181 lines
31 KiB
XML
181 lines
31 KiB
XML
<document id="C025DE77990620B3307D4073A3EF94EB" ID-CLB-Dataset="32718" ID-DOI="10.11646/zootaxa.4290.2.1" ID-GBIF-Dataset="43ebda6c-201a-46af-8b54-e7980b8d79e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="828965" ID-ZooBank="52D6E3C3-BA0A-49C7-865C-00F3F5E5C5D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500110315309" checkinUser="plazi" docAuthor="Gunter, Nicole L. & Weir, Thomas A." docDate="2017" docId="03C887A8FFDDFFD1FF737C5D5D3BB4FF" docLanguage="en" docName="zootaxa.4290.2.1.pdf" docOrigin="Zootaxa 4290 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Monteithocanthon Gunter & Weir, new genus" docType="treatment" docVersion="7" lastPageNumber="223" masterDocId="FFF1FFD0FFCBFFC7FFE47F3A5F6FB348" masterDocTitle="Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species" masterLastPageNumber="243" masterPageNumber="201" pageNumber="223" updateTime="1698465402479" updateUser="ExternalLinkService">
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<mods:title id="D2A2DCB65BA23700BC5445CE04227212">Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species</mods:title>
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<mods:namePart id="B04647E0792A569A3F311CAA84642D14">Gunter, Nicole L.</mods:namePart>
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<mods:roleTerm id="BA2118A992FB8585E7D4031274BD2F1C">Author</mods:roleTerm>
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<mods:namePart id="77231341C95816E487DA3DE83F47E962">Weir, Thomas A.</mods:namePart>
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<mods:title id="BF7E51F54B018ECF3FB95567529A1A63">Zootaxa</mods:title>
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<mods:date id="05C3A177D91E49D8465FCD12523FD040">2017</mods:date>
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<mods:number id="3CE505389F8169927E7126F8E5BBDCA6">4290</mods:number>
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<treatment id="03C887A8FFDDFFD1FF737C5D5D3BB4FF" ID-DOI="http://doi.org/10.5281/zenodo.6033008" ID-GBIF-Taxon="132549203" ID-Zenodo-Dep="6033008" LSID="urn:lsid:plazi:treatment:03C887A8FFDDFFD1FF737C5D5D3BB4FF" httpUri="http://treatment.plazi.org/id/03C887A8FFDDFFD1FF737C5D5D3BB4FF" lastPageNumber="223" pageId="22" pageNumber="223">
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<subSubSection id="C37B6535FFDDFFD1FF737C5D5DA2B0CA" box="[151,717,871,898]" pageId="22" pageNumber="223" type="nomenclature">
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<paragraph id="8BDE36BEFFDDFFD1FF737C5D5DA2B0CA" blockId="22.[151,717,871,898]" box="[151,717,871,898]" pageId="22" pageNumber="223">
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<heading id="D09681D2FFDDFFD1FF737C5D5DA2B0CA" bold="true" box="[151,717,871,898]" fontSize="11" level="1" pageId="22" pageNumber="223" reason="1">
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<emphasis id="B915EAACFFDDFFD1FF737C5D5DA2B0CA" bold="true" box="[151,717,871,898]" pageId="22" pageNumber="223">
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<taxonomicName id="4C614D3DFFDDFFD1FF737C5D5D51B0CA" ID-CoL="63CZ2" authority="Gunter & Weir" authorityName="Gunter & Weir" box="[151,574,871,898]" class="Insecta" family="Scarabaeidae" genus="Monteithocanthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B915EAACFFDDFFD1FF737C5D5E17B0C9" bold="true" box="[151,376,871,897]" italics="true" pageId="22" pageNumber="223">Monteithocanthon</emphasis>
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Gunter & Weir
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22657D7FFDDFFD1FDAF7C525DA2B0CA" box="[587,717,872,898]" pageId="22" pageNumber="223" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B6535FFDDFFD1FF737C945DDBB08F" box="[151,692,942,967]" pageId="22" pageNumber="223" type="materials_examined">
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<paragraph id="8BDE36BEFFDDFFD1FF737C945DDBB08F" blockId="22.[151,1437,942,1975]" box="[151,692,942,967]" pageId="22" pageNumber="223">
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<emphasis id="B915EAACFFDDFFD1FF737C945E5DB08F" bold="true" box="[151,306,942,967]" pageId="22" pageNumber="223">Type species.</emphasis>
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<taxonomicName id="4C614D3DFFDDFFD1FEDD7C955DDBB08F" authority="Matthews, 1974" authorityName="Matthews" authorityYear="1974" box="[313,692,942,967]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="arator">
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<emphasis id="B915EAACFFDDFFD1FEDD7C955E89B08E" box="[313,486,943,966]" italics="true" pageId="22" pageNumber="223">Lepanus arator</emphasis>
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(
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<bibRefCitation id="EFF04B4FFFDDFFD1FE117C945DC3B08F" author="Matthews" box="[501,684,942,967]" pageId="22" pageNumber="223" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini. Australian Journal of Zoology, Supplementary Series, 24, 1 - 211." type="journal article" year="1974">Matthews, 1974</bibRefCitation>
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)
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B6535FFDDFFD1FF237CE85F83B747" pageId="22" pageNumber="223" type="etymology">
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<paragraph id="8BDE36BEFFDDFFD1FF237CE85F83B747" blockId="22.[151,1437,942,1975]" pageId="22" pageNumber="223">
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<emphasis id="B915EAACFFDDFFD1FF237CE85E21B0A3" bold="true" box="[199,334,978,1003]" pageId="22" pageNumber="223">Etymology.</emphasis>
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Named in honour of Geoff Monteith for his contribution to the taxonomy of Australian dung beetles.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B6535FFDDFFD1FF237B205BCEB54F" pageId="22" pageNumber="223" type="diagnosis">
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<paragraph id="8BDE36BEFFDDFFD1FF237B205BCEB54F" blockId="22.[151,1437,942,1975]" pageId="22" pageNumber="223">
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<emphasis id="B915EAACFFDDFFD1FF237B205E2EB77B" bold="true" box="[199,321,1050,1075]" pageId="22" pageNumber="223">Diagnosis.</emphasis>
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Small, total length
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<quantity id="4C999B5BFFDDFFD1FDC67B215DC1B77A" box="[546,686,1051,1075]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.2" metricValueMin="2.1" pageId="22" pageNumber="223" unit="mm" value="2.65" valueMax="3.2" valueMin="2.1">2.1–3.2 mm</quantity>
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. Elytra with 8 striae, 7 on disc and one on edge of epipleura (
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<figureCitation id="135A2A3BFFDDFFD1FA8B7B205FC9B71F" captionStart="FIGURE 7" captionStartId="25.[151,250,1777,1799]" captionTargetBox="[154,1418,200,1756]" captionTargetId="figure@25.[154,1433,193,1756]" captionTargetPageId="25" captionText="FIGURE 7. Key characters of Monteithocanthon new genus, all scale bars 0.25 mm. A, lateral view of elytra (M. arator); B, ventral view of profemur, note the absence of trochantofemoral anterior pit and presence of ornamentation on apex of protibia (M. arator); C, close up of ornamentation on apex of protibia, arrows pointing to comb and brush of setae (M. arator); D, dorsal view of male protibia with three tibial teeth each with only one group of setae dorsally (M. arator ♂); E, ventral view of male protibia (M. arator ♂); F, dorsal view of female protibia, note the apical spur is long and narrow (M. arator ♀); G, view of narrow anterior margin of lateral lobe of mesoventrite (M. arator); H, dorsal view of male mesotibia (M. arator ♂); I, ventral view of male mesotibia (M. arator ♂); J, male metatibia (M. arator ♂); K, view of broad anterior margin of lateral lobe of metaventrite (M. koombooloomba)." httpUri="https://zenodo.org/record/828979/files/figure.png" pageId="22" pageNumber="223">Fig. 7</figureCitation>
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A). Seventh elytral striae obsolete (
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<figureCitation id="135A2A3BFFDDFFD1FDCA7B045D1EB71F" box="[558,625,1086,1111]" captionStart="FIGURE 7" captionStartId="25.[151,250,1777,1799]" captionTargetBox="[154,1418,200,1756]" captionTargetId="figure@25.[154,1433,193,1756]" captionTargetPageId="25" captionText="FIGURE 7. Key characters of Monteithocanthon new genus, all scale bars 0.25 mm. A, lateral view of elytra (M. arator); B, ventral view of profemur, note the absence of trochantofemoral anterior pit and presence of ornamentation on apex of protibia (M. arator); C, close up of ornamentation on apex of protibia, arrows pointing to comb and brush of setae (M. arator); D, dorsal view of male protibia with three tibial teeth each with only one group of setae dorsally (M. arator ♂); E, ventral view of male protibia (M. arator ♂); F, dorsal view of female protibia, note the apical spur is long and narrow (M. arator ♀); G, view of narrow anterior margin of lateral lobe of mesoventrite (M. arator); H, dorsal view of male mesotibia (M. arator ♂); I, ventral view of male mesotibia (M. arator ♂); J, male metatibia (M. arator ♂); K, view of broad anterior margin of lateral lobe of metaventrite (M. koombooloomba)." httpUri="https://zenodo.org/record/828979/files/figure.png" pageId="22" pageNumber="223">Fig. 7</figureCitation>
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A). Pygidium with transverse groove at the base not reaching basal angle (
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<figureCitation id="135A2A3BFFDDFFD1FF7B7B585F8AB733" box="[159,229,1122,1147]" captionStart="FIGURE 9" captionStartId="28.[151,250,1882,1904]" captionTargetBox="[161,1413,200,1843]" captionTargetId="figure@28.[151,1435,193,1860]" captionTargetPageId="28" captionText="FIGURE 9. Pygidium and aedeagus (left view, dorsal view, right view) of species of Monteithocanthon new genus. Scale bars of pygidium 0.5 mm, scale bars of aedeagus 0.2 mm. A, Monteithocanthon arator new combination; B, M. elliotensis new species; C, M. glaber new combination; D, M. koombooloomba new species; E, M. paraarator new species; F, M. peckorum new species." httpUri="https://zenodo.org/record/828983/files/figure.png" pageId="22" pageNumber="223">Fig. 9</figureCitation>
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). Head without basal carina. Dorsal parts of eyes narrow, 2–5 facets wide; narrowest at base of head; eye canthus dividing or nearly dividing eye. Antennae with 3-antennomere club with club antennomeres of approximately equal width. Protibiae of male with comb of stiff setae and ventral brush of setae at inner apex; apical spur small, inconspicuous (
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<figureCitation id="135A2A3BFFDDFFD1FDFE7BF45D31B7AF" box="[538,606,1230,1255]" captionStart="FIGURE 7" captionStartId="25.[151,250,1777,1799]" captionTargetBox="[154,1418,200,1756]" captionTargetId="figure@25.[154,1433,193,1756]" captionTargetPageId="25" captionText="FIGURE 7. Key characters of Monteithocanthon new genus, all scale bars 0.25 mm. A, lateral view of elytra (M. arator); B, ventral view of profemur, note the absence of trochantofemoral anterior pit and presence of ornamentation on apex of protibia (M. arator); C, close up of ornamentation on apex of protibia, arrows pointing to comb and brush of setae (M. arator); D, dorsal view of male protibia with three tibial teeth each with only one group of setae dorsally (M. arator ♂); E, ventral view of male protibia (M. arator ♂); F, dorsal view of female protibia, note the apical spur is long and narrow (M. arator ♀); G, view of narrow anterior margin of lateral lobe of mesoventrite (M. arator); H, dorsal view of male mesotibia (M. arator ♂); I, ventral view of male mesotibia (M. arator ♂); J, male metatibia (M. arator ♂); K, view of broad anterior margin of lateral lobe of metaventrite (M. koombooloomba)." httpUri="https://zenodo.org/record/828979/files/figure.png" pageId="22" pageNumber="223">Fig. 7</figureCitation>
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C–D). Protibiae with 3 teeth on outer edge, not equidistant with apical 2 slightly closer together; serration between teeth and proximal to them (
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<figureCitation id="135A2A3BFFDDFFD1FC557BC85C9CB643" box="[945,1011,1266,1291]" captionStart="FIGURE 7" captionStartId="25.[151,250,1777,1799]" captionTargetBox="[154,1418,200,1756]" captionTargetId="figure@25.[154,1433,193,1756]" captionTargetPageId="25" captionText="FIGURE 7. Key characters of Monteithocanthon new genus, all scale bars 0.25 mm. A, lateral view of elytra (M. arator); B, ventral view of profemur, note the absence of trochantofemoral anterior pit and presence of ornamentation on apex of protibia (M. arator); C, close up of ornamentation on apex of protibia, arrows pointing to comb and brush of setae (M. arator); D, dorsal view of male protibia with three tibial teeth each with only one group of setae dorsally (M. arator ♂); E, ventral view of male protibia (M. arator ♂); F, dorsal view of female protibia, note the apical spur is long and narrow (M. arator ♀); G, view of narrow anterior margin of lateral lobe of mesoventrite (M. arator); H, dorsal view of male mesotibia (M. arator ♂); I, ventral view of male mesotibia (M. arator ♂); J, male metatibia (M. arator ♂); K, view of broad anterior margin of lateral lobe of metaventrite (M. koombooloomba)." httpUri="https://zenodo.org/record/828979/files/figure.png" pageId="22" pageNumber="223">Fig. 7</figureCitation>
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D–F). Each tooth has 1 group of setae dorsally (
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<figureCitation id="135A2A3BFFDDFFD1FEE57A2C5E2AB667" box="[257,325,1302,1327]" captionStart="FIGURE 7" captionStartId="25.[151,250,1777,1799]" captionTargetBox="[154,1418,200,1756]" captionTargetId="figure@25.[154,1433,193,1756]" captionTargetPageId="25" captionText="FIGURE 7. Key characters of Monteithocanthon new genus, all scale bars 0.25 mm. A, lateral view of elytra (M. arator); B, ventral view of profemur, note the absence of trochantofemoral anterior pit and presence of ornamentation on apex of protibia (M. arator); C, close up of ornamentation on apex of protibia, arrows pointing to comb and brush of setae (M. arator); D, dorsal view of male protibia with three tibial teeth each with only one group of setae dorsally (M. arator ♂); E, ventral view of male protibia (M. arator ♂); F, dorsal view of female protibia, note the apical spur is long and narrow (M. arator ♀); G, view of narrow anterior margin of lateral lobe of mesoventrite (M. arator); H, dorsal view of male mesotibia (M. arator ♂); I, ventral view of male mesotibia (M. arator ♂); J, male metatibia (M. arator ♂); K, view of broad anterior margin of lateral lobe of metaventrite (M. koombooloomba)." httpUri="https://zenodo.org/record/828979/files/figure.png" pageId="22" pageNumber="223">Fig. 7</figureCitation>
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D). Distal edge of protibiae straight and trochantofemoral anterior pit absent (
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<figureCitation id="135A2A3BFFDDFFD1FB507A2C5B97B667" box="[1204,1272,1302,1327]" captionStart="FIGURE 7" captionStartId="25.[151,250,1777,1799]" captionTargetBox="[154,1418,200,1756]" captionTargetId="figure@25.[154,1433,193,1756]" captionTargetPageId="25" captionText="FIGURE 7. Key characters of Monteithocanthon new genus, all scale bars 0.25 mm. A, lateral view of elytra (M. arator); B, ventral view of profemur, note the absence of trochantofemoral anterior pit and presence of ornamentation on apex of protibia (M. arator); C, close up of ornamentation on apex of protibia, arrows pointing to comb and brush of setae (M. arator); D, dorsal view of male protibia with three tibial teeth each with only one group of setae dorsally (M. arator ♂); E, ventral view of male protibia (M. arator ♂); F, dorsal view of female protibia, note the apical spur is long and narrow (M. arator ♀); G, view of narrow anterior margin of lateral lobe of mesoventrite (M. arator); H, dorsal view of male mesotibia (M. arator ♂); I, ventral view of male mesotibia (M. arator ♂); J, male metatibia (M. arator ♂); K, view of broad anterior margin of lateral lobe of metaventrite (M. koombooloomba)." httpUri="https://zenodo.org/record/828979/files/figure.png" pageId="22" pageNumber="223">Fig. 7</figureCitation>
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B). Trochanter excavated dorsally for reception of inner apical edge of protibia. Protarsi with 5 tarsomeres, tarsomere 5 the longest. Metatarsomere 1 wider than 2–5, metatarsomere 2 equal to tarsomere
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<quantity id="4C999B5BFFDDFFD1FBB07A645BE9B63F" box="[1108,1158,1374,1399]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="22" pageNumber="223" unit="in" value="5.0">5 in</quantity>
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length (
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<figureCitation id="135A2A3BFFDDFFD1FB147A645A55B63F" box="[1264,1338,1374,1399]" captionStart="FIGURE 7" captionStartId="25.[151,250,1777,1799]" captionTargetBox="[154,1418,200,1756]" captionTargetId="figure@25.[154,1433,193,1756]" captionTargetPageId="25" captionText="FIGURE 7. Key characters of Monteithocanthon new genus, all scale bars 0.25 mm. A, lateral view of elytra (M. arator); B, ventral view of profemur, note the absence of trochantofemoral anterior pit and presence of ornamentation on apex of protibia (M. arator); C, close up of ornamentation on apex of protibia, arrows pointing to comb and brush of setae (M. arator); D, dorsal view of male protibia with three tibial teeth each with only one group of setae dorsally (M. arator ♂); E, ventral view of male protibia (M. arator ♂); F, dorsal view of female protibia, note the apical spur is long and narrow (M. arator ♀); G, view of narrow anterior margin of lateral lobe of mesoventrite (M. arator); H, dorsal view of male mesotibia (M. arator ♂); I, ventral view of male mesotibia (M. arator ♂); J, male metatibia (M. arator ♂); K, view of broad anterior margin of lateral lobe of metaventrite (M. koombooloomba)." httpUri="https://zenodo.org/record/828979/files/figure.png" pageId="22" pageNumber="223">Fig. 7</figureCitation>
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J). Male mesotibiae with dense setae on inner edge (
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<figureCitation id="135A2A3BFFDDFFD1FD667AB85DA8B6D3" box="[642,711,1410,1435]" captionStart="FIGURE 7" captionStartId="25.[151,250,1777,1799]" captionTargetBox="[154,1418,200,1756]" captionTargetId="figure@25.[154,1433,193,1756]" captionTargetPageId="25" captionText="FIGURE 7. Key characters of Monteithocanthon new genus, all scale bars 0.25 mm. A, lateral view of elytra (M. arator); B, ventral view of profemur, note the absence of trochantofemoral anterior pit and presence of ornamentation on apex of protibia (M. arator); C, close up of ornamentation on apex of protibia, arrows pointing to comb and brush of setae (M. arator); D, dorsal view of male protibia with three tibial teeth each with only one group of setae dorsally (M. arator ♂); E, ventral view of male protibia (M. arator ♂); F, dorsal view of female protibia, note the apical spur is long and narrow (M. arator ♀); G, view of narrow anterior margin of lateral lobe of mesoventrite (M. arator); H, dorsal view of male mesotibia (M. arator ♂); I, ventral view of male mesotibia (M. arator ♂); J, male metatibia (M. arator ♂); K, view of broad anterior margin of lateral lobe of metaventrite (M. koombooloomba)." httpUri="https://zenodo.org/record/828979/files/figure.png" pageId="22" pageNumber="223">
|
||
Fig.
|
||
<date id="FFDF107EFFDDFFD1FD537AB95DA8B6D3" box="[695,711,1411,1435]" pageId="22" pageNumber="223">7</date>
|
||
</figureCitation>
|
||
I). Claws toothed (
|
||
<figureCitation id="135A2A3BFFDDFFD1FC7C7AB85CB3B6D3" box="[920,988,1410,1435]" captionStart="FIGURE 7" captionStartId="25.[151,250,1777,1799]" captionTargetBox="[154,1418,200,1756]" captionTargetId="figure@25.[154,1433,193,1756]" captionTargetPageId="25" captionText="FIGURE 7. Key characters of Monteithocanthon new genus, all scale bars 0.25 mm. A, lateral view of elytra (M. arator); B, ventral view of profemur, note the absence of trochantofemoral anterior pit and presence of ornamentation on apex of protibia (M. arator); C, close up of ornamentation on apex of protibia, arrows pointing to comb and brush of setae (M. arator); D, dorsal view of male protibia with three tibial teeth each with only one group of setae dorsally (M. arator ♂); E, ventral view of male protibia (M. arator ♂); F, dorsal view of female protibia, note the apical spur is long and narrow (M. arator ♀); G, view of narrow anterior margin of lateral lobe of mesoventrite (M. arator); H, dorsal view of male mesotibia (M. arator ♂); I, ventral view of male mesotibia (M. arator ♂); J, male metatibia (M. arator ♂); K, view of broad anterior margin of lateral lobe of metaventrite (M. koombooloomba)." httpUri="https://zenodo.org/record/828979/files/figure.png" pageId="22" pageNumber="223">Fig. 7</figureCitation>
|
||
C). Male metatibiae generally bent with an extension but almost straight in
|
||
<taxonomicName id="4C614D3DFFDDFFD1FDFA7A9D5DDEB6F7" box="[542,689,1446,1471]" class="Insecta" family="Scarabaeidae" genus="Monteithocanthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="peckorum">
|
||
<emphasis id="B915EAACFFDDFFD1FDFA7A9D5DDEB6F7" box="[542,689,1446,1471]" italics="true" pageId="22" pageNumber="223">M. peckorum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="135A2A3BFFDDFFD1FD257A9C5C6BB6F7" box="[705,772,1446,1471]" captionStart="FIGURE 8" captionStartId="26.[151,250,955,977]" captionTargetBox="[192,1404,193,933]" captionTargetId="figure@26.[182,1404,193,933]" captionTargetPageId="26" captionText="FIGURE 8. Male metatibia of species of Monteithocanthon new genus, all scale bars 0.25 mm. A, Monteithocanthon arator new combination; B, M. elliotensis new species; C, M. glaber new combination; D, M. koombooloomba new species; E, M. paraarator new species; F, M. peckorum new species; G, female metathoracic leg of M. arator." httpUri="https://zenodo.org/record/828981/files/figure.png" pageId="22" pageNumber="223">Fig. 8</figureCitation>
|
||
). Mesometasternal suture procurved to strongly procurved. Mesepimera impunctate. Aedeagus with parameres asymmetrical, right paramere apically bent downwards and more-or-less expanded apically as in
|
||
<figureCitation id="135A2A3BFFDDFFD1FDDC7AD45D13B54F" box="[568,636,1518,1543]" captionStart="FIGURE 9" captionStartId="28.[151,250,1882,1904]" captionTargetBox="[161,1413,200,1843]" captionTargetId="figure@28.[151,1435,193,1860]" captionTargetPageId="28" captionText="FIGURE 9. Pygidium and aedeagus (left view, dorsal view, right view) of species of Monteithocanthon new genus. Scale bars of pygidium 0.5 mm, scale bars of aedeagus 0.2 mm. A, Monteithocanthon arator new combination; B, M. elliotensis new species; C, M. glaber new combination; D, M. koombooloomba new species; E, M. paraarator new species; F, M. peckorum new species." httpUri="https://zenodo.org/record/828983/files/figure.png" pageId="22" pageNumber="223">Fig. 9</figureCitation>
|
||
. Hind wings fully developed, reduced, or absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B6535FFDDFFD1FF2379285D91B403" pageId="22" pageNumber="223" type="discussion">
|
||
<paragraph id="8BDE36BEFFDDFFD1FF2379285D91B403" blockId="22.[151,1437,942,1975]" pageId="22" pageNumber="223">
|
||
<emphasis id="B915EAACFFDDFFD1FF2379285E54B563" bold="true" box="[199,315,1554,1579]" pageId="22" pageNumber="223">Remarks.</emphasis>
|
||
<bibRefCitation id="EFF04B4FFFDDFFD1FEA779285D6BB563" author="Matthews" box="[323,516,1554,1579]" pageId="22" pageNumber="223" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini. Australian Journal of Zoology, Supplementary Series, 24, 1 - 211." type="journal article" year="1974">Matthews (1974)</bibRefCitation>
|
||
retained the genus
|
||
<taxonomicName id="4C614D3DFFDDFFD1FD06792E5C2EB563" box="[738,833,1556,1579]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EAACFFDDFFD1FD06792E5C2EB563" box="[738,833,1556,1579]" italics="true" pageId="22" pageNumber="223">Lepanus</emphasis>
|
||
</taxonomicName>
|
||
for practical purposes noting that adhering to a strict generic concept in such a morphologically diverse genus would lead to fragmentation into several small, difficult groups. All species of
|
||
<taxonomicName id="4C614D3DFFDDFFD1FE7F79605D06B53B" box="[411,617,1626,1651]" class="Insecta" family="Scarabaeidae" genus="Monteithocanthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="230" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B915EAACFFDDFFD1FE7F79605D06B53B" box="[411,617,1626,1651]" italics="true" pageId="22" pageNumber="223">Monteithocanthon</emphasis>
|
||
</taxonomicName>
|
||
share many morphological affinities with
|
||
<taxonomicName id="4C614D3DFFDDFFD1FBB579665BDFB53B" box="[1105,1200,1628,1651]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EAACFFDDFFD1FBB579665BDFB53B" box="[1105,1200,1628,1651]" italics="true" pageId="22" pageNumber="223">Lepanus</emphasis>
|
||
</taxonomicName>
|
||
so it is unsurprising that
|
||
<taxonomicName id="4C614D3DFFDDFFD1FF2C79455E5AB5DE" box="[200,309,1663,1686]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="arator">
|
||
<emphasis id="B915EAACFFDDFFD1FF2C79455E5AB5DE" box="[200,309,1663,1686]" italics="true" pageId="22" pageNumber="223">M. arator</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C614D3DFFDDFFD1FE8879455EB4B5DF" box="[364,475,1662,1687]" class="Insecta" family="Scarabaeidae" genus="Monteithocanthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="glaber">
|
||
<emphasis id="B915EAACFFDDFFD1FE8879455EB4B5DF" box="[364,475,1662,1687]" italics="true" pageId="22" pageNumber="223">M. glaber</emphasis>
|
||
</taxonomicName>
|
||
were originally described within the genus
|
||
<taxonomicName id="4C614D3DFFDDFFD1FC2379455B49B5DE" box="[967,1062,1663,1686]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EAACFFDDFFD1FC2379455B49B5DE" box="[967,1062,1663,1686]" italics="true" pageId="22" pageNumber="223">Lepanus</emphasis>
|
||
</taxonomicName>
|
||
. A new genus is established here primarily on the basis of genetic results, with the two
|
||
<taxonomicName id="4C614D3DFFDDFFD1FD1379985CABB5F3" box="[759,964,1698,1723]" class="Insecta" family="Scarabaeidae" genus="Monteithocanthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="230" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B915EAACFFDDFFD1FD1379985CABB5F3" box="[759,964,1698,1723]" italics="true" pageId="22" pageNumber="223">Monteithocanthon</emphasis>
|
||
</taxonomicName>
|
||
species sequenced to date always distinct from the clade containing the majority of
|
||
<taxonomicName id="4C614D3DFFDDFFD1FD9679FD5DBEB596" box="[626,721,1735,1758]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915EAACFFDDFFD1FD9679FD5DBEB596" box="[626,721,1735,1758]" italics="true" pageId="22" pageNumber="223">Lepanus</emphasis>
|
||
</taxonomicName>
|
||
species. The phylogenetic grouping and the establishment of a new genus are supported by morphology. The six species assigned here to
|
||
<taxonomicName id="4C614D3DFFDDFFD1FC1179D05BADB44B" box="[1013,1218,1770,1795]" class="Insecta" family="Scarabaeidae" genus="Monteithocanthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="230" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B915EAACFFDDFFD1FC1179D05BADB44B" box="[1013,1218,1770,1795]" italics="true" pageId="22" pageNumber="223">Monteithocanthon</emphasis>
|
||
</taxonomicName>
|
||
share a number of morphological features in combination, although many of these diagnostic characters are also individually found in other Australian ‘deltochiline-like’ genera and species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B6535FFDDFFD1FF23786C5D3BB4FF" pageId="22" pageNumber="223" type="distribution">
|
||
<paragraph id="8BDE36BEFFDDFFD1FF23786C5D3BB4FF" blockId="22.[151,1437,942,1975]" pageId="22" pageNumber="223">
|
||
<emphasis id="B915EAACFFDDFFD1FF23786C5E0FB427" bold="true" box="[199,352,1878,1903]" pageId="22" pageNumber="223">Distribution.</emphasis>
|
||
The genus is known only from
|
||
<collectingCountry id="F376762EFFDDFFD1FD25786C5C44B427" box="[705,811,1878,1903]" name="Australia" pageId="22" pageNumber="223">Australia</collectingCountry>
|
||
. Four species occur in northern
|
||
<collectingRegion id="49A5F85CFFDDFFD1FB69786C5A7EB427" box="[1165,1297,1878,1903]" country="Australia" name="Queensland" pageId="22" pageNumber="223">Queensland</collectingRegion>
|
||
primarily in the Wet Tropics, one species is distributed at high altitude in southeastern
|
||
<collectingRegion id="49A5F85CFFDDFFD1FBC778405BC5B4DB" box="[1059,1194,1914,1939]" country="Australia" name="Queensland" pageId="22" pageNumber="223">Queensland</collectingRegion>
|
||
, and one species in southeastern
|
||
<collectingRegion id="49A5F85CFFDDFFD1FECE78A55E9CB4FF" box="[298,499,1950,1975]" country="Australia" name="Western Australia" pageId="22" pageNumber="223">
|
||
<collectingRegion id="49A5F85CFFDDFFD1FECE78A55EEAB4FE" box="[298,389,1951,1974]" country="American Samoa" name="Western" pageId="22" pageNumber="223">Western</collectingRegion>
|
||
<collectingCountry id="F376762EFFDDFFD1FE6878A45E9CB4FF" box="[396,499,1950,1975]" name="Australia" pageId="22" pageNumber="223">Australia</collectingCountry>
|
||
</collectingRegion>
|
||
(
|
||
<figureCitation id="135A2A3BFFDDFFD1FDE678A45D29B4FF" box="[514,582,1950,1975]" captionStart="FIGURE 5" captionStartId="13.[151,250,1554,1576]" captionTargetBox="[170,1385,589,1501]" captionTargetId="figure@13.[151,1436,589,1532]" captionTargetPageId="13" captionText="FIGURE 5. Distributional maps of species belonging to the new genera Matthewsius and Monteithocanthon including close up insets of A, Pilbara region, Western Australia; B, northeastern Queensland; C, southeastern Queensland; D, southwestern Western Australia; E, southeastern South Australia; F, central to southeastern New South Wales." httpUri="https://zenodo.org/record/828975/files/figure.png" pageId="22" pageNumber="223">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |