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<mods:title id="3E562C9AE28858D4D5240C550AFC357F">A new genus for the freshwater crab Potamonemus asylos Cumberlidgeı 1993 ı (Brachyura: Potamoidea: Potamonautidae) from Cameroonı Central Africaı with a key to the genera of the Potamonautinae</mods:title>
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<mods:namePart id="CF03FAE311BB2A2E157192F6FE0E180F">Cumberlidge, Neil</mods:namePart>
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(
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(
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(a,b), 4(a), 5(a,b,g), 6(a,b), 7(a,d) and 8(a,d,g))
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species.
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, by present designation.
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<emphasis id="B966EAC41031392842A9F934FE0FFEA1" box="[160,391,278,302]" italics="true" pageId="7" pageNumber="665">Potamonemus asylos</emphasis>
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Cumberlidge, 1993: 576 
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584
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, figs 3, 4, 5a 
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b, 6a 
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c, 8a 
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b; tables 2, 3; Cumberlidge 1999: figs 40C, 41C,F, 42C,F, 43C,F, 44C, 53G, 54 
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, 65F, table IX; 
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; 
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<bibRefCitation id="EF834B27103139284356F979FDE2FEFC" author="Cumberlidge N" box="[351,618,347,371]" pageId="7" pageNumber="665" pagination="71 - 78" refId="ref8358" refString="Cumberlidge N. 2011 a. Chapter 6 The status and distribution of freshwater crabs. In: Brooks EGE, Allen DJ, editors Darwall WRT compilers. The status and distribution of freshwater biodiversity in central Africa. Gland, Switzerland and Cambridge (UK): IUCN. p. 71 - 78." type="book chapter" year="2011">Cumberlidge 2011a: 78</bibRefCitation>
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, 80, 82, 86, table 6.1; 
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; 
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<bibRefCitation id="EF834B271031392842A9F95CFD88FE19" author="Mvogo Ndongo PA &amp; Schubart CD &amp; von Rintelen T &amp; Tamesse JL &amp; Cumberlidge N" box="[160,512,382,406]" pageId="7" pageNumber="665" pagination="161 - 173" refId="ref9554" refString="Mvogo Ndongo PA, Schubart CD, von Rintelen T, Tamesse JL, Cumberlidge N. 2017 b. Morphological and molecular evidence for a new species of freshwater crab of the genus Sudanonautes Bott, 1955 (Brachyura: Potamoidea: Potamonautidae) from Cameroon, with notes on its ecology. Zootaxa. 4242: 161 - 173. doi: 10.11646 / zootaxa. 4242.1.8" type="journal article" year="2017">Mvogo Ndongo et al. 2017b: 3</bibRefCitation>
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, table 1; 
<bibRefCitation id="EF834B27103139284063F95CFCB4FE19" author="Daniels SR &amp; Phiri EE &amp; Klaus S &amp; Albrecht C &amp; Cumberlidge N" box="[618,828,382,406]" pageId="7" pageNumber="665" pagination="549 - 567" refId="ref8897" refString="Daniels SR, Phiri EE, Klaus S, Albrecht C, Cumberlidge N. 2015. Multilocus phylogeny of the Afrotropical freshwater crab fauna reveals historical drainage connectivity and transoceanic dispersal since the Eocene. Syst Biol. 64: 549 - 567. doi: 10.1093 / sysbio / syv 011" type="journal article" year="2015">Daniels et al. 2015</bibRefCitation>
, table 1.
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: 
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(
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adult 
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9.0, FW 
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, between 
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<location id="8ECD600D1031392842A9FA0EFF67FDCB" LSID="urn:lsid:plazi:treatment:03BB87C010313927438DF8AEFC39F930:8ECD600D1031392842A9FA0EFF67FDCB" box="[160,239,556,580]" country="Switzerland" latitude="4.638727" longLatPrecision="1" longitude="9.441354" name="Kumba" pageId="7" pageNumber="665">Kumba</location>
(
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, 
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), 
<collectorName id="26E75300103139284025FA0EFD26FDCB" box="[556,686,556,580]" pageId="7" pageNumber="665">South-West</collectorName>
Region, coll
</materialsCitation>
. 
<collectionCode id="ED03AE1310313928415FFA0EFCE1FDCB" box="[854,873,556,580]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="665">R</collectionCode>
.H.L. 
<materialsCitation id="3B7A3C8B1031392841A8FA0EFDF4FDE8" ID-GBIF-Occurrence="2569344767" collectionCode="NHM, NMU" country="Switzerland" pageId="7" pageNumber="665" specimenCode="NHM 1994.587, NMU 1969" specimenCount="1" typeStatus="holotype">
Disney, 1969 (
<specimenCode id="DBB49EAD103139284640FA0EFE8DFDE8" collectionCode="NHM" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="7" pageNumber="665" type="Herbarium">NHM 1994.587</specimenCode>
donation from 
<specimenCode id="DBB49EAD1031392843B2FA6DFDBFFDE8" box="[443,567,591,615]" collectionCode="NMU" pageId="7" pageNumber="665">NMU 1969</specimenCode>
/1991)
</materialsCitation>
. 
<materialsCitation id="3B7A3C8B103139284081FA6DFBAEFD06" ID-GBIF-Occurrence="2571117008" collectorName="South-West" country="Switzerland" latitude="4.638727" location="Kumba" longLatPrecision="1" longitude="9.441354" pageId="7" pageNumber="665" specimenCount="6" specimenCount-female="3" specimenCount-male="3" typeStatus="paratype">
<typeStatus id="54A98874103139284081FA6DFD70FDE8" box="[648,760,591,615]" pageId="7" pageNumber="665" type="paratype">Paratypes</typeStatus>
, 
<specimenCount id="9D14FD5F10313928410DFA6DFCC9FDE8" box="[772,833,591,615]" pageId="7" pageNumber="665" type="female">3 ♀♀</specimenCount>
, CWs 25.4, 23.7, 
<quantity id="4CEA9B3310313928460AFA6DFBE2FDE8" box="[1027,1130,591,615]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="7" pageNumber="665" unit="mm" value="18.7">18.7 mm</quantity>
, 
<specimenCount id="9D14FD5F10313928467CFA6DFF4FFD06" pageId="7" pageNumber="665" type="male">3 ♂♂</specimenCount>
, CWs 22.3, 20.1, 
<quantity id="4CEA9B33103139284397FA53FD81FD06" box="[414,521,625,649]" metricMagnitude="-2" metricUnit="m" metricValue="1.98" pageId="7" pageNumber="665" unit="mm" value="19.8">19.8 mm</quantity>
, between 
<taxonomicName id="4C124D55103139284082FA53FD4AFD06" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" box="[651,706,625,649]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="665" phylum="Arthropoda" rank="genus">Buea</taxonomicName>
(
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, 
<geoCoordinate id="EE265011103139284162FA53FC6FFD06" box="[875,999,625,649]" degrees="9.299551" direction="east" orientation="longitude" pageId="7" pageNumber="665" precision="1" value="9.299551">9.299551°E</geoCoordinate>
) and
</materialsCitation>
<materialsCitation id="3B7A3C8B10313928463DFA53FD52FD23" ID-GBIF-Occurrence="2571117009" collectorName="South-West" country="Switzerland" latitude="4.638727" location="Kumba" longLatPrecision="1" longitude="9.441354" pageId="7" pageNumber="665" specimenCount="6" specimenCount-female="3" specimenCount-male="3" typeStatus="paratype">
<location id="8ECD600D10313928463DFA53FB0BFD06" LSID="urn:lsid:plazi:treatment:03BB87C010313927438DF8AEFC39F930:8ECD600D10313928463DFA53FB0BFD06" box="[1076,1155,625,649]" country="Switzerland" latitude="4.638727" longLatPrecision="1" longitude="9.441354" name="Kumba" pageId="7" pageNumber="665">Kumba</location>
(
<geoCoordinate id="EE2650111031392842AFFAB6FEA1FD23" box="[166,297,660,684]" degrees="4.638727" direction="north" orientation="latitude" pageId="7" pageNumber="665" precision="1" value="4.638727">4.638727°N</geoCoordinate>
, 
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), 
<collectorName id="26E753001031392843CEFAB6FDC1FD23" box="[455,585,660,684]" pageId="7" pageNumber="665">South-West</collectorName>
Region, coll
</materialsCitation>
. 
<collectionCode id="ED03AE131031392840EFFAB6FD71FD23" box="[742,761,660,684]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="665">R</collectionCode>
.H.L. 
<materialsCitation id="3B7A3C8B103139284125FAB6FCA5FD40" ID-GBIF-Occurrence="2569344771" collectionCode="NHM" country="Switzerland" pageId="7" pageNumber="665" specimenCode="NHM 1994.588, 591" specimenCount="1" typeStatus="paratype">
Disney, 1969 (
<specimenCode id="DBB49EAD1031392841C4FAB6FBFEFD23" box="[973,1142,660,684]" collectionCode="NHM" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="7" pageNumber="665" type="Herbarium">NHM 1994.588</specimenCode>
<emphasis id="B966EAC410313928467FFAB6FB0BFD23" box="[1142,1155,660,684]" italics="true" pageId="7" pageNumber="665">–</emphasis>
<specimenCode id="DBB49EAD1031392842A9FA95FF44FD40" box="[160,204,695,719]" collectionCode="NHM" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="7" pageNumber="665" type="Herbarium">591</specimenCode>
), DNA voucher specimens (
<bibRefCitation id="EF834B271031392843F5FA95FD43FD40" author="Daniels SR &amp; Phiri EE &amp; Klaus S &amp; Albrecht C &amp; Cumberlidge N" box="[508,715,695,719]" pageId="7" pageNumber="665" pagination="549 - 567" refId="ref8897" refString="Daniels SR, Phiri EE, Klaus S, Albrecht C, Cumberlidge N. 2015. Multilocus phylogeny of the Afrotropical freshwater crab fauna reveals historical drainage connectivity and transoceanic dispersal since the Eocene. Syst Biol. 64: 549 - 567. doi: 10.1093 / sysbio / syv 011" type="journal article" year="2015">Daniels et al. 2015</bibRefCitation>
, table 1)
</materialsCitation>
. 
<materialsCitation id="3B7A3C8B103139284131FA95FBECFD7E" ID-GBIF-Occurrence="2569344768" collectorName="South-West" country="Switzerland" latitude="4.153484" location="Adult" longLatPrecision="1" longitude="9.299551" municipality="Kumba area" pageId="7" pageNumber="665" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="8ECD600D103139284131FA95FCFDFD40" LSID="urn:lsid:plazi:treatment:03BB87C010313927438DF8AEFC39F930:8ECD600D103139284131FA95FCFDFD40" box="[824,885,695,719]" country="Switzerland" latitude="4.153484" longLatPrecision="1" longitude="9.299551" municipality="Kumba area" name="Adult" pageId="7" pageNumber="665">Adult</location>
<specimenCount id="9D14FD5F103139284175FA95FC18FD40" box="[892,912,695,719]" pageId="7" pageNumber="665" type="male">♂</specimenCount>
, CW 22.4, CL 15.8, 
<collectingCountry id="F305764610313928466BFA95FB0BFD40" box="[1122,1155,695,719]" name="Switzerland" pageId="7" pageNumber="665">CH</collectingCountry>
6.8, FW 
<quantity id="4CEA9B331031392842F2FAFBFEDBFD7E" box="[251,339,729,753]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="7" pageNumber="665" unit="mm" value="6.9">6.9 mm</quantity>
, 
<taxonomicName id="4C124D55103139284356FAFBFE1EFD7E" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" box="[351,406,729,753]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="665" phylum="Arthropoda" rank="genus">Buea</taxonomicName>
(
<geoCoordinate id="EE2650111031392843ADFAFBFDA0FD7E" box="[420,552,729,753]" degrees="4.153484" direction="north" orientation="latitude" pageId="7" pageNumber="665" precision="1" value="4.153484">4.153484°N</geoCoordinate>
, 
<geoCoordinate id="EE26501110313928403AFAFBFD38FD7E" box="[563,688,729,753]" degrees="9.299551" direction="east" orientation="longitude" pageId="7" pageNumber="665" precision="1" value="9.299551">9.299551°E</geoCoordinate>
) 
<collectingMunicipality id="6BC9ACAC1031392840B7FAFBFCC1FD7E" box="[702,841,729,753]" pageId="7" pageNumber="665">Kumba area</collectingMunicipality>
, 
<collectorName id="26E7530010313928415DFAFBFC5EFD7E" box="[852,982,729,753]" pageId="7" pageNumber="665">South-West</collectorName>
Region, coll
</materialsCitation>
. 
<collectionCode id="ED03AE13103139284666FAFBFB08FD7E" box="[1135,1152,729,753]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="665">R</collectionCode>
. H.L. 
<materialsCitation id="3B7A3C8B1031392842DBFADEFC39FC9B" ID-GBIF-Occurrence="2569344775" box="[210,945,764,788]" collectionCode="NMU" country="Switzerland" pageId="7" pageNumber="665" specimenCode="TRW 1969.22" specimenCount="1" typeStatus="paratype">
Disney, 1969 (
<collectionCode id="ED03AE13103139284378FADEFE23FC9B" box="[369,427,764,788]" pageId="7" pageNumber="665">NMU</collectionCode>
<specimenCode id="DBB49EAD1031392843BDFADEFDC3FC9B" box="[436,587,764,788]" collectionCode="TRW" pageId="7" pageNumber="665">TRW 1969.22</specimenCode>
), specimen photographed here
</materialsCitation>
.
</paragraph>
<paragraph id="8BAD36D61031392842A9FB63FCEFFBE0" blockId="7.[160,1156,833,1135]" pageId="7" pageNumber="665">
<emphasis id="B966EAC41031392842A9FB63FE56FCD5" bold="true" box="[160,478,833,858]" italics="true" pageId="7" pageNumber="665">Other material examined.</emphasis>
<materialsCitation id="3B7A3C8B1031392843E6FB60FCF0FCF2" ID-GBIF-Occurrence="2569344765" collectorName="Adult" country="Switzerland" latitude="4.153484" location="Kumba area" longLatPrecision="1" longitude="9.299551" municipality="South-West Region" pageId="7" pageNumber="665" specimenCount="1" specimenCount-female="1">
<collectorName id="26E753001031392843E6FB60FDA4FCD5" box="[495,556,834,858]" pageId="7" pageNumber="665">Adult</collectorName>
<specimenCount id="9D14FD5F103139284033FB60FDC6FCD5" box="[570,590,834,858]" pageId="7" pageNumber="665" type="female">♀</specimenCount>
, CW 23.2, CL 17.2, 
<collectingCountry id="F305764610313928414FFB60FCEFFCD5" box="[838,871,834,858]" name="Switzerland" pageId="7" pageNumber="665">CH</collectingCountry>
7.4, FW 
<quantity id="4CEA9B331031392841D5FB60FBB2FCD5" box="[988,1082,834,858]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="7" pageNumber="665" unit="mm" value="7.4">7.4 mm</quantity>
, 
<taxonomicName id="4C124D55103139284645FB60FB0BFCD5" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" box="[1100,1155,834,858]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="665" phylum="Arthropoda" rank="genus">Buea</taxonomicName>
(
<geoCoordinate id="EE2650111031392842AFFB47FEA1FCF2" box="[166,297,869,893]" degrees="4.153484" direction="north" orientation="latitude" pageId="7" pageNumber="665" precision="1" value="4.153484">4.153484°N</geoCoordinate>
, 
<geoCoordinate id="EE26501110313928433EFB47FE3EFCF2" box="[311,438,869,893]" degrees="9.299551" direction="east" orientation="longitude" pageId="7" pageNumber="665" precision="1" value="9.299551">9.299551°E</geoCoordinate>
), 
<location id="8ECD600D1031392843C3FB47FDDFFCF2" LSID="urn:lsid:plazi:treatment:03BB87C010313927438DF8AEFC39F930:8ECD600D1031392843C3FB47FDDFFCF2" box="[458,599,869,893]" country="Switzerland" latitude="4.153484" longLatPrecision="1" longitude="9.299551" municipality="South-West Region" name="Kumba area" pageId="7" pageNumber="665">Kumba area</location>
, 
<collectingMunicipality id="6BC9ACAC10313928406DFB47FCC9FCF2" box="[612,833,869,893]" pageId="7" pageNumber="665">South-West Region</collectingMunicipality>
, coll
</materialsCitation>
. 
<collectionCode id="ED03AE1310313928418CFB47FC10FCF2" box="[901,920,869,893]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="665">R</collectionCode>
.H.L. 
<materialsCitation id="3B7A3C8B1031392841C4FB47FE5DFC10" ID-GBIF-Occurrence="2569344774" collectingDate="1969-04-29" collectionCode="NMU" collectorName="Disney" country="Switzerland" pageId="7" pageNumber="665" specimenCode="TRW 1969.13" specimenCount="1">
<collectorName id="26E753001031392841C4FB47FB93FCF2" box="[973,1051,869,893]" pageId="7" pageNumber="665">Disney</collectorName>
, 
<date id="FFAC1016103139284621FB47FF5FFC10" pageId="7" pageNumber="665" value="1969-04-29">
<collectingDate id="EFE8E9FE103139284621FB47FF5FFC10" pageId="7" pageNumber="665" value="1969-04-29">29 April 1969</collectingDate>
</date>
(
<collectionCode id="ED03AE131031392842EFFBA5FEA8FC10" box="[230,288,903,927]" pageId="7" pageNumber="665">NMU</collectionCode>
<specimenCode id="DBB49EAD103139284323FBA5FE4AFC10" box="[298,450,903,927]" collectionCode="TRW" pageId="7" pageNumber="665">TRW 1969.13</specimenCode>
a)
</materialsCitation>
. 
<materialsCitation id="3B7A3C8B1031392843E8FBA5FCDDFC4D" ID-GBIF-Occurrence="2569344776" collectorName="Adult" country="Switzerland" location="CH" pageId="7" pageNumber="665" specimenCount="6" specimenCount-female="1" specimenCount-male="2" specimenCount-subadult="3">
<collectorName id="26E753001031392843E8FBA5FD96FC10" box="[481,542,903,927]" pageId="7" pageNumber="665">Adult</collectorName>
<specimenCount id="9D14FD5F103139284021FBA5FDB4FC10" box="[552,572,903,927]" pageId="7" pageNumber="665" type="male">♂</specimenCount>
, CW 19.8, CL 14.5, 
<collectingCountry id="F3057646103139284115FBA5FCB5FC10" box="[796,829,903,927]" name="Switzerland" pageId="7" pageNumber="665">CH</collectingCountry>
6.8, FW 
<quantity id="4CEA9B331031392841AAFBA5FC75FC10" box="[931,1021,903,927]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="7" pageNumber="665" unit="mm" value="6.9">6.9 mm</quantity>
, 
<specimenCount id="9D14FD5F103139284603FBA5FB0BFC10" box="[1034,1155,903,927]" pageId="7" pageNumber="665" type="subadult">3 subadult</specimenCount>
<specimenCount id="9D14FD5F1031392842A9FB88FF4FFC4D" box="[160,199,938,962]" pageId="7" pageNumber="665" type="male">♂♂</specimenCount>
, CWs 16.4, 15.6, 
<quantity id="4CEA9B3310313928438AFB88FE60FC4D" box="[387,488,938,962]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="7" pageNumber="665" unit="mm" value="14.9">14.9 mm</quantity>
, subadult 
<specimenCount id="9D14FD5F103139284055FB88FDF8FC4D" box="[604,624,938,962]" pageId="7" pageNumber="665" type="female">♀</specimenCount>
, CW 
<quantity id="4CEA9B331031392840A0FB88FC86FC4D" box="[681,782,938,962]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="7" pageNumber="665" unit="mm" value="15.3">15.3 mm</quantity>
, 
<specimenCount id="9D14FD5F103139284110FB88FCDDFC4D" box="[793,853,938,962]" pageId="7" pageNumber="665" type="generic">9 juv</specimenCount>
</materialsCitation>
<materialsCitation id="3B7A3C8B10313928415DFB88FE3DFC6A" ID-GBIF-Occurrence="2569344766" collectorName="Okia Stream" country="Switzerland" location="Kumba area" municipality="South-West Region" pageId="7" pageNumber="665" specimenCount="1">
., 
<collectorName id="26E7530010313928416BFB88FC65FC4D" box="[866,1005,938,962]" pageId="7" pageNumber="665">Okia Stream</collectorName>
, 
<location id="8ECD600D1031392841FEFB88FB09FC4D" LSID="urn:lsid:plazi:treatment:03BB87C010313927438DF8AEFC39F930:8ECD600D1031392841FEFB88FB09FC4D" box="[1015,1153,938,962]" country="Switzerland" municipality="South-West Region" name="Kumba area" pageId="7" pageNumber="665">Kumba area</location>
, 
<collectingMunicipality id="6BC9ACAC1031392842A9FBEFFEF5FC6A" box="[160,381,973,997]" pageId="7" pageNumber="665">South-West Region</collectingMunicipality>
, coll
</materialsCitation>
. 
<collectionCode id="ED03AE131031392843CAFBEFFE5EFC6A" box="[451,470,973,997]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="665">R</collectionCode>
.H.L. 
<materialsCitation id="3B7A3C8B103139284003FBEFFC6BFC6A" ID-GBIF-Occurrence="2569344772" box="[522,995,973,997]" collectingDate="1969-05-09" collectionCode="NMU" collectorName="Disney" country="Switzerland" pageId="7" pageNumber="665" specimenCode="TRW 1969.27" specimenCount="1">
<collectorName id="26E75300103139284003FBEFFDD0FC6A" box="[522,600,973,997]" pageId="7" pageNumber="665">Disney</collectorName>
, 
<date id="FFAC101610313928406FFBEFFD67FC6A" box="[614,751,973,997]" pageId="7" pageNumber="665" value="1969-05-09">
<collectingDate id="EFE8E9FE10313928406FFBEFFD67FC6A" box="[614,751,973,997]" pageId="7" pageNumber="665" value="1969-05-09">9 May 1969</collectingDate>
</date>
(
<collectionCode id="ED03AE13103139284109FBEFFCB2FC6A" box="[768,826,973,997]" pageId="7" pageNumber="665">NMU</collectionCode>
<specimenCode id="DBB49EAD10313928414CFBEFFC55FC6A" box="[837,989,973,997]" collectionCode="TRW" pageId="7" pageNumber="665">TRW 1969.27</specimenCode>
)
</materialsCitation>
. Adult 
<specimenCount id="9D14FD5F103139284630FBEEFBC5FC6B" box="[1081,1101,972,996]" pageId="7" pageNumber="665" type="male">♂</specimenCount>
, CW 19.0 mm, 
<specimenCount id="9D14FD5F103139284306FBCDFE0EFB88" box="[271,390,1007,1031]" pageId="7" pageNumber="665" type="subadult">3 subadult</specimenCount>
<specimenCount id="9D14FD5F103139284387FBCDFE3DFB88" box="[398,437,1007,1031]" pageId="7" pageNumber="665" type="male">♂♂</specimenCount>
, CWs 18.4, 15.8, 
<quantity id="4CEA9B33103139284078FBCDFD5EFB88" box="[625,726,1007,1031]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="7" pageNumber="665" unit="mm" value="12.6">12.6 mm</quantity>
, subadult 
<specimenCount id="9D14FD5F103139284143FBCDFCD6FB88" box="[842,862,1007,1031]" pageId="7" pageNumber="665" type="female">♀</specimenCount>
, CW 
<quantity id="4CEA9B33103139284191FBCDFC75FB88" box="[920,1021,1007,1031]" metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" pageId="7" pageNumber="665" unit="mm" value="17.9">17.9 mm</quantity>
, 
<specimenCount id="9D14FD5F103139284601FBCDFBCFFB88" box="[1032,1095,1007,1031]" pageId="7" pageNumber="665" type="generic">
2 juv
<materialsCitation id="3B7A3C8B10313928464AFBCDFBCFFB88" ID-GBIF-Occurrence="2569344763" box="[1091,1095,1007,1031]" collectorName="Okia Stream" country="Switzerland" location="Kumba area" municipality="South-West Region" pageId="7" pageNumber="665" specimenCount="1">.</materialsCitation>
</specimenCount>
, 
<collectorName id="26E75300103139284658FBCDFF7AFBA5" pageId="7" pageNumber="665">Okia Stream</collectorName>
, 
<location id="8ECD600D1031392842F7FC30FE02FBA5" LSID="urn:lsid:plazi:treatment:03BB87C010313927438DF8AEFC39F930:8ECD600D1031392842F7FC30FE02FBA5" box="[254,394,1042,1066]" country="Switzerland" municipality="South-West Region" name="Kumba area" pageId="7" pageNumber="665">Kumba area</location>
, 
<collectingMunicipality id="6BC9ACAC10313928439CFC30FDF9FBA5" box="[405,625,1042,1066]" pageId="7" pageNumber="665">South-West Region</collectingMunicipality>
, coll. 
<collectionCode id="ED03AE131031392840BDFC30FD4FFBA5" box="[692,711,1042,1066]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="665">R</collectionCode>
.H.L. 
<materialsCitation id="3B7A3C8B1031392840F3FC30FE8AFBC2" ID-GBIF-Occurrence="2569344770" collectingDate="1969-03-27" collectionCode="NMU" collectorName="Disney" country="Switzerland" pageId="7" pageNumber="665" specimenCode="TRW 1969.29" specimenCount="1">
<collectorName id="26E753001031392840F3FC30FCC0FBA5" box="[762,840,1042,1066]" pageId="7" pageNumber="665">Disney</collectorName>
, 
<date id="FFAC101610313928415DFC30FC77FBA5" box="[852,1023,1042,1066]" pageId="7" pageNumber="665" value="1969-03-27">
<collectingDate id="EFE8E9FE10313928415DFC30FC77FBA5" box="[852,1023,1042,1066]" pageId="7" pageNumber="665" value="1969-03-27">27 March 1969</collectingDate>
</date>
(
<collectionCode id="ED03AE13103139284607FC30FBC0FBA5" box="[1038,1096,1042,1066]" pageId="7" pageNumber="665">NMU</collectionCode>
<specimenCode id="DBB49EAD103139284658FC30FF74FBC2" collectionCode="TRW" pageId="7" pageNumber="665">TRW 1969.29</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="3B7A3C8B103139284305FC17FBEFFBC2" ID-GBIF-Occurrence="2569344773" box="[268,1127,1076,1101]" collectorName="Limbe &amp; Victoria" country="Switzerland" location="South-West Region" pageId="7" pageNumber="665" specimenCount="2" specimenCount-male="2">
<collectorName id="26E75300103139284305FC17FEDAFBC2" box="[268,338,1077,1101]" pageId="7" pageNumber="665">Limbé</collectorName>
(formerly 
<collectorName id="26E753001031392843CFFC17FD97FBC2" box="[454,543,1077,1101]" pageId="7" pageNumber="665">Victoria</collectorName>
), 
<location id="8ECD600D103139284027FC17FC80FBC2" LSID="urn:lsid:plazi:treatment:03BB87C010313927438DF8AEFC39F930:8ECD600D103139284027FC17FC80FBC2" box="[558,776,1077,1101]" country="Switzerland" name="South-West Region" pageId="7" pageNumber="665">South-West Region</location>
, 
<specimenCount id="9D14FD5F10313928411BFC17FCC6FBC3" box="[786,846,1076,1101]" pageId="7" pageNumber="665" type="male">2 ♂♂</specimenCount>
, CWs 21.3, 
<quantity id="4CEA9B331031392841C7FC17FBBBFBC2" box="[974,1075,1077,1101]" metricMagnitude="-2" metricUnit="m" metricValue="2.01" pageId="7" pageNumber="665" unit="mm" value="20.1">20.1 mm</quantity>
, coll
</materialsCitation>
<materialsCitation id="3B7A3C8B10313928466FFC17FB09FBC2" ID-GBIF-Occurrence="2569344777" box="[1126,1153,1077,1101]" collectorName="Limbe &amp; Victoria" country="Switzerland" isTailEnumeration="true" location="South-West Region" pageId="7" pageNumber="665" specimenCount="1">. E</materialsCitation>
<materialsCitation id="3B7A3C8B103139284689FC17FCEBFBE0" ID-GBIF-Occurrence="2569344778" collectingDate="1912-11-06" collectionCode="ZIM" collectorName="Fickenday" country="Switzerland" pageId="7" pageNumber="665" specimenCount="1">
. 
<collectorName id="26E753001031392842A9FC75FE9BFBE0" box="[160,275,1111,1135]" pageId="7" pageNumber="665">Fickenday</collectorName>
, 
<date id="FFAC1016103139284316FC75FE65FBE0" box="[287,493,1111,1135]" pageId="7" pageNumber="665" value="1912-11-06">
<collectingDate id="EFE8E9FE103139284316FC75FE65FBE0" box="[287,493,1111,1135]" pageId="7" pageNumber="665" value="1912-11-06">6 November 1912</collectingDate>
</date>
, 
<emphasis id="B966EAC41031392843F0FC75FE77FBE0" box="[505,511,1111,1135]" italics="true" pageId="7" pageNumber="665">‘</emphasis>
edible land crabs 
<emphasis id="B966EAC41031392840CBFC75FD40FBE0" box="[706,712,1111,1135]" italics="true" pageId="7" pageNumber="665">’</emphasis>
(
<collectionCode id="ED03AE131031392840DFFC75FC8AFBE0" box="[726,770,1111,1135]" country="Slovenia" httpUri="http://grbio.org/cool/3263-rkr7" name="ZIM Culture Collection of Industrial Microorganisms" pageId="7" pageNumber="665">ZIM</collectionCode>
K-3607)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C308655D1031392842A9FCBEFDDEFAEC" pageId="7" pageNumber="665" type="diagnosis">
<paragraph id="8BAD36D61031392842A9FCBEFE9CFB3A" blockId="7.[160,1156,1180,1379]" box="[160,276,1180,1205]" pageId="7" pageNumber="665">
<emphasis id="B966EAC41031392842A9FCBEFE9CFB3A" bold="true" box="[160,276,1180,1205]" italics="true" pageId="7" pageNumber="665">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BAD36D61031392842A9FCE2FDDEFAEC" blockId="7.[160,1156,1180,1379]" pageId="7" pageNumber="665">
Exorbital tooth absent where anterolateral margin meets lateral orbital margin (
<figureCitation id="13292A531031392841FAFCE2FBD8FB57" box="[1011,1104,1216,1240]" captionStart="Figure 1" captionStartId="3.[160,225,1645,1668]" captionTargetBox="[163,1153,507,1615]" captionTargetId="figure@3.[163,1153,507,1615]" captionTargetPageId="3" captionText="Figure 1. Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon (NHM 1994.587). (a) Whole animal, dorsal view; (b) whole animal, ventral view. Photographs by Phillip Crabb, NHM, London. Scale bar: a = 14 mm; b = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3675496" httpUri="https://zenodo.org/record/3675496/files/figure.png" pageId="7" pageNumber="665">Figures 1</figureCitation>
(a,b) and 4(a)); major cheliped dactylus broad, flattened, not arched (
<figureCitation id="13292A5310313928414FFCC1FC2BFB74" box="[838,931,1251,1275]" captionStart="Figure 1" captionStartId="3.[160,225,1645,1668]" captionTargetBox="[163,1153,507,1615]" captionTargetId="figure@3.[163,1153,507,1615]" captionTargetPageId="3" captionText="Figure 1. Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon (NHM 1994.587). (a) Whole animal, dorsal view; (b) whole animal, ventral view. Photographs by Phillip Crabb, NHM, London. Scale bar: a = 14 mm; b = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3675496" httpUri="https://zenodo.org/record/3675496/files/figure.png" pageId="7" pageNumber="665">Figures 1</figureCitation>
(b) and 5(a,b)); lower margin of cheliped merus with four large jagged pointed teeth (
<figureCitation id="13292A53103139284172FD27FC54FA92" box="[891,988,1285,1309]" captionStart="Figure 1" captionStartId="3.[160,225,1645,1668]" captionTargetBox="[163,1153,507,1615]" captionTargetId="figure@3.[163,1153,507,1615]" captionTargetPageId="3" captionText="Figure 1. Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon (NHM 1994.587). (a) Whole animal, dorsal view; (b) whole animal, ventral view. Photographs by Phillip Crabb, NHM, London. Scale bar: a = 14 mm; b = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3675496" httpUri="https://zenodo.org/record/3675496/files/figure.png" pageId="7" pageNumber="665">Figures 1</figureCitation>
(b) and 6(a,b)); anterior corners of carapace surface smooth; carapace grooves shallow to absent (
<figureCitation id="13292A5310313928460EFD0AFBEBFACF" box="[1031,1123,1320,1344]" captionStart="Figure 1" captionStartId="3.[160,225,1645,1668]" captionTargetBox="[163,1153,507,1615]" captionTargetId="figure@3.[163,1153,507,1615]" captionTargetPageId="3" captionText="Figure 1. Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon (NHM 1994.587). (a) Whole animal, dorsal view; (b) whole animal, ventral view. Photographs by Phillip Crabb, NHM, London. Scale bar: a = 14 mm; b = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3675496" httpUri="https://zenodo.org/record/3675496/files/figure.png" pageId="7" pageNumber="665">Figures 1</figureCitation>
(a) and 4(a); 
<bibRefCitation id="EF834B2710313928430DFD69FE44FAEC" author="Cumberlidge N" box="[260,460,1355,1379]" pageId="7" pageNumber="665" pagination="571 - 584" refId="ref8064" refString="Cumberlidge N. 1993. Two new species of Potamonemus Cumberlidge and Clark, 1992 (Potamoidea, Potamonautidae) from the rain forests of West Africa. J Crustacean Biol. 13: 571 - 584. doi: 10.2307 / 1548798" type="journal article" year="1993">Cumberlidge 1993</bibRefCitation>
, figs. 2a, 3a).
</paragraph>
</subSubSection>
<subSubSection id="C308655D1031392842A9FDB2FBC3FA44" box="[160,1099,1424,1483]" pageId="7" pageNumber="665" type="description">
<paragraph id="8BAD36D61031392842A9FDB2FEAEFA26" blockId="7.[160,1099,1424,1483]" box="[160,294,1424,1449]" pageId="7" pageNumber="665">
<emphasis id="B966EAC41031392842A9FDB2FEAEFA26" bold="true" box="[160,294,1424,1449]" italics="true" pageId="7" pageNumber="665">Description</emphasis>
</paragraph>
<paragraph id="8BAD36D61031392842A9FD91FBC3FA44" blockId="7.[160,1099,1424,1483]" box="[160,1099,1459,1483]" pageId="7" pageNumber="665">
Same as for the genus and for 
<taxonomicName id="4C124D5510313928400CFD91FDEBFA44" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" baseAuthorityName="Cumberlidge" baseAuthorityYear="1993" box="[517,611,1459,1483]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="665" phylum="Arthropoda" rank="species" species="asylos" status="comb. nov.">
<emphasis id="B966EAC410313928400CFD91FDEBFA44" box="[517,611,1459,1483]" italics="true" pageId="7" pageNumber="665">P. asylos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557BF103139284062FD91FD62FA44" box="[619,746,1459,1483]" pageId="7" pageNumber="665" rank="species">comb. nov.</taxonomicNameLabel>
(see 
<bibRefCitation id="EF834B27103139284121FD91FC71FA44" author="Cumberlidge N" box="[808,1017,1459,1483]" pageId="7" pageNumber="665" pagination="571 - 584" refId="ref8064" refString="Cumberlidge N. 1993. Two new species of Potamonemus Cumberlidge and Clark, 1992 (Potamoidea, Potamonautidae) from the rain forests of West Africa. J Crustacean Biol. 13: 571 - 584. doi: 10.2307 / 1548798" type="journal article" year="1993">Cumberlidge 1993</bibRefCitation>
, 1999).
</paragraph>
</subSubSection>
<subSubSection id="C308655D1031392842A9FDDAFC78F9D8" pageId="7" pageNumber="665" type="distribution">
<paragraph id="8BAD36D61031392842A9FDDAFEA4F99E" blockId="7.[160,1155,1528,1623]" box="[160,300,1528,1553]" pageId="7" pageNumber="665">
<emphasis id="B966EAC41031392842A9FDDAFEA4F99E" bold="true" box="[160,300,1528,1553]" italics="true" pageId="7" pageNumber="665">Distribution</emphasis>
</paragraph>
<paragraph id="8BAD36D61031392842A9FE3EFC78F9D8" blockId="7.[160,1155,1528,1623]" pageId="7" pageNumber="665">
<taxonomicName id="4C124D551031392842A9FE3EFEAAF9BB" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" baseAuthorityName="Cumberlidge" baseAuthorityYear="1993" box="[160,290,1564,1588]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="665" phylum="Arthropoda" rank="species" species="asylos" status="comb. nov.">
<emphasis id="B966EAC41031392842A9FE3EFEAAF9BB" box="[160,290,1564,1588]" italics="true" pageId="7" pageNumber="665">Buea asylos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557BF103139284325FE3EFE24F9BB" box="[300,428,1564,1588]" pageId="7" pageNumber="665" rank="species">comb. nov.</taxonomicNameLabel>
is endemic to the rainforest zone of south-western 
<collectingCountry id="F3057646103139284607FE3EFB0BF9BB" box="[1038,1155,1564,1588]" name="Cameroon" pageId="7" pageNumber="665">Cameroon</collectingCountry>
between the towns of Kumba, 
<taxonomicName id="4C124D5510313928400AFE1DFDB2F9D8" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" box="[515,570,1599,1623]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="665" phylum="Arthropoda" rank="genus">Buea</taxonomicName>
and 
<collectingRegion id="49D6F83410313928407EFE1DFD35F9D8" box="[631,701,1599,1623]" country="Cameroon" name="West" pageId="7" pageNumber="665">Limbé</collectingRegion>
(
<bibRefCitation id="EF834B271031392840C2FE1DFC16F9D8" author="Cumberlidge N" box="[715,926,1599,1623]" pageId="7" pageNumber="665" pagination="571 - 584" refId="ref8064" refString="Cumberlidge N. 1993. Two new species of Potamonemus Cumberlidge and Clark, 1992 (Potamoidea, Potamonautidae) from the rain forests of West Africa. J Crustacean Biol. 13: 571 - 584. doi: 10.2307 / 1548798" type="journal article" year="1993">Cumberlidge 1993</bibRefCitation>
, 1999).
</paragraph>
</subSubSection>
<subSubSection id="C308655D1031392842A9FEA6FC89F930" pageId="7" pageNumber="665" type="materials_examined">
<paragraph id="8BAD36D61031392842A9FEA6FEBFF912" blockId="7.[160,769,1668,1727]" box="[160,311,1668,1693]" pageId="7" pageNumber="665">
<emphasis id="B966EAC41031392842A9FEA6FEBFF912" bold="true" box="[160,311,1668,1693]" italics="true" pageId="7" pageNumber="665">
<typeStatus id="54A988741031392842A9FEA6FF50F912" box="[160,216,1668,1693]" pageId="7" pageNumber="665">Type</typeStatus>
locality
</emphasis>
</paragraph>
<paragraph id="8BAD36D61031392842A9FE85FC89F930" blockId="7.[160,769,1668,1727]" box="[160,769,1703,1727]" pageId="7" pageNumber="665">
Between 
<taxonomicName id="4C124D55103139284302FE85FECAF930" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" box="[267,322,1703,1727]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="665" phylum="Arthropoda" rank="genus">Buea</taxonomicName>
and Kumba, south-western 
<collectingCountry id="F305764610313928408FFE85FD75F930" box="[646,765,1703,1727]" name="Cameroon" pageId="7" pageNumber="665">Cameroon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C308655D103E392742A9F8ACFF7EFEBD" pageId="8" pageNumber="666" type="biology_ecology">
<paragraph id="8BAD36D6103E392742A9F8ACFF74FF28" blockId="8.[160,1156,142,306]" box="[160,252,142,167]" pageId="8" pageNumber="666">
<emphasis id="B966EAC4103E392742A9F8ACFF74FF28" bold="true" box="[160,252,142,167]" italics="true" pageId="8" pageNumber="666">Ecology</emphasis>
</paragraph>
<paragraph id="8BAD36D6103E392742A9F890FF7EFEBD" blockId="8.[160,1156,142,306]" pageId="8" pageNumber="666">
<taxonomicName id="4C124D55103E392742A9F890FEAEFF45" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" baseAuthorityName="Cumberlidge" baseAuthorityYear="1993" box="[160,294,178,202]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="species" species="asylos" status="comb. nov.">
<emphasis id="B966EAC4103E392742A9F890FEAEFF45" box="[160,294,178,202]" italics="true" pageId="8" pageNumber="666">Buea asylos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557BF103E39274339F890FE3CFF45" box="[304,436,178,202]" pageId="8" pageNumber="666" rank="species">comb. nov.</taxonomicNameLabel>
is restricted to the humid lowland and montane rainforests of south-western 
<collectingCountry id="F3057646103E39274379F8F6FE60FF63" box="[368,488,212,236]" name="Cameroon" pageId="8" pageNumber="666">Cameroon</collectingCountry>
in the area near Mount 
<collectingCountry id="F3057646103E39274103F8F6FC0AFF63" box="[778,898,212,236]" name="Cameroon" pageId="8" pageNumber="666">Cameroon</collectingCountry>
(
<quantity id="4CEA9B33103E39274198F8F6FC63FF63" box="[913,1003,212,236]" metricMagnitude="3" metricUnit="m" metricValue="4.095" pageId="8" pageNumber="666" unit="m" value="4095.0">4095 m</quantity>
asl) that has an annual average rainfall of more than 
<quantity id="4CEA9B33103E39274092F8D5FC86FE80" box="[667,782,247,271]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="666" unit="mm" value="5000.0">5000 mm</quantity>
(
<bibRefCitation id="EF834B27103E3927412BF8D5FBF6FE80" author="Mvogo Ndongo PA &amp; Cumberlidge N &amp; Poettinger TS &amp; von Rintelen T &amp; Tamesse JL &amp; Schubart CD" box="[802,1150,247,271]" pageId="8" pageNumber="666" pagination="439 - 444" refId="ref9484" refString="Mvogo Ndongo PA, Cumberlidge N, Poettinger TS, von Rintelen T, Tamesse JL, Schubart CD. 2017 a. Molecular evidence for the assignment of the Cameroonian freshwater crab genus Louisea Cumberlidge, 1994, to the Afrotropical subfamily Potamonautinae Bott, 1970 (Crustacea: Potamoidea: Potamonautidae). Zootaxa. 4286: 439 - 444. doi: 10.11646 / zootaxa. 4286.3.12" type="journal article" year="2017">Mvogo Ndongo et al. 2017a</bibRefCitation>
, 
<bibRefCitation id="EF834B27103E392742A9F938FF60FEBD" author="Mvogo Ndongo PA &amp; Schubart CD &amp; von Rintelen T &amp; Tamesse JL &amp; Cumberlidge N" box="[160,232,282,306]" pageId="8" pageNumber="666" pagination="161 - 173" refId="ref9554" refString="Mvogo Ndongo PA, Schubart CD, von Rintelen T, Tamesse JL, Cumberlidge N. 2017 b. Morphological and molecular evidence for a new species of freshwater crab of the genus Sudanonautes Bott, 1955 (Brachyura: Potamoidea: Potamonautidae) from Cameroon, with notes on its ecology. Zootaxa. 4242: 161 - 173. doi: 10.11646 / zootaxa. 4242.1.8" type="journal article" year="2017">2017b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C308655D103E392742A9F97CFC39F930" pageId="8" pageNumber="666" type="discussion">
<paragraph id="8BAD36D6103E392742A9F97CFE8EFEF8" blockId="8.[160,1156,350,756]" box="[160,262,350,375]" pageId="8" pageNumber="666">
<emphasis id="B966EAC4103E392742A9F97CFE8EFEF8" bold="true" box="[160,262,350,375]" italics="true" pageId="8" pageNumber="666">Remarks</emphasis>
</paragraph>
<paragraph id="8BAD36D6103E392742A9F9A0FCFEFD7B" blockId="8.[160,1156,350,756]" pageId="8" pageNumber="666">
Significant morphological differences between 
<taxonomicName id="4C124D55103E392740AFF9A0FC88FE15" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" baseAuthorityName="Cumberlidge" baseAuthorityYear="1993" box="[678,768,386,410]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="species" species="asylos" status="comb. nov.">
<emphasis id="B966EAC4103E392740AFF9A0FC88FE15" box="[678,768,386,410]" italics="true" pageId="8" pageNumber="666">B. asylos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557BF103E39274103F9A0FC0EFE15" box="[778,902,386,410]" pageId="8" pageNumber="666" rank="species">comb. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C124D55103E3927419BF9A0FBBDFE15" authorityName="Cumberlidge and Clark" authorityYear="1992" box="[914,1077,386,410]" class="Malacostraca" family="Potamonautidae" genus="Potamonemus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="species" species="mambilorum">
<emphasis id="B966EAC4103E3927419BF9A0FBBDFE15" box="[914,1077,386,410]" italics="true" pageId="8" pageNumber="666">P. mambilorum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C124D55103E39274678F9A0FF56FE33" authorityName="Cumberlidge" authorityYear="1993" class="Malacostraca" family="Potamonautidae" genus="Potamonemus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="species" species="sachsi">
<emphasis id="B966EAC4103E39274678F9A0FF56FE33" italics="true" pageId="8" pageNumber="666">P. sachsi</emphasis>
</taxonomicName>
were found in the G1, G2 and mandible (
<figureCitation id="13292A53103E39274099F986FD67FE33" box="[656,751,420,444]" captionStart="Figure 7" captionStartId="14.[160,225,1009,1032]" captionTargetBox="[163,1153,142,978]" captionTargetId="figure@14.[163,1153,142,978]" captionTargetPageId="14" captionText="Figure 7. Right G1, ventral view (a–c) and dorsal view (d–f). (a,d) Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon, total G1 length 5.9 mm (TA 2.7 mm, SS 3.2 mm) (NHM 1994.587); (b,e) Potamonemus mambilorum Cumberlidge and Clark, 1992, holotype, adult ♂, CW 34.5 mm, Somié, Cameroon, total G1 length 9.7 mm (TA 3.6 mm, SS 6.1 mm) (NHM 1991:183); (c,f) P. sachsi Cumberlidge, 1993, adult ♂, CW 31.6 mm, Obudu Plateau, Nigeria), total G1 length 9.0 mm (TA 3.4 mm, SS 5.6 mm) (NMU 8.IV.1983A). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3675508" httpUri="https://zenodo.org/record/3675508/files/figure.png" pageId="8" pageNumber="666">Figures 7</figureCitation>
(a 
<emphasis id="B966EAC4103E3927410AF986FC98FE33" box="[771,784,420,444]" italics="true" pageId="8" pageNumber="666">–</emphasis>
f) and 8(a 
<emphasis id="B966EAC4103E3927417BF986FCF7FE33" box="[882,895,420,444]" italics="true" pageId="8" pageNumber="666">–</emphasis>
c,g 
<emphasis id="B966EAC4103E39274194F986FC22FE33" box="[925,938,420,444]" italics="true" pageId="8" pageNumber="666">–</emphasis>
i)) that are used here to define 
<taxonomicName id="4C124D55103E3927430FF9E5FEB2FE50" authority="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels, 2019" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" box="[262,314,455,479]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B966EAC4103E3927430FF9E5FEB2FE50" box="[262,314,455,479]" italics="true" pageId="8" pageNumber="666">Buea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557BF103E39274349F9E5FE2CFE50" box="[320,420,455,479]" pageId="8" pageNumber="666" rank="genus">gen. nov.</taxonomicNameLabel>
Other noteworthy characters of 
<taxonomicName id="4C124D55103E392740F7F9E5FCDDFE50" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" baseAuthorityName="Cumberlidge" baseAuthorityYear="1993" box="[766,853,455,479]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="species" species="asylos" status="comb. nov.">
<emphasis id="B966EAC4103E392740F7F9E5FCDDFE50" box="[766,853,455,479]" italics="true" pageId="8" pageNumber="666">B. asylos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557BF103E39274152F9E5FC5BFE50" box="[859,979,455,479]" pageId="8" pageNumber="666" rank="species">comb. nov.</taxonomicNameLabel>
include those of the chelipeds and carapace of that set it apart from the two species of 
<taxonomicName id="4C124D55103E392741B3F9C8FBD9FD8D" authorityName="Cumberlidge and Clark" authorityYear="1992" box="[954,1105,490,514]" class="Malacostraca" family="Potamonautidae" genus="Potamonemus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC4103E392741B3F9C8FBD9FD8D" box="[954,1105,490,514]" italics="true" pageId="8" pageNumber="666">Potamonemus</emphasis>
</taxonomicName>
: the cheliped merus lower medial margin has large jagged teeth (
<figureCitation id="13292A53103E39274138FA2EFC01FDAB" box="[817,905,524,548]" captionStart="Figure 6" captionStartId="13.[160,225,965,988]" captionTargetBox="[163,1153,142,934]" captionTargetId="figure@13.[163,1153,142,934]" captionTargetPageId="13" captionText="Figure 6. Cheliped merus. (a–b) Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon (NHM 1994.587), ventral view of right and left cheliped merus, respectively; (c–d) Potamonemus mambilorum Cumberlidge and Clark, 1992, holotype, adult ♂, CW 34.5 mm, Somié, Cameroon (NHM 1991:183): (c) ventral view of right cheliped merus; (d) dorsal view of right cheliped merus; (e–f) P. sachsi Cumberlidge, 1993, adult ♂, CW 31.6 mm, Obudu Plateau, Nigeria (NMU 8.IV.1983A), ventral view of right and left cheliped merus respectively. Scale bar: a,b = 3.40 mm; c,d = 4.2 6mm; e,f = 3.90 mm." figureDoi="http://doi.org/10.5281/zenodo.3675506" httpUri="https://zenodo.org/record/3675506/files/figure.png" pageId="8" pageNumber="666">Figure 6</figureCitation>
(a,b)) (vs small granules in 
<taxonomicName id="4C124D55103E392742B0FA0DFED8FDC8" authorityName="Cumberlidge and Clark" authorityYear="1992" box="[185,336,559,583]" class="Malacostraca" family="Potamonautidae" genus="Potamonemus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC4103E392742B0FA0DFED8FDC8" box="[185,336,559,583]" italics="true" pageId="8" pageNumber="666">Potamonemus</emphasis>
</taxonomicName>
, 
<figureCitation id="13292A53103E39274353FA0DFE39FDC8" box="[346,433,559,583]" captionStart="Figure 6" captionStartId="13.[160,225,965,988]" captionTargetBox="[163,1153,142,934]" captionTargetId="figure@13.[163,1153,142,934]" captionTargetPageId="13" captionText="Figure 6. Cheliped merus. (a–b) Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon (NHM 1994.587), ventral view of right and left cheliped merus, respectively; (c–d) Potamonemus mambilorum Cumberlidge and Clark, 1992, holotype, adult ♂, CW 34.5 mm, Somié, Cameroon (NHM 1991:183): (c) ventral view of right cheliped merus; (d) dorsal view of right cheliped merus; (e–f) P. sachsi Cumberlidge, 1993, adult ♂, CW 31.6 mm, Obudu Plateau, Nigeria (NMU 8.IV.1983A), ventral view of right and left cheliped merus respectively. Scale bar: a,b = 3.40 mm; c,d = 4.2 6mm; e,f = 3.90 mm." figureDoi="http://doi.org/10.5281/zenodo.3675506" httpUri="https://zenodo.org/record/3675506/files/figure.png" pageId="8" pageNumber="666">Figure 6</figureCitation>
(c,f)), the cheliped carpus medial margin of 
<taxonomicName id="4C124D55103E3927417EFA0DFC45FDC8" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" baseAuthorityName="Cumberlidge" baseAuthorityYear="1993" box="[887,973,559,583]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="species" species="asylos" status="comb. nov.">
<emphasis id="B966EAC4103E3927417EFA0DFC45FDC8" box="[887,973,559,583]" italics="true" pageId="8" pageNumber="666">B. asylos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557BF103E392741DBFA0DFBC1FDC8" box="[978,1097,559,583]" pageId="8" pageNumber="666" rank="species">comb. nov.</taxonomicNameLabel>
has a small but distinct pointed proximal tooth (
<figureCitation id="13292A53103E3927408BFA70FD56FDE5" box="[642,734,594,618]" captionStart="Figure 5" captionStartId="12.[160,225,965,988]" captionTargetBox="[163,1153,142,934]" captionTargetId="figure@12.[163,1153,142,934]" captionTargetPageId="12" captionText="Figure 5. (a–f) Frontal view of right and left chelipeds; (g–i) dorsal view of right cheliped carpus. Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon (NHM 1994.587), length of right cheliped = 24.2 mm. (c–d) Potamonemus mambilorum Cumberlidge and Clark, 1992, holotype, adult ♂, CW 34.5 mm, Somié, Cameroon (NHM 1991:183), length of right cheliped = 37.0 mm; (e–f) P. sachsi, adult ♂, CW 31.6 mm, Obudu Plateau, Nigeria (NMU 8.IV.1983A), length of right cheliped = 27.6 mm; (g) Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon (NHM 1994.587); (h) P. mambilorum Cumberlidge and Clark, 1992, holotype, adult ♂, CW 34.5 mm, Somié, Cameroon (NHM 1991:183); (i) P. sachsi Cumberlidge, 1993, adult ♂, CW 31.6 mm, Obudu Plateau, Nigeria (NMU 8.IV.1983A). Figures (a–i) scaled for comparison. Photographs (a–f) by Phillip Crabb, NHM, London. Scale bar: a,b = 4.21 mm; c,d = 5.26 mm; e, f = 4.82 mm; g = 3.1 mm; h = 3.83 mm; i = 3.51 mm." figureDoi="http://doi.org/10.5281/zenodo.3675504" httpUri="https://zenodo.org/record/3675504/files/figure.png" pageId="8" pageNumber="666">Figure 5</figureCitation>
(g)) (vs a small granule-sized tooth in 
<taxonomicName id="4C124D55103E392742A9FA56FEBFFD03" authorityName="Cumberlidge and Clark" authorityYear="1992" box="[160,311,628,652]" class="Malacostraca" family="Potamonautidae" genus="Potamonemus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC4103E392742A9FA56FEBFFD03" box="[160,311,628,652]" italics="true" pageId="8" pageNumber="666">Potamonemus</emphasis>
</taxonomicName>
, 
<figureCitation id="13292A53103E3927434AFA56FE15FD03" box="[323,413,628,652]" captionStart="Figure 5" captionStartId="12.[160,225,965,988]" captionTargetBox="[163,1153,142,934]" captionTargetId="figure@12.[163,1153,142,934]" captionTargetPageId="12" captionText="Figure 5. (a–f) Frontal view of right and left chelipeds; (g–i) dorsal view of right cheliped carpus. Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon (NHM 1994.587), length of right cheliped = 24.2 mm. (c–d) Potamonemus mambilorum Cumberlidge and Clark, 1992, holotype, adult ♂, CW 34.5 mm, Somié, Cameroon (NHM 1991:183), length of right cheliped = 37.0 mm; (e–f) P. sachsi, adult ♂, CW 31.6 mm, Obudu Plateau, Nigeria (NMU 8.IV.1983A), length of right cheliped = 27.6 mm; (g) Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon (NHM 1994.587); (h) P. mambilorum Cumberlidge and Clark, 1992, holotype, adult ♂, CW 34.5 mm, Somié, Cameroon (NHM 1991:183); (i) P. sachsi Cumberlidge, 1993, adult ♂, CW 31.6 mm, Obudu Plateau, Nigeria (NMU 8.IV.1983A). Figures (a–i) scaled for comparison. Photographs (a–f) by Phillip Crabb, NHM, London. Scale bar: a,b = 4.21 mm; c,d = 5.26 mm; e, f = 4.82 mm; g = 3.1 mm; h = 3.83 mm; i = 3.51 mm." figureDoi="http://doi.org/10.5281/zenodo.3675504" httpUri="https://zenodo.org/record/3675504/files/figure.png" pageId="8" pageNumber="666">Figure 5</figureCitation>
(h,i)), the cheliped dactylus is distinctly broadened (
<figureCitation id="13292A53103E392741C8FA56FB96FD03" box="[961,1054,628,652]" captionStart="Figure 1" captionStartId="3.[160,225,1645,1668]" captionTargetBox="[163,1153,507,1615]" captionTargetId="figure@3.[163,1153,507,1615]" captionTargetPageId="3" captionText="Figure 1. Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon (NHM 1994.587). (a) Whole animal, dorsal view; (b) whole animal, ventral view. Photographs by Phillip Crabb, NHM, London. Scale bar: a = 14 mm; b = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3675496" httpUri="https://zenodo.org/record/3675496/files/figure.png" pageId="8" pageNumber="666">Figures 1</figureCitation>
(b) and 5 (a)) (vs distinctly slim or arched in 
<taxonomicName id="4C124D55103E39274005FAB5FD2BFD20" authorityName="Cumberlidge and Clark" authorityYear="1992" box="[524,675,663,687]" class="Malacostraca" family="Potamonautidae" genus="Potamonemus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC4103E39274005FAB5FD2BFD20" box="[524,675,663,687]" italics="true" pageId="8" pageNumber="666">Potamonemus</emphasis>
</taxonomicName>
, 
<figureCitation id="13292A53103E392740A6FAB5FC8FFD20" box="[687,775,663,687]" captionStart="Figure 5" captionStartId="12.[160,225,965,988]" captionTargetBox="[163,1153,142,934]" captionTargetId="figure@12.[163,1153,142,934]" captionTargetPageId="12" captionText="Figure 5. (a–f) Frontal view of right and left chelipeds; (g–i) dorsal view of right cheliped carpus. Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon (NHM 1994.587), length of right cheliped = 24.2 mm. (c–d) Potamonemus mambilorum Cumberlidge and Clark, 1992, holotype, adult ♂, CW 34.5 mm, Somié, Cameroon (NHM 1991:183), length of right cheliped = 37.0 mm; (e–f) P. sachsi, adult ♂, CW 31.6 mm, Obudu Plateau, Nigeria (NMU 8.IV.1983A), length of right cheliped = 27.6 mm; (g) Buea asylos (Cumberlidge, 1993) comb. nov., holotype, adult ♂ CW 27.6 mm, between Buea and Kumba, Cameroon (NHM 1994.587); (h) P. mambilorum Cumberlidge and Clark, 1992, holotype, adult ♂, CW 34.5 mm, Somié, Cameroon (NHM 1991:183); (i) P. sachsi Cumberlidge, 1993, adult ♂, CW 31.6 mm, Obudu Plateau, Nigeria (NMU 8.IV.1983A). Figures (a–i) scaled for comparison. Photographs (a–f) by Phillip Crabb, NHM, London. Scale bar: a,b = 4.21 mm; c,d = 5.26 mm; e, f = 4.82 mm; g = 3.1 mm; h = 3.83 mm; i = 3.51 mm." figureDoi="http://doi.org/10.5281/zenodo.3675504" httpUri="https://zenodo.org/record/3675504/files/figure.png" pageId="8" pageNumber="666">Figure 5</figureCitation>
(c,e)), and the carapace proportions of 
<taxonomicName id="4C124D55103E392742B7FA98FE90FD5D" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" baseAuthorityName="Cumberlidge" baseAuthorityYear="1993" box="[190,280,698,722]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="species" species="asylos" status="comb. nov.">
<emphasis id="B966EAC4103E392742B7FA98FE90FD5D" box="[190,280,698,722]" italics="true" pageId="8" pageNumber="666">B. asylos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557BF103E39274329FA98FE13FD5D" box="[288,411,698,722]" pageId="8" pageNumber="666" rank="species">comb. nov.</taxonomicNameLabel>
are wider (CW/FW 3.35, vs 3.08 
<emphasis id="B966EAC4103E392740FCFA98FC8AFD5D" box="[757,770,698,722]" italics="true" pageId="8" pageNumber="666">–</emphasis>
3.12), longer (CW/FW 2.32, vs 2.22 
<emphasis id="B966EAC4103E3927467FFA98FB0BFD5D" box="[1142,1155,698,722]" italics="true" pageId="8" pageNumber="666">–</emphasis>
2.25), and higher (CW/FW 1.10, vs 0.96 
<emphasis id="B966EAC4103E39274037FAFEFDC3FD7B" box="[574,587,732,756]" italics="true" pageId="8" pageNumber="666">–</emphasis>
1.07) than in 
<taxonomicName id="4C124D55103E392740D0FAFEFCF8FD7B" authorityName="Cumberlidge and Clark" authorityYear="1992" box="[729,880,732,756]" class="Malacostraca" family="Potamonautidae" genus="Potamonemus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAC4103E392740D0FAFEFCF8FD7B" box="[729,880,732,756]" italics="true" pageId="8" pageNumber="666">Potamonemus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAD36D6103E392742A9FB03FE05FCB5" blockId="8.[160,1156,801,1727]" box="[160,397,801,826]" pageId="8" pageNumber="666">
<emphasis id="B966EAC4103E392742A9FB03FE05FCB5" bold="true" box="[160,397,801,826]" italics="true" pageId="8" pageNumber="666">Conservation status</emphasis>
</paragraph>
<paragraph id="8BAD36D6103E392742A9FB67FC39F930" blockId="8.[160,1156,801,1727]" pageId="8" pageNumber="666">
The extinction risk status of 
<taxonomicName id="4C124D55103E392743F6FB67FDEBFCD2" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" baseAuthorityName="Cumberlidge" baseAuthorityYear="1993" box="[511,611,837,861]" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="species" species="asylos" status="comb. nov.">
<emphasis id="B966EAC4103E392743F6FB67FDEBFCD2" box="[511,611,837,861]" italics="true" pageId="8" pageNumber="666">B. asylos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557BF103E39274078FB67FD7DFCD2" box="[625,757,837,861]" pageId="8" pageNumber="666" rank="species">comb. nov.</taxonomicNameLabel>
was assessed in 2008 using the International Union for the Conservation of Nature (IUCN) Red List protocols (
<bibRefCitation id="EF834B27103E392742AFFBA8FE00FC2D" author="Cumberlidge N" box="[166,392,906,930]" pageId="8" pageNumber="666" pagination="134656" refId="ref8208" refString="Cumberlidge N 2008 a. Potamonemus asylos. The IUCN Red List of Threatened Species 2008: e. T 134656 A 3993768. [accessed 2018 July 01]. doi: 10.2305 / IUCN. UK. 2008. RLTS. T 134656 A 3993768. en." type="journal article" year="2008">Cumberlidge 2008a</bibRefCitation>
) as Data Deficient (DD) in view of the lack of information on its extent of occurrence (EOO), ecological requirements, population size, population trends and long-term threats (
<bibRefCitation id="EF834B27103E392743AEFBEDFDF2FC68" author="Cumberlidge N" box="[423,634,975,999]" pageId="8" pageNumber="666" pagination="571 - 584" refId="ref8064" refString="Cumberlidge N. 1993. Two new species of Potamonemus Cumberlidge and Clark, 1992 (Potamoidea, Potamonautidae) from the rain forests of West Africa. J Crustacean Biol. 13: 571 - 584. doi: 10.2307 / 1548798" type="journal article" year="1993">Cumberlidge 1993</bibRefCitation>
, 
<bibRefCitation id="EF834B27103E39274081FBEDFD43FC68" author="Cumberlidge N" box="[648,715,975,999]" pageId="8" pageNumber="666" pagination="71 - 78" refId="ref8358" refString="Cumberlidge N. 2011 a. Chapter 6 The status and distribution of freshwater crabs. In: Brooks EGE, Allen DJ, editors Darwall WRT compilers. The status and distribution of freshwater biodiversity in central Africa. Gland, Switzerland and Cambridge (UK): IUCN. p. 71 - 78." type="book chapter" year="2011">2011a</bibRefCitation>
, 
<bibRefCitation id="EF834B27103E392740D0FBEDFC96FC68" author="Cumberlidge N" box="[729,798,975,999]" pageId="8" pageNumber="666" pagination="178 - 199" refId="ref8415" refString="Cumberlidge N. 2011 b. Freshwater crabs of Africa: diversity, distribution, and conservation Chapter 6. In: Darwall WRT, Smith KG, Allen DJ, Holland RA, Harrison IJ, Brooks EGE, editors. The diversity of life in African freshwaters: under water, under threat. An analysis of the status and distribution of freshwater species throughout Mainland Africa. Gland Switzerland and Cambridge (UK): IUCN; p. 178 - 199." type="book chapter" year="2011">2011b</bibRefCitation>
; 
<bibRefCitation id="EF834B27103E39274124FBEDFBB5FC68" author="Cumberlidge N &amp; Ng PKL &amp; Yeo DCJ &amp; Magalhaes C &amp; Campos MR &amp; Alvarez F &amp; Naruse T &amp; Daniels SR &amp; Esser LJ &amp; Attipoe FYK" box="[813,1085,975,999]" pageId="8" pageNumber="666" pagination="1665 - 1673" refId="ref8548" refString="Cumberlidge N, Ng PKL, Yeo DCJ, Magalhaes C, Campos MR, Alvarez F, Naruse T, Daniels SR, Esser LJ, Attipoe FYK, et al. 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biol Cons. 142: 1665 - 1673. doi: 10.1016 / j. biocon. 2009.02.038" type="journal article" year="2009">Cumberlidge et al. 2009</bibRefCitation>
; 
<bibRefCitation id="EF834B27103E39274642FBEDFF5FFB85" author="IUCN" pageId="8" pageNumber="666" refId="ref8986" refString="IUCN. 2012. IUCN red list categories and criteria: version 3.1. 2 nd ed. Gland Switzerland and Cambridge (UK): IUCN; p. iv + p. 32." type="journal volume" year="2012">IUCN 2012</bibRefCitation>
). The three locations available in this study give a recalculated EOO of 
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, and an area of occupancy (AOO) of 
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, using GeoCAT (http://geocat.kew.org; 
<bibRefCitation id="EF834B27103E39274612FC37FE9EFBC0" author="Bachman S &amp; Moat J &amp; Hill AW &amp; de Torre J &amp; Scott B" pageId="8" pageNumber="666" pagination="117 - 126" refId="ref7820" refString="Bachman S, Moat J, Hill AW, de Torre J, Scott B. 2011. Supporting red list threat assessments with GeoCAT: geospatial conservation assessment tool. ZooKeys. 150: 117 - 126. doi: 10.3897 / zookeys. 150.2109" type="journal article" year="2011">Bachman et al. 2011</bibRefCitation>
), but these are still probably underestimates given the paucity of the locality data (but if taken alone would point to a Red List threatened category). The extinction risk status of 
<taxonomicName id="4C124D55103E3927433CFC5FFE55FB1A" authorityName="Cumberlidge and Clark" authorityYear="1992" box="[309,477,1149,1173]" class="Malacostraca" family="Potamonautidae" genus="Potamonemus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="species" species="mambilorum">
<emphasis id="B966EAC4103E3927433CFC5FFE55FB1A" box="[309,477,1149,1173]" italics="true" pageId="8" pageNumber="666">P. mambilorum</emphasis>
</taxonomicName>
(seven locations, EOO 
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; 
<bibRefCitation id="EF834B27103E39274162FC5FFBC2FB1A" author="Cumberlidge N" box="[875,1098,1149,1173]" pageId="8" pageNumber="666" pagination="134438" refId="ref8258" refString="Cumberlidge N 2008 b. Potamonemus mambilorum. The IUCN Red List of Threatened Species 2008: e. T 134438 A 3956693. [accessed 2018 Oct 10]. doi: 10.2305 / IUCN. UK. 2008. RLTS. T 134438 A 3956693. en." type="journal article" year="2008">Cumberlidge 2008b</bibRefCitation>
) and 
<taxonomicName id="4C124D55103E392742A9FCBDFF77FB38" authorityName="Cumberlidge" authorityYear="1993" box="[160,255,1183,1207]" class="Malacostraca" family="Potamonautidae" genus="Potamonemus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="species" species="sachsi">
<emphasis id="B966EAC4103E392742A9FCBDFF77FB38" box="[160,255,1183,1207]" italics="true" pageId="8" pageNumber="666">P. sachsi</emphasis>
</taxonomicName>
(four locations, EOO 
<quantity id="4CEA9B33103E392743F5FCBDFDFCFB38" box="[508,628,1183,1207]" metricMagnitude="7" metricUnit="m" metricValue="2.4219" pageId="8" pageNumber="666" unit="km" value="24219.0">24,219 km</quantity>
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; 
<bibRefCitation id="EF834B27103E39274087FCBDFCE4FB38" author="Cumberlidge N" box="[654,876,1183,1207]" pageId="8" pageNumber="666" pagination="134901" refId="ref8308" refString="Cumberlidge N 2008 c. Potamonemus sachsi. The IUCN Red List of Threatened Species 2008: e. T 134901 A 4033268. [accessed 2018 Oct 10]. doi: 10.2305 / IUCN. UK. 2008. RLTS. T 134901 A 4033268. en." type="journal article" year="2008">Cumberlidge 2008c</bibRefCitation>
) were assessed in 2008 (
<bibRefCitation id="EF834B27103E392742AFFCE0FE03FB55" author="Cumberlidge N" box="[166,395,1218,1242]" pageId="8" pageNumber="666" pagination="134438" refId="ref8258" refString="Cumberlidge N 2008 b. Potamonemus mambilorum. The IUCN Red List of Threatened Species 2008: e. T 134438 A 3956693. [accessed 2018 Oct 10]. doi: 10.2305 / IUCN. UK. 2008. RLTS. T 134438 A 3956693. en." type="journal article" year="2008">Cumberlidge 2008b</bibRefCitation>
, 
<bibRefCitation id="EF834B27103E39274395FCE0FE56FB55" author="Cumberlidge N" box="[412,478,1218,1242]" pageId="8" pageNumber="666" pagination="134901" refId="ref8308" refString="Cumberlidge N 2008 c. Potamonemus sachsi. The IUCN Red List of Threatened Species 2008: e. T 134901 A 4033268. [accessed 2018 Oct 10]. doi: 10.2305 / IUCN. UK. 2008. RLTS. T 134901 A 4033268. en." type="journal article" year="2008">2008c</bibRefCitation>
) as Least Concern (LC) and Vulnerable (VU), respectively, (
<bibRefCitation id="EF834B27103E392742AFFCC7FEF2FB72" author="Cumberlidge N" box="[166,378,1253,1277]" pageId="8" pageNumber="666" pagination="571 - 584" refId="ref8064" refString="Cumberlidge N. 1993. Two new species of Potamonemus Cumberlidge and Clark, 1992 (Potamoidea, Potamonautidae) from the rain forests of West Africa. J Crustacean Biol. 13: 571 - 584. doi: 10.2307 / 1548798" type="journal article" year="1993">Cumberlidge 1993</bibRefCitation>
, 
<bibRefCitation id="EF834B27103E39274383FCC7FE45FB72" author="Cumberlidge N" box="[394,461,1253,1277]" pageId="8" pageNumber="666" pagination="71 - 78" refId="ref8358" refString="Cumberlidge N. 2011 a. Chapter 6 The status and distribution of freshwater crabs. In: Brooks EGE, Allen DJ, editors Darwall WRT compilers. The status and distribution of freshwater biodiversity in central Africa. Gland, Switzerland and Cambridge (UK): IUCN. p. 71 - 78." type="book chapter" year="2011">2011a</bibRefCitation>
, 
<bibRefCitation id="EF834B27103E392743D7FCC7FDABFB72" author="Cumberlidge N" box="[478,547,1253,1277]" pageId="8" pageNumber="666" pagination="178 - 199" refId="ref8415" refString="Cumberlidge N. 2011 b. Freshwater crabs of Africa: diversity, distribution, and conservation Chapter 6. In: Darwall WRT, Smith KG, Allen DJ, Holland RA, Harrison IJ, Brooks EGE, editors. The diversity of life in African freshwaters: under water, under threat. An analysis of the status and distribution of freshwater species throughout Mainland Africa. Gland Switzerland and Cambridge (UK): IUCN; p. 178 - 199." type="book chapter" year="2011">2011b</bibRefCitation>
; 
<bibRefCitation id="EF834B27103E3927403AFCC7FCC2FB72" author="Cumberlidge N &amp; Ng PKL &amp; Yeo DCJ &amp; Magalhaes C &amp; Campos MR &amp; Alvarez F &amp; Naruse T &amp; Daniels SR &amp; Esser LJ &amp; Attipoe FYK" box="[563,842,1253,1277]" pageId="8" pageNumber="666" pagination="1665 - 1673" refId="ref8548" refString="Cumberlidge N, Ng PKL, Yeo DCJ, Magalhaes C, Campos MR, Alvarez F, Naruse T, Daniels SR, Esser LJ, Attipoe FYK, et al. 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biol Cons. 142: 1665 - 1673. doi: 10.1016 / j. biocon. 2009.02.038" type="journal article" year="2009">Cumberlidge et al. 2009</bibRefCitation>
; 
<bibRefCitation id="EF834B27103E39274150FCC7FC5BFB72" author="IUCN" box="[857,979,1253,1277]" pageId="8" pageNumber="666" refId="ref8986" refString="IUCN. 2012. IUCN red list categories and criteria: version 3.1. 2 nd ed. Gland Switzerland and Cambridge (UK): IUCN; p. iv + p. 32." type="journal volume" year="2012">IUCN 2012</bibRefCitation>
). Although the recalculations of the EOO and the AOO for these three species do not in themselves warrant a reassessment of their extinction risk, there is reason to believe that the threat status of these species may have intensified since the last assessment. For example, recent field work in the wetland ecosystems of the 
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and Littoral regions of 
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by the second author indicates that these habitats are being impacted by deforestation and by intensive agricultural practices that are severely altering the flow patterns of small streams and impacting the aquatic biodiversity (
<bibRefCitation id="EF834B27103E39274186FDF5FF6CF99D" author="Mvogo Ndongo PA &amp; Cumberlidge N &amp; Poettinger TS &amp; von Rintelen T &amp; Tamesse JL &amp; Schubart CD" pageId="8" pageNumber="666" pagination="439 - 444" refId="ref9484" refString="Mvogo Ndongo PA, Cumberlidge N, Poettinger TS, von Rintelen T, Tamesse JL, Schubart CD. 2017 a. Molecular evidence for the assignment of the Cameroonian freshwater crab genus Louisea Cumberlidge, 1994, to the Afrotropical subfamily Potamonautinae Bott, 1970 (Crustacea: Potamoidea: Potamonautidae). Zootaxa. 4286: 439 - 444. doi: 10.11646 / zootaxa. 4286.3.12" type="journal article" year="2017">Mvogo Ndongo et al. 2017a</bibRefCitation>
, 
<bibRefCitation id="EF834B27103E392742FCFDD8FEB3F99D" author="Mvogo Ndongo PA &amp; Schubart CD &amp; von Rintelen T &amp; Tamesse JL &amp; Cumberlidge N" box="[245,315,1530,1554]" pageId="8" pageNumber="666" pagination="161 - 173" refId="ref9554" refString="Mvogo Ndongo PA, Schubart CD, von Rintelen T, Tamesse JL, Cumberlidge N. 2017 b. Morphological and molecular evidence for a new species of freshwater crab of the genus Sudanonautes Bott, 1955 (Brachyura: Potamoidea: Potamonautidae) from Cameroon, with notes on its ecology. Zootaxa. 4242: 161 - 173. doi: 10.11646 / zootaxa. 4242.1.8" type="journal article" year="2017">2017b</bibRefCitation>
, 
<bibRefCitation id="EF834B27103E39274345FDD8FE06F99D" author="Mvogo Ndongo PA &amp; von Rintelen T &amp; Schubart CD &amp; Albrecht C &amp; Tamesse JL &amp; Cumberlidge N" box="[332,398,1530,1554]" pageId="8" pageNumber="666" pagination="273 - 280" refId="ref9690" refString="Mvogo Ndongo PA, von Rintelen T, Schubart CD, Albrecht C, Tamesse JL, Cumberlidge N. 2017 c. New data on the taxonomy, ecology, and conservation of the rediscovered Louisea edeaensis (Bott, 1969) (Brachyura: Potamoidea: Potamonautidae), an endangered freshwater crab from Cameroon. Zootaxa. 4231: 273 - 280. doi: 10.11646 / zootaxa. 4231.2.9" type="journal article" year="2017">2017c</bibRefCitation>
, 
<bibRefCitation id="EF834B27103E392743A9FDD8FE5FF99D" author="Mvogo Ndongo PA &amp; von Rintelen T &amp; Albrecht C &amp; Tamesse JL &amp; Cumberlidge N" box="[416,471,1530,1554]" pageId="8" pageNumber="666" pagination="407 - 416" refId="ref9623" refString="Mvogo Ndongo PA, von Rintelen T, Albrecht C, Tamesse JL, Cumberlidge N. 2018. Lost species in Cameroon: rediscovery of the endangered freshwater crab, Louisea balssi (Bott, 1959) (Brachyura: Potamonautidae), with notes on its ecology and conservation. Zootaxa. 4394: 407 - 416. doi: 10.11646 / zootaxa. 4394.3.6" type="journal article" year="2018">2018</bibRefCitation>
). Awareness of these current threats to the habitats of 
<taxonomicName id="4C124D55103E39274679FDD8FF6AF9BA" authorityName="Cumberlidge &amp; Mvogo Ndongo &amp; Clark &amp; Daniels" authorityYear="2019" baseAuthorityName="Cumberlidge" baseAuthorityYear="1993" class="Malacostraca" family="Potamonautidae" genus="Buea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="species" species="asylos" status="comb. nov.">
<emphasis id="B966EAC4103E39274679FDD8FF6AF9BA" italics="true" pageId="8" pageNumber="666">B. asylos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557BF103E392742E0FE3FFEE1F9BA" box="[233,361,1565,1589]" pageId="8" pageNumber="666" rank="species">comb. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C124D55103E3927437AFE3FFD92F9BA" authorityName="Cumberlidge and Clark" authorityYear="1992" box="[371,538,1565,1589]" class="Malacostraca" family="Potamonautidae" genus="Potamonemus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="species" species="mambilorum">
<emphasis id="B966EAC4103E3927437AFE3FFD92F9BA" box="[371,538,1565,1589]" italics="true" pageId="8" pageNumber="666">P. mambilorum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C124D55103E3927405DFE3FFD38F9BA" authorityName="Cumberlidge" authorityYear="1993" box="[596,688,1565,1589]" class="Malacostraca" family="Potamonautidae" genus="Potamonemus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="666" phylum="Arthropoda" rank="species" species="sachsi">
<emphasis id="B966EAC4103E3927405DFE3FFD38F9BA" box="[596,688,1565,1589]" italics="true" pageId="8" pageNumber="666">P. sachsi</emphasis>
</taxonomicName>
means that each of these taxa is likely to be reassigned to a more threatened category once new IUCN Red List extinction risk assessments have been carried out. There is a need for further field research specifically aimed at gathering the data needed for an extinction risk assessment of these littleknown endemic species from this understudied biodiversity hotspot.
</paragraph>
</subSubSection>
</treatment>
</document>