treatments-xml/data/03/C8/87/03C887E5FFF7FF8EFF12A399FC57F884.xml
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<mods:title id="ADACFF3AAC9FAD8D7D4CCCA3AE9016EB">The genital area of Halacaridae (Acari), life stages and development of morphological characters and implication on the classification</mods:title>
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<mods:namePart id="1B4CF76EB3B461CE489CA1E00722A3EC">Bartsch, Ilse</mods:namePart>
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<paragraph id="8BDE36F3FFF7FF8EFF12A399FDF2FB26" blockId="28.[151,538,1158,1218]" box="[151,538,1158,1184]" pageId="28" pageNumber="229">
<heading id="D096819FFFF7FF8EFF12A399FDF2FB26" bold="true" box="[151,538,1158,1184]" fontSize="11" level="1" pageId="28" pageNumber="229" reason="1">
<taxonomicName id="4C614D70FFF7FF8EFF12A399FDF2FB26" authority="Monniot, 1972" authorityName="Monniot" authorityYear="1972" box="[151,538,1158,1184]" class="Arachnida" family="Halacaridae" genus="Scaptognathides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="229" phylum="Arthropoda" rank="genus">
<emphasis id="B915EAE1FFF7FF8EFF12A399FDF2FB26" bold="true" box="[151,538,1158,1184]" pageId="28" pageNumber="229">
<emphasis id="B915EAE1FFF7FF8EFF12A399FEB5FB26" bold="true" box="[151,349,1158,1184]" italics="true" pageId="28" pageNumber="229">Scaptognathides</emphasis>
<bibRefCitation id="EFF04B02FFF7FF8EFEE0A399FDF2FB26" author="Monniot" box="[357,538,1158,1184]" pageId="28" pageNumber="229" refString="Monniot, F. (1972) Scaptognathides, un nouveau genre d'acariens marins (Halacaridae) dans l'Atlantique ouest. Acarologia, 13, 361 - 366." type="journal article" year="1972">Monniot, 1972</bibRefCitation>
</emphasis>
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<paragraph id="8BDE36F3FFF7FF8EFF12A3B6FEA7FB44" blockId="28.[151,538,1158,1218]" box="[151,335,1193,1218]" pageId="28" pageNumber="229">(Figs 69 and 70)</paragraph>
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<paragraph id="8BDE36F3FFF7FF8EFF12A3EEFD11FA8C" blockId="28.[151,1437,1265,1794]" box="[151,761,1265,1290]" pageId="28" pageNumber="229">
<emphasis id="B915EAE1FFF7FF8EFF12A3EEFEDAFA8C" bold="true" box="[151,306,1265,1290]" pageId="28" pageNumber="229">
<typeStatus id="54DA8851FFF7FF8EFF12A3EEFF3AFA8C" box="[151,210,1265,1290]" pageId="28" pageNumber="229">Type</typeStatus>
species.
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<taxonomicName id="4C614D70FFF7FF8EFEBCA3EEFD1DFA8C" authority="Monniot, 1972" authorityName="Monniot" authorityYear="1972" box="[313,757,1265,1290]" class="Arachnida" family="Halacaridae" genus="Scaptognathides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="B915EAE1FFF7FF8EFEBCA3EEFDACFA8C" box="[313,580,1265,1290]" italics="true" pageId="28" pageNumber="229">Scaptognathides planus</emphasis>
<bibRefCitation id="EFF04B02FFF7FF8EFDCEA3EEFD1DFA8C" author="Monniot" box="[587,757,1265,1290]" pageId="28" pageNumber="229" refString="Monniot, F. (1972) Scaptognathides, un nouveau genre d'acariens marins (Halacaridae) dans l'Atlantique ouest. Acarologia, 13, 361 - 366." type="journal article" year="1972">Monniot, 1972</bibRefCitation>
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.
</paragraph>
<paragraph id="8BDE36F3FFF7FF8EFF42A20AFBFBF9AC" blockId="28.[151,1437,1265,1794]" pageId="28" pageNumber="229">
<emphasis id="B915EAE1FFF7FF8EFF42A20AFEF5FAA8" bold="true" box="[199,285,1301,1326]" pageId="28" pageNumber="229">Adults.</emphasis>
In both female and male genital and anal plates fused. Female GA with two pairs of pgs, and one to zero sgs on each genital sclerite. Two pairs of partly fused internal gac in posterior to middle part of GO (Fig. 69;
<bibRefCitation id="EFF04B02FFF7FF8EFF12A242FEA8FAF0" author="Bartsch" box="[151,320,1373,1398]" pageId="28" pageNumber="229" refString="Bartsch, I. (2003 d) Psammophilous halacarids (Halacaridae: Acari) from Dampier, Western Australia. Description of species and faunal comparison of the mesopsammal halacarid fauna of western and eastern Australia. Records of the Western Australian Museum, 22, 23 - 45." type="journal article" year="2003" yearSuffix="d">Bartsch 2003d</bibRefCitation>
: fig.
<date id="FFDF1033FFF7FF8EFEFFA241FE7DFAF0" box="[378,405,1374,1398]" pageId="28" pageNumber="229">7I</date>
). Ovipositor at rest short, not extending beyond anterior margin of GO; when everted five pairs of lamellar, weakly sclerotized genital spines recognizable (
<bibRefCitation id="EFF04B02FFF7FF8EFCFBA29EFB60FA1C" author="Bartsch" box="[894,1160,1409,1434]" pageId="28" pageNumber="229" refString="Bartsch, I. (2003 d) Psammophilous halacarids (Halacaridae: Acari) from Dampier, Western Australia. Description of species and faunal comparison of the mesopsammal halacarid fauna of western and eastern Australia. Records of the Western Australian Museum, 22, 23 - 45." type="journal article" year="2003" yearSuffix="d">Bartsch 2003d: fig. 7D</bibRefCitation>
). Male GO smaller than female GO and distinctly removed from AP. GA with up to 16 pairs of slender pgs, one pair of setae in anterior half of GA, the others lateral and posterior
<collectingCountry id="F3767663FFF7FF8EFDD3A2D5FD48FA64" box="[598,672,1482,1506]" name="Togo" pageId="28" pageNumber="229">to GO</collectingCountry>
, arranged around a genital groove. Each genital sclerite with three short, setiform sgs. Genital acetabula in posterior part of GO, one (
<bibRefCitation id="EFF04B02FFF7FF8EFC1AA2F2FB42F980" author="Monniot" box="[927,1194,1517,1542]" pageId="28" pageNumber="229" refString="Monniot, F. (1972) Scaptognathides, un nouveau genre d'acariens marins (Halacaridae) dans l'Atlantique ouest. Acarologia, 13, 361 - 366." type="journal article" year="1972">Monniot 1972: fig. 2A</bibRefCitation>
), generally two pairs present. AE of adults and juveniles with pair of epimeral tubes on AE (Fig. 70).
</paragraph>
<paragraph id="8BDE36F3FFF7FF8EFF42A12AFE08F93C" blockId="28.[151,1437,1265,1794]" pageId="28" pageNumber="229">
<emphasis id="B915EAE1FFF7FF8EFF42A12AFED4F9C8" bold="true" box="[199,316,1589,1614]" pageId="28" pageNumber="229">Juveniles.</emphasis>
Deutonymph, protonymph and larva known. Deutonymph with GP and AP either fused (
<bibRefCitation id="EFF04B02FFF7FF8EFAC0A12AFEAEF9F4" author="Bartsch" pageId="28" pageNumber="229" refString="Bartsch, I. (1977 b) Interstitielle Fauna von Galapagos. XX. Halacaridae (Acari). Mikrofauna des Meeresbodens, 65, 1 - 108." type="journal article" year="1977" yearSuffix="b">Bartsch 1977b: fig. 365</bibRefCitation>
) or separated (Abé 2011: fig. 1H). Area representing GP with two pairs of small internal gac, two pairs of pgs but zero sgs. GP and AP of protonymph fused; protonymphal GA with one pair of gac (
<bibRefCitation id="EFF04B02FFF7FF8EFB77A162FF01F93C" author="Bartsch" pageId="28" pageNumber="229" refString="Bartsch, I. (1991 c) Arenicolous Halacaridae (Acari) from Hong Kong. Asian Marine Biology, 8, 57 - 75." type="journal article" year="1991" yearSuffix="c">Bartsch 1991c: fig. 1H</bibRefCitation>
); pgs and sgs lacking.
</paragraph>
</subSubSection>
<subSubSection id="C37B6578FFF7FF8EFF42A1DAFC57F884" pageId="28" pageNumber="229" type="discussion">
<paragraph id="8BDE36F3FFF7FF8EFF42A1DAFC57F884" blockId="28.[151,1437,1265,1794]" pageId="28" pageNumber="229">
<emphasis id="B915EAE1FFF7FF8EFF42A1DAFED3F958" bold="true" box="[199,315,1733,1758]" pageId="28" pageNumber="229">Remarks.</emphasis>
The presently eleven described species are psammobionts and live in warm-temperate and tropical sandy deposits. Records are from all oceans (
<bibRefCitation id="EFF04B02FFF7FF8EFD15A1F6FCDFF884" author="Bartsch" box="[656,823,1769,1794]" pageId="28" pageNumber="229" refString="Bartsch, I. (2009 a) Checklist of marine and freshwater halacarid mite genera and species (Halacaridae: Acari) with notes on synonyms, habitats, distribution and descriptions of the taxa. Zootaxa, 1998, 1 - 170." type="journal article" year="2009" yearSuffix="a">Bartsch 2009a</bibRefCitation>
; Abé 2011).
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