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<document id="B28EB6253AC3801932B13DF5230D4EE4" ID-DOI="10.26028/cybium/2020-442-008" ID-ISSN="2101-0315" ID-Zenodo-Dep="12711652" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720644681295" checkinUser="felipe" docAuthor="Okamoto, Makoto, Chen, Wei-Jen &amp; Motomura, Hiroyuki" docDate="2020" docId="03C25D25FD4AFFE1FF0BFD17C553FA3D" docLanguage="en" docName="Cybium.44.2.165-168.pdf" docOrigin="Cybium 44 (2)" docSource="http://dx.doi.org/10.26028/cybium/2020-442-008" docStyle="DocumentStyle:EAB95795642928CB986F0923DE58F0B7.1:Cybium.2019-.journal_article" docStyleId="EAB95795642928CB986F0923DE58F0B7" docStyleName="Cybium.2019-.journal_article" docStyleVersion="1" docTitle="Epigonus exodon Okamoto &amp; Motomura 2012" docType="treatment" docVersion="3" lastPageNumber="167" masterDocId="FFFB255DFD48FFE3FFD3FFECC02AFFC7" masterDocTitle="New distributional records of three deepwater cardinalfishes Epigonus angustifrons, E. denticulatus, and E. exodon (Perciformes: Epigonidae) in the South Indian Ocean" masterLastPageNumber="168" masterPageNumber="165" pageNumber="167" updateTime="1720653836308" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6A085E5BA07A625223D12083F75C2C84">New distributional records of three deepwater cardinalfishes Epigonus angustifrons, E. denticulatus, and E. exodon (Perciformes: Epigonidae) in the South Indian Ocean</mods:title>
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<mods:namePart id="172266B6546FD8041C037E90513609BF">Okamoto, Makoto</mods:namePart>
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<mods:namePart id="9F2D22FBEDADB92971944ECFFE08C524">Chen, Wei-Jen</mods:namePart>
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<mods:namePart id="7176CB8726928FD7BFEE3F1077D348FD">Motomura, Hiroyuki</mods:namePart>
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<treatment id="03C25D25FD4AFFE1FF0BFD17C553FA3D" ID-DOI="http://doi.org/10.5281/zenodo.12713520" ID-Zenodo-Dep="12713520" LSID="urn:lsid:plazi:treatment:03C25D25FD4AFFE1FF0BFD17C553FA3D" httpUri="http://treatment.plazi.org/id/03C25D25FD4AFFE1FF0BFD17C553FA3D" lastPageNumber="167" pageId="2" pageNumber="167">
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<paragraph id="8BD4EC33FD4AFFE1FF0BFD17C299FCD7" blockId="2.[216,691,763,785]" box="[216,691,763,785]" pageId="2" pageNumber="167">
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<taxonomicName id="4C6B97B0FD4AFFE1FF0BFD17C299FCD7" ID-CoL="6FTJG" authority="Okamoto &amp; Motomura, 2012" authorityName="Okamoto &amp; Motomura" authorityYear="2012" box="[216,691,763,785]" family="Epigonidae" genus="Epigonus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B91F3021FD4AFFE1FF0BFD17C299FCD7" bold="true" box="[216,691,763,785]" pageId="2" pageNumber="167">
<emphasis id="B91F3021FD4AFFE1FF0BFD17C1AEFCD6" bold="true" box="[216,388,763,785]" italics="true" pageId="2" pageNumber="167">Epigonus exodon</emphasis>
<bibRefCitation id="EFFA91C2FD4AFFE1FE59FD17C299FCD7" author="OKAMOTO M. &amp; MOTOMURA H." box="[394,691,763,785]" pageId="2" pageNumber="167" pagination="84 - 88" refId="ref3313" refString="OKAMOTO M. &amp; MOTOMURA H., 2012. - Epigonus exodon, a new species of deepwater cardinalfish (Teleostei: Perciformes: Epigonidae) from Reunion, western Indian Ocean. Zootaxa, 3453: 84 - 88. DOI: 10.11646 / zootaxa. 3453.1.6" type="journal article" year="2012">Okamoto &amp; Motomura, 2012</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C371BFB8FD4AFFE1FE52FCFBC220FCEB" box="[385,522,791,813]" pageId="2" pageNumber="167" type="description">
<paragraph id="8BD4EC33FD4AFFE1FE52FCFBC220FCEB" blockId="2.[385,522,791,813]" box="[385,522,791,813]" pageId="2" pageNumber="167">
(
<figureCitation id="1350F0B6FD4AFFE1FE5BFCFBC1E8FCEB" box="[392,450,791,813]" captionStart="Figure 4" captionStartId="2.[132,190,1863,1882]" captionTargetBox="[132,775,1408,1835]" captionTargetId="figure-870@2.[132,775,1408,1842]" captionTargetPageId="2" captionText="Figure 4. Epigonus exodon, NTUM 13370, south of Mayotte, western South Indian Ocean.A: Fresh specimen; B: After fixation." figureDoi="http://doi.org/10.5281/zenodo.12711662" httpUri="https://zenodo.org/record/12711662/files/figure.png" pageId="2" pageNumber="167">Fig. 4</figureCitation>
;
<tableCitation id="C6E9D988FD4AFFE1FE1FFCFBC22EFCEB" box="[460,516,791,813]" captionStart="Table I" captionStartId="1.[132,179,217,236]" captionText="Table I. Meristic counts and measurements (% of standard length) of Epigonus angustifrons, E. denticulatus, and E. exodon specimens examined in this study." pageId="2" pageNumber="167">Tab. I</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C371BFB8FD4AFFE1FF57FCA9C156FC0F" pageId="2" pageNumber="167" type="materials_examined">
<paragraph id="8BD4EC33FD4AFFE1FF57FCA9C16CFC9C" blockId="2.[132,775,837,968]" box="[132,326,837,859]" pageId="2" pageNumber="167">
<emphasis id="B91F3021FD4AFFE1FF57FCA9C16CFC9C" bold="true" box="[132,326,837,859]" pageId="2" pageNumber="167">Material examined</emphasis>
</paragraph>
<paragraph id="8BD4EC33FD4AFFE1FF79FC8CC156FC0F" blockId="2.[132,775,837,968]" pageId="2" pageNumber="167">
<materialsCitation id="3B03E66EFD4AFFE1FF79FC8CC152FC0F" collectingDate="2017-01-27" collectionCode="RV" collectorName="St. CP &amp; French &amp; Antea &amp; Expedition" country="Mayotte" elevation="504" latitude="-13.05" location="western South Indian Ocean" longLatPrecision="1291" longitude="45.016666" pageId="2" pageNumber="167" specimenCode="NTUM 13370" specimenCount="1">
<specimenCount id="9D6D27BAFD4AFFE1FF79FC8CC11DFCB1" box="[170,311,864,886]" count="1" pageId="2" pageNumber="167" type="generic">One specimen</specimenCount>
:
<specimenCode id="DBCD4448FD4AFFE1FE93FC8DC1E1FCB2" box="[320,459,864,886]" collectionCode="NTUM" pageId="2" pageNumber="167">NTUM 13370</specimenCode>
, 131.0 mm SL, female,
<geoCoordinate id="EE5F8AF4FD4AFFE1FD61FC8CC329FCB2" box="[690,771,864,885]" degrees="13" direction="south" minutes="03" orientation="latitude" pageId="2" pageNumber="167" precision="925" value="-13.05">13°03S</geoCoordinate>
,
<geoCoordinate id="EE5F8AF4FD4AFFE1FF57FC97C0F0FC56" box="[132,218,891,913]" degrees="45" direction="east" minutes="01" orientation="longitude" pageId="2" pageNumber="167" precision="925" value="45.016666">45°01E</geoCoordinate>
, south of
<collectingCountry id="F37CACA3FD4AFFE1FEEDFC90C1BEFC56" box="[318,404,892,913]" name="Mayotte" pageId="2" pageNumber="167">Mayotte</collectingCountry>
,
<location id="8EB4BAE8FD4AFFE1FE4DFC90C297FC57" LSID="urn:lsid:plazi:treatment:03C25D25FD4AFFE1FF0BFD17C553FA3D:8EB4BAE8FD4AFFE1FE4DFC90C297FC57" box="[414,701,891,913]" country="Mayotte" latitude="-13.05" longLatPrecision="1291" longitude="45.016666" name="western South Indian Ocean" pageId="2" pageNumber="167">western South Indian Ocean</location>
,
<elevation id="00460B00FD4AFFE1FD1BFC97C095FC6A" metricMagnitude="2" metricUnit="m" metricValue="5.04" pageId="2" pageNumber="167" unit="m" value="504.0">
<quantity id="4C9341D6FD4AFFE1FD1BFC97C32CFC56" box="[712,774,891,913]" metricMagnitude="2" metricUnit="m" metricValue="5.04" pageId="2" pageNumber="167" unit="m" value="504.0">504 m</quantity>
depth
</elevation>
,
<collectorName id="269E89E5FD4AFFE1FF19FC7BC121FC6B" box="[202,267,919,940]" pageId="2" pageNumber="167">St. CP</collectorName>
4829,
<collectorName id="269E89E5FD4AFFE1FE9AFC7BC1A4FC6A" box="[329,398,919,941]" pageId="2" pageNumber="167">French</collectorName>
beam trawl,
<date id="FFD5CAF3FD4AFFE1FDC0FC7BC2BFFC6B" box="[531,661,919,940]" pageId="2" pageNumber="167" value="2017-01-27">
<collectingDate id="EF91331BFD4AFFE1FDC0FC7BC2BFFC6B" box="[531,661,919,940]" pageId="2" pageNumber="167" value="2017-01-27">27 Jan. 2017</collectingDate>
</date>
,
<collectionCode id="ED7A74F6FD4AFFE1FD73FC7BC2EAFC6B" box="[672,704,919,940]" country="Italy" name="Collection of Leptospira Strains" pageId="2" pageNumber="167">RV</collectionCode>
<emphasis id="B91F3021FD4AFFE1FD14FC7BC32BFC6B" box="[711,769,919,940]" italics="true" pageId="2" pageNumber="167">
<collectorName id="269E89E5FD4AFFE1FD14FC7BC32BFC6B" box="[711,769,919,940]" pageId="2" pageNumber="167">Antéa</collectorName>
</emphasis>
, BIOMAGLO
<collectorName id="269E89E5FD4AFFE1FED8FC5EC152FC0F" box="[267,376,946,968]" pageId="2" pageNumber="167">Expedition</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C371BFB8FD4AFFE1FF57FC04C13DFB1F" pageId="2" pageNumber="167" type="diagnosis">
<paragraph id="8BD4EC33FD4AFFE1FF57FC04C0CDFC39" blockId="2.[132,775,1000,1240]" box="[132,231,1000,1022]" pageId="2" pageNumber="167">
<emphasis id="B91F3021FD4AFFE1FF57FC04C0CDFC39" bold="true" box="[132,231,1000,1022]" pageId="2" pageNumber="167">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BD4EC33FD4AFFE1FF79FBEFC13DFB1F" blockId="2.[132,775,1000,1240]" pageId="2" pageNumber="167">
Dorsal-fin rays VII-I, 10; pectoral-fin rays 16-17; total gill rakers 26-28; vertebrae 10 + 15; pyloric caeca 7-10; pored lateral-line scales 35-40 + 3-5; opercular spine absent; ribs absent on last abdominal vertebra; anteriorly projecting teeth present on each side of symphysis of lower jaw (&gt;
<quantity id="4C9341D6FD4AFFE1FE3AFB9CC21AFB41" box="[489,560,1136,1158]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="167" unit="mm" value="60.0">60 mm</quantity>
SL); tongue narrow; shallow V-shaped tooth patch present on rear two-thirds of tongue (&lt;
<quantity id="4C9341D6FD4AFFE1FF71FB4BC0DFFB7B" box="[162,245,1191,1212]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="167" unit="mm" value="100.0">100 mm</quantity>
SL, reduced with growth), maxillary mustache-like process absent.
</paragraph>
</subSubSection>
<subSubSection id="C371BFB8FD4AFFE1FF57FB14C228FA98" pageId="2" pageNumber="167" type="distribution">
<paragraph id="8BD4EC33FD4AFFE1FF57FB14C12BFAC9" blockId="2.[132,775,1272,1375]" box="[132,257,1272,1294]" pageId="2" pageNumber="167">
<emphasis id="B91F3021FD4AFFE1FF57FB14C12BFAC9" bold="true" box="[132,257,1272,1294]" pageId="2" pageNumber="167">Distribution</emphasis>
</paragraph>
<paragraph id="8BD4EC33FD4AFFE1FF79FAFFC228FA98" blockId="2.[132,775,1272,1375]" pageId="2" pageNumber="167">
<collectingCountry id="F37CACA3FD4AFFE1FF79FAFFC0D5FAEE" box="[170,255,1299,1321]" name="Réunion" pageId="2" pageNumber="167">Réunion</collectingCountry>
and the
<collectingCountry id="F37CACA3FD4AFFE1FE8AFAFFC19FFAEF" box="[345,437,1299,1320]" name="Comoros" pageId="2" pageNumber="167">Comoros</collectingCountry>
Archipelago (south of
<collectingCountry id="F37CACA3FD4AFFE1FD70FAFFC2D1FAEF" box="[675,763,1299,1320]" name="Mayotte" pageId="2" pageNumber="167">Mayotte</collectingCountry>
), western South Indian Ocean, at
<quantity id="4C9341D6FD4AFFE1FE12FAC2C207FA83" box="[449,557,1326,1348]" metricMagnitude="2" metricUnit="m" metricValue="4.77" metricValueMax="5.04" metricValueMin="4.5" pageId="2" pageNumber="167" unit="m" value="477.0" valueMax="504.0" valueMin="450.0">450-504 m</quantity>
depths (
<bibRefCitation id="EFFA91C2FD4AFFE1FD53FAC2C102FA98" author="OKAMOTO M. &amp; MOTOMURA H." pageId="2" pageNumber="167" pagination="84 - 88" refId="ref3313" refString="OKAMOTO M. &amp; MOTOMURA H., 2012. - Epigonus exodon, a new species of deepwater cardinalfish (Teleostei: Perciformes: Epigonidae) from Reunion, western Indian Ocean. Zootaxa, 3453: 84 - 88. DOI: 10.11646 / zootaxa. 3453.1.6" type="journal article" year="2012">Okamoto and Motomura, 2012</bibRefCitation>
; present study:
<figureCitation id="1350F0B6FD4AFFE1FE6FFAA5C1DFFA98" box="[444,501,1353,1375]" captionStart="Figure 2" captionStartId="1.[813,870,857,876]" captionTargetBox="[813,1454,215,832]" captionTargetId="figure-901@1.[813,1456,215,835]" captionTargetPageId="1" captionText="Figure 2. Collection localities of Epigonus angustifrons (present study: Z; Okamoto and Gon, 2018: „), E. denticulatus (present study: Δ; Okamoto and Gon, 2018: A), and E. exodon (present study: C; Okamoto and Motomura, 2012: l) in the central and western South Indian Ocean." figureDoi="http://doi.org/10.5281/zenodo.12711656" httpUri="https://zenodo.org/record/12711656/files/figure.png" pageId="2" pageNumber="167">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C371BFB8FD4AFFE1FCFEFF35C553FA3D" pageId="2" pageNumber="167" type="discussion">
<paragraph id="8BD4EC33FD4AFFE1FCFEFF35C3A0FF29" blockId="2.[812,1456,217,1172]" box="[813,906,217,238]" pageId="2" pageNumber="167">
<emphasis id="B91F3021FD4AFFE1FCFEFF35C3A0FF29" bold="true" box="[813,906,217,238]" pageId="2" pageNumber="167">Remarks</emphasis>
</paragraph>
<paragraph id="8BD4EC33FD4AFFE1FC81FF18C542FCD4" blockId="2.[812,1456,217,1172]" pageId="2" pageNumber="167">
The diagnosis of
<taxonomicName id="4C6B97B0FD4AFFE1FC2FFF19C474FECE" authorityName="Okamoto &amp; Motomura" authorityYear="2012" box="[1020,1118,244,266]" family="Epigonidae" genus="Epigonus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B91F3021FD4AFFE1FC2FFF19C474FECE" box="[1020,1118,244,266]" italics="true" pageId="2" pageNumber="167">E. exodon</emphasis>
</taxonomicName>
given here is based on the present specimen and the original description.
<taxonomicName id="4C6B97B0FD4AFFE1FB72FEFDC562FEE2" authorityName="Okamoto &amp; Motomura" authorityYear="2012" box="[1185,1352,272,294]" family="Epigonidae" genus="Epigonus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B91F3021FD4AFFE1FB72FEFDC562FEE2" box="[1185,1352,272,294]" italics="true" pageId="2" pageNumber="167">Epigonus exodon</emphasis>
</taxonomicName>
was originally described on the basis of
<specimenCount id="9D6D27BAFD4AFFE1FB81FEC0C4F5FE86" box="[1106,1247,299,321]" count="2" pageId="2" pageNumber="167" type="generic">two specimens</specimenCount>
collected from
<collectingCountry id="F37CACA3FD4AFFE1FAA7FEC7C37AFE9A" name="Réunion" pageId="2" pageNumber="167">Réunion</collectingCountry>
, Southwest Indian Ocean (
<bibRefCitation id="EFFA91C2FD4AFFE1FBB1FEABC589FE9B" author="OKAMOTO M. &amp; MOTOMURA H." box="[1122,1443,327,349]" pageId="2" pageNumber="167" pagination="84 - 88" refId="ref3313" refString="OKAMOTO M. &amp; MOTOMURA H., 2012. - Epigonus exodon, a new species of deepwater cardinalfish (Teleostei: Perciformes: Epigonidae) from Reunion, western Indian Ocean. Zootaxa, 3453: 84 - 88. DOI: 10.11646 / zootaxa. 3453.1.6" type="journal article" year="2012">Okamoto and Motomura, 2012</bibRefCitation>
). Although a large number of specimens of
<taxonomicName id="4C6B97B0FD4AFFE1FB12FE8FC536FEBF" authorityName="Rafinesque" authorityYear="1810" box="[1217,1308,355,376]" family="Epigonidae" genus="Epigonus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="genus">
<emphasis id="B91F3021FD4AFFE1FB12FE8FC536FEBF" box="[1217,1308,355,376]" italics="true" pageId="2" pageNumber="167">Epigonus</emphasis>
</taxonomicName>
from the western Indian Ocean were examined in museum collections worldwide by
<bibRefCitation id="EFFA91C2FD4AFFE1FC99FE75C46DFE69" author="OKAMOTO M. &amp; GON O." box="[842,1095,409,431]" pageId="2" pageNumber="167" pagination="261 - 291" refId="ref3199" refString="OKAMOTO M. &amp; GON O., 2018. - A review of the deepwater cardinalfish genus Epigonus (Perciformes: Epigonidae) of the Western Indian Ocean, with description of two new species. Zootaxa, 4382: 261 - 291. DOI: 10.11646 / zootaxa. 4382.2.3" type="journal article" year="2018">Okamoto and Gon (2018)</bibRefCitation>
; no additional specimens of this species were found. In the present study, we report this specimen as the third specimen and the first record of the species from the Comoro Archipelago with a colour photograph of the species when fresh.
<taxonomicName id="4C6B97B0FD4AFFE1FCFEFDEBC3F8FDDB" authorityName="Okamoto &amp; Motomura" authorityYear="2012" box="[813,978,519,540]" family="Epigonidae" genus="Epigonus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B91F3021FD4AFFE1FCFEFDEBC3F8FDDB" box="[813,978,519,540]" italics="true" pageId="2" pageNumber="167">Epigonus exodon</emphasis>
</taxonomicName>
has been recognized as a member of the
<taxonomicName id="4C6B97B0FD4AFFE1FA88FDEBC376FDF0" authorityName="Mayer" authorityYear="1974" family="Epigonidae" genus="Epigonus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="oligolepis">
<emphasis id="B91F3021FD4AFFE1FA88FDEBC376FDF0" italics="true" pageId="2" pageNumber="167">E. oligolepis</emphasis>
</taxonomicName>
group by lacking a pungent opercular spine and the absence of ribs on the last abdominal vertebra, and in having dorsal-fin rays VII-I, 10 and 35-40 pored lateral-line scales to the end of the hypural (
<bibRefCitation id="EFFA91C2FD4AFFE1FC5AFD98C4E0FD4D" author="OKAMOTO M. &amp; MOTOMURA H." box="[905,1226,628,650]" pageId="2" pageNumber="167" pagination="155 - 160" refId="ref3251" refString="OKAMOTO M. &amp; MOTOMURA H., 2011. - Epigonus carbonarius, a new species of deepwater cardinalfish (Perciformes: Epigonidae) from the Marquesas Islands, with a redefinition of the Epigonus oligolepis group. Ichthyol. Res., 58: 155 - 160. DOI: 10.1007 / s 10228 - 011 - 0205 - 7" type="journal article" year="2011">Okamoto and Motomura, 2011</bibRefCitation>
,
<bibRefCitation id="EFFA91C2FD4AFFE1FB06FD99C521FD4D" author="OKAMOTO M. &amp; MOTOMURA H." box="[1237,1291,629,650]" pageId="2" pageNumber="167" pagination="84 - 88" refId="ref3313" refString="OKAMOTO M. &amp; MOTOMURA H., 2012. - Epigonus exodon, a new species of deepwater cardinalfish (Teleostei: Perciformes: Epigonidae) from Reunion, western Indian Ocean. Zootaxa, 3453: 84 - 88. DOI: 10.11646 / zootaxa. 3453.1.6" type="journal article" year="2012">2012</bibRefCitation>
; present study). Among the species of this group, only
<taxonomicName id="4C6B97B0FD4AFFE1FB4DFD7DC4D7FD62" authorityName="Okamoto &amp; Motomura" authorityYear="2012" box="[1182,1277,656,678]" family="Epigonidae" genus="Epigonus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B91F3021FD4AFFE1FB4DFD7DC4D7FD62" box="[1182,1277,656,678]" italics="true" pageId="2" pageNumber="167">E. exodon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B97B0FD4AFFE1FAF8FD7DC39BFD06" authority="Gon 1985" authorityName="Gon" authorityYear="1985" family="Epigonidae" genus="Epigonus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="glossodontus">
<emphasis id="B91F3021FD4AFFE1FAF8FD7DC360FD06" italics="true" pageId="2" pageNumber="167">E. glossodontus</emphasis>
<bibRefCitation id="EFFA91C2FD4AFFE1FC83FD40C39BFD06" author="GON O." box="[848,945,683,705]" pageId="2" pageNumber="167" pagination="221 - 229" refId="ref2966" refString="GON O., 1985. - Two new species of the deepsea cardinalfish genus Epigonus (Perciformes, Apogonidae) from the Hawaiian Islands, with a key to the Hawaiian species. Pac. Sci., 39 (2): 221 - 229." type="journal article" year="1985">Gon 1985</bibRefCitation>
</taxonomicName>
(Hawaiian Islands) have anteriorly projecting teeth on the symphysis of lower jaw (
<bibRefCitation id="EFFA91C2FD4AFFE1FBBAFD2BC4FCFD1A" author="GON O." box="[1129,1238,711,733]" pageId="2" pageNumber="167" pagination="221 - 229" refId="ref2966" refString="GON O., 1985. - Two new species of the deepsea cardinalfish genus Epigonus (Perciformes, Apogonidae) from the Hawaiian Islands, with a key to the Hawaiian species. Pac. Sci., 39 (2): 221 - 229." type="journal article" year="1985">Gon, 1985</bibRefCitation>
); however, the former can be distinguished from the latter in having a narrower tongue with a smaller tooth patch (
<bibRefCitation id="EFFA91C2FD4AFFE1FBE3FD12C577FCD4" author="OKAMOTO M. &amp; MOTOMURA H." box="[1072,1373,766,788]" pageId="2" pageNumber="167" pagination="84 - 88" refId="ref3313" refString="OKAMOTO M. &amp; MOTOMURA H., 2012. - Epigonus exodon, a new species of deepwater cardinalfish (Teleostei: Perciformes: Epigonidae) from Reunion, western Indian Ocean. Zootaxa, 3453: 84 - 88. DOI: 10.11646 / zootaxa. 3453.1.6" type="journal article" year="2012">Okamoto and Motomura, 2012</bibRefCitation>
).
</paragraph>
<paragraph id="8BD4EC33FD4AFFE1FC81FCF5C46AFBE1" blockId="2.[812,1456,217,1172]" pageId="2" pageNumber="167">
According to the original description of
<taxonomicName id="4C6B97B0FD4AFFE1FB31FCF6C42DFC8D" authority="(Okamoto and Motomura, 2012)" baseAuthorityName="Okamoto and Motomura" baseAuthorityYear="2012" family="Epigonidae" genus="Epigonus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B91F3021FD4AFFE1FB31FCF6C56FFCE9" box="[1250,1349,793,815]" italics="true" pageId="2" pageNumber="167">E. exodon</emphasis>
(
<bibRefCitation id="EFFA91C2FD4AFFE1FA81FCF5C42AFC8D" author="OKAMOTO M. &amp; MOTOMURA H." pageId="2" pageNumber="167" pagination="84 - 88" refId="ref3313" refString="OKAMOTO M. &amp; MOTOMURA H., 2012. - Epigonus exodon, a new species of deepwater cardinalfish (Teleostei: Perciformes: Epigonidae) from Reunion, western Indian Ocean. Zootaxa, 3453: 84 - 88. DOI: 10.11646 / zootaxa. 3453.1.6" type="journal article" year="2012">Okamoto and Motomura, 2012</bibRefCitation>
)
</taxonomicName>
, the
<typeStatus id="54D05291FD4AFFE1FBE6FCD8C4A1FC8D" box="[1077,1163,820,842]" pageId="2" pageNumber="167" type="holotype">holotype</typeStatus>
and
<typeStatus id="54D05291FD4AFFE1FB6FFCD9C53BFC8D" box="[1212,1297,821,842]" pageId="2" pageNumber="167" type="paratype">paratype</typeStatus>
had tooth patch on the tongue, which was regarded as an important diagnostic character. However, the present specimen has only a few teeth on the tongue, not forming a tooth patch. Since the standard length of the present specimen (131.0 mm SL) is larger than those of the type specimens (60.2-97.0 mm SL), the difference is regarded here as change with growth. A similar reduction of the teeth on the tongue with growth has been confirmed in
<taxonomicName id="4C6B97B0FD4AFFE1FB71FC19C585FBCD" authority="Mayer, 1974" authorityName="Mayer" authorityYear="1974" box="[1186,1455,1012,1034]" family="Epigonidae" genus="Epigonus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="oligolepis">
<emphasis id="B91F3021FD4AFFE1FB71FC19C50CFBCE" box="[1186,1318,1012,1034]" italics="true" pageId="2" pageNumber="167">E. oligolepis</emphasis>
<bibRefCitation id="EFFA91C2FD4AFFE1FAFEFC19C585FBCD" author="MAYER G. F." box="[1325,1455,1013,1034]" pageId="2" pageNumber="167" pagination="147 - 203" refId="ref3043" refString="MAYER G. F., 1974. - A revision of the cardinalfish genus Epigonus (Perciformes, Apogonidae), with descriptions of two new species. Bull. Mus. Comp. Zool., 146: 147 - 203." type="journal article" year="1974">Mayer, 1974</bibRefCitation>
</taxonomicName>
(M. Okamoto, unpubl. data).
</paragraph>
<paragraph id="8BD4EC33FD4AFFE1FC81FBC7C508FB53" blockId="2.[812,1456,217,1172]" pageId="2" pageNumber="167">
<bibRefCitation id="EFFA91C2FD4AFFE1FC81FBC7C4B4FB86" author="OKAMOTO M. &amp; MOTOMURA H." box="[850,1182,1067,1089]" pageId="2" pageNumber="167" pagination="84 - 88" refId="ref3313" refString="OKAMOTO M. &amp; MOTOMURA H., 2012. - Epigonus exodon, a new species of deepwater cardinalfish (Teleostei: Perciformes: Epigonidae) from Reunion, western Indian Ocean. Zootaxa, 3453: 84 - 88. DOI: 10.11646 / zootaxa. 3453.1.6" type="journal article" year="2012">Okamoto and Motomura (2012)</bibRefCitation>
reported that the sexes of the
<typeStatus id="54D05291FD4AFFE1FC81FBABC382FB9A" box="[850,936,1095,1117]" pageId="2" pageNumber="167" type="holotype">holotype</typeStatus>
and
<typeStatus id="54D05291FD4AFFE1FC0AFBABC404FB9B" box="[985,1070,1095,1116]" pageId="2" pageNumber="167" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C6B97B0FD4AFFE1FB9DFBABC49BFB9B" baseAuthorityName="Okamoto and Motomura" baseAuthorityYear="2012" box="[1102,1201,1095,1116]" family="Epigonidae" genus="Epigonus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B91F3021FD4AFFE1FB9DFBABC49BFB9B" box="[1102,1201,1095,1116]" italics="true" pageId="2" pageNumber="167">E. exodon</emphasis>
</taxonomicName>
were male and unknown, respectively. The present specimen was a female with a large number of developed eggs (ca.
<quantity id="4C9341D6FD4AFFE1FB8BFB92C48BFB54" box="[1112,1185,1150,1171]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="167" unit="mm" value="0.4">0.4 mm</quantity>
in diameter).
</paragraph>
<paragraph id="8BD4EC33FD4AFFE1FCFEFB52C553FA3D" blockId="2.[812,1456,1214,1531]" pageId="2" pageNumber="167">
<emphasis id="B91F3021FD4AFFE1FCFEFB52C3FDFB16" bold="true" box="[813,983,1214,1233]" pageId="2" pageNumber="167">Acknowledgments</emphasis>
. The BIOMAGLO was conducted jointly by French National Museum of Natural History (MNHN) as part of the Tropical DeepSea Benthos program, the French Research Institute for Exploitation of the Sea (IFREMER), the “
<collectingCountry id="F37CACA3FD4AFFE1FC3FFAE5C572FADC" box="[1004,1368,1288,1308]" name="French Southern Territories" pageId="2" pageNumber="167">Terres Australes et Antarctiques Françaises</collectingCountry>
” (
<collectingCountry id="F37CACA3FD4AFFE1FAB8FAE4C58CFADC" box="[1387,1446,1288,1307]" name="French Southern Territories" pageId="2" pageNumber="167">TAAF</collectingCountry>
), the Departmental Council of
<collectingCountry id="F37CACA3FD4AFFE1FBF7FACDC441FAF3" box="[1060,1131,1313,1332]" name="Mayotte" pageId="2" pageNumber="167">Mayotte</collectingCountry>
and the French Development Agency (AFD), with the financial support of the European Union (Xe FED). The expedition was operated under permits delivered by the prefecture of
<collectingCountry id="F37CACA3FD4AFFE1FAA8FABFC3A9FAB9" name="Reunion Island" pageId="2" pageNumber="167">Reunion Island</collectingCountry>
(5000044/CZM
<collectingCountry id="F37CACA3FD4AFFE1FBC5FA87C453FAB9" box="[1046,1145,1387,1406]" name="Réunion" pageId="2" pageNumber="167">REUNION</collectingCountry>
/AEM) and
<collectingCountry id="F37CACA3FD4AFFE1FB0CFA80C53CFAB8" box="[1247,1302,1388,1407]" name="French Southern Territories" pageId="2" pageNumber="167">TAAF</collectingCountry>
(arrêté 2017-01). We are grateful to the crew of RV
<emphasis id="B91F3021FD4AFFE1FB8DFA68C4BBFA50" box="[1118,1169,1412,1431]" italics="true" pageId="2" pageNumber="167">Antéa</emphasis>
, the PIs of BIOMAGLO (Laure Corbari; Karine Olu, and Sarah Samadi), crews of the RV
<emphasis id="B91F3021FD4AFFE1FAC2FA71C5BAFA77" box="[1297,1424,1437,1456]" italics="true" pageId="2" pageNumber="167">Austral Leader</emphasis>
for organizing the surveys and the capture of the samples. We are most grateful to the following persons and institutions for specimen loans and assistance: A. Graham (CSIRO); M.-Y. Lee, H.-S. Lin, and J.-N. Chen (NTUM).
</paragraph>
</subSubSection>
</treatment>
</document>