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<document id="BAC49893BE1B495AD2BF4B5B189F50CB" ID-DOI="10.11646/zootaxa.4755.1.6" ID-GBIF-Dataset="9f6752d6-7c9a-4756-acfe-badb2a0301d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3724291" ID-ZooBank="D0B3FFA8-C262-4CD6-98AB-B46C55FCEF6D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584968592865" checkinUser="plazi" docAuthor="Busanello, Dilson G. C., Irmler, Ulrich &amp; Caron, Edilson" docDate="2020" docId="03C887D5FF8AFF8783C6F9CDC45BFE35" docLanguage="en" docName="zootaxa.4755.1.6.pdf" docOrigin="Zootaxa 4755 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudastenus ribeirocostae Busanello &amp; Irmler &amp; Caron 2020, sp. nov." docType="treatment" docVersion="5" lastPageNumber="148" masterDocId="FFF1FFADFF8DFF8E8351FF82C634FFE6" masterDocTitle="Revision of Pseudastenus Bernhauer, synonym with Xenasterides Newton and six new species (Coleoptera, Staphylinidae, Paederinae)" masterLastPageNumber="154" masterPageNumber="139" pageNumber="146" updateTime="1698730789478" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="AF3F8216643F46DA647A217547BFF914">Revision of Pseudastenus Bernhauer, synonym with Xenasterides Newton and six new species (Coleoptera, Staphylinidae, Paederinae)</mods:title>
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<mods:namePart id="7ED07871AF0B1255686A936B5639D744">Busanello, Dilson G. C.</mods:namePart>
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<mods:namePart id="B9E846A586F8CFB25D0E22A297E32CFB">Irmler, Ulrich</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="7DF1A20A8C2E898FF4EBD8CFDA8B753F">Caron, Edilson</mods:namePart>
</mods:name>
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<mods:title id="A9CE5BB6FFFAADC204EF14C5A7C9C470">Zootaxa</mods:title>
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<mods:part id="AF40E57993F5A66C6A7A00A3622DA41B">
<mods:date id="99216F2399A303004B5445E92FE5747C">2020</mods:date>
<mods:detail id="3E6C199549931B6E14A94539F544F10A" type="pubDate">
<mods:number id="4B0997632F0A64095A3D0D77D8D69823">2020-03-23</mods:number>
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<mods:number id="9CCCDA92D32C2A65B5ACB8E92DCB778A">4755</mods:number>
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<mods:identifier id="E6C7A9D61B48418055303CA81BF4454C" type="DOI">10.11646/zootaxa.4755.1.6</mods:identifier>
<mods:identifier id="0B01E7BDE2960AB0D9755C844405D89F" type="GBIF-Dataset">9f6752d6-7c9a-4756-acfe-badb2a0301d0</mods:identifier>
<mods:identifier id="B5AB45CE39FEA70814ED294F88B7A543" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5246CFB98C1C9E01FDD06E87D6832826" type="Zenodo-Dep">3724291</mods:identifier>
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<treatment id="03C887D5FF8AFF8783C6F9CDC45BFE35" ID-DOI="http://doi.org/10.5281/zenodo.3730239" ID-GBIF-Taxon="162696936" ID-Zenodo-Dep="3730239" LSID="urn:lsid:plazi:treatment:03C887D5FF8AFF8783C6F9CDC45BFE35" httpUri="http://treatment.plazi.org/id/03C887D5FF8AFF8783C6F9CDC45BFE35" lastPageId="9" lastPageNumber="148" pageId="7" pageNumber="146">
<subSubSection id="C37B6548FF8AFF8983C6F9CDC570F98C" box="[151,836,1615,1642]" pageId="7" pageNumber="146" type="nomenclature">
<paragraph id="8BDE36C3FF8AFF8983C6F9CDC570F98C" blockId="7.[151,836,1615,1677]" box="[151,836,1615,1642]" pageId="7" pageNumber="146">
<heading id="D09681AFFF8AFF8983C6F9CDC570F98C" bold="true" box="[151,836,1615,1642]" fontSize="11" level="1" pageId="7" pageNumber="146" reason="1">
<emphasis id="B915EAD1FF8AFF8983C6F9CDC570F98C" bold="true" box="[151,836,1615,1642]" pageId="7" pageNumber="146">
<taxonomicName id="4C614D40FF8AFF8983C6F9CDC4EDF98F" ID-CoL="9V8WZ" authority="Busanello &amp; Caron" authorityName="Busanello &amp; Irmler &amp; Caron" authorityYear="2020" box="[151,729,1615,1642]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="ribeirocostae" status="sp. nov.">
<emphasis id="B915EAD1FF8AFF8983C6F9CDC7EBF98C" bold="true" box="[151,479,1615,1642]" italics="true" pageId="7" pageNumber="146">Pseudastenus ribeirocostae</emphasis>
Busanello &amp; Caron
</taxonomicName>
,
<taxonomicNameLabel id="A22657AAFF8AFF8981B7F9D2C570F98C" box="[742,836,1616,1642]" pageId="7" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37B6548FF8AFF8983C6F9F1C72FF96B" box="[151,283,1651,1677]" pageId="7" pageNumber="146" type="description">
<paragraph id="8BDE36C3FF8AFF8983C6F9F1C72FF96B" blockId="7.[151,836,1615,1677]" box="[151,283,1651,1677]" pageId="7" pageNumber="146">
<figureCitation id="135A2A46FF8AFF8983C6F9F1C6C0F96B" box="[151,244,1651,1677]" captionStart="FIGURES 40–45" captionStartId="11.[151,264,1932,1957]" captionTargetBox="[197,1381,213,1868]" captionTargetId="figure@11.[161,1427,181,1906]" captionTargetPageId="11" captionText="FIGURES 40–45. Dorsal view. 40. Pseudastenus ribeirocostae sp. nov.; 41. P. ferrugineus sp. nov.; 42. P. latus sp. nov.; 43. P. oculatus sp. nov.; 44. P. amazonicus sp. nov.; 45. P. schubarti sp. nov. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3724303" httpUri="https://zenodo.org/record/3724303/files/figure.png" pageId="7" pageNumber="146">Figs. 40</figureCitation>
,
<figureCitation id="135A2A46FF8AFF8983AEF9F1C72FF96B" box="[255,283,1651,1677]" captionStart="FIGURES 48–54" captionStartId="13.[151,264,1895,1920]" captionTargetBox="[232,1346,220,1805]" captionTargetId="figure@13.[164,1424,181,1869]" captionTargetPageId="13" captionText="FIGURES 48–54. Aedeagus of Pseudastenus, dorsal and lateral view, respectively, figures 51–54 including microscopic detail of basal slerites. 48. P. ribeirocostae sp. nov.; 49. P. ferrugineus sp. nov.; 50. P. latus sp. nov.; 51. P. barretoi; 52. P. oculatus sp. nov.; 53. P. amazonicus sp. nov.; 54. P. schubarti sp. nov. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3724309" httpUri="https://zenodo.org/record/3724309/files/figure.png" pageId="7" pageNumber="146">48</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37B6548FF8AFF8983C6F938C6C7F86F" pageId="7" pageNumber="146" type="materials_examined">
<paragraph id="8BDE36C3FF8AFF8983C6F938C6C7F86F" blockId="7.[151,1436,1722,2038]" pageId="7" pageNumber="146">
<emphasis id="B915EAD1FF8AFF8983C6F938C732F933" bold="true" box="[151,262,1722,1749]" pageId="7" pageNumber="146">Material.</emphasis>
<materialsCitation id="3B093C9EFF8AFF89825EF939C533F883" ID-GBIF-Occurrence="2579576919" collectingDate="19-07-27" collectionCode="DZUP" country="Brazil" county="Reserva do Cachoeira" determinerName="E. Caron" location="Forest" municipality="Guarequecaba Moist Lowland" pageId="7" pageNumber="146" specimenCode="CC2" specimenCount="2" stateProvince="Para" typeStatus="holotype">
<typeStatus id="54DA8861FF8AFF89825EF939C74FF933" box="[271,379,1723,1749]" pageId="7" pageNumber="146" type="holotype">Holotype</typeStatus>
. Male, dissected, deposited in
<collectionCode id="ED70AE06FF8AFF89818BF93EC517F930" box="[730,803,1724,1750]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="7" pageNumber="146" type="University or college">DZUP</collectionCode>
, labels: (1) “Sample-ID:
<specimenCode id="DBC79EB8FF8AFF898712F939C242F933" box="[1091,1142,1723,1749]" collectionCode="CC" country="Australia" name="CSIRO Canberra Rhizobium Collection" pageId="7" pageNumber="146">CC2</specimenCode>
F4F7/
<taxonomicName id="4C614D40FF8AFF8987E8F939C36CF933" box="[1209,1368,1723,1749]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
:
<taxonomicName id="4C614D40FF8AFF898635F93EC6FBF91F" authorityName="Bierig" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Xenaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Xenaster</taxonomicName>
” [white label, printed in black and handwritten]; (2)
<collectingCountry id="F3767653FF8AFF89804EF95DC5BDF91F" box="[799,905,1759,1785]" name="Brazil" pageId="7" pageNumber="146">BRAZIL</collectingCountry>
:
<collectingRegion id="49A5F821FF8AFF8980C2F962C58EF91C" box="[915,954,1760,1786]" country="Brazil" name="Para" pageId="7" pageNumber="146">PR</collectingRegion>
:
<collectingCounty id="62BF4E4FFF8AFF898095F962C282F91F" box="[964,1206,1759,1786]" pageId="7" pageNumber="146">Reserva do Cachoeira</collectingCounty>
/APA
<collectingMunicipality id="6BBAACB9FF8AFF8987ADF95DC772F8FB" pageId="7" pageNumber="146">Guarequeçaba Moist Lowland</collectingMunicipality>
/
<location id="8EBE6018FF8AFF89821EF881C7ACF8FB" LSID="urn:lsid:plazi:treatment:03C887D5FF8AFF8783C6F9CDC45BFE35:8EBE6018FF8AFF89821EF881C7ACF8FB" box="[335,408,1795,1821]" country="Brazil" county="Reserva do Cachoeira" municipality="Guarequecaba Moist Lowland" name="Forest" pageId="7" pageNumber="146" stateProvince="Para">Forest</location>
,sift.
<location id="8EBE6018FF8AFF89829CF881C439F8FB" LSID="urn:lsid:plazi:treatment:03C887D5FF8AFF8783C6F9CDC45BFE35:8EBE6018FF8AFF89829CF881C439F8FB" box="[461,525,1795,1821]" country="Brazil" county="Reserva do Cachoeira" municipality="Guarequecaba Moist Lowland" name="Litter" pageId="7" pageNumber="146" stateProvince="Para">Litter</location>
, winklerextr. coll./
<location id="8EBE6018FF8AFF8981B3F881C528F8FB" LSID="urn:lsid:plazi:treatment:03C887D5FF8AFF8783C6F9CDC45BFE35:8EBE6018FF8AFF8981B3F881C528F8FB" box="[738,796,1795,1821]" country="Brazil" county="Reserva do Cachoeira" municipality="Guarequecaba Moist Lowland" name="Bihn" pageId="7" pageNumber="146" stateProvince="Para">Bihn</location>
; 20.7.-9-8[Aug.].[20]07” [white label, printed in black]; (3) “
<collectionCode id="ED70AE06FF8AFF89839DF8A5C72CF8A7" box="[204,280,1831,1857]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="7" pageNumber="146" type="University or college">DZUP</collectionCode>
/
<date id="FFDF1003FF8AFF898270F8A5C743F8A7" box="[289,375,1831,1857]" pageId="7" pageNumber="146" value="19-07-27">
<collectingDate id="EF9BE9EBFF8AFF898270F8A5C743F8A7" box="[289,375,1831,1857]" pageId="7" pageNumber="146" value="19-07-27">270719</collectingDate>
</date>
” [white label, printed in black]; (3) “
<typeStatus id="54DA8861FF8AFF898061F8A5C5F2F8A7" box="[816,966,1831,1857]" pageId="7" pageNumber="146" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C614D40FF8AFF89809EF8A5C6E9F883" authority="Busanello &amp; Caron" authorityName="Busanello &amp; Irmler &amp; Caron" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="ribeirocostae">Pseudastenus ribeirocostae Busanello &amp; Caron</taxonomicName>
/det.
<determinerName id="7E02DA07FF8AFF898245F8CEC74EF883" box="[276,378,1867,1894]" pageId="7" pageNumber="146">E. Caron</determinerName>
, 2019” [red label/ printed in black]
</materialsCitation>
.
<materialsCitation id="3B093C9EFF8AFF898040F8CEC2E1F883" ID-GBIF-Occurrence="2579576934" box="[785,1237,1867,1894]" country="Brazil" location="BRAZIL" pageId="7" pageNumber="146" specimenCount="22" typeStatus="paratype">
<typeStatus id="54DA8861FF8AFF898040F8CEC5B6F880" box="[785,898,1868,1894]" pageId="7" pageNumber="146" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F3767653FF8AFF8980DCF8C9C5C3F883" box="[909,1015,1867,1893]" name="Brazil" pageId="7" pageNumber="146">BRAZIL</collectingCountry>
:
<specimenCount id="9D67FD4AFF8AFF898750F8C9C2ADF883" box="[1025,1177,1867,1893]" pageId="7" pageNumber="146" type="generic">22 specimens</specimenCount>
from
</materialsCitation>
<materialsCitation id="3B093C9EFF8AFF89878CF8C9C6C7F86F" ID-GBIF-Occurrence="2579576915" collectionCode="DZUP" country="Brazil" location="Parana" pageId="7" pageNumber="146" specimenCount="1" stateProvince="Parana" typeStatus="paratype">
<collectingCountry id="F3767653FF8AFF89878CF8C9C371F883" box="[1245,1349,1867,1893]" name="Brazil" pageId="7" pageNumber="146">BRAZIL</collectingCountry>
,
<collectingRegion id="49A5F821FF8AFF898601F8C9C3A8F883" box="[1360,1436,1867,1893]" country="Brazil" name="Parana" pageId="7" pageNumber="146">Paraná</collectingRegion>
(
<collectionCode id="ED70AE06FF8AFF8983CEF8EDC6D8F86F" box="[159,236,1903,1929]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="7" pageNumber="146" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37B6548FF8AFF898396F810C295F813" pageId="7" pageNumber="146" type="diagnosis">
<paragraph id="8BDE36C3FF8AFF898396F810C295F813" blockId="7.[151,1436,1722,2038]" pageId="7" pageNumber="146">
<emphasis id="B915EAD1FF8AFF898396F810C776F84B" bold="true" box="[199,322,1938,1965]" pageId="7" pageNumber="146">Diagnosis.</emphasis>
Among the species group with short elytra and reduced hind wings
<taxonomicName id="4C614D40FF8AFF89876AF811C35AF84B" authorityName="Busanello &amp; Irmler &amp; Caron" authorityYear="2020" box="[1083,1390,1939,1965]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="ribeirocostae">
<emphasis id="B915EAD1FF8AFF89876AF811C35AF84B" box="[1083,1390,1939,1965]" italics="true" pageId="7" pageNumber="146">Pseudastenus ribeirocostae</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="4C614D40FF8AFF898238F83AC7D4F837" authorityName="Bernhauer" authorityYear="1933" box="[361,480,1975,2001]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="barretoi">
<emphasis id="B915EAD1FF8AFF898238F83AC7D4F837" box="[361,480,1975,2001]" italics="true" pageId="7" pageNumber="146">P. barretoi</emphasis>
</taxonomicName>
by smaller eyes and from
<taxonomicName id="4C614D40FF8AFF89805FF83AC5A0F837" authorityName="Busanello &amp; Irmler &amp; Caron" authorityYear="2020" box="[782,916,1975,2001]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="schubarti">
<emphasis id="B915EAD1FF8AFF89805FF83AC5A0F837" box="[782,916,1975,2001]" italics="true" pageId="7" pageNumber="146">P. schubarti</emphasis>
</taxonomicName>
by the darker anterior half of elytra. Males of these two species are distinguished by the different arrangement of sclerites in the aedeagus.
</paragraph>
</subSubSection>
<caption id="DF1E664BFF85FF8683C6FA5BC724F9DF" ID-DOI="http://doi.org/10.5281/zenodo.3724301" ID-Zenodo-Dep="3724301" httpUri="https://zenodo.org/record/3724301/files/figure.png" pageId="8" pageNumber="147" startId="8.[151,264,1497,1522]" targetBox="[226,1371,203,1410]" targetPageId="8">
<paragraph id="8BDE36C3FF85FF8683C6FA5BC724F9DF" blockId="8.[151,1436,1497,1594]" pageId="8" pageNumber="147">
<emphasis id="B915EAD1FF85FF8683C6FA5BC7B1FA14" bold="true" box="[151,389,1497,1522]" pageId="8" pageNumber="147">FIGURES 36–39. 36.</emphasis>
<taxonomicName id="4C614D40FF85FF8682C1FA5BC443FA17" authorityName="Bernhauer" authorityYear="1933" box="[400,631,1497,1521]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="barretoi">
<emphasis id="B915EAD1FF85FF8682C1FA5BC443FA17" box="[400,631,1497,1521]" italics="true" pageId="8" pageNumber="147">Pseudastenus barretoi</emphasis>
</taxonomicName>
, lectotype, dorsal view (FMNH);
<emphasis id="B915EAD1FF85FF8680BAFA5BC23FFA14" bold="true" box="[1003,1035,1497,1522]" pageId="8" pageNumber="147">37.</emphasis>
<taxonomicName id="4C614D40FF85FF868747FA58C294FA17" baseAuthorityName="Bierig" baseAuthorityYear="1939" box="[1046,1184,1497,1521]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis id="B915EAD1FF85FF868747FA58C294FA17" box="[1046,1184,1497,1521]" italics="true" pageId="8" pageNumber="147">P. plaumanni</emphasis>
</taxonomicName>
, lectotype, dorsal view (FMNH);
<emphasis id="B915EAD1FF85FF8683AEFA7FC72BF9F0" bold="true" box="[255,287,1533,1558]" pageId="8" pageNumber="147">38.</emphasis>
Labels from the lectotype of
<taxonomicName id="4C614D40FF85FF86811DFA7CC483F9F3" authorityName="Bernhauer" authorityYear="1933" box="[588,695,1533,1557]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="barretoi">
<emphasis id="B915EAD1FF85FF86811DFA7CC483F9F3" box="[588,695,1533,1557]" italics="true" pageId="8" pageNumber="147">P. barretoi</emphasis>
</taxonomicName>
(FMNH);
<emphasis id="B915EAD1FF85FF868074FA7FC571F9F0" bold="true" box="[805,837,1533,1558]" pageId="8" pageNumber="147">39.</emphasis>
Labels from the lectotype of
<taxonomicName id="4C614D40FF85FF868723FA7CC2C3F9F3" baseAuthorityName="Bierig" baseAuthorityYear="1939" box="[1138,1271,1533,1557]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis id="B915EAD1FF85FF868723FA7CC2C3F9F3" box="[1138,1271,1533,1557]" italics="true" pageId="8" pageNumber="147">P. plaumanni</emphasis>
</taxonomicName>
(FMNH). Scale bars: 2 mm.
</paragraph>
</caption>
<subSubSection id="C37B6548FF85FF878396F9E5C43AFF1D" lastPageId="9" lastPageNumber="148" pageId="8" pageNumber="147" type="description">
<paragraph id="8BDE36C3FF85FF878396F9E5C43AFF1D" blockId="8.[151,1436,1639,2026]" lastBlockId="9.[151,1436,153,467]" lastPageId="9" lastPageNumber="148" pageId="8" pageNumber="147">
<emphasis id="B915EAD1FF85FF868396F9E5C76DF964" bold="true" box="[199,345,1639,1666]" pageId="8" pageNumber="147">Description.</emphasis>
Body length:
<quantity id="4C999B26FF85FF8682A9F9EAC46BF964" box="[504,607,1640,1666]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="8" pageNumber="147" unit="mm" value="3.3">3.30 mm</quantity>
. Coloration: Body yellowish brown, pronotum and appendices lighter, abdomen darker. Head
<quantity id="4C999B26FF85FF8682D0F90EC7D2F940" box="[385,486,1676,1702]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="8" pageNumber="147" unit="mm" value="0.49">0.49 mm</quantity>
long,
<quantity id="4C999B26FF85FF86817CF90EC4A5F940" box="[557,657,1676,1702]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="8" pageNumber="147" unit="mm" value="0.49">0.49 mm</quantity>
wide; eyes small, very slightly prominent; 0.17 times as long as total length of head; 0.33 times as long as postocular length; approximately parallel; posterior margin straight; depressed on central vertex in front of neck. Pronotum
<quantity id="4C999B26FF85FF8681DBF956C4DAF908" box="[650,750,1748,1774]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="8" pageNumber="147" unit="mm" value="0.47">0.47 mm</quantity>
long,
<quantity id="4C999B26FF85FF868065F956C5ACF908" box="[820,920,1748,1774]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="8" pageNumber="147" unit="mm" value="0.49">0.49 mm</quantity>
wide; widest close to anterior angles; anterior margin slightly curved; posterior margin straight; sides convergingposteriad. Elytra
<quantity id="4C999B26FF85FF868716F97AC29FF8F4" box="[1095,1195,1784,1810]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="8" pageNumber="147" unit="mm" value="0.47">0.47 mm</quantity>
long,
<quantity id="4C999B26FF85FF8687A2F97AC363F8F4" box="[1267,1367,1784,1810]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="147" unit="mm" value="0.38">0.38 mm</quantity>
wide; as long as pronotum, but conspicuously narrower than pronotum; sides slightly diverging posteriad; widest near the apex; humeral angles rounded; hind wings not developed; posterior angles curved; posterior margin strongly emarginate, exposing tergite II medially; suture 0.82 times as long as lateral side; microtuberculate punctation larger than on head and pronotum; with yellow pubescence pointing posteriad, longer than head and pronotum; posterior edge with fringe of sparse robust setae. Abdomen with dense punctation; with yellow pubescence pointing posteriad; pubescence conspicuously longer than on the rest of body. Aedeagus elongate;
<quantity id="4C999B26FF85FF86872FF852C2D5F80C" box="[1150,1249,2000,2026]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="8" pageNumber="147" unit="mm" value="0.68">0.68 mm</quantity>
long; 5.50 times as long as wide; median lobe with sides slightly converging towards to the apex, robust basal sclerites and pair of inconspicuous apical sclerites,
<quantity id="4C999B26FF84FF8782A3FF3FC462FF31" box="[498,598,189,215]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="148" unit="mm" value="0.09">0.09 mm</quantity>
long; ejaculatory duct with two trifurcate accessorysclerites, proximal two times longer than distal (
<figureCitation id="135A2A46FF84FF8782FCFF63C435FF1D" box="[429,513,225,251]" captionStart="FIGURES 48–54" captionStartId="13.[151,264,1895,1920]" captionTargetBox="[232,1346,220,1805]" captionTargetId="figure@13.[164,1424,181,1869]" captionTargetPageId="13" captionText="FIGURES 48–54. Aedeagus of Pseudastenus, dorsal and lateral view, respectively, figures 51–54 including microscopic detail of basal slerites. 48. P. ribeirocostae sp. nov.; 49. P. ferrugineus sp. nov.; 50. P. latus sp. nov.; 51. P. barretoi; 52. P. oculatus sp. nov.; 53. P. amazonicus sp. nov.; 54. P. schubarti sp. nov. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3724309" httpUri="https://zenodo.org/record/3724309/files/figure.png" pageId="9" pageNumber="148">Fig. 48</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37B6548FF84FF878396FE87C45BFE35" pageId="9" pageNumber="148" type="etymology">
<paragraph id="8BDE36C3FF84FF878396FE87C47FFE81" blockId="9.[151,1436,153,467]" pageId="9" pageNumber="148">
<emphasis id="B915EAD1FF84FF878396FE87C77AFEF9" bold="true" box="[199,334,261,287]" pageId="9" pageNumber="148">Etymology.</emphasis>
The specific name honors professor Dr. Cibele Stramare Ribeiro-Costa of the Federal University of
<collectingRegion id="49A5F821FF84FF8783E4FEABC735FEA5" box="[181,257,297,323]" country="Brazil" name="Parana" pageId="9" pageNumber="148">Paraná</collectingRegion>
(UFPR), specialist in seed beetles (
<taxonomicName id="4C614D40FF84FF8781DAFEABC50DFEA5" authorityName="Latreille" authorityYear="1802" box="[651,825,297,323]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="148" phylum="Arthropoda" rank="family">Chrysomelidae</taxonomicName>
,
<taxonomicName id="4C614D40FF84FF878012FEABC58FFEA2" authorityName="Latreille" authorityYear="1802" box="[835,955,297,324]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="148" phylum="Arthropoda" rank="subFamily" subFamily="Bruchinae">Bruchinae</taxonomicName>
). She has started and encouraged the recent studies of staphylinid beetles in
<collectingCountry id="F3767653FF84FF8782AEFECFC473FE81" box="[511,583,333,359]" name="Brazil" pageId="9" pageNumber="148">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="8BDE36C3FF84FF878396FEF2C352FE6A" blockId="9.[151,1436,153,467]" box="[199,1382,368,396]" pageId="9" pageNumber="148">
<emphasis id="B915EAD1FF84FF878396FEF2C7E5FE6D" bold="true" box="[199,465,368,395]" pageId="9" pageNumber="148">Geographical records.</emphasis>
<taxonomicName id="4C614D40FF84FF878289FEF3C53FFE6D" authorityName="Busanello &amp; Irmler &amp; Caron" authorityYear="2020" box="[472,779,369,395]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="148" phylum="Arthropoda" rank="species" species="ribeirocostae">
<emphasis id="B915EAD1FF84FF878289FEF3C53FFE6D" box="[472,779,369,395]" italics="true" pageId="9" pageNumber="148">Pseudastenus ribeirocostae</emphasis>
</taxonomicName>
is known from the eastern regions of
<collectingRegion id="49A5F821FF84FF8787E5FEF3C334FE6D" box="[1204,1280,369,395]" country="Brazil" name="Parana" pageId="9" pageNumber="148">Paraná</collectingRegion>
(
<collectingCountry id="F3767653FF84FF87865EFEF3C36DFE6A" box="[1295,1369,369,396]" name="Brazil" pageId="9" pageNumber="148">Brazil</collectingCountry>
).
</paragraph>
<paragraph id="8BDE36C3FF84FF878396FE16C45BFE35" blockId="9.[151,1436,153,467]" pageId="9" pageNumber="148">
<emphasis id="B915EAD1FF84FF878396FE16C7B8FE49" bold="true" box="[199,396,404,431]" pageId="9" pageNumber="148">Biological notes.</emphasis>
The specimens of
<taxonomicName id="4C614D40FF84FF878139FE17C5AAFE49" authorityName="Busanello &amp; Irmler &amp; Caron" authorityYear="2020" box="[616,926,405,431]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="148" phylum="Arthropoda" rank="species" species="ribeirocostae">
<emphasis id="B915EAD1FF84FF878139FE17C5AAFE49" box="[616,926,405,431]" italics="true" pageId="9" pageNumber="148">Pseudastenus ribeirocostae</emphasis>
</taxonomicName>
were collected in leaf litter of cambisol and gleysol soils using the Winkler extraction.
</paragraph>
</subSubSection>
</treatment>
</document>
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