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<document id="42C901472B71850411C3AA2DB6E97249" ID-CLB-Dataset="299407" ID-DOI="10.5281/zenodo.11067242" ID-GBIF-Dataset="36a87cae-beb8-49c4-a561-a9e9a73afdf1" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067242" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052119798" checkinUser="felipe" docAuthor="Maddison, David R." docDate="2023" docId="03C825734419A56988CC1F7F0F94FD07" docLanguage="en" docName="ProcCalifAcadSci.67.19.433-450.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (19)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Bembidion cachagua Maddison 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="447" masterDocId="FFF15D0B4414A566892F1D770F4AFFB4" masterDocTitle="A New Subgenus of Bembidion Latreille from México and Guatemala, with Descriptions of Two New Species (Coleoptera: Carabidae)" masterLastPageNumber="450" masterPageNumber="433" pageNumber="446" updateTime="1719951180206" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="E15F670F8962F2E927BA0C6F0FBFDC1A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="45651052222278FA004FD2CC8438EE67">
<mods:title id="D4380F540DE6720B10CC8708E2BA4C1E">A New Subgenus of Bembidion Latreille from México and Guatemala, with Descriptions of Two New Species (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="58D2C7AAE65061C4864DDF12374A0E99" type="personal">
<mods:role id="EAF58040E21D92ACAE0123A26CAE7575">
<mods:roleTerm id="DD6F98881412522204523F651AD95CAC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="779E3CF9E05814DB7142BA7874E1845B">Maddison, David R.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C4BA20204B87B9DAE2C1E3219CE9149A">text</mods:typeOfResource>
<mods:relatedItem id="D1B64EF31EF7351227AECA0F40B1CF52" type="host">
<mods:titleInfo id="48A8B035FE352609A8D9858945C61104">
<mods:title id="77032007719042335F3C6456787518D5">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="722962029A3605F33129564BA5050F66">
<mods:date id="3C04556BBC55D261B240D5CB3EDAEB44">2023</mods:date>
<mods:detail id="7FCAF0700C1E79421C5D0A98BD81A0F7" type="pubDate">
<mods:number id="BD03886D32C13BE339CEB3C723CA0401">2023-02-15</mods:number>
</mods:detail>
<mods:detail id="7D7ED6A9D0A96CB8E382A3CDB96A5D46" type="volume">
<mods:number id="80D3FA16DA8C36EEA80A3ECCDBD802C4">67</mods:number>
</mods:detail>
<mods:detail id="9E8C6FF90F308A572A9978B8EF4D2B82" type="issue">
<mods:number id="7E2B9E90EBDBCD63CB5345F2E52AD220">19</mods:number>
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<mods:extent id="2A98160775DFC1FC34A04E688123B77C" unit="page">
<mods:start id="F4AB1D6AE7BBFD897B58F86482ED7E3F">433</mods:start>
<mods:end id="0EE17A5AD0B06F6D99F18D951E65675C">450</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="CA7EA3AB7DDCB4B6310D51F4B84B6ECC">journal article</mods:classification>
<mods:identifier id="6490B83E942762E1A20EE19DB2256709" type="CLB-Dataset">299407</mods:identifier>
<mods:identifier id="DFA9A4BC9A9406E8CA8B0FBB2BDF391C" type="DOI">10.5281/zenodo.11067242</mods:identifier>
<mods:identifier id="7D3CDBF6D4D650F1E1D0CE1406299E44" type="GBIF-Dataset">36a87cae-beb8-49c4-a561-a9e9a73afdf1</mods:identifier>
<mods:identifier id="07F5E7253622C85C3BC959DC1CBD7F7E" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="4682DC3F64A76F09FEF307E6A409A67B" type="Zenodo-Dep">11067242</mods:identifier>
</mods:mods>
<treatment id="03C825734419A56988CC1F7F0F94FD07" ID-DOI="http://doi.org/10.5281/zenodo.12627935" ID-GBIF-Taxon="234411869" ID-Zenodo-Dep="12627935" LSID="urn:lsid:plazi:treatment:03C825734419A56988CC1F7F0F94FD07" httpUri="http://treatment.plazi.org/id/03C825734419A56988CC1F7F0F94FD07" lastPageId="15" lastPageNumber="447" pageId="13" pageNumber="446">
<subSubSection id="C37BC7EE4419A56B88CC1F7F0C64FD94" box="[483,814,520,544]" pageId="13" pageNumber="446" type="nomenclature">
<paragraph id="8BDE94654419A56B88CC1F7F0C64FD94" blockId="13.[483,814,520,576]" box="[483,814,520,544]" pageId="13" pageNumber="446">
<heading id="D09623094419A56B88CC1F7F0C64FD94" box="[483,814,520,544]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="446" reason="4">
<taxonomicName id="4C61EFE64419A56B88CC1F7F0D87FD94" authority="Maddison, 2023" authorityName="Maddison" authorityYear="2023" box="[483,717,520,544]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="446" phylum="Arthropoda" rank="species" species="cachagua" status="sp. nov.">
<emphasis id="B91548774419A56B88CC1F7F0D87FD94" bold="true" box="[483,717,520,544]" italics="true" pageId="13" pageNumber="446">Bembidion cachagua</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A226F50C4419A56B8BF41F7E0C64FD94" box="[731,814,521,544]" pageId="13" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37BC7EE4419A56B8B101F5F0D9BFDF4" box="[575,721,552,576]" pageId="13" pageNumber="446" type="description">
<paragraph id="8BDE94654419A56B8B101F5F0D9BFDF4" blockId="13.[483,814,520,576]" box="[575,721,552,576]" pageId="13" pageNumber="446">
(
<figureCitation id="135A88E04419A56B8B691F5F0DE4FDF4" box="[582,686,552,576]" captionStart="FIGURE" captionStartId="14.[155,167,802,821]" captionTargetBox="[209,1077,234,782]" captionTargetId="figure-213@14.[205,1091,216,791]" captionTargetPageId="14" captionText="FIGURE 7. Holotype of Bembidion cachagua. (A) Habitus, scale bar 1 mm. (B) Forebody, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067254" httpUri="https://zenodo.org/record/11067254/files/figure.png" pageId="13" pageNumber="446">Figures 7</figureCitation>
,
<figureCitation id="135A88E04419A56B8B941F5F0D82FDF4" box="[699,712,552,576]" captionStart="FIGURE" captionStartId="14.[635,647,1439,1458]" captionTargetBox="[603,1164,846,1422]" captionTargetId="figure-234@14.[600,1176,856,1425]" captionTargetPageId="14" captionText="FIGURE 8. Aedeagus of holotype of Bembidion cachagua. (A) Left lateral view. (B) Right lateral view. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12627939" httpUri="https://zenodo.org/record/12627939/files/figure.png" pageId="13" pageNumber="446">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37BC7EE4419A56B89841F2B0C2EFB6D" pageId="13" pageNumber="446" type="materials_examined">
<paragraph id="8BDE94654419A56B89841F2B0C01FD41" blockId="13.[123,1174,604,1725]" pageId="13" pageNumber="446">
<materialsCitation id="3B099E384419A56B89841F2B0C01FD41" ID-GBIF-Occurrence="4902825309" collectingDate="1948-05-29" collectionCode="CAS" collectorName="J. M. Linsdale" country="United States of America" county="Monterey Co." location="Hastings Nat. History Reserve" municipality="Hastings Nat. History Reserve" pageId="13" pageNumber="446" specimenCount="1" specimenCount-male="1" stateProvince="CALIFORNIA" typeStatus="holotype">
<emphasis id="B91548774419A56B89841F2B0E59FDC7" bold="true" box="[171,275,604,627]" pageId="13" pageNumber="446">
<typeStatus id="54DA2AC74419A56B89841F2B0E59FDC7" box="[171,275,604,627]" pageId="13" pageNumber="446" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D675FEC4419A56B88351F2B0E1AFDC0" box="[282,336,604,628]" count="1" pageId="13" pageNumber="446" type="male">male</specimenCount>
, in
<collectionCode id="ED700CA04419A56B88571F2B0EE4FDC0" box="[376,430,604,628]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="13" pageNumber="446" type="Museum">CAS</collectionCode>
, labeled: “
<collectingCountry id="F376D4F54419A56B8B0D1F2B0D25FDC0" box="[546,623,604,628]" name="United States of America" pageId="13" pageNumber="446">U.S.A.</collectingCountry>
:
<collectingRegion id="49A55A874419A56B8B551F2B0C6AFDC0" box="[634,800,604,628]" country="United States of America" name="California" pageId="13" pageNumber="446">CALIFORNIA</collectingRegion>
:
<collectingCounty id="62BFECE94419A56B8A041F2B0C88FDC0" box="[811,962,604,628]" pageId="13" pageNumber="446">Monterey Co.</collectingCounty>
,
<location id="8EBEC2BE4419A56B8AE11F2B0E40FD20" LSID="urn:lsid:plazi:treatment:03C825734419A56988CC1F7F0F94FD07:8EBEC2BE4419A56B8AE11F2B0E40FD20" country="United States of America" county="Monterey Co." municipality="Hastings Nat. History Reserve" name="Hastings Nat. History Reserve" pageId="13" pageNumber="446" stateProvince="CALIFORNIA">Hastings Nat. History Reserve</location>
.
<collectorName id="2694F1B34419A56B88361F0A0EE5FD20" box="[281,431,636,660]" pageId="13" pageNumber="446">J.M. Linsdale</collectorName>
Colln.”, “C.A.S. Accession
<date id="FFDFB2A54419A56B8BDF1F0B0C02FD20" box="[752,840,636,660]" pageId="13" pageNumber="446" value="1948-05-29">
<collectingDate id="EF9B4B4D4419A56B8BDF1F0B0C02FD20" box="[752,840,636,660]" pageId="13" pageNumber="446" value="1948-05-29">29-V-48</collectingDate>
</date>
Cachagua Creek”, “David R.
<taxonomicName id="4C61EFE64419A56B89541FEB0FACFD00" authority="Maddison, 2023" authorityName="Maddison" authorityYear="2023" box="[123,230,668,692]" pageId="13" pageNumber="441" rank="subGenus" status="subgen. nov." subGenus="Maddison">Maddison</taxonomicName>
DNA3842 DNA Voucher” [pale green paper], “HOLOTYPE
<taxonomicName id="4C61EFE64419A56B8A811FEB0F96FD60" authority="David R." authorityName="David R." class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="446" phylum="Arthropoda" rank="species" species="cachagua">Bembidion cachagua David R.</taxonomicName>
Maddison” [partly handwritten, on red paper]. Genitalia mounted in Euparal in between coverslips pinned with specimen; extracted DNA stored separately.
</materialsCitation>
</paragraph>
<paragraph id="8BDE94654419A56B89841F890E13FCC2" blockId="13.[123,1174,604,1725]" pageId="13" pageNumber="446">
<materialsCitation id="3B099E384419A56B89841F890C9DFCA1" ID-GBIF-Occurrence="4902825308" box="[171,983,765,789]" collectingDate="1948-05-29" collectionCode="CAS, OSAC" collectorName="Same" country="United States of America" county="Monterey Co." location="Cachagua Creek" municipality="Hastings Nat. History Reserve" pageId="13" pageNumber="446" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="California" typeStatus="paratype">
<emphasis id="B91548774419A56B89841F890E57FCA1" bold="true" box="[171,285,766,789]" pageId="13" pageNumber="446">
<typeStatus id="54DA2AC74419A56B89841F890E57FCA1" box="[171,285,766,789]" pageId="13" pageNumber="446" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D675FEC4419A56B88011F8A0ECDFCA1" box="[302,391,765,789]" count="2" pageId="13" pageNumber="446" type="male">2 males</specimenCount>
,
<specimenCount id="9D675FEC4419A56B88BB1F8A0D49FCA1" box="[404,515,765,789]" count="4" pageId="13" pageNumber="446" type="female">4 females</specimenCount>
). Same data as holotype (3:
<collectionCode id="ED700CA04419A56B8A101F8A0C3FFCA1" box="[831,885,765,789]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="13" pageNumber="446" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED700CA04419A56B8AAC1F8A0C85FCA1" box="[899,975,765,789]" collectionName="USA, Oregon, Corvallis, Oregon State University" country="USA" name="Oregon State Arthropod Collection" pageId="13" pageNumber="446" type="University or college">OSAC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B099E384419A56B8ACC1F8A0E1EFCC2" ID-GBIF-Occurrence="4902825302" collectionCode="EMEC, DNA, MCZ, UIUC" collectorName="S. Cal." country="United States of America" county="Monterey Co." isTailEnumeration="true" location="Toschi" municipality="Pinnacles Nat'l Mon." pageId="13" pageNumber="446" specimenCode="348099" specimenCount="1" stateProvince="Isle of Anglesey" typeStatus="paratype">
“
<collectingMunicipality id="6BBA0E1F4419A56B8AC31F8A0FFEFC81" pageId="13" pageNumber="446">Pinnacles Nat’l Mon.</collectingMunicipality>
<date id="FFDFB2A54419A56B89931E6A0E46FC81" box="[188,268,797,821]" pageId="13" pageNumber="446" value="58-04-19">4.19.58</date>
<location id="8EBEC2BE4419A56B883C1E6A0E16FC81" LSID="urn:lsid:plazi:treatment:03C825734419A56988CC1F7F0F94FD07:8EBEC2BE4419A56B883C1E6A0E16FC81" box="[275,348,797,821]" country="United States of America" county="Monterey Co." municipality="Pinnacles Nat'l Mon." name="Toschi" pageId="13" pageNumber="446" stateProvince="Isle of Anglesey">Toschi</location>
” (1:
<collectionCode id="ED700CA04419A56B88BC1E6A0EABFC81" box="[403,481,797,821]" collectionName="USA, California, Berkeley, University of California, Essig Museum of Entomology" country="USA" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="13" pageNumber="446" type="University or college">EMEC</collectionCode>
; this is D.R.
<taxonomicName id="4C61EFE64419A56B8B401E6A0D90FC81" authority="Maddison, 2023" authorityName="Maddison" authorityYear="2023" box="[623,730,797,821]" pageId="13" pageNumber="441" rank="subGenus" status="subgen. nov." subGenus="Maddison">Maddison</taxonomicName>
<collectionCode id="ED700CA04419A56B8BCD1E6A0C50FC81" box="[738,794,797,821]" pageId="13" pageNumber="446">DNA</collectionCode>
voucher number 4758, and
<collectionCode id="ED700CA04419A56B8D641E6A0BDFFC81" box="[1099,1173,797,821]" collectionName="USA, California, Berkeley, University of California, Essig Museum of Entomology" pageId="13" pageNumber="446">EMEC</collectionCode>
specimen number
<specimenCode id="DBC73C1E4419A56B886F1E490EDFFCE2" box="[320,405,830,854]" pageId="13" pageNumber="446">348099</specimenCode>
). “Santa Barbara Cal. May ’91 L.E.R.” (1:
<collectionCode id="ED700CA04419A56B8A491E490CE9FCE2" box="[870,931,830,854]" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" country="USA" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="13" pageNumber="446" type="Museum">MCZ</collectionCode>
, Hayward Collection). “S. Cal.” (1:
<collectionCode id="ED700CA04419A56B88281E290E07FCC2" box="[263,333,862,886]" pageId="13" pageNumber="446">UIUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDE94654419A56B89841E080D39FB2C" blockId="13.[123,1174,604,1725]" pageId="13" pageNumber="446">
<emphasis id="B91548774419A56B89841E080E0BFC22" bold="true" box="[171,321,895,918]" pageId="13" pageNumber="446">Type locality</emphasis>
.
<collectingMunicipality id="6BBA0E1F4419A56B887C1E090D42FC22" box="[339,520,894,918]" pageId="13" pageNumber="446">Cachagua Creek</collectingMunicipality>
does not flow through
<location id="8EBEC2BE4419A56B8A361E090BD8FC22" LSID="urn:lsid:plazi:treatment:03C825734419A56988CC1F7F0F94FD07:8EBEC2BE4419A56B8A361E090BD8FC22" box="[793,1170,894,918]" country="United States of America" county="Monterey Co." municipality="Cachagua Creek" name="Hastings Natural History Reserve" pageId="13" pageNumber="446" stateProvince="CALIFORNIA">Hastings Natural History Reserve</location>
, although a tributary,
<location id="8EBEC2BE4419A56B884F1EE90EA2FC02" LSID="urn:lsid:plazi:treatment:03C825734419A56988CC1F7F0F94FD07:8EBEC2BE4419A56B884F1EE90EA2FC02" box="[352,488,926,950]" country="United States of America" county="Monterey Co." municipality="Cachagua Creek" name="Finch Creek" pageId="13" pageNumber="446" stateProvince="CALIFORNIA">Finch Creek</location>
, does. At its nearest
<location id="8EBEC2BE4419A56B8BE01EE90CC8FC02" LSID="urn:lsid:plazi:treatment:03C825734419A56988CC1F7F0F94FD07:8EBEC2BE4419A56B8BE01EE90CC8FC02" box="[719,898,926,950]" country="United States of America" county="Monterey Co." municipality="Cachagua Creek" name="Cachagua Creek" pageId="13" pageNumber="446" stateProvince="CALIFORNIA">Cachagua Creek</location>
is approximately
<quantity id="4C9939804419A56B8D661EE90BDFFC02" box="[1097,1173,926,950]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="446" unit="km" value="2.7">
<locationDeviation id="9A6C8D474419A56B8D661EE90BDFFC02" box="[1097,1173,926,950]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="446" unit="km" value="2.7">2.7 km</locationDeviation>
</quantity>
from the boundary of the Reserve. It is reasonable to presume that the upper label on the
<typeStatus id="54DA2AC74419A56B8D1E1EC80BDBFC63" box="[1073,1169,959,983]" pageId="13" pageNumber="446" type="holotype">holotype</typeStatus>
, “
<collectingCountry id="F376D4F54419A56B89A81EA80F99FC43" box="[135,211,991,1015]" name="United States of America" pageId="13" pageNumber="446">U.S.A.</collectingCountry>
:
<collectingRegion id="49A55A874419A56B89F21EA80EC9FC43" box="[221,387,991,1015]" country="United States of America" name="California" pageId="13" pageNumber="446">CALIFORNIA</collectingRegion>
:
<collectingCounty id="62BFECE94419A56B88A31E970D68FC43" box="[396,546,991,1016]" pageId="13" pageNumber="446">Monterey Co.</collectingCounty>
,
<collectorName id="2694F1B34419A56B8B031EA80B12FC43" box="[556,1112,991,1015]" pageId="13" pageNumber="446">Hastings Nat. History Reserve. J.M. Linsdale Colln.</collectorName>
”, was a generic label used for all specimens from the Linsdale collection, as
<collectorName id="2694F1B34419A56B8A4C19770CBFFBA3" box="[867,1013,1023,1047]" pageId="13" pageNumber="446">J.M. Linsdale</collectorName>
was the Director of the
<location id="8EBEC2BE4419A56B89CA19680D05FB83" LSID="urn:lsid:plazi:treatment:03C825734419A56988CC1F7F0F94FD07:8EBEC2BE4419A56B89CA19680D05FB83" box="[229,591,1055,1079]" country="United States of America" county="Monterey Co." municipality="Cachagua Creek" name="Hastings Natural History Reserve" pageId="13" pageNumber="446" stateProvince="CALIFORNIA">Hastings Natural History Reserve</location>
at the time (
<bibRefCitation id="EFF0E9944419A56B8BF719680C36FB83" author="ALAGONA, P. S." box="[728,892,1055,1079]" pageId="13" pageNumber="446" pagination="651 - 680" refId="ref7531" refString="ALAGONA, P. S. 2012. A Sanctuary for Science: The Hastings Natural History Reservation and the origins of the University of California's Natural Reserve System. Journal of the History of Biology 45: 651 - 680." type="journal article" year="2012">Alagona, 2012</bibRefCitation>
). The lower label presumably allowed customization for each specimen, and the locality information on the second label is thus likely more accurate. Thus, the
<typeStatus id="54DA2AC74419A56B88D219160D61FBCD" box="[509,555,1121,1145]" pageId="13" pageNumber="446">type</typeStatus>
locality would more accurately be stated to be “Cachagua Creek near Hastings Natural History Reserve”.
</paragraph>
<paragraph id="8BDE94654419A56B898419D60C2EFB6D" blockId="13.[123,1174,604,1725]" pageId="13" pageNumber="446">
<emphasis id="B91548774419A56B898419D60EBBFB0C" bold="true" box="[171,497,1185,1208]" pageId="13" pageNumber="446">Derivation of specific epithet</emphasis>
. Named after Cachagua Creek, the locality at which four of the seven known specimens were found. Treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C37BC7EE4419A569898419950F94FD07" lastPageId="15" lastPageNumber="448" pageId="13" pageNumber="446" type="description">
<paragraph id="8BDE94654419A568898419950E0CFC64" blockId="13.[123,1174,604,1725]" lastBlockId="14.[123,574,856,1424]" lastPageId="14" lastPageNumber="447" pageId="13" pageNumber="446">
<emphasis id="B91548774419A56B898419950E65FB4D" bold="true" box="[171,303,1250,1273]" pageId="13" pageNumber="446">Description</emphasis>
. Body length
<quantity id="4C9939804419A56B88FC19960D3CFB4D" box="[467,630,1249,1273]" metricMagnitude="-3" metricUnit="m" metricValue="3.675" metricValueMax="3.9" metricValueMin="3.45" pageId="13" pageNumber="446" unit="mm" value="3.675" valueMax="3.9" valueMin="3.45">3.45–3.90 mm</quantity>
. Forebody dark brown or piceous, with metallic greenish-yellow or brassy sheen; elytra pale chestnut brown, with the four testaceous elytral spots contrasting only slightly against the ground color (
<figureCitation id="135A88E04419A56B8B8118550DB9FA8E" box="[686,755,1314,1338]" captionStart="FIGURE" captionStartId="14.[155,167,802,821]" captionTargetBox="[209,1077,234,782]" captionTargetId="figure-213@14.[205,1091,216,791]" captionTargetPageId="14" captionText="FIGURE 7. Holotype of Bembidion cachagua. (A) Habitus, scale bar 1 mm. (B) Forebody, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067254" httpUri="https://zenodo.org/record/11067254/files/figure.png" pageId="13" pageNumber="446">Fig. 7</figureCitation>
). In
<specimenCount id="9D675FEC4419A56B8A0418550C99FA8E" box="[811,979,1314,1338]" count="4" pageId="13" pageNumber="446" type="generic">four specimens</specimenCount>
, the antennae are infuscated throughout; in the other
<specimenCount id="9D675FEC4419A56B88D418350DE1FAEE" box="[507,683,1346,1370]" count="3" pageId="13" pageNumber="446" type="generic">three specimens</specimenCount>
, two of which appear to be teneral, the basal 2.5 antennomeres are testaceous; legs pale reddish brown, slightly infuscated. Frontal furrows shallow, not prolonged onto clypeus. Mentum with anterior lateral regions triangular, large; central tooth of mentum triangular, slightly rounded. Prothorax narrow, only slightly wider than head; hind angles slightly obtuse; posterolateral carina of pronotum long and more or less straight; posterior region of pronotum rugose, wrinkled and with evident punctures (
<figureCitation id="135A88E04419A56B8A6D18940CCFFA4F" box="[834,901,1507,1531]" captionStart="FIGURE" captionStartId="14.[155,167,802,821]" captionTargetBox="[209,1077,234,782]" captionTargetId="figure-213@14.[205,1091,216,791]" captionTargetPageId="14" captionText="FIGURE 7. Holotype of Bembidion cachagua. (A) Habitus, scale bar 1 mm. (B) Forebody, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067254" httpUri="https://zenodo.org/record/11067254/files/figure.png" pageId="13" pageNumber="446">Fig. 7</figureCitation>
). Elytra with lateral bead not prolonged medially at shoulder; striae shallow, poorly defined, with extremely small punctures. Microsculpture evident throughout the dorsal surface of the forebody in most specimens, more or less isodiametric or slightly transversely stretched, in some specimens effaced in the center of the head between the eyes, and in the center of the pronotal disc; microsculpture effaced from most of elytra, but evident behind the second dorsal discal seta (ed5) as defined, slightly transverse sculpticells. Pronotum with two lateral setae on each side; elytron with two setae in third stria. Aedeagus (
<figureCitation id="135A88E0441AA568899B1E2F0FB3FCC4" box="[180,249,856,880]" captionStart="FIGURE" captionStartId="14.[635,647,1439,1458]" captionTargetBox="[603,1164,846,1422]" captionTargetId="figure-234@14.[600,1176,856,1425]" captionTargetPageId="14" captionText="FIGURE 8. Aedeagus of holotype of Bembidion cachagua. (A) Left lateral view. (B) Right lateral view. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12627939" httpUri="https://zenodo.org/record/12627939/files/figure.png" pageId="14" pageNumber="447">Fig. 8</figureCitation>
) with a small brush sclerite similar to that of
<emphasis id="B9154877441AA568881D1E0E0EC3FC24" box="[306,393,888,912]" italics="true" pageId="14" pageNumber="447">B. horni</emphasis>
, but with a long, slightly curved flagellum, most similar to that of
<emphasis id="B9154877441AA56889E71ECE0E75FC64" box="[200,319,952,976]" italics="true" pageId="14" pageNumber="447">B. festivum</emphasis>
.
</paragraph>
<caption id="DF1EC4ED441AA56889B41E550B73FC81" ID-DOI="http://doi.org/10.5281/zenodo.11067254" ID-Zenodo-Dep="11067254" box="[155,1081,801,821]" httpUri="https://zenodo.org/record/11067254/files/figure.png" pageId="14" pageNumber="447" startId="14.[155,167,802,821]" targetBox="[209,1077,234,782]" targetPageId="14" targetType="figure">
<paragraph id="8BDE9465441AA56889B41E550B73FC81" blockId="14.[155,1081,801,821]" box="[155,1081,801,821]" pageId="14" pageNumber="447">
<smallCapsWord id="8D3802B9441AA56889B41E550F92FC80" baselines="816,816" box="[155,216,802,821]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="14" pageNumber="447">FIGURE</smallCapsWord>
7. Holotype of
<taxonomicName id="4C61EFE6441AA568884E1E560D5CFC80" box="[353,534,801,820]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="447" phylum="Arthropoda" rank="species" species="cachagua">
<emphasis id="B9154877441AA568884E1E560D5CFC80" box="[353,534,801,820]" italics="true" pageId="14" pageNumber="447">Bembidion cachagua</emphasis>
</taxonomicName>
. (A) Habitus, scale bar 1 mm. (B) Forebody, scale bar 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BDE9465441AA56889841EAE0E69FB44" blockId="14.[123,574,856,1424]" pageId="14" pageNumber="447">
Most easily distinguished from other
<emphasis id="B9154877441AA56889541E8E0FBCFBA4" box="[123,246,1017,1040]" italics="true" pageId="14" pageNumber="447">Liocosmius</emphasis>
by the stronger and more widespread microsculpture, the greenishyellow metallic sheen on the dorsal surface, and the relatively low-contrast pattern on the elytra. In comparison to
<emphasis id="B9154877441AA5688B2A190E0FB5FB04" italics="true" pageId="14" pageNumber="447">Bembidion horni</emphasis>
in particular, the pronotum in
<emphasis id="B9154877441AA568895419CE0E44FB64" box="[123,270,1208,1232]" italics="true" pageId="14" pageNumber="447">B. cachagua</emphasis>
is much narrower, with straighter sides.
</paragraph>
<paragraph id="8BDE9465441AA5688984198E0FA0FAC4" blockId="14.[123,574,856,1424]" pageId="14" pageNumber="447">
<emphasis id="B9154877441AA5688984198E0E50FAA4" bold="true" box="[171,282,1273,1296]" pageId="14" pageNumber="447">Variation</emphasis>
. Some specimens have numerous punctures on the dorsal surface of the head and in anterior regions of the pronotum.
</paragraph>
<paragraph id="8BDE9465441AA5688984180E0E17FA44" blockId="14.[123,574,856,1424]" lastBlockId="14.[123,1173,1496,1712]" pageId="14" pageNumber="447">
<emphasis id="B9154877441AA5688984180E0E9FFA24" bold="true" box="[171,469,1401,1424]" pageId="14" pageNumber="447">Geographic Distribution</emphasis>
. Known only from California from Monterey and San Benito Counties south to Santa Barbara County (Fig. 9).
</paragraph>
<caption id="DF1EC4ED441AA5688B5418E80B1DFA78" ID-DOI="http://doi.org/10.5281/zenodo.12627939" ID-Zenodo-Dep="12627939" httpUri="https://zenodo.org/record/12627939/files/figure.png" pageId="14" pageNumber="447" startId="14.[635,647,1439,1458]" targetBox="[603,1164,846,1422]" targetPageId="14">
<paragraph id="8BDE9465441AA5688B5418E80B1DFA78" blockId="14.[603,1173,1438,1484]" pageId="14" pageNumber="447">
FIGURE 8. Aedeagus of holotype of
<emphasis id="B9154877441AA5688A9A18E90B21FA05" box="[949,1131,1438,1457]" italics="true" pageId="14" pageNumber="447">Bembidion cachagua</emphasis>
. (A) Left lateral view. (B) Right lateral view. Scale bar 100 µm.
</paragraph>
</caption>
<paragraph id="8BDE9465441AA5688984188E0B76F984" blockId="14.[123,1173,1496,1712]" pageId="14" pageNumber="447">
<emphasis id="B9154877441AA5688984188E0DD5F9A4" bold="true" box="[171,671,1529,1552]" pageId="14" pageNumber="447">Geographic relationships with other species.</emphasis>
Found sympatrically with four other species of subgenus
<emphasis id="B9154877441AA56889CA1B6E0E14F984" box="[229,350,1561,1584]" italics="true" pageId="14" pageNumber="447">Liocosmius</emphasis>
:
<emphasis id="B9154877441AA56888421B6F0D68F984" box="[365,546,1560,1584]" italics="true" pageId="14" pageNumber="447">Bembidion horni</emphasis>
,
<emphasis id="B9154877441AA5688B1F1B6E0DE0F984" box="[560,682,1560,1584]" italics="true" pageId="14" pageNumber="447">B. mundum</emphasis>
,
<emphasis id="B9154877441AA5688B971B6E0CC9F984" box="[696,899,1560,1584]" italics="true" pageId="14" pageNumber="447">B. darlingtonielum</emphasis>
, and
<emphasis id="B9154877441AA5688A911B6E0B7CF984" box="[958,1078,1560,1584]" italics="true" pageId="14" pageNumber="447">B. festivum</emphasis>
.
</paragraph>
<paragraph id="8BDE9465441AA56889841B4E0E07F924" blockId="14.[123,1173,1496,1712]" pageId="14" pageNumber="447">
<emphasis id="B9154877441AA56889841B4E0E48F9E4" bold="true" box="[171,258,1593,1616]" pageId="14" pageNumber="447">Habitat</emphasis>
. Unknown. It may occur in similar habitats as other
<emphasis id="B9154877441AA5688A7B1B4E0C87F9E4" box="[852,973,1593,1616]" italics="true" pageId="14" pageNumber="447">Liocosmius</emphasis>
, that is, on sandy shores of creeks, most often in shaded or partly shaded areas, and in general where the sand banks are relatively steep.
</paragraph>
<paragraph id="8BDE9465441AA56989841BEE0F94FD07" blockId="14.[123,1173,1496,1712]" lastBlockId="15.[123,1174,219,1030]" lastPageId="15" lastPageNumber="448" pageId="14" pageNumber="447">
<emphasis id="B9154877441AA56889841BEE0FA1F904" bold="true" box="[171,235,1689,1712]" pageId="14" pageNumber="447">Notes</emphasis>
. I have seen no specimens of this species collected since 1958. In late April and early May 2014, John Sproul and I searched at multiple known localities for this species, including at Hastings Natural History Reserve, Pinnacles National Park, around Santa Barbara, and at many other localities in between. We searched in typical
<taxonomicName id="4C61EFE6441BA56988C11C0B0D2DFE27" authorityName="Casey" authorityYear="1918" box="[494,615,380,403]" class="Insecta" family="Carabidae" genus="Liocosmius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="B9154877441BA56988C11C0B0D2DFE27" box="[494,615,380,403]" italics="true" pageId="15" pageNumber="448">Liocosmius</emphasis>
</taxonomicName>
microhabitats (including steep sand banks in partial shade), with no success. Along Finch Creek in Hastings Natural History Reserve, the only
<taxonomicName id="4C61EFE6441BA56989C61C8C0E17FDA7" authorityName="Latreille" authorityYear="1802" box="[233,349,507,531]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="B9154877441BA56989C61C8C0E17FDA7" box="[233,349,507,531]" italics="true" pageId="15" pageNumber="448">Bembidion</emphasis>
</taxonomicName>
in those microhabitats was
<taxonomicName id="4C61EFE6441BA56989811F6B0EE8FD87" authority="LeConte" authorityName="LeConte" box="[174,418,539,563]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="448" phylum="Arthropoda" rank="species" species="iridescens">
<emphasis id="B9154877441BA56989811F6B0E70FD87" box="[174,314,539,563]" italics="true" pageId="15" pageNumber="448">B. iridescens</emphasis>
LeConte
</taxonomicName>
; in nearby microhabitats along the creek shore were three species of
<taxonomicName id="4C61EFE6441BA56989DA1F2C0E23FDC7" authorityName="Latreille" authorityYear="1802" box="[245,361,603,627]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="B9154877441BA56989DA1F2C0E23FDC7" box="[245,361,603,627]" italics="true" pageId="15" pageNumber="448">Bembidion</emphasis>
</taxonomicName>
:
<taxonomicName id="4C61EFE6441BA56988531F2B0D6CFDC7" authorityName="Hayward" authorityYear="1897" box="[380,550,603,627]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="448" phylum="Arthropoda" rank="species" species="californicum">
<emphasis id="B9154877441BA56988531F2B0D6CFDC7" box="[380,550,603,627]" italics="true" pageId="15" pageNumber="448">B. californicum</emphasis>
</taxonomicName>
Hayward,
<taxonomicName id="4C61EFE6441BA56989911F0B0EF5FD27" authority="LeConte" authorityName="LeConte" box="[190,447,635,659]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="448" phylum="Arthropoda" rank="species" species="perspicuum">
<emphasis id="B9154877441BA56989911F0B0E12FD27" box="[190,344,636,659]" italics="true" pageId="15" pageNumber="448">B. perspicuum</emphasis>
LeConte
</taxonomicName>
, and
<taxonomicName id="4C61EFE6441BA56988D81F0B0D2DFD27" authorityName="LeConte" authorityYear="1857" box="[503,615,635,659]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="448" phylum="Arthropoda" rank="species" species="lugubre">
<emphasis id="B9154877441BA56988D81F0B0D2DFD27" box="[503,615,635,659]" italics="true" pageId="15" pageNumber="448">B. lugubre</emphasis>
</taxonomicName>
LeConte.
</paragraph>
</subSubSection>
</treatment>
</document>
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