treatments-xml/data/03/C5/18/03C5181CFFA22976FEE1F9DAFCE86290.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.4765867" ID-GBIF-Dataset="85bac109-2041-4a74-8b6b-088e82b0b899" ID-GBIF-Taxon="182165162" ID-ISSN="1854-0392" ID-Zenodo-Dep="4765867" approvalRequired="3" approvalRequired_for_textStreams="3" checkinTime="1620957841889" checkinUser="felipe" docAuthor="Stark, Bill P. &amp; Sivec, Ignac" docDate="2009" docId="03C5181CFFA22976FEE1F9DAFCE86290" docLanguage="en" docName="Illiesia.5.14.156-163.pdf" docOrigin="Illiesia 5 (14)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Sweltsa wui Stark &amp; Sivec 2009, sp. n." docType="treatment" docVersion="5" lastPageId="6" lastPageNumber="162" masterDocId="FFFC6064FFA32970FF9CFFF4FFF66133" masterDocTitle="Sweltsa Wui And Haploperla Valentinae (Plecoptera: Chloroperlidae), Two New Stoneflies From Sichuan Province, China" masterLastPageNumber="163" masterPageNumber="156" pageId="1" pageNumber="157" updateTime="1643565906184" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Sweltsa Wui And Haploperla Valentinae (Plecoptera: Chloroperlidae), Two New Stoneflies From Sichuan Province, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
<mods:affiliation>Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sivec, Ignac</mods:namePart>
<mods:affiliation>Slovenian Museum of Natural History, Prešernova 20, P. O. Box 290, SLO- 1001 Ljubljana, Slovenia E-mail: isivec @ pms-lj. si</mods:affiliation>
<mods:nameIdentifier type="email">isivec@pms-lj.si</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Illiesia</mods:title>
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<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="volume">
<mods:number>5</mods:number>
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<mods:detail type="issue">
<mods:number>14</mods:number>
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<mods:start>156</mods:start>
<mods:end>163</mods:end>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4759387</mods:identifier>
<mods:identifier type="GBIF-Dataset">85bac109-2041-4a74-8b6b-088e82b0b899</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4759387</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4765867" ID-GBIF-Taxon="182165162" ID-Zenodo-Dep="4765867" LSID="urn:lsid:plazi:treatment:03C5181CFFA22976FEE1F9DAFCE86290" httpUri="http://treatment.plazi.org/id/03C5181CFFA22976FEE1F9DAFCE86290" lastPageId="6" lastPageNumber="162" pageId="1" pageNumber="157">
<subSubSection box="[381,606,1582,1606]" pageId="1" pageNumber="157" type="nomenclature">
<paragraph blockId="1.[381,606,1582,1606]" box="[381,606,1582,1606]" pageId="1" pageNumber="157">
<heading bold="true" box="[381,606,1582,1606]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="157" reason="2">
<emphasis bold="true" box="[381,606,1582,1606]" pageId="1" pageNumber="157">
<taxonomicName authority="Stark &amp; Sivec, 2009" authorityName="Stark &amp; Sivec" authorityYear="2009" box="[381,528,1582,1606]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="157" phylum="Arthropoda" rank="species" species="wui" status="sp. n.">
<emphasis bold="true" box="[381,528,1582,1606]" italics="true" pageId="1" pageNumber="157">Sweltsa wui</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[541,606,1582,1606]" pageId="1" pageNumber="157" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[435,551,1618,1642]" pageId="1" pageNumber="157" type="description">
<paragraph blockId="1.[435,551,1618,1642]" box="[435,551,1618,1642]" pageId="1" pageNumber="157">
(
<figureCitation box="[444,542,1618,1642]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="1.[189,241,1475,1499]" captionStartId-1="2.[189,241,1618,1642]" captionTargetBox-0="[400,1209,400,1432]" captionTargetBox-1="[297,1337,231,1538]" captionTargetId-0="figure-57@1.[400,1231,400,1456]" captionTargetId-1="figure-57@2.[287,1345,223,1546]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figs. 1-4. Sweltsa wui. 1. Head and pronotum. 2. Male terminalia, dorsal. 3. Epiproct, dorsal. 4. Male terminalia, lateral." captionText-1="Figs. 5-10. Scanning electron micrographs of male genitalia. Sweltsa wui (5-8). 5. Terminalia, dorsal. 6. Epiproct, dorsal. 7. Epiproct, lateral. 8. Epiproct surface detail, frontal aspect. Haploperla valentinae (9-10). 9. Terminalia, dorsal. 10. Epiproct, dorsoapical margin." figureDoi-0="http://doi.org/10.5281/zenodo.4759391" figureDoi-1="http://doi.org/10.5281/zenodo.4759393" httpUri-0="https://zenodo.org/record/4759391/files/figure.png" httpUri-1="https://zenodo.org/record/4759393/files/figure.png" pageId="1" pageNumber="157">Figs. 1-8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="157" type="materials_examined">
<paragraph blockId="1.[189,798,1688,1927]" pageId="1" pageNumber="157">
<emphasis bold="true" box="[189,426,1689,1713]" pageId="1" pageNumber="157">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111997302" collectingDate="1991-06-01" collectingDateMax="1991-06-04" collectingDateMin="1991-06-01" collectionCode="PMSL" collectorName="B. Horvat &amp; I. Sivec" country="China" county="Pitiao River" elevation="3350" location="Balangshan Pass" municipality="Welong" pageId="1" pageNumber="157" specimenCount="4" specimenCount-male="4" stateProvince="Sichuan Province" typeStatus="holotype">
<typeStatus box="[434,546,1689,1713]" pageId="1" pageNumber="157">Holotype</typeStatus>
<specimenCount box="[554,574,1688,1713]" pageId="1" pageNumber="157" type="male">♂</specimenCount>
and 
<specimenCount box="[633,674,1688,1713]" pageId="1" pageNumber="157" type="male">3 ♂</specimenCount>
<typeStatus box="[681,797,1690,1714]" pageId="1" pageNumber="157">paratypes</typeStatus>
from 
<collectingCountry box="[252,325,1725,1749]" name="China" pageId="1" pageNumber="157">China</collectingCountry>
, 
<collectingRegion box="[334,540,1725,1749]" country="China" name="Sichuan" pageId="1" pageNumber="157">Sichuan Province</collectingRegion>
, 
<collectingCounty box="[549,689,1725,1749]" pageId="1" pageNumber="157">Pitiao River</collectingCounty>
, 
<collectingMunicipality box="[698,793,1725,1749]" pageId="1" pageNumber="157">Welong</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03C5181CFFA22976FEE1F9DAFCE86290:8EB3FFD1FFA22971FF21F915FE7F67CB" box="[189,393,1761,1785]" country="China" county="Pitiao River" municipality="Welong" name="Balangshan Pass" pageId="1" pageNumber="157" stateProvince="Sichuan Province">Balangshan Pass</location>
, 
<quantity box="[410,566,1761,1785]" metricMagnitude="3" metricUnit="m" metricValue="3.35" metricValueMax="4.0" metricValueMin="2.7" pageId="1" pageNumber="157" unit="m" value="3350.0" valueMax="4000.0" valueMin="2700.0">
<elevation box="[410,566,1761,1785]" metricMagnitude="3" metricUnit="m" metricValue="3.35" metricValueMax="4.0" metricValueMin="2.7" pageId="1" pageNumber="157" unit="m" value="3350.0" valueMax="4000.0" valueMin="2700.0">2700-4000 m</elevation>
</quantity>
, 
<date box="[584,756,1761,1785]" pageId="1" pageNumber="157" value="1991-06-01" valueMax="1991-06-04" valueMin="1991-06-01">
<collectingDate box="[584,756,1761,1785]" pageId="1" pageNumber="157" value="1991-06-01" valueMax="1991-06-04" valueMin="1991-06-01">1-4 June 1991</collectingDate>
</date>
, 
<collectorName pageId="1" pageNumber="157">B. Horvat</collectorName>
&amp; 
<collectorName box="[307,391,1796,1820]" pageId="1" pageNumber="157">I. Sivec</collectorName>
(
<collectionCode box="[405,481,1796,1820]" country="Slovenia" httpUri="http://grbio.org/cool/4mab-159w" name="Slovenian Museum of Natural History (Prirodosloveni Muzej Slovenije)" pageId="1" pageNumber="157">PMSL</collectionCode>
, 1 paratype-BPS).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="159" pageId="1" pageNumber="157" type="description">
<paragraph blockId="1.[189,798,1688,1927]" lastBlockId="1.[835,1443,1582,1926]" pageId="1" pageNumber="157">
<emphasis bold="true" box="[189,366,1831,1855]" pageId="1" pageNumber="157">Adult habitus.</emphasis>
General color pale yellow in alcohol but with a distinctive, narrowly divided, median, pronotal band (
<figureCitation box="[368,428,1903,1927]" captionStart="Figs" captionStartId="1.[189,241,1475,1499]" captionTargetBox="[400,1209,400,1432]" captionTargetId="figure-57@1.[400,1231,400,1456]" captionTargetPageId="1" captionText="Figs. 1-4. Sweltsa wui. 1. Head and pronotum. 2. Male terminalia, dorsal. 3. Epiproct, dorsal. 4. Male terminalia, lateral." figureDoi="http://doi.org/10.5281/zenodo.4759391" httpUri="https://zenodo.org/record/4759391/files/figure.png" pageId="1" pageNumber="157">Fig. 1</figureCitation>
) and obscure interocellar brown spot; meso and metathoracic notae with dark brown U-patterns, and abdomen bearing a continuous brown longitudinal band on segments 1-8; tergum 9 with a small, basal dumbbell shaped brown spot. Wings transparent, legs pale.
</paragraph>
<paragraph blockId="1.[835,1443,1582,1926]" lastBlockId="2.[835,1443,1795,1926]" lastPageId="2" lastPageNumber="158" pageId="1" pageNumber="157">
<emphasis bold="true" box="[835,904,1760,1784]" pageId="1" pageNumber="157">Male.</emphasis>
Forewing length 9.5 mm. Tergum 9 without transverse process and without median excavation on posterior margin (
<figureCitation box="[1085,1185,1831,1855]" captionStart="Figs" captionStartId="2.[189,241,1618,1642]" captionTargetBox="[297,1337,231,1538]" captionTargetId="figure-57@2.[287,1345,223,1546]" captionTargetPageId="2" captionText="Figs. 5-10. Scanning electron micrographs of male genitalia. Sweltsa wui (5-8). 5. Terminalia, dorsal. 6. Epiproct, dorsal. 7. Epiproct, lateral. 8. Epiproct surface detail, frontal aspect. Haploperla valentinae (9-10). 9. Terminalia, dorsal. 10. Epiproct, dorsoapical margin." figureDoi="http://doi.org/10.5281/zenodo.4759393" httpUri="https://zenodo.org/record/4759393/files/figure.png" pageId="1" pageNumber="157">Figs. 5-6</figureCitation>
). Epiproct about 2.5 X longer than basal width, rod-like but with lateral margins slightly constricted from near base to apical third. Epiproct length ca. 
<quantity box="[484,570,1795,1820]" metricMagnitude="-1" metricUnit="m" metricValue="3.4299999999999997" pageId="2" pageNumber="158" unit="mm" value="343.0">343 µm</quantity>
, subapical width ca. 
<quantity box="[189,279,1831,1855]" metricMagnitude="-1" metricUnit="m" metricValue="1.29" pageId="2" pageNumber="158" unit="mm" value="129.0">129 µm</quantity>
, basal width ca. 
<quantity box="[472,562,1831,1855]" metricMagnitude="-1" metricUnit="m" metricValue="1.38" pageId="2" pageNumber="158" unit="mm" value="138.0">138 µm</quantity>
, width at narrowest point ca. 
<quantity box="[295,371,1866,1891]" metricMagnitude="-2" metricUnit="m" metricValue="7.1" pageId="2" pageNumber="158" unit="mm" value="71.0">71 µm</quantity>
. Epiproct apex a smooth, sclerotized cap but most of dorsal and ventral surface covered with golden brown, closely appressed setae (
<figureCitation captionStart="Figs" captionStartId="1.[189,241,1475,1499]" captionTargetBox="[400,1209,400,1432]" captionTargetId="figure-57@1.[400,1231,400,1456]" captionTargetPageId="1" captionText="Figs. 1-4. Sweltsa wui. 1. Head and pronotum. 2. Male terminalia, dorsal. 3. Epiproct, dorsal. 4. Male terminalia, lateral." figureDoi="http://doi.org/10.5281/zenodo.4759391" httpUri="https://zenodo.org/record/4759391/files/figure.png" pageId="2" pageNumber="158">Figs. 2- 4</figureCitation>
, 
<figureCitation box="[862,899,1831,1855]" captionStart="Figs" captionStartId="2.[189,241,1618,1642]" captionTargetBox="[297,1337,231,1538]" captionTargetId="figure-57@2.[287,1345,223,1546]" captionTargetPageId="2" captionText="Figs. 5-10. Scanning electron micrographs of male genitalia. Sweltsa wui (5-8). 5. Terminalia, dorsal. 6. Epiproct, dorsal. 7. Epiproct, lateral. 8. Epiproct surface detail, frontal aspect. Haploperla valentinae (9-10). 9. Terminalia, dorsal. 10. Epiproct, dorsoapical margin." figureDoi="http://doi.org/10.5281/zenodo.4759393" httpUri="https://zenodo.org/record/4759393/files/figure.png" pageId="2" pageNumber="158">5-8</figureCitation>
); base with dorsal and lateral bare areas (
<figureCitation captionStart="Figs" captionStartId="2.[189,241,1618,1642]" captionTargetBox="[297,1337,231,1538]" captionTargetId="figure-57@2.[287,1345,223,1546]" captionTargetPageId="2" captionText="Figs. 5-10. Scanning electron micrographs of male genitalia. Sweltsa wui (5-8). 5. Terminalia, dorsal. 6. Epiproct, dorsal. 7. Epiproct, lateral. 8. Epiproct surface detail, frontal aspect. Haploperla valentinae (9-10). 9. Terminalia, dorsal. 10. Epiproct, dorsoapical margin." figureDoi="http://doi.org/10.5281/zenodo.4759393" httpUri="https://zenodo.org/record/4759393/files/figure.png" pageId="2" pageNumber="158">Figs. 6-7</figureCitation>
). Epiproct apex slightly inflated in lateral aspect (
<figureCitation box="[843,921,1902,1926]" captionStart="Figs" captionStartId="1.[189,241,1475,1499]" captionTargetBox="[400,1209,400,1432]" captionTargetId="figure-57@1.[400,1231,400,1456]" captionTargetPageId="1" captionText="Figs. 1-4. Sweltsa wui. 1. Head and pronotum. 2. Male terminalia, dorsal. 3. Epiproct, dorsal. 4. Male terminalia, lateral." figureDoi="http://doi.org/10.5281/zenodo.4759391" httpUri="https://zenodo.org/record/4759391/files/figure.png" pageId="2" pageNumber="158">Figs. 4</figureCitation>
, 
<figureCitation box="[933,949,1902,1926]" captionStart="Figs" captionStartId="2.[189,241,1618,1642]" captionTargetBox="[297,1337,231,1538]" captionTargetId="figure-57@2.[287,1345,223,1546]" captionTargetPageId="2" captionText="Figs. 5-10. Scanning electron micrographs of male genitalia. Sweltsa wui (5-8). 5. Terminalia, dorsal. 6. Epiproct, dorsal. 7. Epiproct, lateral. 8. Epiproct surface detail, frontal aspect. Haploperla valentinae (9-10). 9. Terminalia, dorsal. 10. Epiproct, dorsoapical margin." figureDoi="http://doi.org/10.5281/zenodo.4759393" httpUri="https://zenodo.org/record/4759393/files/figure.png" pageId="2" pageNumber="158">7</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4759393" ID-Zenodo-Dep="4759393" httpUri="https://zenodo.org/record/4759393/files/figure.png" pageId="2" pageNumber="158" startId="2.[189,241,1618,1642]" targetBox="[297,1337,231,1538]" targetPageId="2">
<paragraph blockId="2.[189,1443,1617,1713]" pageId="2" pageNumber="158">
Figs. 5-10. Scanning electron micrographs of male genitalia. 
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2009" box="[934,1069,1618,1642]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="158" phylum="Arthropoda" rank="species" species="wui">
<emphasis box="[934,1069,1618,1642]" italics="true" pageId="2" pageNumber="158">Sweltsa wui</emphasis>
</taxonomicName>
(5-8). 5. Terminalia, dorsal. 6. Epiproct, dorsal. 7. Epiproct, lateral. 8. Epiproct surface detail, frontal aspect. 
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2009" box="[1101,1336,1653,1677]" class="Insecta" family="Chloroperlidae" genus="Haploperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="158" phylum="Arthropoda" rank="species" species="valentinae">
<emphasis box="[1101,1336,1653,1677]" italics="true" pageId="2" pageNumber="158">Haploperla valentinae</emphasis>
</taxonomicName>
(9-10). 9. Terminalia, dorsal. 10. Epiproct, dorsoapical margin.
</paragraph>
</caption>
<paragraph blockId="3.[189,798,197,471]" box="[189,416,197,221]" pageId="3" pageNumber="159">
<emphasis bold="true" box="[189,283,197,221]" pageId="3" pageNumber="159">Female.</emphasis>
Unknown.
</paragraph>
<paragraph blockId="3.[189,798,197,471]" box="[189,396,232,257]" pageId="3" pageNumber="159">
<emphasis bold="true" box="[189,264,233,257]" pageId="3" pageNumber="159">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="159" type="etymology">
<paragraph blockId="3.[189,798,197,471]" pageId="3" pageNumber="159">
<emphasis bold="true" box="[189,327,268,292]" pageId="3" pageNumber="159">Etymology.</emphasis>
We dedicate this species to the late Professor Dr. C.F. Wu in recognition of his enormous contribution to the knowledge of Chinese 
<taxonomicName box="[669,794,340,364]" class="Insecta" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="159" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="162" pageId="3" pageNumber="159" type="diagnosis">
<paragraph blockId="3.[189,798,197,471]" lastBlockId="3.[835,1443,197,469]" pageId="3" pageNumber="159">
<emphasis bold="true" box="[189,319,375,399]" pageId="3" pageNumber="159">Diagnosis.</emphasis>
This species is similar in pronotal pigment pattern to 
<taxonomicName authority="(Zhiltzova &amp; Levanidova 1978)" baseAuthorityName="Zhiltzova &amp; Levanidova" baseAuthorityYear="1978" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="insularis">
<emphasis box="[437,627,411,435]" italics="true" pageId="3" pageNumber="159">Sweltsa insularis</emphasis>
(
<bibRefCitation author="Zhiltzova, L. A. &amp; I. M. Levanidova" pageId="3" pageNumber="159" pagination="3 - 29" refId="ref2485" refString="Zhiltzova, L. A. &amp; I. M. Levanidova. 1978. New species of stoneflies (Plecoptera) from the Far East. Trudy Zoologischeskogo Instituta Akademia Nauk, 61: 3 - 29. [In Russian]." type="journal article" year="1978">Zhiltzova &amp; Levanidova 1978</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName authority="(Okamoto) (Ham 2008" authorityName="Ham" authorityYear="2008" baseAuthorityName="Okamoto" baseAuthorityYear="1912" box="[427,797,446,471]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="nikkoensis">
<emphasis box="[427,573,446,470]" italics="true" pageId="3" pageNumber="159">S. nikkoensis</emphasis>
(Okamoto) (
<bibRefCitation author="Ham, S. A." box="[738,797,447,471]" pageId="3" pageNumber="159" pagination="185 - 189" refId="ref2216" refString="Ham, S. A. 2008. Two species of Chloroperlidae (Insecta: Plecoptera) new to Korea, with adult keys to species of the family in Korea. Korean Journal of Systematic Zoology, 24: 185 - 189." type="journal article" year="2008">Ham</bibRefCitation>
</taxonomicName>
2008), and also to the Japanese species 
<taxonomicName authority="(Okamoto)" baseAuthorityName="Okamoto" baseAuthorityYear="1912" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[1287,1443,197,221]" italics="true" pageId="3" pageNumber="159">S. abdominalis</emphasis>
(Okamoto)
</taxonomicName>
and 
<taxonomicName authority="(Okamoto)" baseAuthorityName="Okamoto" box="[1024,1328,232,256]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="sapporoensis">
<emphasis box="[1024,1190,233,256]" italics="true" pageId="3" pageNumber="159">S. sapporoensis</emphasis>
(Okamoto)
</taxonomicName>
, however the general shape and setal pattern of the epiproct is different among these species [except 
<taxonomicName box="[1290,1443,304,327]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis box="[1290,1443,304,327]" italics="true" pageId="3" pageNumber="159">S. sapporensis</emphasis>
</taxonomicName>
(
<figureCitation box="[843,970,339,363]" captionStart="Figs" captionStartId="4.[189,241,1689,1713]" captionTargetBox="[297,1337,269,1576]" captionTargetId="figure-57@4.[287,1345,259,1586]" captionTargetPageId="4" captionText="Figs. 19-24. Scanning electron micrographs of male genitalia. Sweltsa abdominalis, Japan,Yoki-zawa, Yamakitamachi, Kanagawa, 10 May 1996, T. Shimizu, (19-21). 19. Male terminalia, dorsal. 20. Epiproct, lateral. 21. Epiproct apex, lateral. Sweltsa sapporoensis, streamlets along Tottabetsu-gawa stream, Obihiro City, Hokkaido, Japan, 16 June 1992, T. Shimizu (22-24). 22. Male terminalia, dorsal. 23. Epiproct, lateral. 24. Epiproct apex, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4759399" httpUri="https://zenodo.org/record/4759399/files/figure.png" pageId="3" pageNumber="159">Figs. 22-24</figureCitation>
) and 
<taxonomicName authorityName="Ham" authorityYear="2008" baseAuthorityName="Okamoto" baseAuthorityYear="1912" box="[1035,1174,339,363]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="nikkoensis">
<emphasis box="[1035,1174,339,363]" italics="true" pageId="3" pageNumber="159">S. nikkoensis</emphasis>
</taxonomicName>
(
<figureCitation box="[1188,1314,339,363]" captionStart="Figs" captionStartId="5.[189,241,1689,1713]" captionTargetBox="[297,1337,269,1575]" captionTargetId="figure-57@5.[287,1345,259,1583]" captionTargetPageId="5" captionText="Figs. 25-30. Scanning electron micrographs of male genitalia. Sweltsa nikkoensis, Russia, p. Kedrovaja, Zapovednik Kedrovaja Pad, 14 May 2002, V. Teslenko det. (25-27). 25. Male terminalia, dorsal. 26. Epiproct apex, dorsal. 27. Male 9th tergal process. Sweltsa insularis, Russia, o. Sahalin, p.Belaja, sb. Makarchenko, 29 June-7 July 1986, V. Teslenko det.(28-30). 28. Male terminalia, dorsal. 29. Epiproct apex, dorsal. 30. Male terminalia, oblique dorsolateral." figureDoi="http://doi.org/10.5281/zenodo.4759405" httpUri="https://zenodo.org/record/4759405/files/figure.png" pageId="3" pageNumber="159">Figs. 25-27</figureCitation>
) which are quite similar]. 
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2009" box="[1010,1142,374,398]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="wui">
<emphasis box="[1010,1142,374,398]" italics="true" pageId="3" pageNumber="159">Sweltsa wui</emphasis>
</taxonomicName>
is the only known Asian species in which the epiproct apex is as wide as, or wider than, the base (
<figureCitation box="[1087,1189,445,469]" captionStart="Figs" captionStartId="2.[189,241,1618,1642]" captionTargetBox="[297,1337,231,1538]" captionTargetId="figure-57@2.[287,1345,223,1546]" captionTargetPageId="2" captionText="Figs. 5-10. Scanning electron micrographs of male genitalia. Sweltsa wui (5-8). 5. Terminalia, dorsal. 6. Epiproct, dorsal. 7. Epiproct, lateral. 8. Epiproct surface detail, frontal aspect. Haploperla valentinae (9-10). 9. Terminalia, dorsal. 10. Epiproct, dorsoapical margin." figureDoi="http://doi.org/10.5281/zenodo.4759393" httpUri="https://zenodo.org/record/4759393/files/figure.png" pageId="3" pageNumber="159">Figs. 5-6</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4759397" ID-Zenodo-Dep="4759397" httpUri="https://zenodo.org/record/4759397/files/figure.png" pageId="3" pageNumber="159" startId="3.[189,241,1796,1820]" targetBox="[345,1307,544,1678]" targetPageId="3">
<paragraph blockId="3.[189,1442,1796,1892]" pageId="3" pageNumber="159">
Figs. 11-18. 
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2009" box="[331,569,1796,1820]" class="Insecta" family="Chloroperlidae" genus="Haploperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="159" phylum="Arthropoda" rank="species" species="valentinae">
<emphasis box="[331,569,1796,1820]" italics="true" pageId="3" pageNumber="159">Haploperla valentinae</emphasis>
</taxonomicName>
. 11. Adult head and pronotum. 12. Male terminalia, dorsal. 13. Epiproct, dorsal. 14. Metanotal U-suture. 15. Female sterna 8-9. 16. Larval head and pronotum. 17. Epiproct, lateral. 18. Larval cercal segments 9-10, lateral.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4759399" ID-Zenodo-Dep="4759399" httpUri="https://zenodo.org/record/4759399/files/figure.png" pageId="4" pageNumber="160" startId="4.[189,241,1689,1713]" targetBox="[297,1337,269,1576]" targetPageId="4">
<paragraph blockId="4.[189,1443,1689,1855]" pageId="4" pageNumber="160">
Figs. 19-24. Scanning electron micrographs of male genitalia. 
<taxonomicName baseAuthorityName="Okamoto" baseAuthorityYear="1912" box="[897,1109,1689,1713]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="160" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[897,1109,1689,1713]" italics="true" pageId="4" pageNumber="160">Sweltsa abdominalis</emphasis>
</taxonomicName>
, Japan,Yoki-zawa, Yamakitamachi, Kanagawa, 10 May 1996, T. Shimizu, (19-21). 19. Male terminalia, dorsal. 20. Epiproct, lateral. 21. Epiproct apex, lateral. 
<taxonomicName box="[450,673,1760,1784]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="160" phylum="Arthropoda" rank="species" species="sapporoensis">
<emphasis box="[450,673,1760,1784]" italics="true" pageId="4" pageNumber="160">Sweltsa sapporoensis</emphasis>
</taxonomicName>
, streamlets along Tottabetsu-gawa stream, Obihiro City, Hokkaido, Japan, 16 June 1992, T. Shimizu (22-24). 22. Male terminalia, dorsal. 23. Epiproct, lateral. 24. Epiproct apex, dorsal.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4759405" ID-Zenodo-Dep="4759405" httpUri="https://zenodo.org/record/4759405/files/figure.png" pageId="5" pageNumber="161" startId="5.[189,241,1689,1713]" targetBox="[297,1337,269,1575]" targetPageId="5">
<paragraph blockId="5.[189,1443,1688,1855]" pageId="5" pageNumber="161">
Figs. 25-30. Scanning electron micrographs of male genitalia. 
<taxonomicName authorityName="Ham" authorityYear="2008" baseAuthorityName="Okamoto" baseAuthorityYear="1912" box="[958,1163,1689,1713]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="161" phylum="Arthropoda" rank="species" species="nikkoensis">
<emphasis box="[958,1163,1689,1713]" italics="true" pageId="5" pageNumber="161">Sweltsa nikkoensis</emphasis>
</taxonomicName>
, Russia, p. Kedrovaja, Zapovednik Kedrovaja Pad, 14 May 2002, V. Teslenko det. (25-27). 25. Male terminalia, dorsal. 26. Epiproct apex, dorsal. 27. Male 9 
<superScript attach="left" box="[470,485,1759,1773]" fontSize="6" pageId="5" pageNumber="161">th</superScript>
tergal process. 
<taxonomicName baseAuthorityName="Zhiltzova &amp; Levanidova" baseAuthorityYear="1978" box="[671,856,1760,1784]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="161" phylum="Arthropoda" rank="species" species="insularis">
<emphasis box="[671,856,1760,1784]" italics="true" pageId="5" pageNumber="161">Sweltsa insularis</emphasis>
</taxonomicName>
, Russia, o. Sahalin, p.Belaja, sb. Makarchenko, 29 June-7 July 1986, V. Teslenko det.(28-30). 28. Male terminalia, dorsal. 29. Epiproct apex, dorsal. 30. Male terminalia, oblique dorsolateral.
</paragraph>
</caption>
<paragraph blockId="6.[189,798,197,932]" pageId="6" pageNumber="162">
The species of genus 
<taxonomicName box="[475,557,197,221]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis box="[475,557,197,221]" italics="true" pageId="6" pageNumber="162">Sweltsa</emphasis>
</taxonomicName>
are quite variable in epiproct structure and several species complexes in the group may at some time be given generic rank. The structure of tergum 
<quantity box="[475,518,303,327]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="6" pageNumber="162" unit="in" value="9.0">9 in</quantity>
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2009" box="[525,594,304,327]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="wui">
<emphasis box="[525,594,304,327]" italics="true" pageId="6" pageNumber="162">S. wui</emphasis>
</taxonomicName>
suggests it could be placed in another genus. A few, unrelated Nearctic species are also known to lack the transverse process (
<bibRefCitation author="Kondratieff, B. C. &amp; R. W. Baumann" box="[297,662,410,434]" pageId="6" pageNumber="162" pagination="20 - 29" refId="ref2320" refString="Kondratieff, B. C. &amp; R. W. Baumann. 2009. A contribution to the knowledge of Sweltsa exquisita (Frison) and S. occidens (Frison) and description of a new species of Sweltsa from the northern Rocky Mountains, U. S. A. (Plecoptera: Chloroperlidae). Illiesia, 5: 20 - 29." type="journal article" year="2009">Kondratieff &amp; Baumann 2009</bibRefCitation>
), but these species have the posterior margin of tergum 9 strongly concave as in other Nearctic members of the genus. Our study of Asian 
<taxonomicName box="[538,620,516,540]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis box="[538,620,516,540]" italics="true" pageId="6" pageNumber="162">Sweltsa</emphasis>
</taxonomicName>
indicates two Japanese species, 
<taxonomicName baseAuthorityName="Okamoto" baseAuthorityYear="1912" box="[401,560,552,576]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[401,560,552,576]" italics="true" pageId="6" pageNumber="162">S. abdominalis</emphasis>
</taxonomicName>
(
<figureCitation box="[577,706,552,576]" captionStart="Figs" captionStartId="4.[189,241,1689,1713]" captionTargetBox="[297,1337,269,1576]" captionTargetId="figure-57@4.[287,1345,259,1586]" captionTargetPageId="4" captionText="Figs. 19-24. Scanning electron micrographs of male genitalia. Sweltsa abdominalis, Japan,Yoki-zawa, Yamakitamachi, Kanagawa, 10 May 1996, T. Shimizu, (19-21). 19. Male terminalia, dorsal. 20. Epiproct, lateral. 21. Epiproct apex, lateral. Sweltsa sapporoensis, streamlets along Tottabetsu-gawa stream, Obihiro City, Hokkaido, Japan, 16 June 1992, T. Shimizu (22-24). 22. Male terminalia, dorsal. 23. Epiproct, lateral. 24. Epiproct apex, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4759399" httpUri="https://zenodo.org/record/4759399/files/figure.png" pageId="6" pageNumber="162">Figs. 19-21</figureCitation>
) and 
<taxonomicName class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="sapporoensis">
<emphasis italics="true" pageId="6" pageNumber="162">S. sapporoensis</emphasis>
</taxonomicName>
(
<figureCitation box="[356,499,587,611]" captionStart="Figs" captionStartId="4.[189,241,1689,1713]" captionTargetBox="[297,1337,269,1576]" captionTargetId="figure-57@4.[287,1345,259,1586]" captionTargetPageId="4" captionText="Figs. 19-24. Scanning electron micrographs of male genitalia. Sweltsa abdominalis, Japan,Yoki-zawa, Yamakitamachi, Kanagawa, 10 May 1996, T. Shimizu, (19-21). 19. Male terminalia, dorsal. 20. Epiproct, lateral. 21. Epiproct apex, lateral. Sweltsa sapporoensis, streamlets along Tottabetsu-gawa stream, Obihiro City, Hokkaido, Japan, 16 June 1992, T. Shimizu (22-24). 22. Male terminalia, dorsal. 23. Epiproct, lateral. 24. Epiproct apex, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4759399" httpUri="https://zenodo.org/record/4759399/files/figure.png" pageId="6" pageNumber="162">Figs. 22-24</figureCitation>
) share the tergum 9 modifications with Nearctic members of the group but also share a similar color pattern with 
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2009" box="[689,759,659,682]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="wui">
<emphasis box="[689,759,659,682]" italics="true" pageId="6" pageNumber="162">S. wui</emphasis>
</taxonomicName>
. In addition, 
<taxonomicName baseAuthorityName="Zhiltzova &amp; Levanidova" baseAuthorityYear="1978" box="[303,425,694,718]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="insularis">
<emphasis box="[303,425,694,718]" italics="true" pageId="6" pageNumber="162">S. insularis</emphasis>
</taxonomicName>
, a similarly pigmented Russian species, also lacks the tergal 9 modification (
<figureCitation captionStart="Figs" captionStartId="5.[189,241,1689,1713]" captionTargetBox="[297,1337,269,1575]" captionTargetId="figure-57@5.[287,1345,259,1583]" captionTargetPageId="5" captionText="Figs. 25-30. Scanning electron micrographs of male genitalia. Sweltsa nikkoensis, Russia, p. Kedrovaja, Zapovednik Kedrovaja Pad, 14 May 2002, V. Teslenko det. (25-27). 25. Male terminalia, dorsal. 26. Epiproct apex, dorsal. 27. Male 9th tergal process. Sweltsa insularis, Russia, o. Sahalin, p.Belaja, sb. Makarchenko, 29 June-7 July 1986, V. Teslenko det.(28-30). 28. Male terminalia, dorsal. 29. Epiproct apex, dorsal. 30. Male terminalia, oblique dorsolateral." figureDoi="http://doi.org/10.5281/zenodo.4759405" httpUri="https://zenodo.org/record/4759405/files/figure.png" pageId="6" pageNumber="162">Figs. 28- 30</figureCitation>
), but 
<taxonomicName authorityName="Zhiltzova &amp; Levanidova" authorityYear="1978" box="[288,403,765,789]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="colorata">
<emphasis box="[288,403,765,789]" italics="true" pageId="6" pageNumber="162">S. colorata</emphasis>
</taxonomicName>
(= 
<taxonomicName authorityName="Ham" authorityYear="2008" baseAuthorityName="Okamoto" baseAuthorityYear="1912" box="[442,582,765,789]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="nikkoensis">
<emphasis box="[442,582,765,789]" italics="true" pageId="6" pageNumber="162">S. nikkoensis</emphasis>
</taxonomicName>
), another Russian species with similar pigment, has a well developed process on tergum 9 (
<figureCitation box="[446,574,836,860]" captionStart="Figs" captionStartId="5.[189,241,1689,1713]" captionTargetBox="[297,1337,269,1575]" captionTargetId="figure-57@5.[287,1345,259,1583]" captionTargetPageId="5" captionText="Figs. 25-30. Scanning electron micrographs of male genitalia. Sweltsa nikkoensis, Russia, p. Kedrovaja, Zapovednik Kedrovaja Pad, 14 May 2002, V. Teslenko det. (25-27). 25. Male terminalia, dorsal. 26. Epiproct apex, dorsal. 27. Male 9th tergal process. Sweltsa insularis, Russia, o. Sahalin, p.Belaja, sb. Makarchenko, 29 June-7 July 1986, V. Teslenko det.(28-30). 28. Male terminalia, dorsal. 29. Epiproct apex, dorsal. 30. Male terminalia, oblique dorsolateral." figureDoi="http://doi.org/10.5281/zenodo.4759405" httpUri="https://zenodo.org/record/4759405/files/figure.png" pageId="6" pageNumber="162">Figs. 25-27</figureCitation>
). Consequently, in the absence of data from other life stages, we are tentatively placing the new species in genus 
<taxonomicName box="[708,790,907,931]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis box="[708,790,907,931]" italics="true" pageId="6" pageNumber="162">Sweltsa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>