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<document id="A23025D671959D5983B1E24B6BFDA37D" ID-CLB-Dataset="32031" ID-DOI="10.11646/zootaxa.4323.2.3" ID-GBIF-Dataset="a2533170-b900-4d5a-9ce4-feb70e66daa3" ID-ISSN="1175-5326" ID-Zenodo-Dep="898900" ID-ZooBank="Ddd775C4-1A34-4034-A65A-3D9C6E038Fbd" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505970751217" checkinUser="plazi" docAuthor="Chen, Hui-Ming" docDate="2017" docId="03C887A1FE14FFF0FF31E928FD3FFE43" docLanguage="en" docName="zootaxa.4323.2.3.pdf" docOrigin="Zootaxa 4323 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glyphiulus latus Chen 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="200" masterDocId="FFF1FFD9FE15FFF3FFA6EA7EFF81FF93" masterDocTitle="Two new species of the millipede genus Glyphiulus Gervais, 1847 from Southwest China (Diplopoda: Spirostreptida: Cambalopsidae)" masterLastPageNumber="208" masterPageNumber="197" pageNumber="198" updateTime="1698468815119" updateUser="ExternalLinkService">
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<mods:title id="EC32EE7F5B373E410D75F89FCE5A6EE5">Two new species of the millipede genus Glyphiulus Gervais, 1847 from Southwest China (Diplopoda: Spirostreptida: Cambalopsidae)</mods:title>
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<mods:namePart id="59DB7CB6568B4975F59BBDCB48511ADC">Chen, Hui-Ming</mods:namePart>
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<treatment id="03C887A1FE14FFF0FF31E928FD3FFE43" ID-DOI="http://doi.org/10.5281/zenodo.6034184" ID-GBIF-Taxon="134837848" ID-Zenodo-Dep="6034184" LSID="urn:lsid:plazi:treatment:03C887A1FE14FFF0FF31E928FD3FFE43" httpUri="http://treatment.plazi.org/id/03C887A1FE14FFF0FF31E928FD3FFE43" lastPageId="3" lastPageNumber="200" pageId="1" pageNumber="198">
<subSubSection id="C37B653CFE14FFF2FF31E928FDE7FC01" pageId="1" pageNumber="198" type="nomenclature">
<paragraph id="8BDE36B7FE14FFF2FF31E928FE42FCE3" blockId="1.[151,614,854,914]" box="[151,451,854,880]" pageId="1" pageNumber="198">
<heading id="D09681DBFE14FFF2FF31E928FE42FCE3" bold="true" box="[151,451,854,880]" fontSize="11" level="1" pageId="1" pageNumber="198" reason="1">
<emphasis id="B915EAA5FE14FFF2FF31E928FE42FCE3" bold="true" box="[151,451,854,880]" pageId="1" pageNumber="198">
<taxonomicName id="4C614D34FE14FFF2FF31E928FEDEFCE3" authority="Chen, 2017" authorityName="Chen" authorityYear="2017" box="[151,351,854,880]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
<emphasis id="B915EAA5FE14FFF2FF31E928FEDEFCE3" bold="true" box="[151,351,854,880]" italics="true" pageId="1" pageNumber="198">Glyphiulus latus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22657DEFE14FFF2FEC3E928FE42FCE3" box="[357,451,854,880]" pageId="1" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDE36B7FE14FFF2FF31E907FDE7FC01" blockId="1.[151,614,854,914]" box="[151,614,889,914]" pageId="1" pageNumber="198">
<figureCitation id="135A2A32FE14FFF2FF31E907FE8FFC01" box="[151,270,889,914]" captionStart="FIGURE 1" captionStartId="2.[151,250,1953,1975]" captionTargetBox="[151,1436,483,1928]" captionTargetId="figure@2.[151,1436,483,1928]" captionTargetPageId="2" captionText="FIGURE 1 A F. Glyphiulus latus sp. nov., holotype. A. anterior part of body, lateral view; B. same, dorsal view; C. midbody segments, lateral view; D. same, dorsal view; E. posterior part of body, lateral view; F. same, ventral view." httpUri="https://zenodo.org/record/898902/files/figure.png" pageId="1" pageNumber="198">Figs 1AF</figureCitation>
,
<figureCitation id="135A2A32FE14FFF2FEBCE904FEE0FC01" box="[282,353,890,914]" captionStart="FIGURE 2" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[163,1423,511,1927]" captionTargetId="figure@3.[163,1423,511,1927]" captionTargetPageId="3" captionText="FIGURE 2 A D. Glyphiulus latus sp. nov., anterior part of body, lateral view. A. female paratype; B. male holotype; C. another female paratype; D. male paratype. Green arrows show the variation of the lateralmost crest of collum. Blue arrows indicate the presence of the ectoparasitic fungi, order Entomophthorales." httpUri="https://zenodo.org/record/898904/files/figure.png" pageId="1" pageNumber="198">2AD</figureCitation>
,
<figureCitation id="135A2A32FE14FFF2FECAE904FE2DFC01" box="[364,428,890,914]" captionStart="FIGURE 3" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[151,1434,193,1944]" captionTargetId="figure@4.[151,1436,193,1944]" captionTargetPageId="4" captionText="FIGURE 3 A F. Glyphiulus latus sp. nov., holotype. A. gnathochilarium, ventral view; B. legs 1, anterior view; C. legs 2, caudal view; D. legs 3, caudal view; E. cross-section of a midbody segment, caudal view; F. midbody leg, lateral view." httpUri="https://zenodo.org/record/898906/files/figure.png" pageId="1" pageNumber="198">3AF</figureCitation>
,
<figureCitation id="135A2A32FE14FFF2FE11E904FE7CFC01" box="[439,509,890,914]" captionStart="FIGURE 4" captionStartId="5.[151,250,743,765]" captionTargetBox="[313,1275,184,738]" captionTargetId="figure@5.[312,1276,183,739]" captionTargetPageId="5" captionText="FIGURE 4 A B. Glyphiulus latus sp. nov., holotype. A. anterior gonopods, caudal view; B. posterior gonopods, caudal view." httpUri="https://zenodo.org/record/898908/files/figure.png" pageId="1" pageNumber="198">4AB</figureCitation>
,
<figureCitation id="135A2A32FE14FFF2FDAEE907FDCDFC01" box="[520,588,889,914]" captionStart="FIGURE 5" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[335,1273,830,1888]" captionTargetId="figure@5.[303,1291,810,1907]" captionTargetPageId="5" captionText="FIGURE 5 A E. Glyphiulus latus sp. nov., holotype. A. anterior gonopods, caudal view; B. posterior gonopods, caudal view; C. clypeus, anterior view; D. antenna, lateral view; E. legs 2, caudal view. Abbreviations: BS, bacilliform sensilla; c, coxosterna; cmp, coxosternal mesal processes; F, flagellum; L, lamelliform lobe; P, penis; T, telopodite. Scar bars: A, B, E = 0.2 mm; C = 0.1 mm; D = 0.5 mm." httpUri="https://zenodo.org/record/898910/files/figure.png" pageId="1" pageNumber="198">5AE</figureCitation>
,
<figureCitation id="135A2A32FE14FFF2FDFEE907FDE7FC01" box="[600,614,889,914]" captionStart="FIGURE 6" captionStartId="6.[151,250,1171,1193]" captionTargetBox="[151,1436,156,1159]" captionTargetId="figure@6.[151,1436,156,1159]" captionTargetPageId="6" captionText="FIGURE 6. Habitus of Glyphiulus latus sp. nov.." httpUri="https://zenodo.org/record/898912/files/figure.png" pageId="1" pageNumber="198">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37B653CFE14FFF2FF31E9BFFD83FBB1" pageId="1" pageNumber="198" type="materials_examined">
<paragraph id="8BDE36B7FE14FFF2FF31E9BFFD83FBB1" blockId="1.[151,1437,961,2030]" pageId="1" pageNumber="198">
<materialsCitation id="3B093CEAFE14FFF2FF31E9BFFB93FC6D" ID-GBIF-Occurrence="1653530305" collectingDate="2015-11-04" collectionCode="IBGAS" collectorName="H. M. Chen" country="China" county="Muchuan County" latitude="29.197002" location="Longgong Cave" longLatPrecision="1" longitude="103.69312" municipality="Leshan City" pageId="1" pageNumber="198" specimenCount="2" stateProvince="Sichuan Province" typeStatus="holotype">
<emphasis id="B915EAA5FE14FFF2FF31E9BFFE49FC49" bold="true" box="[151,456,961,986]" pageId="1" pageNumber="198">
<typeStatus id="54DA8815FE14FFF2FF31E9BFFF53FC49" box="[151,210,961,986]" pageId="1" pageNumber="198">Type</typeStatus>
material.
<typeStatus id="54DA8815FE14FFF2FEFDE9BFFE49FC49" box="[347,456,961,986]" pageId="1" pageNumber="198" type="holotype">Holotype</typeStatus>
</emphasis>
male,
<collectingCountry id="F3767627FE14FFF2FD85E9BFFDEBFC49" box="[547,618,961,986]" name="China" pageId="1" pageNumber="198">China</collectingCountry>
:
<collectingRegion id="49A5F855FE14FFF2FDDAE9BFFCCDFC49" box="[636,844,961,986]" country="China" name="Sichuan" pageId="1" pageNumber="198">Sichuan Province</collectingRegion>
,
<collectingMunicipality id="6BBAACCDFE14FFF2FCF8E9BFFC6EFC49" box="[862,1007,961,986]" pageId="1" pageNumber="198">Leshan City</collectingMunicipality>
,
<collectingCounty id="62BF4E3BFE14FFF2FBA4E9BFFB4FFC49" box="[1026,1230,961,986]" pageId="1" pageNumber="198">Muchuan County</collectingCounty>
,
<location id="8EBE606CFE14FFF2FB47E9BCFA1DFC49" LSID="urn:lsid:plazi:treatment:03C887A1FE14FFF0FF31E928FD3FFE43:8EBE606CFE14FFF2FB47E9BCFA1DFC49" box="[1249,1436,962,986]" country="China" county="Muchuan County" latitude="29.197002" longLatPrecision="1" longitude="103.69312" municipality="Leshan City" name="Longgong Cave" pageId="1" pageNumber="198" stateProvince="Sichuan Province">Longgong Cave</location>
(
<geoCoordinate id="EE555070FE14FFF2FF39E998FEC8FC6D" box="[159,329,998,1022]" direction="north" orientation="latitude" pageId="1" pageNumber="198" precision="1" value="29.197002">29°11'49.21&quot;N</geoCoordinate>
,
<geoCoordinate id="EE555070FE14FFF2FEF2E99BFD88FC6D" box="[340,521,997,1022]" direction="east" orientation="longitude" pageId="1" pageNumber="198" precision="1" value="103.69312">103°41'35.24&quot;E</geoCoordinate>
),
<collectingDate id="EF9BE99FFE14FFF2FDBAE998FD68FC6D" box="[540,745,997,1022]" pageId="1" pageNumber="198" value="2015-11-04">
<date id="FFDF1077FE14FFF2FDBAE998FD68FC6D" box="[540,745,997,1022]" pageId="1" pageNumber="198" value="2015-11-04">4 November 2015</date>
</collectingDate>
,
<collectorName id="26945361FE14FFF2FD52E998FCF0FC6D" box="[756,881,997,1022]" pageId="1" pageNumber="198">H.M. Chen</collectorName>
leg. (
<collectionCode id="ED70AE72FE14FFF2FC16E998FB8BFC6D" box="[944,1034,998,1022]" pageId="1" pageNumber="198">IBGAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B093CEAFE14FFF2FBBAE99BFE7FFBB1" ID-GBIF-Occurrence="1653530306" collectingDate="2015-11-04" collectionCode="IBGAS" collectorName="H. M. Chen" country="China" county="Muchuan County" latitude="29.197002" location="Longgong Cave" longLatPrecision="1" longitude="103.69312" municipality="Leshan City" pageId="1" pageNumber="198" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Sichuan Province" typeStatus="paratype">
<emphasis id="B915EAA5FE14FFF2FBBAE99BFB1CFC6D" bold="true" box="[1052,1181,997,1022]" pageId="1" pageNumber="198">
<typeStatus id="54DA8815FE14FFF2FBBAE99BFB18FC6D" box="[1052,1177,997,1022]" pageId="1" pageNumber="198" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D67FD3EFE14FFF2FB02E99BFB72FC6D" box="[1188,1267,997,1022]" pageId="1" pageNumber="198" type="male">2males</specimenCount>
and
<specimenCount id="9D67FD3EFE14FFF2FA8FE99BFA19FC6D" box="[1321,1432,997,1022]" pageId="1" pageNumber="198" type="female">6 females</specimenCount>
, same data as holotype (
<collectionCode id="ED70AE72FE14FFF2FE3AEE74FE77FBB1" box="[412,502,1034,1058]" pageId="1" pageNumber="198">IBGAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B653CFE14FFF2FF61EE53FCB5FBF9" pageId="1" pageNumber="198" type="etymology">
<paragraph id="8BDE36B7FE14FFF2FF61EE53FCB5FBF9" blockId="1.[151,1437,961,2030]" pageId="1" pageNumber="198">
<emphasis id="B915EAA5FE14FFF2FF61EE53FECFFBD5" bold="true" box="[199,334,1069,1094]" pageId="1" pageNumber="198">Etymology.</emphasis>
This specific name is derived from the Latin word
<taxonomicName id="4C614D34FE14FFF2FC33EE53FC4AFBD6" authorityName="Chen" authorityYear="2017" box="[917,971,1069,1093]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B915EAA5FE14FFF2FC33EE53FC4AFBD6" box="[917,971,1069,1093]" italics="true" pageId="1" pageNumber="198">latus</emphasis>
</taxonomicName>
, meaning broad and refers to the broad mesal process of the coxosternum of the anterior gonopods.
</paragraph>
</subSubSection>
<subSubSection id="C37B653CFE14FFF2FF61EE0BFF49FAD1" pageId="1" pageNumber="198" type="diagnosis">
<paragraph id="8BDE36B7FE14FFF2FF61EE0BFF49FAD1" blockId="1.[151,1437,961,2030]" pageId="1" pageNumber="198">
<emphasis id="B915EAA5FE14FFF2FF61EE0BFEC0FB1D" bold="true" box="[199,321,1141,1166]" pageId="1" pageNumber="198">Diagnosis.</emphasis>
The new species differ from other species in this genus except
<taxonomicName id="4C614D34FE14FFF2FBADEE0BFD0CFB21" authority="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel, 2011" authorityName="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel" authorityYear="2011" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="submediator">
<emphasis id="B915EAA5FE14FFF2FBADEE0BFA98FB1E" box="[1035,1305,1141,1165]" italics="true" pageId="1" pageNumber="198">Glyphiulus submediator</emphasis>
Golovatch, Geoffroy, Mauriès &amp; VandenSpiegel, 2011
</taxonomicName>
(
<bibRefCitation id="EFF04B46FE14FFF2FD04EEE7FC3BFB21" author="Golovatch" box="[674,954,1177,1202]" pageId="1" pageNumber="198" refString="Golovatch, S. I., Geoffroy, J. - J., Mauries, J. - P. &amp; VandenSpiegel, D. (2011 b) New species of the millipede genus Glyphiulus Gervais, 1847 from the javanicus - group (Diplopoda: Spirostreptida: Cambalopsidae). Arthropoda Selecta, 20 (3), 149 - 165." type="journal article" year="2011" yearSuffix="b">
Golovatch
<emphasis id="B915EAA5FE14FFF2FC83EEE4FCDCFB22" box="[805,861,1177,1201]" italics="true" pageId="1" pageNumber="198">et al</emphasis>
. 2011b
</bibRefCitation>
: 164, figs 1114) from
<collectingCountry id="F3767627FE14FFF2FB78EEE7FAC1FB21" box="[1246,1344,1177,1202]" name="Vietnam" pageId="1" pageNumber="198">Vietnam</collectingCountry>
, by the presence a prolonged and broad coxosternal mesal processes of the anterior gonopods. It can be differentiated from
<taxonomicName id="4C614D34FE14FFF2FF31EE9FFE28FB6A" authorityName="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel" authorityYear="2011" box="[151,425,1249,1273]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="submediator">
<emphasis id="B915EAA5FE14FFF2FF31EE9FFE28FB6A" box="[151,425,1249,1273]" italics="true" pageId="1" pageNumber="198">Glyphiulus submediator</emphasis>
</taxonomicName>
by the elongated flagellum of the posterior gonopods, male legs 1 with 5-segmented telopodites, metatergal carinotaxic formula: 2/2+I/i+3/3+I/i+2/2, and clypeus with four teeth anteromesally (
<figureCitation id="135A2A32FE14FFF2FAC9EF7BFF3AFAD1" captionStart="FIGURE 5" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[335,1273,830,1888]" captionTargetId="figure@5.[303,1291,810,1907]" captionTargetPageId="5" captionText="FIGURE 5 A E. Glyphiulus latus sp. nov., holotype. A. anterior gonopods, caudal view; B. posterior gonopods, caudal view; C. clypeus, anterior view; D. antenna, lateral view; E. legs 2, caudal view. Abbreviations: BS, bacilliform sensilla; c, coxosterna; cmp, coxosternal mesal processes; F, flagellum; L, lamelliform lobe; P, penis; T, telopodite. Scar bars: A, B, E = 0.2 mm; C = 0.1 mm; D = 0.5 mm." httpUri="https://zenodo.org/record/898910/files/figure.png" pageId="1" pageNumber="198">Fig. 5C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37B653CFE14FFF1FF61EF33FD20FE3E" lastPageId="2" lastPageNumber="199" pageId="1" pageNumber="198" type="description">
<paragraph id="8BDE36B7FE14FFF2FF61EF33FCA3FA1A" blockId="1.[151,1437,961,2030]" pageId="1" pageNumber="198">
<emphasis id="B915EAA5FE14FFF2FF61EF33FED8FAF5" bold="true" box="[199,345,1357,1382]" pageId="1" pageNumber="198">Description.</emphasis>
<typeStatus id="54DA8815FE14FFF2FEC5EF33FE4AFAF5" box="[355,459,1357,1382]" pageId="1" pageNumber="198" type="holotype">Holotype</typeStatus>
with 66p + 1a + T,
<quantity id="4C999B52FE14FFF2FD63EF30FC99FAF6" box="[709,792,1358,1382]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="198" unit="mm" value="47.0">47 mm</quantity>
long and
<quantity id="4C999B52FE14FFF2FC35EF30FC6CFAF6" box="[915,1005,1358,1382]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="198" unit="mm" value="2.3">2.3 mm</quantity>
wide;
<typeStatus id="54DA8815FE14FFF2FBE7EF30FB2DFAF5" box="[1089,1196,1358,1382]" pageId="1" pageNumber="198" type="paratype">paratypes</typeStatus>
(male, female) with 6067p + 14a + T, length
<quantity id="4C999B52FE14FFF2FE61EF0FFDC3FA1A" box="[455,578,1393,1418]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" metricValueMax="5.1" metricValueMin="4.1" pageId="1" pageNumber="198" unit="mm" value="46.0" valueMax="51.0" valueMin="41.0">4151 mm</quantity>
, width
<quantity id="4C999B52FE14FFF2FD32EF0FFC9FFA1A" box="[660,798,1393,1418]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="1" pageNumber="198" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.22.5 mm</quantity>
.
</paragraph>
<paragraph id="8BDE36B7FE14FFF2FF61EFEBFC1DF9AD" blockId="1.[151,1437,961,2030]" pageId="1" pageNumber="198">
Coloration yellow-brown to red-brown, tergal crests dark red-brown to castaneous brown (
<figureCitation id="135A2A32FE14FFF2FB56EFEBFAEFFA3D" box="[1264,1390,1429,1454]" captionStart="FIGURE 1" captionStartId="2.[151,250,1953,1975]" captionTargetBox="[151,1436,483,1928]" captionTargetId="figure@2.[151,1436,483,1928]" captionTargetPageId="2" captionText="FIGURE 1 A F. Glyphiulus latus sp. nov., holotype. A. anterior part of body, lateral view; B. same, dorsal view; C. midbody segments, lateral view; D. same, dorsal view; E. posterior part of body, lateral view; F. same, ventral view." httpUri="https://zenodo.org/record/898902/files/figure.png" pageId="1" pageNumber="198">Figs 1AF</figureCitation>
,
<figureCitation id="135A2A32FE14FFF2FAD8EFEBFA0FFA3D" box="[1406,1422,1429,1454]" captionStart="FIGURE 6" captionStartId="6.[151,250,1171,1193]" captionTargetBox="[151,1436,156,1159]" captionTargetId="figure@6.[151,1436,156,1159]" captionTargetPageId="6" captionText="FIGURE 6. Habitus of Glyphiulus latus sp. nov.." httpUri="https://zenodo.org/record/898912/files/figure.png" pageId="1" pageNumber="198">6</figureCitation>
). Clypeus with four teeth anteromedially (
<figureCitation id="135A2A32FE14FFF2FD34EFC7FD74FA41" box="[658,757,1465,1490]" captionStart="FIGURE 5" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[335,1273,830,1888]" captionTargetId="figure@5.[303,1291,810,1907]" captionTargetPageId="5" captionText="FIGURE 5 A E. Glyphiulus latus sp. nov., holotype. A. anterior gonopods, caudal view; B. posterior gonopods, caudal view; C. clypeus, anterior view; D. antenna, lateral view; E. legs 2, caudal view. Abbreviations: BS, bacilliform sensilla; c, coxosterna; cmp, coxosternal mesal processes; F, flagellum; L, lamelliform lobe; P, penis; T, telopodite. Scar bars: A, B, E = 0.2 mm; C = 0.1 mm; D = 0.5 mm." httpUri="https://zenodo.org/record/898910/files/figure.png" pageId="1" pageNumber="198">Fig. 5C</figureCitation>
). Eyes barely discernible as slightly convex tubercles, unpigmented, 510 on each side of head (
<figureCitation id="135A2A32FE14FFF2FD21EFA3FD69FA65" box="[647,744,1501,1526]" captionStart="FIGURE 1" captionStartId="2.[151,250,1953,1975]" captionTargetBox="[151,1436,483,1928]" captionTargetId="figure@2.[151,1436,483,1928]" captionTargetPageId="2" captionText="FIGURE 1 A F. Glyphiulus latus sp. nov., holotype. A. anterior part of body, lateral view; B. same, dorsal view; C. midbody segments, lateral view; D. same, dorsal view; E. posterior part of body, lateral view; F. same, ventral view." httpUri="https://zenodo.org/record/898902/files/figure.png" pageId="1" pageNumber="198">Figs 1A</figureCitation>
,
<figureCitation id="135A2A32FE14FFF2FD5EEFA0FCC0FA65" box="[760,833,1502,1526]" captionStart="FIGURE 2" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[163,1423,511,1927]" captionTargetId="figure@3.[163,1423,511,1927]" captionTargetPageId="3" captionText="FIGURE 2 A D. Glyphiulus latus sp. nov., anterior part of body, lateral view. A. female paratype; B. male holotype; C. another female paratype; D. male paratype. Green arrows show the variation of the lateralmost crest of collum. Blue arrows indicate the presence of the ectoparasitic fungi, order Entomophthorales." httpUri="https://zenodo.org/record/898904/files/figure.png" pageId="1" pageNumber="198">2AD</figureCitation>
). Antennae slender, terminal part of antennomere 5 expanded, antennomeres 5 and 6 each with a distodorsal group of bacilliform sensilla (
<figureCitation id="135A2A32FE14FFF2FB06EC7FFA83F989" box="[1184,1282,1537,1562]" captionStart="FIGURE 1" captionStartId="2.[151,250,1953,1975]" captionTargetBox="[151,1436,483,1928]" captionTargetId="figure@2.[151,1436,483,1928]" captionTargetPageId="2" captionText="FIGURE 1 A F. Glyphiulus latus sp. nov., holotype. A. anterior part of body, lateral view; B. same, dorsal view; C. midbody segments, lateral view; D. same, dorsal view; E. posterior part of body, lateral view; F. same, ventral view." httpUri="https://zenodo.org/record/898902/files/figure.png" pageId="1" pageNumber="198">Figs 1A</figureCitation>
,
<figureCitation id="135A2A32FE14FFF2FAB4EC7CFADBF989" box="[1298,1370,1538,1562]" captionStart="FIGURE 2" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[163,1423,511,1927]" captionTargetId="figure@3.[163,1423,511,1927]" captionTargetPageId="3" captionText="FIGURE 2 A D. Glyphiulus latus sp. nov., anterior part of body, lateral view. A. female paratype; B. male holotype; C. another female paratype; D. male paratype. Green arrows show the variation of the lateralmost crest of collum. Blue arrows indicate the presence of the ectoparasitic fungi, order Entomophthorales." httpUri="https://zenodo.org/record/898904/files/figure.png" pageId="1" pageNumber="198">2AD</figureCitation>
,
<figureCitation id="135A2A32FE14FFF2FACCEC7FFA0EF989" box="[1386,1423,1537,1562]" captionStart="FIGURE 5" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[335,1273,830,1888]" captionTargetId="figure@5.[303,1291,810,1907]" captionTargetPageId="5" captionText="FIGURE 5 A E. Glyphiulus latus sp. nov., holotype. A. anterior gonopods, caudal view; B. posterior gonopods, caudal view; C. clypeus, anterior view; D. antenna, lateral view; E. legs 2, caudal view. Abbreviations: BS, bacilliform sensilla; c, coxosterna; cmp, coxosternal mesal processes; F, flagellum; L, lamelliform lobe; P, penis; T, telopodite. Scar bars: A, B, E = 0.2 mm; C = 0.1 mm; D = 0.5 mm." httpUri="https://zenodo.org/record/898910/files/figure.png" pageId="1" pageNumber="198">5D</figureCitation>
). Gnathochilarium with a separate promentum, polytrichous (
<figureCitation id="135A2A32FE14FFF2FC93EC5BFC0EF9AD" box="[821,911,1573,1598]" captionStart="FIGURE 3" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[151,1434,193,1944]" captionTargetId="figure@4.[151,1436,193,1944]" captionTargetPageId="4" captionText="FIGURE 3 A F. Glyphiulus latus sp. nov., holotype. A. gnathochilarium, ventral view; B. legs 1, anterior view; C. legs 2, caudal view; D. legs 3, caudal view; E. cross-section of a midbody segment, caudal view; F. midbody leg, lateral view." httpUri="https://zenodo.org/record/898906/files/figure.png" pageId="1" pageNumber="198">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="8BDE36B7FE14FFF2FF61EC37FD05F8A9" blockId="1.[151,1437,961,2030]" pageId="1" pageNumber="198">
Collum strongly crested, lateralmost crest continuous or split, carinotaxic formula (1a+2c+IIIIV+5c+6a+pc+ma+pc+6a+5c+IVIII+2c+1a) or (IIII+4c+5a+pc+ma+pc+5a+4c+IIII) (
<figureCitation id="135A2A32FE14FFF2FAA8EC13FA11F915" box="[1294,1424,1645,1670]" captionStart="FIGURE 2" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[163,1423,511,1927]" captionTargetId="figure@3.[163,1423,511,1927]" captionTargetPageId="3" captionText="FIGURE 2 A D. Glyphiulus latus sp. nov., anterior part of body, lateral view. A. female paratype; B. male holotype; C. another female paratype; D. male paratype. Green arrows show the variation of the lateralmost crest of collum. Blue arrows indicate the presence of the ectoparasitic fungi, order Entomophthorales." httpUri="https://zenodo.org/record/898904/files/figure.png" pageId="1" pageNumber="198">Figs 2AD</figureCitation>
). Subsequent metaterga noticeably crested as well. Postcollum constriction modest (
<figureCitation id="135A2A32FE14FFF2FB88ECEFFB05F939" box="[1070,1156,1681,1706]" captionStart="FIGURE 1" captionStartId="2.[151,250,1953,1975]" captionTargetBox="[151,1436,483,1928]" captionTargetId="figure@2.[151,1436,483,1928]" captionTargetPageId="2" captionText="FIGURE 1 A F. Glyphiulus latus sp. nov., holotype. A. anterior part of body, lateral view; B. same, dorsal view; C. midbody segments, lateral view; D. same, dorsal view; E. posterior part of body, lateral view; F. same, ventral view." httpUri="https://zenodo.org/record/898902/files/figure.png" pageId="1" pageNumber="198">Fig. 1B</figureCitation>
). Ozopores starting from body ring 5 (
<figureCitation id="135A2A32FE14FFF2FE88ECCBFE30F95D" box="[302,433,1717,1742]" captionStart="FIGURE 1" captionStartId="2.[151,250,1953,1975]" captionTargetBox="[151,1436,483,1928]" captionTargetId="figure@2.[151,1436,483,1928]" captionTargetPageId="2" captionText="FIGURE 1 A F. Glyphiulus latus sp. nov., holotype. A. anterior part of body, lateral view; B. same, dorsal view; C. midbody segments, lateral view; D. same, dorsal view; E. posterior part of body, lateral view; F. same, ventral view." httpUri="https://zenodo.org/record/898902/files/figure.png" pageId="1" pageNumber="198">Figs 1A, B</figureCitation>
). Ozoporiferous tubercles round, wider than high; midbody metatergal crests complete. Crests divided into two transverse rows of tubercles, the anterior tubercle round and upright, the posterior one triangular, directed caudally. Lateral crests rather small. Carinotaxic formula 2/2+I/i+3/3+I/i+2/2 (
<figureCitation id="135A2A32FE14FFF2FAABEC83FA11F885" box="[1293,1424,1789,1814]" captionStart="FIGURE 1" captionStartId="2.[151,250,1953,1975]" captionTargetBox="[151,1436,483,1928]" captionTargetId="figure@2.[151,1436,483,1928]" captionTargetPageId="2" captionText="FIGURE 1 A F. Glyphiulus latus sp. nov., holotype. A. anterior part of body, lateral view; B. same, dorsal view; C. midbody segments, lateral view; D. same, dorsal view; E. posterior part of body, lateral view; F. same, ventral view." httpUri="https://zenodo.org/record/898902/files/figure.png" pageId="1" pageNumber="198">Figs 1AE</figureCitation>
). Body rings round in cross-section (
<figureCitation id="135A2A32FE14FFF2FD86ED5FFDF6F8A9" box="[544,631,1825,1850]" captionStart="FIGURE 3" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[151,1434,193,1944]" captionTargetId="figure@4.[151,1436,193,1944]" captionTargetPageId="4" captionText="FIGURE 3 A F. Glyphiulus latus sp. nov., holotype. A. gnathochilarium, ventral view; B. legs 1, anterior view; C. legs 2, caudal view; D. legs 3, caudal view; E. cross-section of a midbody segment, caudal view; F. midbody leg, lateral view." httpUri="https://zenodo.org/record/898906/files/figure.png" pageId="1" pageNumber="198">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph id="8BDE36B7FE14FFF2FF61ED3BFD07F835" blockId="1.[151,1437,961,2030]" pageId="1" pageNumber="198">
Epiproct simple, devoid of tuberculation, with a rounded ridge in caudal part and an evident axial rib dorsally. Paraprocts convex, with only a shallow but evident depression beside caudal edge, polytrichous. Hypoproct crescent shaped, weakly setose (
<figureCitation id="135A2A32FE14FFF2FDA7EDF3FDF8F835" box="[513,633,1933,1958]" captionStart="FIGURE 1" captionStartId="2.[151,250,1953,1975]" captionTargetBox="[151,1436,483,1928]" captionTargetId="figure@2.[151,1436,483,1928]" captionTargetPageId="2" captionText="FIGURE 1 A F. Glyphiulus latus sp. nov., holotype. A. anterior part of body, lateral view; B. same, dorsal view; C. midbody segments, lateral view; D. same, dorsal view; E. posterior part of body, lateral view; F. same, ventral view." httpUri="https://zenodo.org/record/898902/files/figure.png" pageId="1" pageNumber="198">Figs 1E, F</figureCitation>
).
</paragraph>
<paragraph id="8BDE36B7FE14FFF1FF61EDCFFD89FF6B" blockId="1.[151,1437,961,2030]" lastBlockId="2.[151,1436,151,429]" lastPageId="2" lastPageNumber="199" pageId="1" pageNumber="198">
Male legs 1 5-segmented, with considerably shortened telopodites and a pair of large, subdigitiform, medially contiguous, forward directed coxal processes with groups of long, robust setae at base (
<figureCitation id="135A2A32FE14FFF2FBC4EDABFB3BF87D" box="[1122,1210,2005,2030]" captionStart="FIGURE 3" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[151,1434,193,1944]" captionTargetId="figure@4.[151,1436,193,1944]" captionTargetPageId="4" captionText="FIGURE 3 A F. Glyphiulus latus sp. nov., holotype. A. gnathochilarium, ventral view; B. legs 1, anterior view; C. legs 2, caudal view; D. legs 3, caudal view; E. cross-section of a midbody segment, caudal view; F. midbody leg, lateral view." httpUri="https://zenodo.org/record/898906/files/figure.png" pageId="1" pageNumber="198">Fig. 3B</figureCitation>
). Male legs 2 nearly normal. Penes broad, rounded, each with 4 robust setae distolaterally (
<figureCitation id="135A2A32FE17FFF1FC16EAE9FB8DFF23" box="[944,1036,151,176]" captionStart="FIGURE 3" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[151,1434,193,1944]" captionTargetId="figure@4.[151,1436,193,1944]" captionTargetPageId="4" captionText="FIGURE 3 A F. Glyphiulus latus sp. nov., holotype. A. gnathochilarium, ventral view; B. legs 1, anterior view; C. legs 2, caudal view; D. legs 3, caudal view; E. cross-section of a midbody segment, caudal view; F. midbody leg, lateral view." httpUri="https://zenodo.org/record/898906/files/figure.png" pageId="2" pageNumber="199">Figs 3C</figureCitation>
,
<figureCitation id="135A2A32FE17FFF1FBBFEAE9FBBAFF23" box="[1049,1083,151,176]" captionStart="FIGURE 5" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[335,1273,830,1888]" captionTargetId="figure@5.[303,1291,810,1907]" captionTargetPageId="5" captionText="FIGURE 5 A E. Glyphiulus latus sp. nov., holotype. A. anterior gonopods, caudal view; B. posterior gonopods, caudal view; C. clypeus, anterior view; D. antenna, lateral view; E. legs 2, caudal view. Abbreviations: BS, bacilliform sensilla; c, coxosterna; cmp, coxosternal mesal processes; F, flagellum; L, lamelliform lobe; P, penis; T, telopodite. Scar bars: A, B, E = 0.2 mm; C = 0.1 mm; D = 0.5 mm." httpUri="https://zenodo.org/record/898910/files/figure.png" pageId="2" pageNumber="199">5E</figureCitation>
). Male legs 3 modified, having coxa especially slender and elongate (
<figureCitation id="135A2A32FE17FFF1FDF2EAC2FD33FF47" box="[596,690,188,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[151,1434,193,1944]" captionTargetId="figure@4.[151,1436,193,1944]" captionTargetPageId="4" captionText="FIGURE 3 A F. Glyphiulus latus sp. nov., holotype. A. gnathochilarium, ventral view; B. legs 1, anterior view; C. legs 2, caudal view; D. legs 3, caudal view; E. cross-section of a midbody segment, caudal view; F. midbody leg, lateral view." httpUri="https://zenodo.org/record/898906/files/figure.png" pageId="2" pageNumber="199">Fig. 3D</figureCitation>
). Midbody legs obviously longer than body diameter (
<figureCitation id="135A2A32FE17FFF1FA92EAC2FA11FF47" box="[1332,1424,188,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[151,1434,193,1944]" captionTargetId="figure@4.[151,1436,193,1944]" captionTargetPageId="4" captionText="FIGURE 3 A F. Glyphiulus latus sp. nov., holotype. A. gnathochilarium, ventral view; B. legs 1, anterior view; C. legs 2, caudal view; D. legs 3, caudal view; E. cross-section of a midbody segment, caudal view; F. midbody leg, lateral view." httpUri="https://zenodo.org/record/898906/files/figure.png" pageId="2" pageNumber="199">Fig. 3E</figureCitation>
). Accessory claw absent (
<figureCitation id="135A2A32FE17FFF1FE03EAA1FE7BFF6B" box="[421,506,223,248]" captionStart="FIGURE 3" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[151,1434,193,1944]" captionTargetId="figure@4.[151,1436,193,1944]" captionTargetPageId="4" captionText="FIGURE 3 A F. Glyphiulus latus sp. nov., holotype. A. gnathochilarium, ventral view; B. legs 1, anterior view; C. legs 2, caudal view; D. legs 3, caudal view; E. cross-section of a midbody segment, caudal view; F. midbody leg, lateral view." httpUri="https://zenodo.org/record/898906/files/figure.png" pageId="2" pageNumber="199">Fig. 3F</figureCitation>
).
</paragraph>
<paragraph id="8BDE36B7FE17FFF1FF61EB7AFD20FE3E" blockId="2.[151,1436,151,429]" pageId="2" pageNumber="199">
Anterior gonopods with a pair of shield-like coxosterna, sunken medially from caudal view. Prolonged and broad coxosternal mesal processes of anterior gonopods present. Telopodites curved, moveable, lateral, 1- segmented and each with 5 strong setae distally (
<figureCitation id="135A2A32FE17FFF1FD6EEB32FCA6FEF7" box="[712,807,332,357]" captionStart="FIGURE 4" captionStartId="5.[151,250,743,765]" captionTargetBox="[313,1275,184,738]" captionTargetId="figure@5.[312,1276,183,739]" captionTargetPageId="5" captionText="FIGURE 4 A B. Glyphiulus latus sp. nov., holotype. A. anterior gonopods, caudal view; B. posterior gonopods, caudal view." httpUri="https://zenodo.org/record/898908/files/figure.png" pageId="2" pageNumber="199">Figs 4A</figureCitation>
,
<figureCitation id="135A2A32FE17FFF1FC92EB32FCDBFEF6" box="[820,858,332,357]" captionStart="FIGURE 5" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[335,1273,830,1888]" captionTargetId="figure@5.[303,1291,810,1907]" captionTargetPageId="5" captionText="FIGURE 5 A E. Glyphiulus latus sp. nov., holotype. A. anterior gonopods, caudal view; B. posterior gonopods, caudal view; C. clypeus, anterior view; D. antenna, lateral view; E. legs 2, caudal view. Abbreviations: BS, bacilliform sensilla; c, coxosterna; cmp, coxosternal mesal processes; F, flagellum; L, lamelliform lobe; P, penis; T, telopodite. Scar bars: A, B, E = 0.2 mm; C = 0.1 mm; D = 0.5 mm." httpUri="https://zenodo.org/record/898910/files/figure.png" pageId="2" pageNumber="199">5A</figureCitation>
). Posterior gonopods compact, coxite with medial, apically lamelliform lobe and slender, elongated, curved flagellum; the mediolateral margin of coxite with two rows of strong and curved setae (
<figureCitation id="135A2A32FE17FFF1FDAFEBEAFDE5FE3F" box="[521,612,404,429]" captionStart="FIGURE 4" captionStartId="5.[151,250,743,765]" captionTargetBox="[313,1275,184,738]" captionTargetId="figure@5.[312,1276,183,739]" captionTargetPageId="5" captionText="FIGURE 4 A B. Glyphiulus latus sp. nov., holotype. A. anterior gonopods, caudal view; B. posterior gonopods, caudal view." httpUri="https://zenodo.org/record/898908/files/figure.png" pageId="2" pageNumber="199">Figs 4B</figureCitation>
,
<figureCitation id="135A2A32FE17FFF1FDD6EBEAFD15FE3E" box="[624,660,404,429]" captionStart="FIGURE 5" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[335,1273,830,1888]" captionTargetId="figure@5.[303,1291,810,1907]" captionTargetPageId="5" captionText="FIGURE 5 A E. Glyphiulus latus sp. nov., holotype. A. anterior gonopods, caudal view; B. posterior gonopods, caudal view; C. clypeus, anterior view; D. antenna, lateral view; E. legs 2, caudal view. Abbreviations: BS, bacilliform sensilla; c, coxosterna; cmp, coxosternal mesal processes; F, flagellum; L, lamelliform lobe; P, penis; T, telopodite. Scar bars: A, B, E = 0.2 mm; C = 0.1 mm; D = 0.5 mm." httpUri="https://zenodo.org/record/898910/files/figure.png" pageId="2" pageNumber="199">5B</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF1E663FFE17FFF1FF31EDDFFB4DF846" httpUri="https://zenodo.org/record/898902/files/figure.png" pageId="2" pageNumber="199" targetBox="[151,1436,483,1928]" targetPageId="2">
<paragraph id="8BDE36B7FE17FFF1FF31EDDFFB4DF846" blockId="2.[151,1436,1953,2005]" pageId="2" pageNumber="199">
<emphasis id="B915EAA5FE17FFF1FF31EDDFFEC1F825" bold="true" box="[151,320,1953,1975]" pageId="2" pageNumber="199">FIGURE 1AF.</emphasis>
<taxonomicName id="4C614D34FE17FFF1FEE0EDDFFE6CF825" authority="Chen, 2017" authorityName="Chen" authorityYear="2017" box="[326,493,1953,1974]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
<emphasis id="B915EAA5FE17FFF1FEE0EDDFFE6CF825" box="[326,493,1953,1974]" italics="true" pageId="2" pageNumber="199">Glyphiulus latus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22657DEFE17FFF1FE53EDDFFDC6F825" box="[501,583,1953,1974]" pageId="2" pageNumber="199" rank="species">
<emphasis id="B915EAA5FE17FFF1FE53EDDFFDC6F825" bold="true" box="[501,583,1953,1974]" pageId="2" pageNumber="199">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype. A. anterior part of body, lateral view; B. same, dorsal view; C. midbody segments, lateral view; D. same, dorsal view; E. posterior part of body, lateral view; F. same, ventral view.
</paragraph>
</caption>
<subSubSection id="C37B653CFE16FFF0FF61EAE9FD3FFE43" pageId="3" pageNumber="200" type="distribution">
<paragraph id="8BDE36B7FE16FFF0FF61EAE9FC00FF23" blockId="3.[151,1436,151,464]" box="[199,897,151,176]" pageId="3" pageNumber="200">
<emphasis id="B915EAA5FE16FFF0FF61EAE9FEE1FF23" bold="true" box="[199,352,151,176]" pageId="3" pageNumber="200">Distribution.</emphasis>
Known only from
<typeStatus id="54DA8815FE16FFF0FD91EAE6FDE6FF23" box="[567,615,152,176]" pageId="3" pageNumber="200">type</typeStatus>
locality,
<collectingRegion id="49A5F855FE16FFF0FD68EAE9FCAAFF23" box="[718,811,151,176]" country="China" name="Sichuan" pageId="3" pageNumber="200">Sichuan</collectingRegion>
,
<collectingCountry id="F3767627FE16FFF0FC90EAE9FCFCFF23" box="[822,893,151,176]" name="China" pageId="3" pageNumber="200">China</collectingCountry>
.
</paragraph>
<paragraph id="8BDE36B7FE16FFF0FF61EAC2FCDEFF6B" blockId="3.[151,1436,151,464]" pageId="3" pageNumber="200">
<emphasis id="B915EAA5FE16FFF0FF61EAC2FE90FF46" bold="true" box="[199,273,188,213]" pageId="3" pageNumber="200">Notes.</emphasis>
Based on its evident troglomorphic traits, like eye reduction, elongation of legs and antennae and general pallid coloration, this species seems to be a troglobiont.
</paragraph>
<paragraph id="8BDE36B7FE16FFF0FF61EB7AFD3FFE43" blockId="3.[151,1436,151,464]" pageId="3" pageNumber="200">
The carinotaxic formula of the collum and metazona in
<taxonomicName id="4C614D34FE16FFF0FCE1EB7AFC43FE8F" authorityName="Chen" authorityYear="2017" box="[839,962,260,284]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="200" phylum="Arthropoda" rank="genus">
<emphasis id="B915EAA5FE16FFF0FCE1EB7AFC43FE8F" box="[839,962,260,284]" italics="true" pageId="3" pageNumber="200">Glyphiulus</emphasis>
</taxonomicName>
are considered stable within species, and highly useful in classification (
<bibRefCitation id="EFF04B46FE16FFF0FDA5EB59FCA1FED3" author="Golovatch" box="[515,800,295,320]" pageId="3" pageNumber="200" refString="Golovatch, S. I., Geoffroy, J. - J., Mauries, J. - P. &amp; VandenSpiege, D. (2007 b) Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2. The javanicus - group. Zoosystema, 29 (3), 417 - 456." type="journal article" year="2007" yearSuffix="b">
Golovatch
<emphasis id="B915EAA5FE16FFF0FD23EB57FD3AFED3" box="[645,699,296,320]" italics="true" pageId="3" pageNumber="200">et al</emphasis>
., 2007b
</bibRefCitation>
). However, this species shows noticeable intraspecific variation of the carinotaxic formula of the collum. This variation is reflected in the lateralmost crest which may twisted or divided from the middle (
<figureCitation id="135A2A32FE16FFF0FD94EB11FD35FE1B" box="[562,692,367,392]" captionStart="FIGURE 2" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[163,1423,511,1927]" captionTargetId="figure@3.[163,1423,511,1927]" captionTargetPageId="3" captionText="FIGURE 2 A D. Glyphiulus latus sp. nov., anterior part of body, lateral view. A. female paratype; B. male holotype; C. another female paratype; D. male paratype. Green arrows show the variation of the lateralmost crest of collum. Blue arrows indicate the presence of the ectoparasitic fungi, order Entomophthorales." httpUri="https://zenodo.org/record/898904/files/figure.png" pageId="3" pageNumber="200">Figs 2BD</figureCitation>
). When divided, the lateralmost crest gives rise to two differently shaped, shorter crests, one derived from the anterior part, one from the posterior part of the original crest. The number of crests thus increases by one (
<figureCitation id="135A2A32FE16FFF0FDF0EBC9FD30FE43" box="[598,689,439,464]" captionStart="FIGURE 2" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[163,1423,511,1927]" captionTargetId="figure@3.[163,1423,511,1927]" captionTargetPageId="3" captionText="FIGURE 2 A D. Glyphiulus latus sp. nov., anterior part of body, lateral view. A. female paratype; B. male holotype; C. another female paratype; D. male paratype. Green arrows show the variation of the lateralmost crest of collum. Blue arrows indicate the presence of the ectoparasitic fungi, order Entomophthorales." httpUri="https://zenodo.org/record/898904/files/figure.png" pageId="3" pageNumber="200">Fig. 2D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>