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<document id="BBEE9E3EE3CA46B71E23259D020A9575" ID-CLB-Dataset="8613" ID-DOI="10.5281/zenodo.4468137" ID-GBIF-Dataset="8db060a7-8799-4203-b2d3-10af003456f1" ID-ISSN="ISSN 0374-1036" ID-Zenodo-Dep="4468137" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611663721085" checkinUser="felipe" docAuthor="Bulirsch, Petr" docDate="2009" docId="03BB87F7143AFFD57C01CDE2FE29FD10" docLanguage="en" docName="ActaEntMusNatPra.49.2.559-576.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (suppl.) 49 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Dyschiriodes (Paradyschirius) sabahensis Bulirsch 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="562" masterDocId="FF82FF8F1438FFD67D6ACD3AFFE4FFE4" masterDocTitle="Contribution to the Asian and Afrotropical species of the genus Dyschiriodes (Coleoptera: Carabidae: Scaritinae)" masterLastPageNumber="576" masterPageNumber="559" pageNumber="561" updateTime="1698925426155" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1A70741C61E110452CFBECED7BC799DD">Contribution to the Asian and Afrotropical species of the genus Dyschiriodes (Coleoptera: Carabidae: Scaritinae)</mods:title>
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<mods:name id="F2A0E0F445BA1AF063F579B1C97ED032" type="personal">
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<mods:namePart id="322EA58628935650B9AFF2684E343957">Bulirsch, Petr</mods:namePart>
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<mods:title id="CC5C03620F93D1D9C509729319B85ED0">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="87F2B5C2F644A273B036665A5152C71D">
<mods:date id="6DFAA7C530CCBCDEFCCF7C0DBB99F959">2009</mods:date>
<mods:detail id="B8D6C4F952E3FC1D8599ED7F72EA8052" type="series">
<mods:title id="27A67AF660BFE800A23F30C2A9C7EF5C">suppl.</mods:title>
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<mods:number id="99CA47730ED7963A7419A708842A445B">2009-12-15</mods:number>
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<mods:number id="64FAE7221CE6EA82982AA40AD646AA82">49</mods:number>
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<mods:identifier id="5DE5E3436DB7720C6CE09C5EA1A84516" type="DOI">10.5281/zenodo.4468137</mods:identifier>
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<treatment id="03BB87F7143AFFD57C01CDE2FE29FD10" ID-DOI="http://doi.org/10.5281/zenodo.4477029" ID-GBIF-Taxon="177136671" ID-Zenodo-Dep="4477029" LSID="urn:lsid:plazi:treatment:03BB87F7143AFFD57C01CDE2FE29FD10" httpUri="http://treatment.plazi.org/id/03BB87F7143AFFD57C01CDE2FE29FD10" lastPageId="3" lastPageNumber="562" pageId="2" pageNumber="561">
<subSubSection id="C308656A143AFFD47C01CDE2FC77FF15" box="[363,915,216,241]" pageId="2" pageNumber="561" type="nomenclature">
<paragraph id="8BAD36E1143AFFD47C01CDE2FC77FF15" blockId="2.[363,915,216,241]" box="[363,915,216,241]" pageId="2" pageNumber="561">
<heading id="D0E5818D143AFFD47C01CDE2FC77FF15" bold="true" box="[363,915,216,241]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="561" reason="3">
<emphasis id="B966EAF3143AFFD47C01CDE2FC77FF15" bold="true" box="[363,915,216,241]" pageId="2" pageNumber="561">
<taxonomicName id="4C124D62143AFFD47C01CDE2FCD2FF15" ID-CoL="8MDCH" authority="Bulirsch, 2009" authorityName="Bulirsch" authorityYear="2009" box="[363,822,216,241]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="sabahensis" status="sp. nov." subGenus="Paradyschirius">
<emphasis id="B966EAF3143AFFD47C01CDE2FE1DFF15" bold="true" box="[363,505,216,241]" italics="true" pageId="2" pageNumber="561">Dyschiriodes</emphasis>
(
<emphasis id="B966EAF3143AFFD47F63CDE2FD4BFF15" bold="true" box="[521,687,216,241]" italics="true" pageId="2" pageNumber="561">Paradyschirius</emphasis>
)
<emphasis id="B966EAF3143AFFD47FD4CDE2FCD2FF15" bold="true" box="[702,822,216,241]" italics="true" pageId="2" pageNumber="561">sabahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2555788143AFFD47E57CDE3FC77FF15" box="[829,915,217,241]" pageId="2" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C308656A143AFFD47F2ACDCDFD5AFEEF" box="[576,702,247,267]" pageId="2" pageNumber="561" type="description">
<paragraph id="8BAD36E1143AFFD47F2ACDCDFD5AFEEF" blockId="2.[151,1128,247,408]" box="[576,702,247,267]" pageId="2" pageNumber="561">
(
<figureCitation id="13292A64143AFFD47F2CCDCDFD66FEEF" box="[582,642,247,267]" captionStart="Figs" captionStartId="4.[151,190,1532,1551]" captionTargetBox="[151,1116,222,1425]" captionTargetId="figure-10@4.[151,1116,219,1425]" captionTargetPageId="4" captionText="Figs. 15. Habitus of holotypes. 1 Dyschiriodes sabahensis sp. nov. (3.15 mm); 2 D. hajeki sp. nov. (3.75 mm); 3 D. ruthmuellerae sp. nov. (3.70 mm); 4 D. kadleci sp. nov. (3.45 mm); 5 D. facchinii sp. nov. (3.70 mm)." figureDoi="http://doi.org/10.5281/zenodo.4468139" httpUri="https://zenodo.org/record/4468139/files/figure.png" pageId="2" pageNumber="561">Figs. 1</figureCitation>
,
<figureCitation id="13292A64143AFFD47FE6CDC2FD7CFEEF" box="[652,664,248,267]" captionStart="Figs" captionStartId="7.[113,152,1452,1471]" captionTargetBox="[144,1076,229,1414]" captionTargetId="figure-13@7.[113,1088,219,1420]" captionTargetPageId="7" captionText="Figs. 813. Aedeagus, right lateral view. 8 Dyschiriodes sabahensis sp. nov.; 9 D. ruthmuellerae sp. nov.; 10 D. kadleci sp. nov.; 11 D. facchinii sp. nov.; 12 D. jelineki sp. nov.; 13 D. genieri sp. nov. (all holotypes)." figureDoi="http://doi.org/10.5281/zenodo.4468143" httpUri="https://zenodo.org/record/4468143/files/figure.png" pageId="2" pageNumber="561">8</figureCitation>
,
<figureCitation id="13292A64143AFFD47FCBCDC2FD53FEEF" box="[673,695,248,267]" captionStart="Figs" captionStartId="8.[151,190,1087,1106]" captionTargetBox="[164,1110,218,1058]" captionTargetId="figure-152@8.[151,1123,211,1058]" captionTargetPageId="8" captionText="Figs. 1418.Apex of aedeagus, ventral view.14 Dyschiriodes sabahensis sp. nov.; 15 D. ruthmuellerae sp. nov.; 16 D. kadleci sp. nov.; 17 D. facchinii sp. nov.; 18 D. genieri sp. nov. (all holotypes)." figureDoi="http://doi.org/10.5281/zenodo.4468145" httpUri="https://zenodo.org/record/4468145/files/figure.png" pageId="2" pageNumber="561">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C308656A143AFFD47DFDCC20FE95FE7C" pageId="2" pageNumber="561" type="materials_examined">
<paragraph id="8BAD36E1143AFFD47DFDCC20FDBDFEC9" blockId="2.[151,1128,247,408]" box="[151,601,281,301]" pageId="2" pageNumber="561">
<emphasis id="B966EAF3143AFFD47DFDCC20FEF4FEC9" bold="true" box="[151,272,281,301]" pageId="2" pageNumber="561">
<typeStatus id="54A98843143AFFD47DFDCC20FF20FEC9" box="[151,196,282,301]" pageId="2" pageNumber="561">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3057671143AFFD47C7FCC20FE81FEC9" box="[277,357,282,301]" name="Malaysia" pageId="2" pageNumber="561">Malaysia</collectingCountry>
,
<collectingRegion id="49D6F803143AFFD47C04CC20FE40FEC9" box="[366,420,282,301]" country="Malaysia" name="Sabah" pageId="2" pageNumber="561">Sabah</collectingRegion>
,
<quantity id="4CEA9B04143AFFD47CC7CC20FE06FEC9" box="[429,482,282,301]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="561" unit="km" value="25.0">25 km</quantity>
E of Telupid.
</paragraph>
<paragraph id="8BAD36E1143AFFD47DFDCC0FFE95FE7C" blockId="2.[151,1128,247,408]" pageId="2" pageNumber="561">
<emphasis id="B966EAF3143AFFD47DFDCC0FFEFFFEAC" bold="true" box="[151,283,308,328]" pageId="2" pageNumber="561">Type material.</emphasis>
<materialsCitation id="3B7A3CBC143AFFD47C4ACC0FFE57FE86" ID-GBIF-Occurrence="3026664303" collectingDate="1983-08-17" collectionCode="USNM" collectorName="G. F. Hevel &amp; W. E. Steiner" location="Telupid" pageId="2" pageNumber="561" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="54A98843143AFFD47C4ACC0FFE90FEA2" box="[288,372,309,328]" pageId="2" pageNumber="561" type="holotype">HOLOTYPE</typeStatus>
: ♁, MALAYSIA:
<collectingRegion id="49D6F803143AFFD47F73CC0FFD81FEAC" box="[537,613,309,328]" country="Malaysia" name="Sabah" pageId="2" pageNumber="561">SABAH</collectingRegion>
: /
<quantity id="4CEA9B04143AFFD47F1DCC0EFD49FEA3" box="[631,685,308,327]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="561" unit="km" value="25.0">25 km</quantity>
E
<location id="8ECD603A143AFFD47FAECC0EFCE1FEA3" LSID="urn:lsid:plazi:treatment:03BB87F7143AFFD57C01CDE2FE29FD10:8ECD603A143AFFD47FAECC0EFCE1FEA3" box="[708,773,308,327]" name="Telupid" pageId="2" pageNumber="561" stateProvince="Sabah">Telupid</location>
/
<date id="FFAC1021143AFFD47E7ECC0EFC9FFEA3" box="[788,891,308,327]" pageId="2" pageNumber="561" value="1983-08-17">
<collectingDate id="EFE8E9C9143AFFD47E7ECC0EFC9FFEA3" box="[788,891,308,327]" pageId="2" pageNumber="561" value="1983-08-17">17.viii.1983</collectingDate>
</date>
/ at blacklight /
<collectorName id="26E75337143AFFD4796FCC0FFB83FEA3" box="[1029,1127,308,328]" pageId="2" pageNumber="561">G. F. Hevel</collectorName>
&amp;
<collectorName id="26E75337143AFFD47DC4CC6AFEC6FE86" box="[174,290,335,355]" pageId="2" pageNumber="561">W. E. Steiner</collectorName>
(leg.) (
<collectionCode id="ED03AE24143AFFD47C00CC75FE48FE86" box="[362,428,335,354]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="561" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7A3CBC143AFFD47CD6CC6AFF31FE99" ID-GBIF-Occurrence="3026664304" collectingDate="1983-08-17" collectionCode="USNM, PBCP" collectorName="G. F. Hevel &amp; W. E. Steiner" country="Malaysia" location="Telupid" pageId="2" pageNumber="561" specimenCount="15" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="54A98843143AFFD47CD6CC6AFDF0FE85" box="[444,532,336,355]" pageId="2" pageNumber="561" type="paratype">PARATYPES</typeStatus>
:
<emphasis id="B966EAF3143AFFD47F4BCC74FD3BFE86" bold="true" box="[545,735,334,354]" pageId="2" pageNumber="561">
<collectingCountry id="F3057671143AFFD47F4BCC74FD73FE86" box="[545,663,334,354]" name="Malaysia" pageId="2" pageNumber="561">MALAYSIA</collectingCountry>
:
<collectingRegion id="49D6F803143AFFD47FCBCC74FD3CFE85" box="[673,728,334,354]" country="Malaysia" name="Sabah" pageId="2" pageNumber="561">SABAH</collectingRegion>
:
</emphasis>
<specimenCount id="9D14FD68143AFFD47F8CCC75FCCBFE87" box="[742,815,335,355]" pageId="2" pageNumber="561" type="generic">15 spec.</specimenCount>
with the same data as HT (
<collectionCode id="ED03AE24143AFFD47948CC75FB87FE86" box="[1058,1123,335,354]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="561" type="Museum">USNM</collectionCode>
,
<collectionCode id="ED03AE24143AFFD47DFDCC50FF2BFE99" box="[151,207,362,381]" pageId="2" pageNumber="561">PBCP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7A3CBC143AFFD47D8ACC50FE89FE7C" ID-GBIF-Occurrence="3026664309" collectingDate="1983-08-14" collectionCode="USNM, PBPC" collectorName="G. F. Hevel &amp; W. E. Steiner'" country="Malaysia" location="Telupid" pageId="2" pageNumber="561" specimenCount="2" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="9D14FD68143AFFD47D8ACC50FEC4FE99" box="[224,288,362,381]" pageId="2" pageNumber="561" type="generic">2 spec.</specimenCount>
, MALAYSIA:
<collectingRegion id="49D6F803143AFFD47CC6CC50FE1CFE99" box="[428,504,362,381]" country="Malaysia" name="Sabah" pageId="2" pageNumber="561">SABAH</collectingRegion>
: /
<quantity id="4CEA9B04143AFFD47F64CC50FDA2FE99" box="[526,582,362,381]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="561" unit="km" value="20.0">20 km</quantity>
E
<location id="8ECD603A143AFFD47F0BCC50FD46FE99" LSID="urn:lsid:plazi:treatment:03BB87F7143AFFD57C01CDE2FE29FD10:8ECD603A143AFFD47F0BCC50FD46FE99" box="[609,674,362,381]" country="Malaysia" name="Telupid" pageId="2" pageNumber="561" stateProvince="Sabah">Telupid</location>
/
<date id="FFAC1021143AFFD47FDCCC50FCF9FE99" box="[694,797,362,381]" pageId="2" pageNumber="561" value="1983-08-14">
<collectingDate id="EFE8E9C9143AFFD47FDCCC50FCF9FE99" box="[694,797,362,381]" pageId="2" pageNumber="561" value="1983-08-14">14.viii.1983</collectingDate>
</date>
/ at blacklight /
<collectorName id="26E75337143AFFD47EDBCC50FBF3FE99" box="[945,1047,362,381]" pageId="2" pageNumber="561">G. F. Hevel</collectorName>
&amp;
<collectorName id="26E75337143AFFD4795FCC50FF3EFE73" pageId="2" pageNumber="561">W. E. Steiner</collectorName>
(
<collectionCode id="ED03AE24143AFFD47D8FCCBFFEC2FE7C" box="[229,294,389,408]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="561" type="Museum">USNM</collectionCode>
,
<collectionCode id="ED03AE24143AFFD47C45CCBFFE83FE7C" box="[303,359,389,408]" pageId="2" pageNumber="561">PBPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C308656A143AFFD47DFDCC8AFD2DF9E3" pageId="2" pageNumber="561" type="description">
<paragraph id="8BAD36E1143AFFD47DFDCC8AFD2DFDCC" blockId="2.[151,1127,432,1544]" pageId="2" pageNumber="561">
<emphasis id="B966EAF3143AFFD47DFDCC8AFEC4FE2C" bold="true" box="[151,288,432,456]" pageId="2" pageNumber="561">Description.</emphasis>
Habitus as in
<figureCitation id="13292A64143AFFD47CC5CC8AFE08FE2C" box="[431,492,432,456]" captionStart="Figs" captionStartId="4.[151,190,1532,1551]" captionTargetBox="[151,1116,222,1425]" captionTargetId="figure-10@4.[151,1116,219,1425]" captionTargetPageId="4" captionText="Figs. 15. Habitus of holotypes. 1 Dyschiriodes sabahensis sp. nov. (3.15 mm); 2 D. hajeki sp. nov. (3.75 mm); 3 D. ruthmuellerae sp. nov. (3.70 mm); 4 D. kadleci sp. nov. (3.45 mm); 5 D. facchinii sp. nov. (3.70 mm)." figureDoi="http://doi.org/10.5281/zenodo.4468139" httpUri="https://zenodo.org/record/4468139/files/figure.png" pageId="2" pageNumber="561">Fig. 1</figureCitation>
; length
<quantity id="4CEA9B04143AFFD47F57CC8AFD30FE23" box="[573,724,432,456]" metricMagnitude="-3" metricUnit="m" metricValue="3.125" metricValueMax="3.35" metricValueMin="2.9" pageId="2" pageNumber="561" unit="mm" value="3.125" valueMax="3.35" valueMin="2.9">2.903.35 mm</quantity>
(mean
<quantity id="4CEA9B04143AFFD47E4ACC8AFC9BFE23" box="[800,895,432,456]" metricMagnitude="-3" metricUnit="m" metricValue="3.17" pageId="2" pageNumber="561" unit="mm" value="3.17">3.17 mm</quantity>
, n = 18; HT
<quantity id="4CEA9B04143AFFD47E91CC8AFBB8FE2C" box="[1019,1116,432,456]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" pageId="2" pageNumber="561" unit="mm" value="3.15">3.15 mm</quantity>
). Colour brown fuliginous, surface with slight green-bronze metallic lustre, anterior parts of head and base and apical part of elytra slightly lighter, brownish translucent, legs rusty red, antennae and mouth-parts slightly lighter, yellowish.
</paragraph>
<paragraph id="8BAD36E1143AFFD47DDFCF0AFD47FD0C" blockId="2.[151,1127,432,1544]" pageId="2" pageNumber="561">Head. Anterior margin of clypeus with distinctly protruding lateral lobes, between them slightly convex; clypeofrontal area with characteristic T-forming carina and with additional fine, irregular carinae/rugosities between longitudinal carina and facial furrows; facial furrows deep, regularly diverged posteriorly; distance between them larger than length of eye. Surface of vertex even, smooth, with very fine and sparse micropunctures. Eyes moderately large, strongly convex. Antennae submoniliform.</paragraph>
<paragraph id="8BAD36E1143AFFD47DDFCFCAFC3CFC2C" blockId="2.[151,1127,432,1544]" pageId="2" pageNumber="561">Pronotum. Strongly convex; outline regularly and moderately strongly rounded, moderately attenuated anteriorly; 0.971.04 (mean 1.00; HT 0.99) times as wide as long, 1.361.43 (mean 1.41; HT 1.40) times as wide as head; widest in anterior two thirds. Anterior angles blunt, posterior ones moderately rounded. Anterior transverse impression distinct, with sparse cross striae; median line moderately impressed at sides and much finer, superficial on disc; lateral channel moderately broad, reflexed lateral margin extended slightly beyond posterior setiferous puncture. Surface shiny, mirror-like, with very fine micropunctures.</paragraph>
<paragraph id="8BAD36E1143AFFD47DDFCEEAFDF7FAAC" blockId="2.[151,1127,432,1544]" pageId="2" pageNumber="561">Elytra. Elongate, very slightly and broadly concave in anterior fifth in lateral view; 1.721.83 (mean 1.78; HT 1.78) times as long as wide, 1.101.17 (mean 1.14; HT 1.13) times as wide as pronotum; humeri moderately protruded, without humeral teeth; elytral base strongly sloping; outline in basal half slightly broadened on sides, broadest at anterior third, more strongly attenuated towards apex than towards humeri; suture not depressed at base. Base without basal border and tubercles; BSP deeply connected with first stria. Striae characteristic of subgroup: striae 17 deep, moderately punctate, punctures disappearing in third fourth, distinctly narrower than width of intervals; stria 8 very fine, recognisable only in middle third, striae 23 and 67 diminish basally, stria 3 disappearing before anterior DSP; striae 45 distinctly deeper and broader behind base; all striae moderately deep apically; intervals slightly vaulted. Three PHSP (only two in two PTs), three DSP (all in stria 3) and two ASP (both in deep apical stria).</paragraph>
<paragraph id="8BAD36E1143AFFD47DDFC86AFEF9FA4C" blockId="2.[151,1127,432,1544]" pageId="2" pageNumber="561">Protibia.Apical spine broad and blunt, strongly curved backwards but not inwards, slightly shorter than uncinate apical spur; distal marginal tooth large, moderately sharp, proximal one small, blunt.</paragraph>
<paragraph id="8BAD36E1143AFFD47DDFC88AFD2DF9E3" blockId="2.[151,1127,432,1544]" pageId="2" pageNumber="561">
Aedeagus. Shape as in
<figureCitation id="13292A64143AFFD47CC0C88AFE16FA2C" box="[426,498,1456,1480]" captionStart="Figs" captionStartId="7.[113,152,1452,1471]" captionTargetBox="[144,1076,229,1414]" captionTargetId="figure-13@7.[113,1088,219,1420]" captionTargetPageId="7" captionText="Figs. 813. Aedeagus, right lateral view. 8 Dyschiriodes sabahensis sp. nov.; 9 D. ruthmuellerae sp. nov.; 10 D. kadleci sp. nov.; 11 D. facchinii sp. nov.; 12 D. jelineki sp. nov.; 13 D. genieri sp. nov. (all holotypes)." figureDoi="http://doi.org/10.5281/zenodo.4468143" httpUri="https://zenodo.org/record/4468143/files/figure.png" pageId="2" pageNumber="561">Figs. 8</figureCitation>
and
<figureCitation id="13292A64143AFFD47F4FC88AFDA6FA2C" box="[549,578,1456,1480]" captionStart="Figs" captionStartId="8.[151,190,1087,1106]" captionTargetBox="[164,1110,218,1058]" captionTargetId="figure-152@8.[151,1123,211,1058]" captionTargetPageId="8" captionText="Figs. 1418.Apex of aedeagus, ventral view.14 Dyschiriodes sabahensis sp. nov.; 15 D. ruthmuellerae sp. nov.; 16 D. kadleci sp. nov.; 17 D. facchinii sp. nov.; 18 D. genieri sp. nov. (all holotypes)." figureDoi="http://doi.org/10.5281/zenodo.4468145" httpUri="https://zenodo.org/record/4468145/files/figure.png" pageId="2" pageNumber="561">14</figureCitation>
; in HT
<quantity id="4CEA9B04143AFFD47FFBC88AFD0BFA23" box="[657,751,1456,1480]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="2" pageNumber="561" unit="mm" value="0.59">0.59 mm</quantity>
long, median lobe moderately bent down, indistinctly bulging in apical third. Apical lamella medium sized, asymmetric rounded, narrowed apically (
<figureCitation id="13292A64143AFFD47C0CC8CAFE51F9EC" box="[358,437,1520,1544]" captionStart="Figs" captionStartId="8.[151,190,1087,1106]" captionTargetBox="[164,1110,218,1058]" captionTargetId="figure-152@8.[151,1123,211,1058]" captionTargetPageId="8" captionText="Figs. 1418.Apex of aedeagus, ventral view.14 Dyschiriodes sabahensis sp. nov.; 15 D. ruthmuellerae sp. nov.; 16 D. kadleci sp. nov.; 17 D. facchinii sp. nov.; 18 D. genieri sp. nov. (all holotypes)." figureDoi="http://doi.org/10.5281/zenodo.4468145" httpUri="https://zenodo.org/record/4468145/files/figure.png" pageId="2" pageNumber="561">Fig. 14</figureCitation>
). Paramere without setae.
</paragraph>
</subSubSection>
<subSubSection id="C308656A143BFFD57D1BCDE1FC17FD50" pageId="3" pageNumber="562" type="diagnosis">
<paragraph id="8BAD36E1143BFFD57D1BCDE1FC17FD50" blockId="3.[113,1089,219,756]" pageId="3" pageNumber="562">
<emphasis id="B966EAF3143BFFD57D1BCDE1FE82FF17" bold="true" box="[113,358,219,243]" pageId="3" pageNumber="562">Differential diagnosis.</emphasis>
<taxonomicName id="4C124D62143BFFD57C00CDE7FD89FF10" authority="Bulirsch, 2009" authorityName="Bulirsch" authorityYear="2009" box="[362,621,221,244]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="sabahensis" status="sp. nov.">
<emphasis id="B966EAF3143BFFD57C00CDE7FD89FF10" box="[362,621,221,244]" italics="true" pageId="3" pageNumber="562">Dyschiriodes sabahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2555788143BFFD57F18CDE6FD25FF17" box="[626,705,220,243]" pageId="3" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the
<taxonomicName id="4C124D62143BFFD57E30CDE7FC3EFF10" baseAuthorityName="Putzeys" baseAuthorityYear="1878" box="[858,986,221,244]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B966EAF3143BFFD57E30CDE7FC3EFF10" box="[858,986,221,244]" italics="true" pageId="3" pageNumber="562">D. verticalis</emphasis>
</taxonomicName>
subgroup of the
<taxonomicName id="4C124D62143BFFD57DDECDC7FEADFEF0" baseAuthorityName="Duftschmid" baseAuthorityYear="1812" box="[180,329,253,276]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="substriatus">
<emphasis id="B966EAF3143BFFD57DDECDC7FEADFEF0" box="[180,329,253,276]" italics="true" pageId="3" pageNumber="562">D. substriatus</emphasis>
</taxonomicName>
group (sensu
<bibRefCitation id="EF834B10143BFFD57CB7CDC6FD75FEF7" author="FEDORENKO D. N." box="[477,657,251,275]" pageId="3" pageNumber="562" refId="ref10270" refString="FEDORENKO D. N. 1996: Reclassification of world Dyschiriini, with a revision of the Palearctic fauna (Coleoptera, Carabidae). Pensoft Publishers, Sofia-Moscow-St. Petersburg, 224 pp." type="book" year="1996">FEDORENKO 1996</bibRefCitation>
,
<bibRefCitation id="EF834B10143BFFD57FF1CDC6FD05FEF0" author="FEDORENKO D. N." box="[667,737,252,276]" pageId="3" pageNumber="562" pagination="135 - 152" refId="ref10305" refString="FEDORENKO D. N. 1997 a: (Revision of Palearctic and Paleotropical species of the substriatus group, genus Dyschiodes, Jeannel, 1941 (Coleoptera, Carabidae )). Entomologicheskoe Obozrenie 76: 135 - 152 (in Russian, English summary)." type="journal article" year="1997">1997a</bibRefCitation>
). It can be distinguished from the most similar
<taxonomicName id="4C124D62143BFFD57D93CC27FD04FED0" authority="(Putzeys, 1878)" baseAuthorityName="Putzeys" baseAuthorityYear="1878" box="[249,736,284,308]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="verticalis" subGenus="Paradyschirius">
<emphasis id="B966EAF3143BFFD57D93CC27FEF0FED0" box="[249,276,285,308]" italics="true" pageId="3" pageNumber="562">D.</emphasis>
(
<emphasis id="B966EAF3143BFFD57C4BCC27FE20FED0" box="[289,452,285,308]" italics="true" pageId="3" pageNumber="562">Paradyschirius</emphasis>
)
<emphasis id="B966EAF3143BFFD57CB8CC27FDD1FED0" box="[466,565,285,308]" italics="true" pageId="3" pageNumber="562">verticalis</emphasis>
(Putzeys, 1878)
</taxonomicName>
by a more rounded and anteriorly less attenuated pronotum, different number of setiferous punctures on the elytra (two ASP and 23 PHSP in
<taxonomicName id="4C124D62143BFFD57C44CC67FE20FE90" authority="Bulirsch, 2009" authorityName="Bulirsch" authorityYear="2009" box="[302,452,349,372]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="sabahensis" status="sp. nov.">
<emphasis id="B966EAF3143BFFD57C44CC67FE20FE90" box="[302,452,349,372]" italics="true" pageId="3" pageNumber="562">D. sabahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2555788143BFFD57CA0CC66FDFBFE97" box="[458,543,348,371]" pageId="3" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
, one ASP and 12 PHSP in
<taxonomicName id="4C124D62143BFFD57E22CC67FC28FE90" baseAuthorityName="Putzeys" baseAuthorityYear="1878" box="[840,972,349,372]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B966EAF3143BFFD57E22CC67FC28FE90" box="[840,972,349,372]" italics="true" pageId="3" pageNumber="562">D. verticalis</emphasis>
</taxonomicName>
), finer and distinctly punctate elytral striae, especially more distinctly punctate stria 7, and anteriorly more abbreviated stria 3. From
<taxonomicName id="4C124D62143BFFD57CEDCCA7FCE5FE50" authority="(Andrewes, 1929)" baseAuthorityName="Andrewes" baseAuthorityYear="1929" box="[391,769,412,436]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Paradyschirius">
<emphasis id="B966EAF3143BFFD57CEDCCA7FE45FE50" box="[391,417,413,436]" italics="true" pageId="3" pageNumber="562">D.</emphasis>
(
<emphasis id="B966EAF3143BFFD57CDBCCA7FE22FE50" box="[433,454,413,436]" italics="true" pageId="3" pageNumber="562">P.</emphasis>
)
<emphasis id="B966EAF3143BFFD57CBFCCA7FDDCFE50" box="[469,568,413,436]" italics="true" pageId="3" pageNumber="562">hingstoni</emphasis>
(
<bibRefCitation id="EF834B10143BFFD57F2DCCA6FD1EFE50" author="ANDREWES H. E." box="[583,762,412,436]" pageId="3" pageNumber="562" refId="ref9878" refString="ANDREWES H. E. 1929: The Fauna of British India, including Ceylon and Burma. Coleoptera. Carabidae. Vol. 1 - Carabinae. Taylor &amp; Francis, London, xviii + 431 pp., 10 pls. + 1 map." type="book" year="1929">Andrewes, 1929</bibRefCitation>
)
</taxonomicName>
it differs mainly by a smaller body (
<quantity id="4CEA9B04143BFFD57DD9CC81FEA9FE37" box="[179,333,443,467]" metricMagnitude="-3" metricUnit="m" metricValue="3.125" metricValueMax="3.35" metricValueMin="2.9" pageId="3" pageNumber="562" unit="mm" value="3.125" valueMax="3.35" valueMin="2.9">2.903.35 mm</quantity>
in
<taxonomicName id="4C124D62143BFFD57C06CC87FDE4FE30" authority="Bulirsch, 2009" authorityName="Bulirsch" authorityYear="2009" box="[364,512,445,468]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="sabahensis" status="sp. nov.">
<emphasis id="B966EAF3143BFFD57C06CC87FDE4FE30" box="[364,512,445,468]" italics="true" pageId="3" pageNumber="562">D. sabahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2555788143BFFD57F6ECC86FDB3FE37" box="[516,599,444,467]" pageId="3" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
,
<quantity id="4CEA9B04143BFFD57F0ACC81FD1CFE37" box="[608,760,443,467]" metricMagnitude="-3" metricUnit="m" metricValue="3.875" metricValueMax="4.15" metricValueMin="3.6" pageId="3" pageNumber="562" unit="mm" value="3.875" valueMax="4.15" valueMin="3.6">3.604.15 mm</quantity>
in
<taxonomicName id="4C124D62143BFFD57E7DCC87FC7DFE30" baseAuthorityName="Andrewes" baseAuthorityYear="1929" box="[791,921,445,468]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="hingstoni">
<emphasis id="B966EAF3143BFFD57E7DCC87FC7DFE30" box="[791,921,445,468]" italics="true" pageId="3" pageNumber="562">D. hingstoni</emphasis>
</taxonomicName>
), more rounded and anteriorly less attenuated pronotum and different number of setiferous punctures on the elytra (one ASP and two PHSP in
<taxonomicName id="4C124D62143BFFD57C8DCCC7FD88FDF0" baseAuthorityName="Andrewes" baseAuthorityYear="1929" box="[487,620,509,532]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="hingstoni">
<emphasis id="B966EAF3143BFFD57C8DCCC7FD88FDF0" box="[487,620,509,532]" italics="true" pageId="3" pageNumber="562">D. hingstoni</emphasis>
</taxonomicName>
). Finally, it differs from
<taxonomicName id="4C124D62143BFFD57E17CCC7FED1FDD0" authority="(Andrewes, 1929)" baseAuthorityName="Andrewes" baseAuthorityYear="1929" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="tenuescens" subGenus="Paradyschirius">
<emphasis id="B966EAF3143BFFD57E17CCC7FC7CFDF0" box="[893,920,509,532]" italics="true" pageId="3" pageNumber="562">D.</emphasis>
(
<emphasis id="B966EAF3143BFFD57EC2CCC7FC59FDF0" box="[936,957,509,532]" italics="true" pageId="3" pageNumber="562">P.</emphasis>
)
<emphasis id="B966EAF3143BFFD57EA7CCC7FBA5FDF0" box="[973,1089,509,532]" italics="true" pageId="3" pageNumber="562">tenuescens</emphasis>
(
<bibRefCitation id="EF834B10143BFFD57D13CF26FECAFDD0" author="ANDREWES H. E." box="[121,302,540,564]" pageId="3" pageNumber="562" refId="ref9878" refString="ANDREWES H. E. 1929: The Fauna of British India, including Ceylon and Burma. Coleoptera. Carabidae. Vol. 1 - Carabinae. Taylor &amp; Francis, London, xviii + 431 pp., 10 pls. + 1 map." type="book" year="1929">Andrewes, 1929</bibRefCitation>
)
</taxonomicName>
by larger body (
<quantity id="4CEA9B04143BFFD57C9BCF26FD74FDD7" box="[497,656,540,564]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.7" metricValueMin="2.1" pageId="3" pageNumber="562" unit="mm" value="2.4" valueMax="2.7" valueMin="2.1">2.102.70 mm</quantity>
in
<taxonomicName id="4C124D62143BFFD57FD3CF27FCB5FDD0" baseAuthorityName="Andrewes" baseAuthorityYear="1929" box="[697,849,541,564]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="tenuescens">
<emphasis id="B966EAF3143BFFD57FD3CF27FCB5FDD0" box="[697,849,541,564]" italics="true" pageId="3" pageNumber="562">D. tenuescens</emphasis>
</taxonomicName>
), different number of setiferous punctures on the elytra (one ASP and one PHSP in
<taxonomicName id="4C124D62143BFFD57E60CF07FC44FDB0" baseAuthorityName="Andrewes" baseAuthorityYear="1929" box="[778,928,573,596]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="tenuescens">
<emphasis id="B966EAF3143BFFD57E60CF07FC44FDB0" box="[778,928,573,596]" italics="true" pageId="3" pageNumber="562">D. tenuescens</emphasis>
</taxonomicName>
), basally more abbreviated elytral stria 3 and distinctly punctate stria 7. Moreover,
<taxonomicName id="4C124D62143BFFD57E25CF67FC02FD90" authority="Bulirsch, 2009" authorityName="Bulirsch" authorityYear="2009" box="[847,998,605,628]" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="sabahensis" status="sp. nov.">
<emphasis id="B966EAF3143BFFD57E25CF67FC02FD90" box="[847,998,605,628]" italics="true" pageId="3" pageNumber="562">D. sabahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2555788143BFFD57E87CF66FBA5FD97" box="[1005,1089,604,627]" pageId="3" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from all these species by the shape of the median lobe of the aedeagus and its apical lamella (see
<figureCitation id="13292A64143BFFD57CE1CFA6FE30FD50" box="[395,468,668,692]" captionStart="Figs" captionStartId="7.[113,152,1452,1471]" captionTargetBox="[144,1076,229,1414]" captionTargetId="figure-13@7.[113,1088,219,1420]" captionTargetPageId="7" captionText="Figs. 813. Aedeagus, right lateral view. 8 Dyschiriodes sabahensis sp. nov.; 9 D. ruthmuellerae sp. nov.; 10 D. kadleci sp. nov.; 11 D. facchinii sp. nov.; 12 D. jelineki sp. nov.; 13 D. genieri sp. nov. (all holotypes)." figureDoi="http://doi.org/10.5281/zenodo.4468143" httpUri="https://zenodo.org/record/4468143/files/figure.png" pageId="3" pageNumber="562">Figs. 8</figureCitation>
and
<figureCitation id="13292A64143BFFD57F62CFA6FDC7FD50" box="[520,547,668,692]" captionStart="Figs" captionStartId="8.[151,190,1087,1106]" captionTargetBox="[164,1110,218,1058]" captionTargetId="figure-152@8.[151,1123,211,1058]" captionTargetPageId="8" captionText="Figs. 1418.Apex of aedeagus, ventral view.14 Dyschiriodes sabahensis sp. nov.; 15 D. ruthmuellerae sp. nov.; 16 D. kadleci sp. nov.; 17 D. facchinii sp. nov.; 18 D. genieri sp. nov. (all holotypes)." figureDoi="http://doi.org/10.5281/zenodo.4468145" httpUri="https://zenodo.org/record/4468145/files/figure.png" pageId="3" pageNumber="562">14</figureCitation>
versus Figs.
<quantity id="4CEA9B04143BFFD57FC5CFA1FCEAFD50" box="[687,782,667,692]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="6.35" metricValueMin="5.334" pageId="3" pageNumber="562" unit="in" value="23.0" valueMax="25.0" valueMin="21.0">2125 in</quantity>
<bibRefCitation id="EF834B10143BFFD57E7ECFA6FC0BFD50" author="FEDORENKO D. N." box="[788,1007,668,692]" pageId="3" pageNumber="562" pagination="135 - 152" refId="ref10305" refString="FEDORENKO D. N. 1997 a: (Revision of Palearctic and Paleotropical species of the substriatus group, genus Dyschiodes, Jeannel, 1941 (Coleoptera, Carabidae )). Entomologicheskoe Obozrenie 76: 135 - 152 (in Russian, English summary)." type="journal article" year="1997">FEDORENKO (1997a))</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C308656A143BFFD57D1BCF86FBC3FD30" box="[113,1063,700,724]" pageId="3" pageNumber="562" type="etymology">
<paragraph id="8BAD36E1143BFFD57D1BCF86FBC3FD30" blockId="3.[113,1089,219,756]" box="[113,1063,700,724]" pageId="3" pageNumber="562">
<emphasis id="B966EAF3143BFFD57D1BCF86FF16FD30" bold="true" box="[113,242,700,724]" pageId="3" pageNumber="562">Etymology.</emphasis>
The species epithet is derived from the state in which the species was found.
</paragraph>
</subSubSection>
<subSubSection id="C308656A143BFFD57D1BCFE1FE29FD10" box="[113,461,731,756]" pageId="3" pageNumber="562" type="distribution">
<paragraph id="8BAD36E1143BFFD57D1BCFE1FE29FD10" blockId="3.[113,1089,219,756]" box="[113,461,731,756]" pageId="3" pageNumber="562">
<emphasis id="B966EAF3143BFFD57D1BCFE1FEE0FD17" bold="true" box="[113,260,731,755]" pageId="3" pageNumber="562">Distribution.</emphasis>
<collectingCountry id="F3057671143BFFD57C60CFE6FE88FD10" box="[266,364,732,756]" name="Malaysia" pageId="3" pageNumber="562">Malaysia</collectingCountry>
(
<collectingRegion id="49D6F803143BFFD57C10CFE6FE25FD10" box="[378,449,732,756]" country="Malaysia" name="Sabah" pageId="3" pageNumber="562">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>