147 lines
17 KiB
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147 lines
17 KiB
XML
<document id="80C081C8BFEF6561CE1C4F941F25FBFE" ID-DOI="10.37828/em.2023.63.2" ID-ISSN="2336-9744" ID-ZooBank="urn:lsid:zoobank.org:pub:24309667-AD59-4E64-BD11-F5216394126A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1686850451515" checkinUser="felipe" docAuthor="Volynkin, Anton V." docDate="2023" docId="03BB87BEFF9AD5463F85FC77FD67A957" docLanguage="en" docName="EcolMontenegrina.63.15-23.pdf" docOrigin="Ecologica Montenegrina 63" docSource="http://dx.doi.org/10.37828/em.2023.63.2" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Teulisna pallida Volynkin 2023" docType="treatment" docVersion="1" lastPageNumber="17" masterDocId="FF82FFC6FF9BD5443F12FFDCFFDAAB06" masterDocTitle="Taxonomic review of the Teulisna pallida (Moore) species-group with descriptions of two new species from the Himalayas (Lepidoptera: Erebidae: Arctiinae: Lithosiini)" masterLastPageNumber="23" masterPageNumber="15" pageNumber="16" updateTime="1686850452878" updateUser="felipe" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BDF23E3E02A3ACE878DC87ACF1C738BB">Taxonomic review of the Teulisna pallida (Moore) species-group with descriptions of two new species from the Himalayas (Lepidoptera: Erebidae: Arctiinae: Lithosiini)</mods:title>
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<mods:namePart id="CC377CF179D5863EB04E289D98CFDBB3">Volynkin, Anton V.</mods:namePart>
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<mods:title id="1F1799BF7399E94D76E6BCCAB19A0F92">Ecologica Montenegrina</mods:title>
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<mods:part id="7364B5FBA749A9D8B0909C30A08917A2">
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<mods:date id="D9F35437BB10205FED1132AEDB9F23A5">2023</mods:date>
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<mods:number id="5F938BE868656A9847E31C38EC573CBC">2023-05-31</mods:number>
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<mods:number id="6E32BFE879DC0979C57A2425930F4574">63</mods:number>
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<mods:url id="5D46E2CE4B59A65A181D5A722CFEE778">http://dx.doi.org/10.37828/em.2023.63.2</mods:url>
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<mods:identifier id="1142189C2BB7F94E504A204E1D1098DD" type="DOI">10.37828/em.2023.63.2</mods:identifier>
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<mods:identifier id="225D04801A8AA3841A419BB5EA347BF0" type="ISSN">2336-9744</mods:identifier>
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<mods:identifier id="3550B7C93072F7CE5D60E21708A6DE9A" type="ZooBank">urn:lsid:zoobank.org:pub:24309667-AD59-4E64-BD11-F5216394126A</mods:identifier>
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<treatment id="03BB87BEFF9AD5463F85FC77FD67A957" LSID="urn:lsid:plazi:treatment:03BB87BEFF9AD5463F85FC77FD67A957" httpUri="http://treatment.plazi.org/id/03BB87BEFF9AD5463F85FC77FD67A957" lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16">
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<subSubSection id="C3086523FF9AD5453F85FC77FD9CA8C3" box="[151,582,939,965]" pageId="1" pageNumber="16" type="nomenclature">
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<paragraph id="8BAD36A8FF9AD5453F85FC77FD9CA8C3" blockId="1.[151,582,939,965]" box="[151,582,939,965]" pageId="1" pageNumber="16">
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<heading id="D0E581C4FF9AD5453F85FC77FD9CA8C3" box="[151,582,939,965]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="16" reason="2">
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<emphasis id="B966EABAFF9AD5453F85FC77FD9CA8C3" bold="true" box="[151,582,939,965]" pageId="1" pageNumber="16">
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The
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<taxonomicName id="4C124D2BFF9AD5453FDDFC77FE4EA8C3" authorityName="Volynkin" authorityYear="2023" baseAuthorityName="Volynkin" baseAuthorityYear="2023" box="[207,404,939,965]" class="Insecta" family="Arctiidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="species" species="pallida">
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<emphasis id="B966EABAFF9AD5453FDDFC77FE4EA8C3" bold="true" box="[207,404,939,965]" italics="true" pageId="1" pageNumber="16">Teulisna pallida</emphasis>
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</taxonomicName>
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species-group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3086523FF9AD5463F85FC31FDAEAA06" lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16" type="diagnosis">
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<paragraph id="8BAD36A8FF9AD5453F85FC31FB1CAF6B" blockId="1.[151,1437,1005,2010]" pageId="1" pageNumber="16">
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<emphasis id="B966EABAFF9AD5453F85FC31FECAAF01" bold="true" box="[151,272,1005,1031]" pageId="1" pageNumber="16">Diagnosis</emphasis>
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. The species-group is characterised by the relatively limited sexual dimorphism. In the male genitalia, unlike a number of other
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<taxonomicName id="4C124D2BFF9AD5453D2AFBD3FD27AF2C" box="[568,765,1039,1066]" pageId="1" pageNumber="16">
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<emphasis id="B966EABAFF9AD5453D2AFBD3FD47AF2C" box="[568,669,1039,1066]" italics="true" pageId="1" pageNumber="16">Teulisna</emphasis>
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species
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</taxonomicName>
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, the vincular coremata are small, the vincular medial rod is reduced, and the distal saccular process is broad, flattened and distally bilobate. In the female genitalia, the corpus and the appendix bursae are gelatinous, which is characteristic of the species-group.
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</paragraph>
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<paragraph id="8BAD36A8FF9AD5463FE4FBA8FDAEAA06" blockId="1.[151,1437,1005,2010]" lastBlockId="2.[151,1436,195,323]" lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16">
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<emphasis id="B966EABAFF9AD5453FE4FBA8FDBFAF88" bold="true" box="[246,613,1140,1166]" pageId="1" pageNumber="16">Morphological characteristic</emphasis>
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.
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<emphasis id="B966EABAFF9AD5453D6BFBA8FD16AF88" bold="true" box="[633,716,1140,1166]" pageId="1" pageNumber="16">Adults</emphasis>
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.
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<emphasis id="B966EABAFF9AD5453DF2FBA8FCFAAF88" bold="true" box="[736,800,1140,1166]" pageId="1" pageNumber="16">Male</emphasis>
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. Antenna ciliate. Body from greyish-brown to dark brown. Forewing elongate and narrow with postmedially convex costa. Forewing ground colour from pale ochreous to dark brown. Costa with narrow, dash-shaped black spot antemedially and broader trapezoidal black spot medially. Discal spot black, dot-shaped. Additional, postero-medial spot present in certain species. Medial and postmedial areas with intense greyish-brown suffusion in cell and along anal margin forming shade distally terminating by irregularly dentate subterminal line. Terminal area with weak greyish-brown suffusion. Hindwing pale ochreous with dark grey suffusion medially. Abdomen from ochreous brown to brown.
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<emphasis id="B966EABAFF9AD5453E63FABCFDF2AE7C" bold="true" box="[369,552,1376,1402]" pageId="1" pageNumber="16">Male genitalia</emphasis>
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. Uncus elongate, slender, laterally flattened, smoothly downcurved and apically pointed. Tegumen short with broad arms fused in posterior three-quarters. Vinculum longer than tegumen, with thin and flexible arms anteriorly connected by thin commissure. Intravincular membrane with two small anterio-lateral coremata. Valva broad, lobular with almost parallel margins and apically rounded dorsal section. Clasper heavily sclerotised and having short spike-like ampulla in certain species. Sacculi broad and connected to each other by wide commissure fused with juxta. Distal saccular process broad, flattened, with two triangular distal lobes. Juxta pyramidal and medially membranous. Phallus broad, cylindrical, somewhat downcurved medially and with short and rounded coecum. Vesica broad, somewhat longer than phallus, directed distally, having several diverticula. Proximal ventral diverticulum large and heavily scobinated. Proximal dorsal diverticulum present in certain species, membranous. Medial ventral diverticulum conical and weakly granulose. Medial dorsal diverticulum conical or utricular, heavily granulose. Distal ventral diverticulum membranous, sack-like, with vesica ejaculatorius originating from its dorsal side. Distal dorsal diverticulum short, utricular and granulose, and having long and robust nail-like terminal cornutus.
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<emphasis id="B966EABAFF9AD5453E64F8CBFE0BAC37" bold="true" box="[374,465,1815,1841]" pageId="1" pageNumber="16">Female</emphasis>
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. Antenna ciliate. Sexual dimorphism limited: in comparison to male, female with more elongate forewing lacking proximal costal spot and, in most species, having less intense dark suffusion medially.
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<emphasis id="B966EABAFF9AD5453E1FF886FE06AC72" bold="true" box="[269,476,1882,1908]" pageId="1" pageNumber="16">Female genitalia</emphasis>
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. Papilla analis trapezoidal with rounded corners, setose. Apophyses elongate and thin, equal in length. Ductus bursae short, dorso-ventrally flattened, with convex lateral margins. Posterior section of corpus bursae heavily gelatinous, dilated, with semiglobular postero-lateral diverticulum, and broad dorsal sclerotised pocket in certain species. Anterior section of corpus bursae weakly gelatinous, shorter and narrower than posterior one, teardrop-shaped. Appendix bursae heavily gelatinous, conical or sack-like, situated laterally on right side.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3086523FF99D5463FE4FEDBFD67A957" pageId="2" pageNumber="17" type="distribution">
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<paragraph id="8BAD36A8FF99D5463FE4FEDBFEFBAA45" blockId="2.[151,1436,195,323]" pageId="2" pageNumber="17">
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<emphasis id="B966EABAFF99D5463FE4FEDBFE4AAA27" bold="true" box="[246,400,263,289]" pageId="2" pageNumber="17">Distribution</emphasis>
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. Species of the species-group are known from Himalaya (North and Northeast
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<collectingCountry id="F3057638FF99D5463A4DFEDBFA41AA24" box="[1375,1435,263,290]" name="India" pageId="2" pageNumber="17">India</collectingCountry>
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and
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<collectingCountry id="F3057638FF99D5463FDBFEF4FECEAA45" box="[201,276,296,323]" name="Nepal" pageId="2" pageNumber="17">Nepal</collectingCountry>
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).
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</paragraph>
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<paragraph id="8BAD36A8FF99D5463F85FE53FD8DAAAF" blockId="2.[151,701,399,593]" box="[151,599,399,425]" pageId="2" pageNumber="17">
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<emphasis id="B966EABAFF99D5463F85FE53FD8DAAAF" bold="true" box="[151,599,399,425]" pageId="2" pageNumber="17">Species content of the species-group</emphasis>
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</paragraph>
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<paragraph id="8BAD36A8FF99D5463F85FE6CFD96AACC" blockId="2.[151,701,399,593]" box="[151,588,431,458]" pageId="2" pageNumber="17">
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–
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<taxonomicName id="4C124D2BFF99D5463FBFFE6DFE0FAACC" authority="(Moore, 1878)" authorityName="Volynkin" authorityYear="2023" baseAuthorityName="Volynkin" baseAuthorityYear="2023" box="[173,469,431,458]" class="Insecta" family="Arctiidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="pallida" status="comb. n.">
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<emphasis id="B966EABAFF99D5463FBFFE6DFEC5AACC" box="[173,287,431,458]" italics="true" pageId="2" pageNumber="17">T. pallida</emphasis>
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(
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<bibRefCitation id="EF834B59FF99D5463E3DFE73FE11AACC" author="Moore, F." box="[303,459,431,458]" pageId="2" pageNumber="17" pagination="3 - 37" refId="ref4162" refString="Moore, F. (1878) A revision of certain genera of European and Asiatic Lithosiinae, with characters of new genera and species. Proceedings of the general meetings for scientific business of the Zoological Society of London, 1878, 3 - 37, pls. 1 - 3. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 07927. x" type="journal article" year="1878">Moore, 1878</bibRefCitation>
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)
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</taxonomicName>
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,
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<emphasis id="B966EABAFF99D5463EF2FE6CFD96AACC" bold="true" box="[480,588,432,458]" pageId="2" pageNumber="17">
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<taxonomicNameLabel id="A25557C1FF99D5463EF2FE6CFD96AACC" box="[480,588,432,458]" pageId="2" pageNumber="17" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8BAD36A8FF99D5463F85FE0EFEA2AAEA" blockId="2.[151,701,399,593]" box="[151,376,465,492]" pageId="2" pageNumber="17">
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–
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<taxonomicName id="4C124D2BFF99D5463FBFFE0EFEF4AAEA" authority="Volynkin, 2023" authorityName="Volynkin" authorityYear="2023" box="[173,302,465,492]" class="Insecta" family="Arctiidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="ganesha" status="sp. n.">
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<emphasis id="B966EABAFF99D5463FBFFE0EFEF4AAEA" box="[173,302,465,492]" italics="true" pageId="2" pageNumber="17">T. ganesha</emphasis>
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</taxonomicName>
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<emphasis id="B966EABAFF99D5463E27FE0EFEA2AAEA" bold="true" box="[309,376,466,492]" pageId="2" pageNumber="17">
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<taxonomicNameLabel id="A25557C1FF99D5463E27FE0EFEA2AAEA" box="[309,376,466,492]" pageId="2" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8BAD36A8FF99D5463F85FE28FEA9A90B" blockId="2.[151,701,399,593]" box="[151,371,499,526]" pageId="2" pageNumber="17">
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–
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<taxonomicName id="4C124D2BFF99D5463FBFFE28FEF3A908" authority="Volynkin, 2023" authorityName="Volynkin" authorityYear="2023" box="[173,297,499,526]" class="Insecta" family="Arctiidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="thomasi" status="sp. n.">
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<emphasis id="B966EABAFF99D5463FBFFE28FEF3A908" box="[173,297,499,526]" italics="true" pageId="2" pageNumber="17">T. thomasi</emphasis>
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</taxonomicName>
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<emphasis id="B966EABAFF99D5463E23FE2FFEA9A90B" bold="true" box="[305,371,499,525]" pageId="2" pageNumber="17">
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<taxonomicNameLabel id="A25557C1FF99D5463E23FE2FFEA9A90B" box="[305,371,499,525]" pageId="2" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8BAD36A8FF99D5463F85FDCAFD47A929" blockId="2.[151,701,399,593]" box="[151,669,533,560]" pageId="2" pageNumber="17">
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–
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<taxonomicName id="4C124D2BFF99D5463FBFFDCAFDFCA936" authority="(Moore, 1878)" authorityName="Volynkin" authorityYear="2023" baseAuthorityName="Moore" baseAuthorityYear="1878" box="[173,550,533,560]" class="Insecta" family="Erebidae" genus="Cossa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="quadrisignata" status="comb. n.">
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<emphasis id="B966EABAFF99D5463FBFFDCAFEABA936" box="[173,369,533,560]" italics="true" pageId="2" pageNumber="17">T. quadrisignata</emphasis>
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(
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<bibRefCitation id="EF834B59FF99D5463E92FDC9FDC4A936" author="Moore, F." box="[384,542,533,560]" pageId="2" pageNumber="17" pagination="3 - 37" refId="ref4162" refString="Moore, F. (1878) A revision of certain genera of European and Asiatic Lithosiinae, with characters of new genera and species. Proceedings of the general meetings for scientific business of the Zoological Society of London, 1878, 3 - 37, pls. 1 - 3. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 07927. x" type="journal article" year="1878">Moore, 1878</bibRefCitation>
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)
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</taxonomicName>
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,
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<emphasis id="B966EABAFF99D5463D20FDC9FD47A929" bold="true" box="[562,669,533,559]" pageId="2" pageNumber="17">
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<taxonomicNameLabel id="A25557C1FF99D5463D20FDC9FD47A929" box="[562,669,533,559]" pageId="2" pageNumber="17" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8BAD36A8FF99D5463FE4FDEBFD67A957" blockId="2.[151,701,399,593]" box="[246,701,566,593]" pageId="2" pageNumber="17">
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=
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<taxonomicName id="4C124D2BFF99D5463E1CFDEBFDBAA957" authority="Moore, 1878" authorityName="Moore" authorityYear="1878" box="[270,608,566,593]" class="Insecta" family="Erebidae" genus="Cossa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="brunnea" status="syn. n.">
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<emphasis id="B966EABAFF99D5463E1CFDEBFE65A957" box="[270,447,566,593]" italics="true" pageId="2" pageNumber="17">Cossa brunnea</emphasis>
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<bibRefCitation id="EF834B59FF99D5463ED4FDEBFDBAA957" author="Moore, F." box="[454,608,566,593]" pageId="2" pageNumber="17" pagination="3 - 37" refId="ref4162" refString="Moore, F. (1878) A revision of certain genera of European and Asiatic Lithosiinae, with characters of new genera and species. Proceedings of the general meetings for scientific business of the Zoological Society of London, 1878, 3 - 37, pls. 1 - 3. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 07927. x" type="journal article" year="1878">Moore, 1878</bibRefCitation>
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</taxonomicName>
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,
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<emphasis id="B966EABAFF99D5463D7EFDEBFD67A957" bold="true" box="[620,701,567,593]" pageId="2" pageNumber="17">
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<taxonomicNameLabel id="A25557C1FF99D5463D7EFDEBFD67A957" box="[620,701,567,593]" pageId="2" pageNumber="17">syn. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |