148 lines
17 KiB
XML
148 lines
17 KiB
XML
<document id="A4641EA5D478A56579DD26F4550B7933" ID-DOI="10.11646/zootaxa.3741.1.2" ID-GBIF-Dataset="6a6ba78e-a9c5-4d11-997a-3d06e5e31520" ID-ISSN="1175-5326" ID-Zenodo-Dep="223773" ID-ZooBank="BF5EA9B8-C6F4-448A-BEF9-1976AB4EC308" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460702326177" checkinUser="plazi" docAuthor="Lazányi, Eszter & Vagalinski, Boyan" docDate="2013" docId="03C887D3FFD9FF97FF3498FAFEC4A9BC" docLanguage="en" docName="zt03741p100.pdf" docOrigin="Zootaxa 3741 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaphyllum spathulatum Lohmander 1936" docType="treatment" docVersion="8" lastPageNumber="87" masterDocId="FFF1FFABFFF9FFB7FFA39926FFF2AD11" masterDocTitle="Redefinition of the millipede subgenus Megaphyllum sensu stricto Verhoeff, 1894 and neotype designation for Megaphyllum austriacum (Latzel, 1884) (Myriapoda: Diplopoda: Julida: Julidae)" masterLastPageNumber="100" masterPageNumber="55" pageNumber="87" updateTime="1698356536038" updateUser="plazi">
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<mods:titleInfo id="9FF67DA742005908DF808536F0162758">
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<mods:title id="D388B26A0CCA94FFE4EDE93B8A40C999">Redefinition of the millipede subgenus Megaphyllum sensu stricto Verhoeff, 1894 and neotype designation for Megaphyllum austriacum (Latzel, 1884) (Myriapoda: Diplopoda: Julida: Julidae)</mods:title>
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<mods:namePart id="5EB9726FE46726454D3167AD5CDBE9B3">Lazányi, Eszter</mods:namePart>
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<mods:roleTerm id="17B8C244D612DE82753DFDD91EA6ABB1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="89BE24797612F318EDD5FFB9C42D199C">Vagalinski, Boyan</mods:namePart>
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<mods:typeOfResource id="6BCB8A42259D0D0FCA752BCBC75F4CA3">text</mods:typeOfResource>
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<mods:title id="C8DB6BB9DE4285D0BF1CA1FC64E68A3C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="812C4C606C7D3005B04E4E6D5EF822C6">
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<mods:date id="484838336FD39D727AC69D28FF7A1347">2013</mods:date>
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<mods:detail id="00A4472D164DE3A3C374B74DF22D2FD2" type="volume">
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<mods:number id="E1D627543F2C1A3DF01B1E20244AF560">3741</mods:number>
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</mods:detail>
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<mods:detail id="C93CF039C763FCC533E907B57B09051D" type="issue">
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<mods:number id="49961D9A0CD4711CC2F04401F66B98A1">1</mods:number>
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<mods:classification id="81845414C09D301F96F3A5DD1113F7E0">journal article</mods:classification>
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<mods:identifier id="DB7610C25EF6BDE741EE20090D290121" type="DOI">10.11646/zootaxa.3741.1.2</mods:identifier>
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<mods:identifier id="EB63722CE168F6C6B4DA42E2F75F8BC3" type="GBIF-Dataset">6a6ba78e-a9c5-4d11-997a-3d06e5e31520</mods:identifier>
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<mods:identifier id="AF08EA61D28195047DF72CF428A612A6" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="37DCE37AE48F729933513CC0A06C0B8A" type="Zenodo-Dep">223773</mods:identifier>
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<treatment id="03C887D3FFD9FF97FF3498FAFEC4A9BC" ID-DOI="http://doi.org/10.5281/zenodo.6151744" ID-GBIF-Taxon="119583558" ID-Zenodo-Dep="6151744" LSID="urn:lsid:plazi:treatment:03C887D3FFD9FF97FF3498FAFEC4A9BC" httpUri="http://treatment.plazi.org/id/03C887D3FFD9FF97FF3498FAFEC4A9BC" lastPageNumber="87" pageId="32" pageNumber="87">
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<subSubSection id="C37B654EFFD9FF97FF3498FAFCD2AF6A" pageId="32" pageNumber="87" type="nomenclature">
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<paragraph id="8BDE36C5FFD9FF97FF3498FAFD20ACE7" blockId="32.[151,722,476,535]" box="[151,722,476,502]" pageId="32" pageNumber="87">
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<heading id="D09681A9FFD9FF97FF3498FAFD20ACE7" bold="true" box="[151,722,476,502]" fontSize="11" level="1" pageId="32" pageNumber="87" reason="1">
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<taxonomicName id="4C614D46FFD9FF97FF3498FAFD20ACE7" ID-CoL="8KR84" authority="Lohmander, 1936" authorityName="Lohmander" authorityYear="1936" box="[151,722,476,502]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="32" pageNumber="87" phylum="Arthropoda" rank="species" species="spathulatum">
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<emphasis id="B915EAD7FFD9FF97FF3498FAFD20ACE7" bold="true" box="[151,722,476,502]" pageId="32" pageNumber="87">
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<emphasis id="B915EAD7FFD9FF97FF3498FAFE2CACE7" bold="true" box="[151,478,476,502]" italics="true" pageId="32" pageNumber="87">Megaphyllum spathulatum</emphasis>
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(Lohmander, 1936)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BDE36C5FFD9FF97FF3498D8FEE6AF06" blockId="32.[151,722,476,535]" box="[151,276,510,535]" pageId="32" pageNumber="87">
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<figureCitation id="135A2A40FFD9FF97FF3498D8FEE6AF06" box="[151,276,510,535]" captionStart="FIGURES 18 b – d" captionStartId="32.[151,264,1855,1877]" captionTargetBox="[310,1276,1254,1831]" captionTargetId="figure@32.[307,1280,1253,1834]" captionTargetPageId="32" captionText="FIGURES 18 b – d. Megaphyllum spathulatum after Lohmander (1936): (18 b) right gonopods, mesal view; (18 c) left promere, meso-posterior view; (18 d) probably left opisthomere, lateral view. Abbreviations: ah: anterior hump; ap: anterior process, asp: anterior process of the solenomere, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, sh: serrated hump, P: promere." httpUri="https://zenodo.org/record/223790/files/figure.png" pageId="32" pageNumber="87">Figs 18b–d</figureCitation>
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</paragraph>
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<paragraph id="8BDE36C5FFD9FF97FF349B60FCD2AF6A" blockId="32.[151,1019,582,666]" pageId="32" pageNumber="87">
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<taxonomicName id="4C614D46FFD9FF97FF349B60FCB6AF4D" authority="Lohmander, 1936: 104" authorityName="Lohmander" authorityPageNumber="104" authorityYear="1936" box="[151,836,582,604]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="32" pageNumber="87" phylum="Arthropoda" rank="species" species="spathulatus" subGenus="Chromatoiulus">
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<emphasis id="B915EAD7FFD9FF97FF349B60FDA5AF4D" box="[151,599,582,604]" italics="true" pageId="32" pageNumber="87">Chromatoiulus (Chromatoiulus) spathulatus</emphasis>
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Lohmander, 1936: 104
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</taxonomicName>
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–109, Figs 80–84.
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<taxonomicName id="4C614D46FFD9FF97FF349B43FCEEAF6A" authority="Attems 1940: 306" authorityName="Attems" authorityPageNumber="306" authorityYear="1940" box="[151,796,613,635]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="32" pageNumber="87" phylum="Arthropoda" rank="species" species="spathulatus" subGenus="Chromatoiulus">
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<emphasis id="B915EAD7FFD9FF97FF349B43FDA5AF6A" box="[151,599,613,635]" italics="true" pageId="32" pageNumber="87">Chromatoiulus (Chromatoiulus) spathulatus</emphasis>
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: Attems 1940: 306
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B654EFFD9FF97FF349BA2FCF1AF8B" box="[151,771,644,666]" pageId="32" pageNumber="87" type="description">
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<paragraph id="8BDE36C5FFD9FF97FF349BA2FCF1AF8B" blockId="32.[151,1019,582,666]" box="[151,771,644,666]" pageId="32" pageNumber="87">
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<taxonomicName id="4C614D46FFD9FF97FF349BA2FD79AF8B" authority="Golovatch 1990: 364" authorityName="Golovatch" authorityPageNumber="364" authorityYear="1990" box="[151,651,644,666]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="32" pageNumber="87" phylum="Arthropoda" rank="species" species="spathulatum">
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<emphasis id="B915EAD7FFD9FF97FF349BA2FE57AF8B" box="[151,421,644,666]" italics="true" pageId="32" pageNumber="87">Megaphyllum spathulatum</emphasis>
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: Golovatch 1990: 364
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</taxonomicName>
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; 1992: 381.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B654EFFD9FF97FF349BE7FBE3AFCB" box="[151,1041,705,730]" pageId="32" pageNumber="87" type="distribution">
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<paragraph id="8BDE36C5FFD9FF97FF349BE7FBE3AFCB" blockId="32.[151,1436,705,1198]" box="[151,1041,705,730]" pageId="32" pageNumber="87">
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<emphasis id="B915EAD7FFD9FF97FF349BE7FEC2AFCB" bold="true" box="[151,304,705,730]" pageId="32" pageNumber="87">Distribution.</emphasis>
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Western Caucasus (unspecified
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<typeStatus id="54DA8867FFD9FF97FD389BE4FD39AFCB" box="[667,715,706,730]" pageId="32" pageNumber="87">type</typeStatus>
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locality) (Lohmander 1936).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B654EFFD9FF97FF649BC3FCF4A90F" pageId="32" pageNumber="87" type="diagnosis">
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<paragraph id="8BDE36C5FFD9FF97FF649BC3FCAEAE57" blockId="32.[151,1436,705,1198]" pageId="32" pageNumber="87">
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<emphasis id="B915EAD7FFD9FF97FF649BC3FEB3AFEF" bold="true" box="[199,321,741,766]" pageId="32" pageNumber="87">Diagnosis.</emphasis>
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The species is easily recognized from other consubgeners by a combination of a strongly tapering promere (
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<figureCitation id="135A2A40FFD9FF97FEA59A2FFE75AE33" box="[262,391,777,802]" captionStart="FIGURES 17 a – g" captionStartId="31.[151,264,1371,1393]" captionTargetBox="[268,1318,446,1349]" captionTargetId="figure@31.[263,1323,445,1350]" captionTargetPageId="31" captionText="FIGURES 17 a – g. Megaphyllum sjaelandicum male and female from Dorpat, Estonia, ZMB 8734: (17 a) gonopods, posterior view; (17 b) left gonopods, mesal view; (17 c) right promere, meso-posterior view; (17 d) right opisthomere, mesal view; penis, (17 e) posterior and right lateral views; (17 g) right vulva, antero-mesal view (setae omitted). Abbreviations: ah: anterior hump; ap: anterior process, asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; h: hump, l: lobe, op: operculum, pa: posterior ampulla, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/223789/files/figure.png" pageId="32" pageNumber="87">Figs 17b–c</figureCitation>
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) and a tagged posterior opisthomere process (
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<emphasis id="B915EAD7FFD9FF97FC029A2CFC4FAE30" bold="true" box="[929,957,778,801]" italics="true" pageId="32" pageNumber="87">pp</emphasis>
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) connecting to the opisthomere at higher angles (around 50°–60°, see
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<figureCitation id="135A2A40FFD9FF97FE7A9A0BFD80AE57" box="[473,626,813,838]" captionStart="FIGURES 18 b – d" captionStartId="32.[151,264,1855,1877]" captionTargetBox="[310,1276,1254,1831]" captionTargetId="figure@32.[307,1280,1253,1834]" captionTargetPageId="32" captionText="FIGURES 18 b – d. Megaphyllum spathulatum after Lohmander (1936): (18 b) right gonopods, mesal view; (18 c) left promere, meso-posterior view; (18 d) probably left opisthomere, lateral view. Abbreviations: ah: anterior hump; ap: anterior process, asp: anterior process of the solenomere, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, sh: serrated hump, P: promere." httpUri="https://zenodo.org/record/223790/files/figure.png" pageId="32" pageNumber="87">Figs 18b, 18d</figureCitation>
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) (Lohmander 1936).
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</paragraph>
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<paragraph id="8BDE36C5FFD9FF97FF649A77FBC7AE9F" blockId="32.[151,1436,705,1198]" pageId="32" pageNumber="87">
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Male gonopods: Promere (
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<emphasis id="B915EAD7FFD9FF97FE549A74FDFAAE78" bold="true" box="[503,520,850,873]" italics="true" pageId="32" pageNumber="87">P</emphasis>
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) (
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<figureCitation id="135A2A40FFD9FF97FD819A77FD50AE7B" box="[546,674,849,874]" captionStart="FIGURES 18 b – d" captionStartId="32.[151,264,1855,1877]" captionTargetBox="[310,1276,1254,1831]" captionTargetId="figure@32.[307,1280,1253,1834]" captionTargetPageId="32" captionText="FIGURES 18 b – d. Megaphyllum spathulatum after Lohmander (1936): (18 b) right gonopods, mesal view; (18 c) left promere, meso-posterior view; (18 d) probably left opisthomere, lateral view. Abbreviations: ah: anterior hump; ap: anterior process, asp: anterior process of the solenomere, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, sh: serrated hump, P: promere." httpUri="https://zenodo.org/record/223790/files/figure.png" pageId="32" pageNumber="87">Figs 18b–c</figureCitation>
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) strongly tapering, medially with an apical hump with spines (
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<emphasis id="B915EAD7FFD9FF97FAD29A77FA79AE7B" bold="true" box="[1393,1419,849,874]" italics="true" pageId="32" pageNumber="87">sh</emphasis>
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). Posterior solenomere process (
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<emphasis id="B915EAD7FFD9FF97FE4C9A50FDE7AE9C" bold="true" box="[495,533,886,909]" italics="true" pageId="32" pageNumber="87">psp</emphasis>
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) well-developed, longer than anterior one (
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<emphasis id="B915EAD7FFD9FF97FC5E9A50FBD6AE9C" bold="true" box="[1021,1060,886,909]" italics="true" pageId="32" pageNumber="87">asp</emphasis>
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).
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</paragraph>
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<paragraph id="8BDE36C5FFD9FF97FF649ABFFCF4A90F" blockId="32.[151,1436,705,1198]" pageId="32" pageNumber="87">
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According to Lohmander (1936): body colour probably faded due to the conservation; but both sexes seemed to be brownish, yellow-grey, sometimes with a dark dorsal band medially; the ozopore line is also dark. Thus, it resembles
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<taxonomicName id="4C614D46FFD9FF97FEB19AC4FE23AEE8" box="[274,465,993,1017]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="32" pageNumber="87" phylum="Arthropoda" rank="species" species="sjaelandicum">
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<emphasis id="B915EAD7FFD9FF97FEB19AC4FE23AEE8" box="[274,465,993,1017]" italics="true" pageId="32" pageNumber="87">M. sjaelandicum</emphasis>
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</taxonomicName>
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more than other species in the group. Body length and height (Lohmander 1936): males:
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<quantity id="4C999B20FFD9FF97FF449D23FEC3A90F" box="[231,305,1029,1054]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="32" pageNumber="87" unit="mm" value="35.0">35mm</quantity>
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,
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<quantity id="4C999B20FFD9FF97FE9F9D20FE7DA90F" box="[316,399,1030,1054]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="32" pageNumber="87" unit="mm" value="2.7">2.7mm</quantity>
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; females:
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<quantity id="4C999B20FFD9FF97FDA39D20FD86A90F" box="[512,628,1030,1054]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="32" pageNumber="87" unit="mm" value="40.0" valueMax="42.0" valueMin="38.0">38–42mm</quantity>
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,
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<quantity id="4C999B20FFD9FF97FDDC9D20FCF0A90F" box="[639,770,1030,1054]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.8" metricValueMin="3.4" pageId="32" pageNumber="87" unit="mm" value="3.5999999999999996" valueMax="3.8" valueMin="3.4">3.4–3.8mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B654EFFD9FF97FF649D0FFEC4A9BC" pageId="32" pageNumber="87" type="discussion">
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<paragraph id="8BDE36C5FFD9FF97FF649D0FFEC4A9BC" blockId="32.[151,1436,705,1198]" pageId="32" pageNumber="87">
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<emphasis id="B915EAD7FFD9FF97FF649D0FFEC9A953" bold="true" box="[199,315,1065,1090]" pageId="32" pageNumber="87">Remarks.</emphasis>
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<taxonomicName id="4C614D46FFD9FF97FEE69D0CFE0AA950" box="[325,504,1065,1089]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="32" pageNumber="87" phylum="Arthropoda" rank="species" species="spathulatum">
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<emphasis id="B915EAD7FFD9FF97FEE69D0CFE0AA950" box="[325,504,1065,1089]" italics="true" pageId="32" pageNumber="87">M. spathulatum</emphasis>
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</taxonomicName>
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is regarded as the only neoendemic
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<emphasis id="B915EAD7FFD9FF97FC0B9D0FFB72A950" box="[936,1152,1065,1089]" italics="true" pageId="32" pageNumber="87">
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<taxonomicName id="4C614D46FFD9FF97FC0B9D0FFBB0A950" box="[936,1090,1065,1089]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="32" pageNumber="87" phylum="Arthropoda" rank="genus" sensu="stricto">Megaphyllum</taxonomicName>
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<taxonomicNameLabel id="A22657ACFFD9FF97FBEE9D0CFB72A950" box="[1101,1152,1066,1089]" pageId="32" pageNumber="87" sensu="stricto">s.str.</taxonomicNameLabel>
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</emphasis>
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species in the Caucasus (Golovatch 1992). However, there are doubtful records of
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<taxonomicName id="4C614D46FFD9FF97FCE69D68FC3EA974" box="[837,972,1101,1125]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="32" pageNumber="87" phylum="Arthropoda" rank="species" species="hercules">
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<emphasis id="B915EAD7FFD9FF97FCE69D68FC3EA974" box="[837,972,1101,1125]" italics="true" pageId="32" pageNumber="87">M. hercules</emphasis>
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</taxonomicName>
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from the region (Lignau 1903; Issajew 1911). Lohmander (1936) stated that mistaking the two species would not have been possible, so the two species may co-occur.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |