" pageNumber="55" type="multiple">
<subSubSection id="C36CFAE9FF89FFCCFF5082A6FCFEFF67" box="[151,791,151,180]" pageId="3" pageNumber="55" type="multiple">
<paragraph id="8BC9A962FF89FFCCFF5082A6FCFEFF67" blockId="3.[151,791,151,180]" box="[151,791,151,180]" pageId="3" pageNumber="55">
<heading id="D0811E0EFF89FFCCFF5082A6FCFEFF67" bold="true" box="[151,791,151,180]" fontSize="12" level="1" pageId="3" pageNumber="55" reason="1">
<emphasis id="B9027570FF89FFCCFF5082A6FCFEFF67" bold="true" box="[151,791,151,180]" pageId="3" pageNumber="55">
Diagnostic key to the species of
<taxonomicName id="4C76D2E1FF89FFCCFD8282A6FCFEFF67" authorityName="Yesilyurt &amp; H. Schneider" authorityYear="2010" box="[581,791,151,179]" class="Polypodiopsida" family="Pteridaceae" genus="Calciphilopteris" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="55" phylum="Tracheophyta" rank="genus">
<emphasis id="B9027570FF89FFCCFD8282A6FCFEFF67" bold="true" box="[581,791,151,179]" italics="true" pageId="3" pageNumber="55">Calciphilopteris</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36CFAE9FF89FFCCFF5082D5FA75FBF0" pageId="3" pageNumber="55" type="key">
<key id="FFE7FE53FF89FFCCFF5082D5FA75FBF0" pageId="3" pageNumber="55">
<keyStep id="3082B1C7FF89FFCCFF5082D5FA72FE28" pageId="3" pageNumber="55">
<paragraph id="8BC9A962FF89FFCCFF5082D5FA75FEBD" blockId="3.[151,1436,228,1060]" pageId="3" pageNumber="55">
<keyLead id="30870A57FF89FFCCFF5082D5FA75FEBD" pageId="3" pageNumber="55">
1. Basal pinna/segment with 24 basiscopic segments, all ascendant; first basiscopic segment with 13 lobes (
<figureCitation id="134DB5E7FF89FFCCFAFB82D5FA64FF2F" box="[1340,1421,228,251]" captionStart="FIGURE 1" captionStartId="2.[151,255,1674,1698]" captionTargetBox="[195,1395,307,1652]" captionTargetId="figure-134@2.[164,1433,269,1705]" captionTargetPageId="2" captionText="FIGURE 1. Sketches of fertile leaves showing their general attributes and morphological variation: A. Calciphilopteris ludens. B. Calciphilopteris papuana. C. Calciphilopteris wallichii." figureDoi="http://doi.org/10.5281/zenodo.4893514" httpUri="https://zenodo.org/record/4893514/files/figure.png" pageId="3" pageNumber="55">Fig. 1B</figureCitation>
), conspicuous or not, ascendant; sinus (between pinnae/segments) usually narrow, acute or round (seldom slightly rectangular); fertile lamina pedate-pinnatifid to pedate-2(3)-pinnatifid, broadly ovate to cordate-ovate, pinnae/segments 37 (
<figureCitation id="134DB5E7FF89FFCCFE848363FE78FEBD" box="[323,401,338,361]" captionStart="FIGURE 1" captionStartId="2.[151,255,1674,1698]" captionTargetBox="[195,1395,307,1652]" captionTargetId="figure-134@2.[164,1433,269,1705]" captionTargetPageId="2" captionText="FIGURE 1. Sketches of fertile leaves showing their general attributes and morphological variation: A. Calciphilopteris ludens. B. Calciphilopteris papuana. C. Calciphilopteris wallichii." figureDoi="http://doi.org/10.5281/zenodo.4893514" httpUri="https://zenodo.org/record/4893514/files/figure.png" pageId="3" pageNumber="55">Fig 1A</figureCitation>
), all equally ascendant, long lanceolate to linear lanceolate (except basal pinna)
<emphasis id="B9027570FF89FFCCFB238363FA75FEBD" box="[1252,1436,338,361]" italics="true" pageId="3" pageNumber="55">
........
<taxonomicName id="4C76D2E1FF89FFCCFAE78363FA75FEBD" ID-CoL="PHN6" authorityName="Yesilyurt &amp; H. Schneider" authorityYear="2010" baseAuthorityName="Yesilyurt &amp; H. Schneider" baseAuthorityYear="2010" box="[1312,1436,338,361]" class="Polypodiopsida" family="Pteridaceae" genus="Calciphilopteris" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="55" phylum="Tracheophyta" rank="species" species="papuana">C. papuana</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC9A962FF89FFCCFF508346FA72FE28" blockId="3.[151,1436,228,1060]" pageId="3" pageNumber="55">
<keyLead id="30870A57FF89FFCCFF508346FA72FE28" pageId="3" pageNumber="55">- Basal pinnae/segment with 13 basiscopic segments, ascendant or not; first segment very rarely lobed; sinus (between pinnae/segments) rounded to slightly rectangular; fertile lamina pedate to pedate-pinnatifid, hastate to trilobed, ovate-lanceolate, slightly pentagonal or obovate, pinnae/segments 14, all equally ascendant or not, usually lanceolate (except basal one) ........................................................................................................................................ 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="3082B1C7FF89FFCCFF50803BFA75FD0D" pageId="3" pageNumber="55">
<paragraph id="8BC9A962FF89FFCCFF50803BFA75FDBF" blockId="3.[151,1436,228,1060]" pageId="3" pageNumber="55">
<keyLead id="30870A57FF89FFCCFF50803BFA75FDBF" pageId="3" pageNumber="55">
2. Scales of rhizome and base of petiole long lanceolate to linear lanceolate, caudate, margin (half way towards apex) dentate; fertile lamina generally sagittate to trilobed (rarely pedate), hydathodes absent; marginal veins regularly anastomosing along the leaf margin
<emphasis id="B9027570FF89FFCCFD8D8065FA75FDBF" box="[586,1436,596,619]" italics="true" pageId="3" pageNumber="55">
.............................................................................................................
<taxonomicName id="4C76D2E1FF89FFCCFAE08065FA75FDBF" ID-CoL="PHN4" authorityName="Yesilyurt &amp; H. Schneider" authorityYear="2010" baseAuthorityName="Yesilyurt &amp; H. Schneider" baseAuthorityYear="2010" box="[1319,1436,596,619]" class="Polypodiopsida" family="Pteridaceae" genus="Calciphilopteris" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="55" phylum="Tracheophyta" rank="species" species="alleniae">C. alleniae</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC9A962FF89FFCCFF508049FA75FD0D" blockId="3.[151,1436,228,1060]" pageId="3" pageNumber="55">
<keyLead id="30870A57FF89FFCCFF508049FA75FD0D" pageId="3" pageNumber="55">- Scales of rhizome and base of petiole lanceolate to long lanceolate, not caudate, margin (half way towards apex) remotely dentate to entire; fertile lamina hastate, pedate to pedate-bipinnatifid, hydathodes present (sometimes obscure); marginal veins ending free along the leaf margin ......................................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="3082B1C7FF89FFCCFF5080D6FA75FBF0" pageId="3" pageNumber="55">
<paragraph id="8BC9A962FF89FFCCFF5080D6FA7DFCB8" blockId="3.[151,1436,228,1060]" pageId="3" pageNumber="55">
<keyLead id="30870A57FF89FFCCFF5080D6FA7DFCB8" pageId="3" pageNumber="55">
3. Basal pinnae with acroscopic and basiscopic segments; scales margin (half way towards apex) entire; fertile lamina commonly pedate-pinnatifid to pedate-2-pinnatifid, pentagonal, pinnae/segments 24, all equally ascendant; basal pinna/segment with 1(2) acroscopic segment and 23 basiscopic segments/lobes (
<figureCitation id="134DB5E7FF89FFCCFBF08101FB63FC93" box="[1079,1162,816,839]" captionStart="FIGURE 1" captionStartId="2.[151,255,1674,1698]" captionTargetBox="[195,1395,307,1652]" captionTargetId="figure-134@2.[164,1433,269,1705]" captionTargetPageId="2" captionText="FIGURE 1. Sketches of fertile leaves showing their general attributes and morphological variation: A. Calciphilopteris ludens. B. Calciphilopteris papuana. C. Calciphilopteris wallichii." figureDoi="http://doi.org/10.5281/zenodo.4893514" httpUri="https://zenodo.org/record/4893514/files/figure.png" pageId="3" pageNumber="55">Fig. 1A</figureCitation>
), all ascendant; first basiscopic segment/lobe generally unlobed
<emphasis id="B9027570FF89FFCCFDAC8164FA7DFCB8" box="[619,1428,853,876]" italics="true" pageId="3" pageNumber="55">
..........................................................................................................
<taxonomicName id="4C76D2E1FF89FFCCFAF78164FA7DFCB8" ID-CoL="PHN5" authorityName="Yesilyurt &amp; H. Schneider" authorityYear="2010" baseAuthorityName="Yesilyurt &amp; H. Schneider" baseAuthorityYear="2010" box="[1328,1428,853,876]" class="Polypodiopsida" family="Pteridaceae" genus="Calciphilopteris" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="55" phylum="Tracheophyta" rank="species" species="ludens">C. ludens</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC9A962FF89FFCCFF50814BFA75FBF0" blockId="3.[151,1436,228,1060]" pageId="3" pageNumber="55">
<keyLead id="30870A57FF89FFCCFF50814BFA75FBF0" pageId="3" pageNumber="55">
- Basal pinnae with acroscopic pinnae only; scales margin (half way towards apex) dentate to occasionally dentate; fertile lamina commonly hastate to pedate (pedate-pinnatifid), ovate-lanceolate, slightly pentagonal, pinnae/segments 12(3), basal pinna/segment ascendant, other segments/lobes patent to slightly ascendant; basal pinna/segment with 1(2) basiscopic segment/lobe, which is usually unlobed and generally parallel to the petiole (
<figureCitation id="134DB5E7FF89FFCCFAE481D9FA9DFC2B" box="[1315,1396,1000,1023]" captionStart="FIGURE 1" captionStartId="2.[151,255,1674,1698]" captionTargetBox="[195,1395,307,1652]" captionTargetId="figure-134@2.[164,1433,269,1705]" captionTargetPageId="2" captionText="FIGURE 1. Sketches of fertile leaves showing their general attributes and morphological variation: A. Calciphilopteris ludens. B. Calciphilopteris papuana. C. Calciphilopteris wallichii." figureDoi="http://doi.org/10.5281/zenodo.4893514" httpUri="https://zenodo.org/record/4893514/files/figure.png" pageId="3" pageNumber="55">Fig. 1C</figureCitation>
) ... ..................................................................................................................................................................
<taxonomicName id="4C76D2E1FF89FFCCFAD8863CFA75FBF0" ID-CoL="PHN7" authorityName="Yesilyurt &amp; H. Schneider" authorityYear="2010" baseAuthorityName="Yesilyurt &amp; H. Schneider" baseAuthorityYear="2010" box="[1311,1436,1037,1060]" class="Polypodiopsida" family="Pteridaceae" genus="Calciphilopteris" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="55" phylum="Tracheophyta" rank="species" species="wallichii">
<emphasis id="B9027570FF89FFCCFAD8863CFA75FBF0" box="[1311,1436,1037,1060]" italics="true" pageId="3" pageNumber="55">C. wallichii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</subSection>
</treatment>
</document>